Late Cretaceous Strata and Vertebrate Fossils of North Texas

Total Page:16

File Type:pdf, Size:1020Kb

Late Cretaceous Strata and Vertebrate Fossils of North Texas The Geological Society of America Field Guide 30 2013 Late Cretaceous strata and vertebrate fossils of North Texas Louis L. Jacobs* Michael J. Polcyn Dale A. Winkler Timothy S. Myers Jamell G. Kennedy John B. Wagner Roy M. Huffi ngton Department of Earth Sciences, Southern Methodist University, Dallas, Texas 75275, USA ABSTRACT Outcrops of Late Cretaceous Gulf Series strata (Woodbine, Eagle Ford, and Aus- tin) in the Dallas area expose middle Cenomanian to the early Campanian (96 to ~83 Ma) rocks, which are well known in the subsurface of the oil-rich East Texas Basin. Together with the underlying Comanche Series and overlying younger Gulf Series, this set of strata provides a record of the last 50 million years of the Creta- ceous. Although both marine and terrestrial vertebrates are known in this interval, the Late Cretaceous record is primarily marine. On this fi eld trip, sites are visited that have yielded sharks, bony fi sh, turtles, dinosaurs, crocodiles, pterosaurs, mam- mals, long- and short-necked plesiosaurs, and a classic record of mosasaur evolution. INTRODUCTION of the Gulf Series (Woodbine, Eagle Ford, Austin, Taylor, and Navarro) are generally considered third-order cycles (e.g., Adams The stratigraphic section (Fig. 1) between Lake Grapevine and Carr, 2010; Liro et al., 1994), ranging from ~2 million years west of the Dallas–Fort Worth International Airport and the North in duration (Woodbine) to ~12 million (Taylor). Because of time Sulphur River (Fig. 2), 130 km to the east of Dallas, records the and logistical constraints, this fi eld trip will focus on the Wood- past 30 million years of Cretaceous time and displays in out- bine, Eagle Ford, and Austin units between Lake Grapevine and crop the highly petroliferous rock units that produce from the downtown Dallas. subsurface in the East Texas Basin. The rocks comprise the Gulf Structurally, the rocks visited on this fi eld trip lie on the west Series of Hill (1887, 1901; Jacobs et al., 2012). Hill divided the side of the East Texas Basin, which extends between the San Mar- Gulf Series into the Woodbine Formation, Eagle Ford Formation, cos Arch and the Sabine Uplift (Fig. 3), both of which are Late Austin Chalk, Taylor Formation, and Navarro Formation. Each Cretaceous topographic highs (Adams 2006, 2009). The western of these units was subdivided further by Hill and other authors. boundary of the basin from Dallas toward the east is marked by The placement of Hill’s original units and their subdivisions into two parallel fault systems (Fig. 2). In Dallas, the Balcones fault a lithostratigraphic hierarchy is utilitarian but inconsistent among zone of central Texas extends along the White Rock escarpment various authors. More importantly, however, the major units (Reaser, 1961) capped by the Austin Chalk, and according to *[email protected] Jacobs, L.L., Polcyn, M.J., Winkler, D.A., Myers, T.S., Kennedy, J.G., and Wagner, J.B., 2013, Late Cretaceous strata and vertebrate fossils of North Texas, in Hunt, B.B., and Catlos, E.J., eds., Late Cretaceous to Quaternary Strata and Fossils of Texas: Field Excursions Celebrating 125 Years of GSA and Texas Geology, GSA South-Central Section Meeting, Austin, Texas, April 2013: Geological Society of America Field Guide 30, p. 1–13, doi:10.1130/2013.0030(01). For permission to copy, contact [email protected]. © 2013 The Geological Society of America. All rights reserved. 1 2 Jacobs et al. Reaser (2002), behaves as a true hinge. [These faults are not to be confused with Paleozoic faults of the Ouachita trend slightly to the west in the Fort Worth Basin that may have been activated by fl uid injection related to Barnett Shale production (Frohlich, 2012).] East of Dallas lies the Mexia fault zone, which is a pull- apart structure that formed at the edge of the mobile salt deposits of the Jurassic Luann Formation, which underlies the East Texas Basin (Jackson, 1982). Faults of the Mexia fault zone were active during the Cretaceous and into the Paleogene. The regressive Woodbine Formation at the base of the Gulf Series represents fl uvial, strand plain, deltaic, and paludal envi- ronments (Oliver, 1971). In the Dallas area the Woodbine was emergent as shown by its angiosperm fl ora, including leaves and