Key to the Females of the New World Genera + Ctenocerinae (Epipompilus) (modified from Banks 1947, Evans 1966, Coloma de Correa 1981, & Wahis & Rojas 2003)

(The sexes have separate keys for these subfamilies. The male key is presented elsewhere.)

a bb b aa

b bb

c c cc

1. a. Pronotum with the collar not well differentiated from the disc, the streptaulus absent medially and the collar on nearly the same plane as the disc; or if on a lower plane the streptaulus is absent altogether. b. Pronotum longer than mesoscutum in most genera. c. Eyes sometimes wholly covered with short setae (Aporini)...... 2

aa. Pronotum with the collar separated from the disc by a complete streptaulus, the disc sloping upward strongly from the collar. bb. Pronotum shorter than mesoscutum along the midline, or at least not notably longer. cc. Eyes never setose ()...... 8

a aa

b bb

c c cc cc

2. a. Fore wing with three SMCs. b. Front femora not swollen (incrassate). c. or if somewhat so the labrum is strongly exerted and the maxillary palpi very long...... 3

aa. Fore wing with two SMCs. bb. Front femora usually swollen (incrassate). cc. Labrum not strongly exerted nor the palpi unusually long...... 4

aa a b a

bb

c cc

3. a. Eyes setose. b. Pronotum relatively short, the streptaulus absent. c. Mandibles without a fimbriate groove beneath...... Epipompilus Kohl

aa. Eyes not setose. bb. Pronotum elongate, the streptaulus present except medially. cc. Mandibles with a strong fimbriate groove beneath...... Banks

a

aa

4. a. SMC2 receiving only one recurrent vein (1m-cu), the second recurrent vein (2m- cu) meeting M beyond (or occasionally interstitial with) with second intercubital vein (2r-m)...... 5

aa. SMC2 receiving two recurrent veins (both 1m-cu and 2m-cu)...... 6

aa a

b bb

5. a. Claws bifid. b. Hind wing with the transverse median vein (cu-a) leaving the anal vein at an angle, meeting media basad of origin of cubitus (Cu) by about or slightly more or less than its own length...... Notoplaniceps Bradley

aa. Claws dentate. bb. Hind wing variable, rarely as above in American species...... Spinola

aa aa

bb bb bb bb

6. a. Front tibiae with stout, decurved spines at apex. b. Middle and hind coxae well separated, the metasternum visible.….Chelaporus Bradley

aa. Front tibiae without such spines at apex. bb. Middle and hind coxae close together, the metasternum not readily visible...... 7

aa

a

b bb

7. a. Transverse median vein (cu-a) of hind wing leaving anal vein at an angle, short and straight. b. SMC2 slightly if at all wider than high...... Allaporus Banks

aa. Transverse median vein (cu-a) of hind wing not leaving anal vein at an angle, in fact not clearly separated from it, long and strongly arched. bb. SMC2 nearly or quite twice as wide as high……………………... Euplaniceps Haupt

a

aa

b bb bb b

c c cc cc

cc c c cc

8. a. Propodeum not produced posterolaterally into sharp, conical processes (such processes present on the middle of the slope in some ). b. Legs more or less spinose. c. Postnotum a transverse band of variable width...... 9

aa. Propodeum bearing distinct, more or less conical processes posterolaterally. bb. Either the legs smooth and almost devoid of spines: cc. or the postnotum concealed dorsally...... 28

aa a

bb b

9. a. Transverse median vein (cu-a) of hind wing leaving anal vein at an angle, short and straight. b. Posterior rim of propodeum virtually absent...... Tastiotenia Evans

aa. Anal (A) and transverse median veins (cu-a) of hind wing confluent and forming an arch. bb. Posterior rim of propodeum well developed...... 10

a

aa

bb

b b

10. a. Anal lobe of hind wing very large, about three-fourths the length of the submedian cell. b. Spines beneath apical tarsal segments in irregular double row...... Chalcochares Banks

