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Different Speciation Types Meet in a Mediterranean Genus: The
This is an Accepted Manuscript of an article published in Taxon on 4 May 2017, available online: https://doi.org/10.12705/662.7 Different speciation types meet in a Mediterranean genus: the biogeographic history of Cymbalaria (Plantaginaceae). Running head: Phylogeny and biogeographic history of Cymbalaria Pau Carnicero1, Llorenç Sáez1, 2, Núria Garcia-Jacas3 and Mercè Galbany- Casals1 1 Departament de Biologia Animal, Biologia Vegetal i Ecologia, Facultat de Biociències, Universitat Autònoma de Barcelona, 08193 Bellaterra, Spain. 2 Societat d’Història Natural de les Balears (SHNB), C/ Margarida Xirgu 16, E-07011 Palma de Mallorca, Balearic Islands, Spain. 3 Institut Botànic de Barcelona (IBB-CSIC-ICUB), Pg. del Migdia s/n, ES-08038 Barcelona, Spain Author for correspondence: Pau Carnicero, [email protected] ORCID: P.C., http://orcid.org/0000-0002-8345-3309 Abstract Cymbalaria comprises ten species and six subspecies growing in rocky habitats in the Mediterranean Basin. Several features, such as the genus’ highly fragmented distribution as well as noticeable ecological differentiation between partially sympatric species and presence of ploidy barriers between species suggest the involvement of different speciation types in its evolution. The aims of this study were to test the monophyly of Cymbalaria and to reconstruct infrageneric phylogenetic relationships, to infer the genus’ biogeographic history by estimating divergence times and ancestral distribution areas of lineages, and to disentangle the role of different speciation types. To address these issues, we constructed a phylogeny with a complete taxon sampling based on ITS, 3'ETS, ndhF and rpl32-trnL sequences. We used the nuclear ribosomal DNA data to produce a time-calibrated phylogeny, which served as basis for estimating ploidy level evolution and biogeographic history. -
Vascular Plants and a Brief History of the Kiowa and Rita Blanca National Grasslands
United States Department of Agriculture Vascular Plants and a Brief Forest Service Rocky Mountain History of the Kiowa and Rita Research Station General Technical Report Blanca National Grasslands RMRS-GTR-233 December 2009 Donald L. Hazlett, Michael H. Schiebout, and Paulette L. Ford Hazlett, Donald L.; Schiebout, Michael H.; and Ford, Paulette L. 2009. Vascular plants and a brief history of the Kiowa and Rita Blanca National Grasslands. Gen. Tech. Rep. RMRS- GTR-233. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 44 p. Abstract Administered by the USDA Forest Service, the Kiowa and Rita Blanca National Grasslands occupy 230,000 acres of public land extending from northeastern New Mexico into the panhandles of Oklahoma and Texas. A mosaic of topographic features including canyons, plateaus, rolling grasslands and outcrops supports a diverse flora. Eight hundred twenty six (826) species of vascular plant species representing 81 plant families are known to occur on or near these public lands. This report includes a history of the area; ethnobotanical information; an introductory overview of the area including its climate, geology, vegetation, habitats, fauna, and ecological history; and a plant survey and information about the rare, poisonous, and exotic species from the area. A vascular plant checklist of 816 vascular plant taxa in the appendix includes scientific and common names, habitat types, and general distribution data for each species. This list is based on extensive plant collections and available herbarium collections. Authors Donald L. Hazlett is an ethnobotanist, Director of New World Plants and People consulting, and a research associate at the Denver Botanic Gardens, Denver, CO. -