logs up to 30 cm or more in diameter, bones, teeth, and foot- prints of terrestrial vertebrates, trace fossils of intertidal crabs, and occasional Protosols and Histosols (sensu Mack et al., 1993). The remainder of the Gulf Series is marine. The Wood- bine and the Eagle Ford are completely truncated by erosion at the Sabine Uplift on the eastern margin of the East Texas Basin, indicating post-Woodbine uplift followed by subsidence prior to the deposition of the Austin Chalk, which is continuous across the Basin (Adams and Carr, 2010; Fig. 3). The structural setting of the East Texas Basin indicates that depositional systems of the Late Cretaceous were not solely controlled by eustatic sea level (Adams, 2009). Chronological control of the Gulf Series is provided primar- ily through biostratigraphy, although tuffaceous units and ben- tonites have long been reported in the Woodbine, Eagle Ford, and Austin (e.g., Ross et al., 1929). Radiometric dates of 86 Ma have been determined for some Austin bentonites and have been recorded in unpublished theses (e.g., Collins, 1997). A δ13C curve based on plant remains spanning the Comanche Series through the base of the Austin Chalk has been published, but it is impre- cise (Jacobs et al., 2005). A more precise biostratigraphic and chemostratigraphic study of the Austin Chalk in Dallas County is provided by Gale et al. (2007) who propose the section as a global stratotype for the base of the Santonian Stage. Chalk-marl couplets in the lower Austin may record Milankovitch signals, possibly the 412 Ka eccentricity cycle and the 25 Ka precession cycle (Larson et al., 1997). The transition from the Comanche to the Gulf Series involves major reorganization of marine vertebrate communities, includ- ing the mid-Cretaceous extinction of ichthyosaurs, the decline of pliosaurs and the ascension of elasmosaur and polycotylid plesiosaurs, the fi rst radiation of marine snakes, and the evolu- tion of lizard-like marine dolichosaurs and mosasaurs, the latter of which rapidly diversify and become the apex predators of the Upper Cretaceous oceans. Marine rocks in north Texas preserve a nearly continuous record of these marine predators throughout Figure 1. Late Cretaceous stratigraphic section of north Texas. Hutchins formation (*) of the Austin Group has not been formally designated. Late Cretaceous strata and vertebrate fossils of North Texas 3 Figure 2. Geologic map of Upper Cre- taceous strata of north Texas (data from Stoeser et al., 2005). Figure 3. Cross section of East Texas Basin Mississippi Embayment after Adams and Carr (2010). This section represents strata to the south of the Dallas–Fort Worth area, across the San Marcos Swell. 4 Jacobs et al. the remainder of the Austin, Taylor and Navarro series, or essen- and Carr (2010) recognized two major units in the Woodbine, tially all of the Late Cretaceous. the Dexter and the Lewisville. Locally, Reaser (2002) recog- On occasion, terrestrial taxa are preserved in marine beds. nized the Rush Creek, Dexter, Lewisville, and Arlington forma- When these isolated occurrences are combined with terrestrial tions, and the Tarrant and Templeton facies within the Wood- fossils reported from marginal marine and terrestrial deposits bine. The Woodbine is considered to be middle Cenomanian in lower in the sequence, the Upper Cretaceous record of terres- age, or ~96 Ma. trial ecosystems in the Dallas–Fort Worth (DFW) area reveals At Stop 1, Woodbine foreshore deposits include cross- evidence of thriving terrestrial communities comprising a bedded and bioturbated sands. There is an extensive bioturbated variety of dinosaurs, pterosaurs, crocodiles, turtles, birds, and unit characterized by crab feeding balls and burrows (e.g., Allen mammals. Discoveries of new material in the 14 years since the and Curran, 1974). A channel is seen cutting across dune sands, last review of the north Texas fossil record (Jacobs and Win- with its margin continuing into the bioturbated unit. Rare frag- kler, 1998) have signifi cantly expanded our understanding of ments of fossil wood are found here. the taxonomic composition of Late Cretaceous terrestrial com- Hadrosaur (duckbill) dinosaur bones and several spectacular munities in the DFW area. trackways are also found in Murrell Park (Lee, 1997a, 1997b). Our objectives on this fi eld trip (Fig. 4) are to review the The holotype of the early hadrosaur Protohadros byrdi