aa. Anal lobe of hind wing smaller, at most slightly more than half the length of the submedian cell. bb. Spines beneath the apical tarsal segments, when present, in a median row (occasionally also with one or two lateral ones)...... 11

a a a aa aa aa a a a aa aa aa

b bb b bb

11. a. Postnotum arcuately broadened on each side of the median line, then constricted again opposite the propodeal spiracles. b. Transverse median vein (cu-a) of hind wing meeting media beyond the origin of cubitus (Cu): c. Except some-times in , which possesses a scale-like pubescence on Tl: d. and Austrochares, in which foretibiae are spined dorsally most of their length ………………………………………………………………………………………….12

aa. Postnotum a transverse band with nearly parallel margins, or broadened at the midline. bb. Hind wing variable. cc. Without scale-like pubescence. dd. Fore tibiae spined dorsally only apically...... 15

aa a

b bb

12. a. Claws bifid. b. Transverse medial vein (cu-a) of hind wing meeting media near or slightly beyond origin of cubitus (Cu)………………………………………………………………………...... 13

aa. Claws dentate (except front claws sometimes bifid). bb. Transverse medial vein (cu-a) of hind wing meeting media well beyond origin of cubitus (Cu)…………………………………………………………………………….....…...... 14

aa a

a a a

b bb bb b

13. a. Tl, and usually parts of the propodeum and thorax, bearing scale-like pubescence. b. Clypeus transverse, its upper margin slightly irregular toward the sides…………...... … .………………………………………………………………………… Episyron Schiødte

aa. Body without scale-like pubescence. bb. Clypeus large, its upper margin sinuate...... 13a

a

aa

b

bb c cc

bb b

cc c

13a. a. Tarsal comb absent. b. Malar space well developed. c. Labrum fully exerted. d. Apex of metasoma compressed….……...... Gribodo

a. Tarsal comb present. b. Malar space not well developed. c. Labrum only partially exerted, if at all. d. Apex of metasoma not compressed...... Austrochares Banks

a

aa

bb b

b

bb

b b

14. a. Apical tarsal segments not spined beneath. b. Spines of the front tibiae mostly on the outer side………...... Sericopompilus Howard

aa. Apical tarsal segments spined beneath. bb. Front tibiae with a series of strong spines above…………... Howard

a

aa

b

bb

15. a. Second recurrent vein (2m-cu) of fore wing arising on the subdiscoidal vein much more than half the distance from the base of the subdiscoidal vein to the outer wing margin. b. Apical tergite densely bristly only in a few species of Priochilus……...... 16

aa. Second recurrent vein (2m-cu) of fore wing arising on the subdiscoidal vein about half or somewhat less than half the distance from the base of the subdiscoidal vein to the outer wing margin, or if somewhat more than half (some ): bb. Apical tergite densely bristly………………………………………...... 20

a aa

b bb

c cc

16. a. Pulvillar pad large, the comb consisting of 18 or more short parallel setulae. b. Apical lateral spines on the penultimate (4th) tarsal segments more than half as long as the ultimate (5th) segment. c. Clypeus with a strong, sharply defined median emargination...... Aplochares Banks

aa. Pulvillar pad very small, the comb consisting of a few divergent setulae. bb. Apical lateral spines of penultimate (4th) tarsal segments much shorter than above. cc. American species with the clypeus more shallowly and broadly emarginate if at all…. ………………………………………...... 17

a aa

bb

b

17. a. Claws dentate. b. Third discoidal cell with a pocket at its lower, inner corner...... 18

aa. Claws bifid, the inner ray broader than the outer ray. bb. Third discoidal cell without a pocket at its lower, inner corner...... 19

a aa

b bb

18. a. Front without a tubercle as below. b. Propodeum with the slope rather low and even, often striate or with short, pale, semi- erect pubescence. c. Integument of various colors ...... Ashmead