Alien Flora of Europe: Species Diversity, Temporal Trends, Geographical Patterns and Research Needs
Preslia 80: 101–149, 2008 101 Alien flora of Europe: species diversity, temporal trends, geographical patterns and research needs Zavlečená flóra Evropy: druhová diverzita, časové trendy, zákonitosti geografického rozšíření a oblasti budoucího výzkumu Philip W. L a m b d o n1,2#, Petr P y š e k3,4*, Corina B a s n o u5, Martin H e j d a3,4, Margari- taArianoutsou6, Franz E s s l7, Vojtěch J a r o š í k4,3, Jan P e r g l3, Marten W i n t e r8, Paulina A n a s t a s i u9, Pavlos A n d r i opoulos6, Ioannis B a z o s6, Giuseppe Brundu10, Laura C e l e s t i - G r a p o w11, Philippe C h a s s o t12, Pinelopi D e l i p e t - rou13, Melanie J o s e f s s o n14, Salit K a r k15, Stefan K l o t z8, Yannis K o k k o r i s6, Ingolf K ü h n8, Hélia M a r c h a n t e16, Irena P e r g l o v á3, Joan P i n o5, Montserrat Vilà17, Andreas Z i k o s6, David R o y1 & Philip E. H u l m e18 1Centre for Ecology and Hydrology, Hill of Brathens, Banchory, Aberdeenshire AB31 4BW, Scotland, e-mail; [email protected], [email protected]; 2Kew Herbarium, Royal Botanic Gardens Kew, Richmond, Surrey, TW9 3AB, United Kingdom; 3Institute of Bot- any, Academy of Sciences of the Czech Republic, CZ-252 43 Průhonice, Czech Republic, e-mail: [email protected], [email protected], [email protected], [email protected]; 4Department of Ecology, Faculty of Science, Charles University, Viničná 7, CZ-128 01 Praha 2, Czech Republic; e-mail: [email protected]; 5Center for Ecological Research and Forestry Applications, Universitat Autònoma de Barcelona, 08193 Bellaterra, Spain, e-mail: [email protected], [email protected]; 6University of Athens, Faculty of Biology, Department of Ecology & Systematics, 15784 Athens, Greece, e-mail: [email protected], [email protected], [email protected], [email protected], [email protected]; 7Federal Environment Agency, Department of Nature Conservation, Spittelauer Lände 5, 1090 Vienna, Austria, e-mail: [email protected]; 8Helmholtz Centre for Environmental Research – UFZ, Department of Community Ecology, Theodor-Lieser- Str. -
Flora Mediterranea 26
FLORA MEDITERRANEA 26 Published under the auspices of OPTIMA by the Herbarium Mediterraneum Panormitanum Palermo – 2016 FLORA MEDITERRANEA Edited on behalf of the International Foundation pro Herbario Mediterraneo by Francesco M. Raimondo, Werner Greuter & Gianniantonio Domina Editorial board G. Domina (Palermo), F. Garbari (Pisa), W. Greuter (Berlin), S. L. Jury (Reading), G. Kamari (Patras), P. Mazzola (Palermo), S. Pignatti (Roma), F. M. Raimondo (Palermo), C. Salmeri (Palermo), B. Valdés (Sevilla), G. Venturella (Palermo). Advisory Committee P. V. Arrigoni (Firenze) P. Küpfer (Neuchatel) H. M. Burdet (Genève) J. Mathez (Montpellier) A. Carapezza (Palermo) G. Moggi (Firenze) C. D. K. Cook (Zurich) E. Nardi (Firenze) R. Courtecuisse (Lille) P. L. Nimis (Trieste) V. Demoulin (Liège) D. Phitos (Patras) F. Ehrendorfer (Wien) L. Poldini (Trieste) M. Erben (Munchen) R. M. Ros Espín (Murcia) G. Giaccone (Catania) A. Strid (Copenhagen) V. H. Heywood (Reading) B. Zimmer (Berlin) Editorial Office Editorial assistance: A. M. Mannino Editorial secretariat: V. Spadaro & P. Campisi Layout & Tecnical editing: E. Di Gristina & F. La Sorte Design: V. Magro & L. C. Raimondo Redazione di "Flora Mediterranea" Herbarium Mediterraneum Panormitanum, Università di Palermo Via Lincoln, 2 I-90133 Palermo, Italy [email protected] Printed by Luxograph s.r.l., Piazza Bartolomeo da Messina, 2/E - Palermo Registration at Tribunale di Palermo, no. 27 of 12 July 1991 ISSN: 1120-4052 printed, 2240-4538 online DOI: 10.7320/FlMedit26.001 Copyright © by International Foundation pro Herbario Mediterraneo, Palermo Contents V. Hugonnot & L. Chavoutier: A modern record of one of the rarest European mosses, Ptychomitrium incurvum (Ptychomitriaceae), in Eastern Pyrenees, France . 5 P. Chène, M. -