was found stratigraphic section from the Woodbine through the Austin to within a few kilometers of here (Head, 1998). Footprint ichno- become better informed as to the stratigraphy and to recognize taxa from Murrell Park have been attributed to theropod (Fusci- the truly exceptional sequence of marine amniote vertebrate fos- napedis woodbinensis) and hadrosaur (Caririchnium protohad- sils—the marine reptiles—found here. The Gulf Series as a whole rosaurichnos) dinosaurs and to a large bird (Magnoavipes lowei) documents the past 30 million years of life in Cretaceous seas. (Lee, 1997b). The major trackways are now covered (Fig. 6). No other section in North America can boast better coverage, and The only marine amniote described thus far from the Wood- taken with the underlying Comanche Series, it is unique in the bine is the pholidosaurid crocodile Terminonaris cf. T. robusta record it provides for the past 50 million years of the Cretaceous (Adams et al., 2011) from the uppermost part of the Woodbine period. Time and interest permitting, we will close this fi eld trip Formation. The ammonite Conlinoceras tarrantense is found by a visit to the fossil hall of the Perot Museum of Nature and within the Lewisville member and in the Tarrant member of the Science, which presents a magnifi cent portrayal of the fossil ver- overlying Eagle Ford.
Recommended publications
  • For Peer Review
    Biological Journal of the Linnean Society Marine tethysuchian c rocodyliform from the ?Aptian -Albian (Early Cretaceous) of the Isle of Wight, England Journal:For Biological Peer Journal of theReview Linnean Society Manuscript ID: BJLS-3227.R1 Manuscript Type: Research Article Date Submitted by the Author: 05-May-2014 Complete List of Authors: Young, Mark; University of Edinburgh, Biological Sciences; University of Southampton, School of Ocean and Earth Science Steel, Lorna; Natural History Museum, Earth Sciences Foffa, Davide; University of Bristol, Department of Earth Sciences Price, Trevor; Dinosaur Isle Museum, Naish, Darren; University of Southampton, School of Ocean and Earth Science Tennant, Jon; Imperial College London, Department of Earth Science and Engineering Albian, Aptian, Cretaceous, Dyrosauridae, England, Ferruginous Sands Keywords: Formation, Isle of Wight, Pholidosauridae, Tethysuchia, Upper Greensand Formation Biological Journal of the Linnean Society Page 1 of 50 Biological Journal of the Linnean Society 1 2 3 Marine tethysuchian crocodyliform from the ?Aptian-Albian (Early Cretaceous) 4 5 6 of the Isle of Wight, England 7 8 9 10 by MARK T. YOUNG 1,2 *, LORNA STEEL 3, DAVIDE FOFFA 4, TREVOR PRICE 5 11 12 2 6 13 DARREN NAISH and JONATHAN P. TENNANT 14 15 16 1 17 Institute of Evolutionary Biology, School of Biological Sciences, The King’s Buildings, University 18 For Peer Review 19 of Edinburgh, Edinburgh, EH9 3JW, United Kingdom 20 21 2 School of Ocean and Earth Science, National Oceanography Centre, University of Southampton,
    [Show full text]
  • SPRING 1963 Bulletin 4 BURR A. SILVER
    SPRING 1963 Bulletin 4 The Bluehonnet Lake Waco Formation {Upper Central Lagoonal Deposit BURR A. SILVER thinking is more important than elaborate equipment-" Frank Carney, Ph.D. Professor of Geology Baylor University 1929-1934 Objectives of Geological at Baylor The of a geologist in a university covers but a few years; his education continues throughout his active life. The purposes of training geologists at Baylor University are to provide a sound basis of un­ derstanding and to foster a truly geological point of view, both of which are essential for continued professional growth. The staff considers geology to be unique among sciences since it is primarily a field science. All geologic research including that done in laboratories must be firmly supported by field observations. The student is encouraged to develop an inquiring objective attitude and to examine critically all geological concepts and principles. The development of a mature and professional attitude toward geology and geological research is a principal concern of the department. PRESS WACO, TEXAS BAYLOR GEOLOGICAL STUDIES BULLETIN NO. 4 The Member, Lake Waco Formation {Upper Central Texas - - A Lagoonal Deposit BURR A. SILVER BAYLOR UNIVERSITY Department of Geology Waco, Texas Spring, 1963 Baylor Geological Studies EDITORIAL STAFF F. Brown, Jr., Ph.D., Editor stratigraphy, paleontology O. T. Hayward, Ph.D., Adviser stratigraphy-sedimentation, structure, geophysics-petroleum, groundwater R. M. A., Business Manager archeology, geomorphology, vertebrate paleontology James W. Dixon, Jr., Ph.D. stratigraphy, paleontology, structure Walter T. Huang, Ph.D. mineralogy, petrology, metallic minerals Jean M. Spencer, B.S., Associate Editor Bulletin No. 4 The Baylor Geological Studies Bulletin is published Spring and Fall, by the Department of Geology at Baylor University.