aa. Front with a blunt tubercle between and slightly above the antennal sockets. bb. Propodeum strongly convex, more or less humped laterally, not striate or with semi- erect pubescence. cc. Integument normally dark orange ….…………………….... Tachypompilus Ashmead

aa b

a

bb

19. a. Pronotum short, sloping evenly in front. b. Antennae very slender throughout, the outer flagellar segments elongate; cc. Metasoma not compressed...... Priochilus Banks

aa. Pronotum with the disc subangularly produced anteromedially, the anterior face nearly or quite perpendicular to the disc. bb. Outer segments of flagellum rather short and thick in most species. cc. Metasoma weakly to fairly strongly compressed apically...... Balboana Banks

aa a

b

bb

20. a. Apical tergite (pygidium, T6) with numerous stiff, bristly setae, usually quite densely bristly. b. Pulvillar comb strong, of from 12 to 26 subparallel setae...... Anoplius Dufour

aa. Apical tergite (pygidium, T6) without bristles, often with some strong setae, but these are sparse and not bristly. bb. Pulvillar comb variable…...... 21

aa a

21. a. Claws bifid……...………………………………………………………………….… 22

aa. Claws dentate...... …………………………………………………………………... 23

a

aa

b bb

aa a

bb b

22. a. Malar space well developed. b. Labrum fully exerted. c. Apex of metasoma compressed…..……...... Paracyphononyx Gribodo

aa. Malar space not developed. bb. Labrum not usually exerted. cc. Apex of metasoma not compressed…………………..………….. Dicranoplius Haupt

a aa

b bb b b

23. a. Antennae very short, the middle flagellar segments tending to be slightly broadened and flattened on one side, third segment at most 3.5X as long as thick, usually less than 3X as long as thick. b. Pulvillar comb of not more than seven weak setulae………..…….…………………. 24

aa. Antennae long and slender, third segment generally more than 3.5 X as long as thick. bb. Pulvillar comb variable, when reduced the antennae quite long and slender...... 25

aa a

24. a. Pronotum in lateral view shorter than the mesonotum and surface almost complete oblique without dorsal face…….………………………………...... …………… Aridestus Banks

aa. Pronotum in lateral view as long as the mesonotum and having a distinct dorsal face... ………………………………………………………………………... Lepeletier

a aa

aa

25. a. Apical tarsal segment (5th) without a median row of ventral spines similar in size to preceeding segment; either no ventral spines or with no more than 3 spines near base or with a complete row of very minute spines.…………….……………………………………………... 26

aa. Apical tarsal segment (5th) with a median row of ventral spines similar in size to preceeding segment, which form a row that at least extends past middle of segment…………..… ……………………………………………….…………………….. Kincaid

a a aa

26. a. Front basitarsus (1st) with two comb-spines, one at middle and one at apex…………..... …………………………………………………………….……… Perissopompilus Evans

aa. Front basitarsus (1st) with three, sometimes four, comb-spines….………………… 27

a aa

bb b

bb b

cc c

c c

27. a. Apical tarsal segment (5th) without ventral spines. b. Propodeum and mesopleura bare or with dark setae. c. Pulvillar comb weakly developed…….…………………..….. Hesperopompilus Evans

aa. Apical tarsal segment (5th) with 1-3 ventral spines near base. bb. Propodeum and mesopleura, as well as other portions of body, with white setae. cc. Pulvillar comb strongly developed with 20 setulae….…..………… Xerochares Evans

a aa

bb bb

b b bb bb

b b

cc c

dd

28. a. Front tarsus with a comb and legs otherwise rather spinose. b. Postnotum concealed dorsally, the metanotum and propodeum in broad contact. c. Frequently with only two SMCs dd. Malar space variable…………...... ………………………… Banks

aa. Front tarsus without a comb, legs unusually smooth and devoid of spines. bb. Postnotum complete. cc. Three SMCs present. dd. Malar space fairly long...... …………..……………….. Allochares Banks