    [Show full text]
  • Evaluation of the Depositional Environment of the Eagle Ford
    Louisiana State University LSU Digital Commons LSU Master's Theses Graduate School 2012 Evaluation of the depositional environment of the Eagle Ford Formation using well log, seismic, and core data in the Hawkville Trough, LaSalle and McMullen counties, south Texas Zachary Paul Hendershott Louisiana State University and Agricultural and Mechanical College, [email protected] Follow this and additional works at: https://digitalcommons.lsu.edu/gradschool_theses Part of the Earth Sciences Commons Recommended Citation Hendershott, Zachary Paul, "Evaluation of the depositional environment of the Eagle Ford Formation using well log, seismic, and core data in the Hawkville Trough, LaSalle and McMullen counties, south Texas" (2012). LSU Master's Theses. 863. https://digitalcommons.lsu.edu/gradschool_theses/863 This Thesis is brought to you for free and open access by the Graduate School at LSU Digital Commons. It has been accepted for inclusion in LSU Master's Theses by an authorized graduate school editor of LSU Digital Commons. For more information, please contact [email protected]. EVALUATION OF THE DEPOSITIONAL ENVIRONMENT OF THE EAGLE FORD FORMATION USING WELL LOG, SEISMIC, AND CORE DATA IN THE HAWKVILLE TROUGH, LASALLE AND MCMULLEN COUNTIES, SOUTH TEXAS A Thesis Submitted to the Graduate Faculty of the Louisiana State University Agricultural and Mechanical College in partial fulfillment of the requirements for degree of Master of Science in The Department of Geology and Geophysics by Zachary Paul Hendershott B.S., University of the South – Sewanee, 2009 December 2012 ACKNOWLEDGEMENTS I would like to thank my committee chair and advisor, Dr. Jeffrey Nunn, for his constant guidance and support during my academic career at LSU.
    [Show full text]
  • Estimating the Evolutionary Rates in Mosasauroids and Plesiosaurs: Discussion of Niche Occupation in Late Cretaceous Seas
    Estimating the evolutionary rates in mosasauroids and plesiosaurs: discussion of niche occupation in Late Cretaceous seas Daniel Madzia1 and Andrea Cau2 1 Department of Evolutionary Paleobiology, Institute of Paleobiology, Polish Academy of Sciences, Warsaw, Poland 2 Independent, Parma, Italy ABSTRACT Observations of temporal overlap of niche occupation among Late Cretaceous marine amniotes suggest that the rise and diversification of mosasauroid squamates might have been influenced by competition with or disappearance of some plesiosaur taxa. We discuss that hypothesis through comparisons of the rates of morphological evolution of mosasauroids throughout their evolutionary history with those inferred for contemporary plesiosaur clades. We used expanded versions of two species- level phylogenetic datasets of both these groups, updated them with stratigraphic information, and analyzed using the Bayesian inference to estimate the rates of divergence for each clade. The oscillations in evolutionary rates of the mosasauroid and plesiosaur lineages that overlapped in time and space were then used as a baseline for discussion and comparisons of traits that can affect the shape of the niche structures of aquatic amniotes, such as tooth morphologies, body size, swimming abilities, metabolism, and reproduction. Only two groups of plesiosaurs are considered to be possible niche competitors of mosasauroids: the brachauchenine pliosaurids and the polycotylid leptocleidians. However, direct evidence for interactions between mosasauroids and plesiosaurs is scarce and limited only to large mosasauroids as the Submitted 31 July 2019 predators/scavengers and polycotylids as their prey. The first mosasauroids differed Accepted 18 March 2020 from contemporary plesiosaurs in certain aspects of all discussed traits and no evidence Published 13 April 2020 suggests that early representatives of Mosasauroidea diversified after competitions with Corresponding author plesiosaurs.
    [Show full text]
  • Mollusks from the Pepper Shale Member of the Woodbine Formation Mclennan County, Texas
    Mollusks From the Pepper Shale Member of the Woodbine Formation McLennan County, Texas GEOLOGICAL SURVEY PROFESSIONAL PAPER 243-E Mollusks From the Pepper Shale Member of the Woodbine Formation McLennan County, Texas By LLOYD WILLIAM STEPHENSON SHORTER CONTRIBUTIONS TO GENERAL GEOLOGY, 1952, PAGES 57-68 GEOLOGICAL SURVEY PROFESSIONAL PAPER 243-E Descriptions and illustrations of new species offossils of Cenomanian age UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1953 UNITED STATES DEPARTMENT OF THE INTERIOR Douglas McKay, Secretary GEOLOGICAL SURVEY W. E. Wrather, Director For sale by the Superintendent of Documents, U. S. Government Printing Office Washington 25, D. C. - Price 25 cents (paper cover) CONTENTS Page Abstract________________________________________________________________ 57 Historical sketch__-___-_-__--__--_l________________-__-_-__---__--_-______ 57 Type section of the Pepper shale member.____________________________________ 58 Section of Pepper shale at Haunted Hill._______________-____-__-___---_------- 58 Systematic descriptions._______.._______________-_-__-_-_-____---__-_-_-_----_ 59 Pelecypoda_ ________________________________________________________ 59 Gastropoda.______-____-_-_____________--_____-___--__-___--__-______ 64 Cephalopoda-_______-_______________________-___--__---_---_-__-— 65 References......___________________________________________________________ 65 Index.__________________________________________________ 67 ILLUSTRATIONS Plate 13. Molluscan fossils, mainly from the Pepper shale__________
    [Show full text]
  • DFW Industrial Marketview
    MARKETVIEW Dallas/Fort Worth Industrial, Q4 2018 Demand tops 20 million sq. ft. for third consecutive year. 6.0% 14,513,927 SF 8,273,325 SF 7,108,176 SF Figure 1: DFW Industrial Market *Arrows indicate trend from previous quarter. MSF Vacancy Rate (%) 30 7 25 20 15 6 10 5 0 5 2013 2014 2015 2016 2017 2018 Total Net Absorption Delivered Construction Construction Starts Total Vacancy Source: CBRE Research, Q4 2018. SUPPLY AND DEMAND REMAIN BALANCED PRODUCT PIPELINE CONTRACTS SUBSTANTIALLY MARKET-WIDE VACANCY RATE REMAINS SNUG Q4 2018 CBRE Research © 2019 CBRE, Inc. | 1 MARKETVIEW DALLAS/FORT WORTH INDUSTRIAL Figure 2: DFW Industrial Market Snapshot Net Direct Total Q4 2018 Q4 2018 2018 Avail. Direct Total Rentable Available Vacancy Vacancy Direct Net Total Net Total Market Rate Vacancy Vacancy Area (SF) Rate Rate Absorption Absorption Absorption (%) (SF) (SF) (SF) (%) (%) (SF) (SF) (SF) DFW Airport 76,537,446 7,143,995 9.3 4,981,889 6.5 5,496,992 7.2 1,021,613 1,093,964 2,816,521 East Dallas 36,711,061 3,247,617 8.8 1,640,650 4.5 1,674,582 4.6 455,405 421,473 323,735 Northeast Dallas 98,857,581 7,721,313 7.8 4,920,189 5.0 5,634,661 5.7 749,356 666,538 1,314,114 Northwest Dallas 110,278,941 6,873,465 6.2 4,158,527 3.8 4,433,034 4.0 360,646 333,645 1,007,725 South Dallas 77,278,616 13,465,000 17.4 12,086,981 15.6 12,226,952 15.8 1,206,291 1,254,279 5,847,155 South Stemmons 116,757,096 7,800,940 6.7 4,460,334 3.8 4,759,216 4.1 (375,470) (376,688) (1,290,388) Great SW/Arlington 111,266,798 8,018,274 7.2 5,043,620 4.5 5,140,768 4.6 2,566,998 2,601,478 7,293,671 North Ft.
    [Show full text]
  • Oral History of Bonton and Ideal Neighborhoods in Dallas, Texas
    ORAL HISTORY OF BONTON AND IDEAL NEIGHBORHOODS IN DALLAS, TEXAS Briana Payne Thesis Prepared for the Degree of MASTER OF ARTS UNIVERSITY OF NORTH TEXAS December 2015 APPROVED: Andrew Nelson, Committee Chair Beverly Ann Davenport, Committee Member Matthew Durington, Committee Member Lisa Henry, Chair of the Department of Anthropology Costas Tsatsoulis, Dean of the Toulouse Graduate School Payne, Briana. Oral History of Bonton and Ideal Neighborhoods in Dallas, Texas. Master of Arts (Applied Anthropology), December 2015, 145 pp., 3 tables, 67 figures, references, 223 titles. The Bonton and Ideal neighborhoods in Dallas Texas, developed in the early 1900s, experienced physical and social decay throughout the 1980s. Neighborhood organizations and resident activism were vital to the rebirth of the community in the 1990s. Current revitalization efforts taking place there have been a source of contention as the neighborhood continues to overcome inequalities created by decades of racialized city planning initiatives. This thesis focuses on how the structuring structure of whiteness has historically affected, and continues to affect, the neighborhoods of Ideal and Bonton, as well as acts to identify how black residents have navigated their landscape and increased their collective capital through neighborhood activism. Copyright 2015 By Briana Payne ii ACKNOWLEDGEMENTS First, I would like to thank the Neighborhood Investment Program for providing me with the opportunity to work with them as they continue to improve the Bonton and Ideal neighborhoods in South Dallas. Without their acknowledgement of the importance of conserving the history of this community, I would not have been able to share the important life stories of the residents.
    [Show full text]
  • An Early Bothremydid from the Arlington Archosaur Site of Texas Brent Adrian1*, Heather F
    www.nature.com/scientificreports OPEN An early bothremydid from the Arlington Archosaur Site of Texas Brent Adrian1*, Heather F. Smith1, Christopher R. Noto2 & Aryeh Grossman1 Four turtle taxa are previously documented from the Cenomanian Arlington Archosaur Site (AAS) of the Lewisville Formation (Woodbine Group) in Texas. Herein, we describe a new side-necked turtle (Pleurodira), Pleurochayah appalachius gen. et sp. nov., which is a basal member of the Bothremydidae. Pleurochayah appalachius gen. et sp. nov. shares synapomorphic characters with other bothremydids, including shared traits with Kurmademydini and Cearachelyini, but has a unique combination of skull and shell traits. The new taxon is signifcant because it is the oldest crown pleurodiran turtle from North America and Laurasia, predating bothremynines Algorachelus peregrinus and Paiutemys tibert from Europe and North America respectively. This discovery also documents the oldest evidence of dispersal of crown Pleurodira from Gondwana to Laurasia. Pleurochayah appalachius gen. et sp. nov. is compared to previously described fossil pleurodires, placed in a modifed phylogenetic analysis of pelomedusoid turtles, and discussed in the context of pleurodiran distribution in the mid-Cretaceous. Its unique combination of characters demonstrates marine adaptation and dispersal capability among basal bothremydids. Pleurodira, colloquially known as “side-necked” turtles, form one of two major clades of turtles known from the Early Cretaceous to present 1,2. Pleurodires are Gondwanan in origin, with the oldest unambiguous crown pleurodire dated to the Barremian in the Early Cretaceous2. Pleurodiran fossils typically come from relatively warm regions, and have a more limited distribution than Cryptodira (hidden-neck turtles)3–6. Living pleurodires are restricted to tropical regions once belonging to Gondwana 7,8.
    [Show full text]
  • Pterosaur Distribution in Time and Space: an Atlas 61
    Zitteliana An International Journal of Palaeontology and Geobiology Series B/Reihe B Abhandlungen der Bayerischen Staatssammlung für Pa lä on to lo gie und Geologie B28 DAVID W. E. HONE & ERIC BUFFETAUT (Eds) Flugsaurier: pterosaur papers in honour of Peter Wellnhofer CONTENTS/INHALT Dedication 3 PETER WELLNHOFER A short history of pterosaur research 7 KEVIN PADIAN Were pterosaur ancestors bipedal or quadrupedal?: Morphometric, functional, and phylogenetic considerations 21 DAVID W. E. HONE & MICHAEL J. BENTON Contrasting supertree and total-evidence methods: the origin of the pterosaurs 35 PAUL M. BARRETT, RICHARD J. BUTLER, NICHOLAS P. EDWARDS & ANDREW R. MILNER Pterosaur distribution in time and space: an atlas 61 LORNA STEEL The palaeohistology of pterosaur bone: an overview 109 S. CHRISTOPHER BENNETT Morphological evolution of the wing of pterosaurs: myology and function 127 MARK P. WITTON A new approach to determining pterosaur body mass and its implications for pterosaur fl ight 143 MICHAEL B. HABIB Comparative evidence for quadrupedal launch in pterosaurs 159 ROSS A. ELGIN, CARLOS A. GRAU, COLIN PALMER, DAVID W. E. HONE, DOUGLAS GREENWELL & MICHAEL J. BENTON Aerodynamic characters of the cranial crest in Pteranodon 167 DAVID M. MARTILL & MARK P. WITTON Catastrophic failure in a pterosaur skull from the Cretaceous Santana Formation of Brazil 175 MARTIN LOCKLEY, JERALD D. HARRIS & LAURA MITCHELL A global overview of pterosaur ichnology: tracksite distribution in space and time 185 DAVID M. UNWIN & D. CHARLES DEEMING Pterosaur eggshell structure and its implications for pterosaur reproductive biology 199 DAVID M. MARTILL, MARK P. WITTON & ANDREW GALE Possible azhdarchoid pterosaur remains from the Coniacian (Late Cretaceous) of England 209 TAISSA RODRIGUES & ALEXANDER W.
    [Show full text]
  • (Squamata: Mosasauridae) from the Late Cretaceous Of
    C. R. Palevol 14 (2015) 483–493 Contents lists available at ScienceDirect Comptes Rendus Palevol www.sci encedirect.com General Palaeontology, Systematics and Evolution (Vertebrate Palaeontology) An halisaurine (Squamata: Mosasauridae) from the Late Cretaceous of Patagonia, with a preserved tympanic disc: Insights into the mosasaur middle ear Un halisauriné (Squamata : Mosasauridae) du Crétacé supérieur de Patagonie, à disque tympanique conservé : un aperc¸ u de l’oreille moyenne des mosasaures a,∗ b Marta S. Fernández , Marianella Talevi a CONICET - División Paleontología Vertebrados, Museo de La Plata, Paseo del Bosque s/n, 1900 La Plata, Argentina b CONICET - Instituto de Investigación en Paleobiología y Geología, Universidad Nacional de Río Negro, Isidro Lobo y Belgrano, 8332 General Roca, Río Negro, Argentina a b s t r a c t a r t i c l e i n f o Article history: Halisaurinae is a subfamily of enigmatic, small- to medium-sized mosasauroids, which Received 15 September 2014 retain a mosaic of primitive and derived features. The first record of a South American Hal- Accepted after revision 13 May 2015 isaurus with precise stratigraphic information includes a quadrate carrying a tympanic disc together with twelve vertebrae, collected in the Late Maastrichtian of Jagüel Formation Handled by Nathalie Bardet in northern Patagonia (Argentina). The preservation of a tympanic disc allows exploring and discussing the mechanisms of sound transmission in these mosasauroids. The loca- Keywords: tion of the tympanic disc resembles that one formed by the extracolumella of aquatic Halisaurus turtles and at least one extant lizard. Based on morphological comparison of the middle Patagonia ear we discuss previous hypotheses on the modification of the tympanic middle ear system Late Maastrichtian of mosasauroids for underwater hearing, in a manner similar to that observed in aquatic Cretaceous turtles.
    [Show full text]
  • Signature Redacted Signature of Author
    Multifamily Site Development - Bishop Arts District, Dallas, TX by Justin L Rice B.A., Finance and Real Estate, 2005 Texas Tech University Submitted to the Program in Real Estate Development in Conjunction with the Center for Real Estate in Partial Fulfillment of the Requirements for the Degree of Master of Science in Real Estate Development at the Massachusetts Institute of Technology September, 2017 @2017 Justin L Rice All rights reserved The author hereby grants to MIT permission to reproduce and to distribute publicly paper and electronic copies of this thesis document in whole !op in any medium now known or hereafter created. Signature redacted Signature of Author Cente- Rpl Estate ( July 28, 2017 Certified by Signature redacted ;400 John Kennedy r Lecturer, MIT Center for Real Estate Thesis Supervisor Accepted by Signature redacted Albert Saiz Director Center for Real Estate MASAHSET INTTUTE OF TECHNOLOGY SEP 13 2017 LIBRARIES ARCIGNE. Multifamily Site Development - Bishop Arts District, Dallas, TX by Justin Rice Submitted to the Program in Real Estate Development in Conjunction with the Center for Real Estate on July 28, 2017 in Partial Fulfillment of the Requirements for the Degree of Master of Science in Real Estate Development ABSTRACT This thesis explores the development potential and land value of a vacant parcel of land listed for sale in Dallas, TX. Further, this thesis proposes three different multifamily rental site plan designs for the parcel in an effort to maximize the value of both the development and the land. The site is located just two blocks from Dallas' popular Bishop Arts retail district.
    [Show full text]
  • Craniofacial Morphology of Simosuchus Clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar
    Society of Vertebrate Paleontology Memoir 10 Journal of Vertebrate Paleontology Volume 30, Supplement to Number 6: 13–98, November 2010 © 2010 by the Society of Vertebrate Paleontology CRANIOFACIAL MORPHOLOGY OF SIMOSUCHUS CLARKI (CROCODYLIFORMES: NOTOSUCHIA) FROM THE LATE CRETACEOUS OF MADAGASCAR NATHAN J. KLEY,*,1 JOSEPH J. W. SERTICH,1 ALAN H. TURNER,1 DAVID W. KRAUSE,1 PATRICK M. O’CONNOR,2 and JUSTIN A. GEORGI3 1Department of Anatomical Sciences, Stony Brook University, Stony Brook, New York, 11794-8081, U.S.A., [email protected]; [email protected]; [email protected]; [email protected]; 2Department of Biomedical Sciences, Ohio University College of Osteopathic Medicine, Athens, Ohio 45701, U.S.A., [email protected]; 3Department of Anatomy, Arizona College of Osteopathic Medicine, Midwestern University, Glendale, Arizona 85308, U.S.A., [email protected] ABSTRACT—Simosuchus clarki is a small, pug-nosed notosuchian crocodyliform from the Late Cretaceous of Madagascar. Originally described on the basis of a single specimen including a remarkably complete and well-preserved skull and lower jaw, S. clarki is now known from five additional specimens that preserve portions of the craniofacial skeleton. Collectively, these six specimens represent all elements of the head skeleton except the stapedes, thus making the craniofacial skeleton of S. clarki one of the best and most completely preserved among all known basal mesoeucrocodylians. In this report, we provide a detailed description of the entire head skeleton of S. clarki, including a portion of the hyobranchial apparatus. The two most complete and well-preserved specimens differ substantially in several size and shape variables (e.g., projections, angulations, and areas of ornamentation), suggestive of sexual dimorphism.
    [Show full text]