Evolution, Biogeography and Systematics of the Genus Cymbalaria Hill Evolució, Biogeografia I Sistemàtica Del Gènere Cymbalaria Hill Ph.D
ADVERTIMENT. Lʼaccés als continguts dʼaquesta tesi queda condicionat a lʼacceptació de les condicions dʼús establertes per la següent llicència Creative Commons: http://cat.creativecommons.org/?page_id=184 ADVERTENCIA. El acceso a los contenidos de esta tesis queda condicionado a la aceptación de las condiciones de uso establecidas por la siguiente licencia Creative Commons: http://es.creativecommons.org/blog/licencias/ WARNING. The access to the contents of this doctoral thesis it is limited to the acceptance of the use conditions set by the following Creative Commons license: https://creativecommons.org/licenses/?lang=en Evolution, biogeography and systematics of the genus Cymbalaria Hill Evolució, biogeografia i sistemàtica del gènere Cymbalaria Hill Ph.D. Thesis Pau Carnicero Campmany Unitat de Botànica Departament de Biologia Animal, Biolo- gia Vegetal i Ecologia Facultat de Biociències Universitat Autònoma de Barcelona Evolution, biogeography and systematics of the genus Cymbalaria Hill Ph.D. Thesis Pau Carnicero Campmany Bellaterra, 2017 Programa de doctorat en Ecologia Terrestre Unitat de Botànica Departament de Biologia Animal, Biolo- gia Vegetal i Ecologia Facultat de Biociències Universitat Autònoma de Barcelona Evolution, biogeography and systematics of the genus Cymbalaria Hill Memòria presentada per: Pau Carnicero Campmany per optar al grau de Doctor amb el vist-i-plau dels directors de tesi: Dra. Mercè Galbany Casals Dr. Llorenç Sáez Gonyalons (Directora i Tutora acadèmica) Unitat de Botànica Unitat de Botànica Departament de Biologia Departament de Biologia Animal, Vegetal i Ecologia Animal, Vegetal i Ecologia Facultat de Biociències Facultat de Biociències Universitat Autònoma de Barcelona Universitat Autònoma de Barcelona Dra. Núria Garcia Jacas Institut Botànic de Barcelona (IBB-CSIC-ICUB) Programa de doctorat en Ecologia Terrestre “When on board of H. -
GENETIC OBSERVATIONS on the GENUS LINARIA a Few Years Ago, I
GENETIC OBSERVATIONS ON THE GENUS LINARIA E. M. EAST Harvard University, Bussey Institution, Jamaica Plain, Massachusetts Received January 9, 1933 A few years ago, I obtained seeds from eighteen presumably different species of the genus Linaria-chiefly through the kindness of Professor Doctor E. BAURand of HAAGEund ScHMIm-in order to determine the value' of this group for genetical investigation. The list of species follows, together with some notes on their compatibility with each other. 1. L. bipartita Willd. Hab. northern Africa. Erect, branching, annual type. Fls. large, violet-purple, with orange palates above, becoming whitish toward the base. Spurs long and curved. Closely related to Nos. 7,9, 10, 14, and probably will cross with them and give fertile hybrids. No crosses were obtained when the plants were used as female with Nos. 3 (12 pol.) and 17 (16 pol.). 2. L. canadensis Dumont. Hab. New Brunswick, New England, and south to southwest. Slender, erect, annual. Lvs. linear. Fls. small, violet-blue to purple. Late flowering. No crosses tried because of this point. 3. L. Cymbalaria Mill. (Kenilworth ivy). Hab. Europe. Four types grown, received under the names vulgare (trailing), alba (trailing with white flowers), globosa (bushy), and compacta (bushy). A trailing, glabrous plant, with reniform-orbicular, 5-9 lobed leaves. Fls. small, axillary, of various shades of purple above and of yellow at the lip. Spurs short. No crosses Earlier, I have made similar surveys of other genera; but, as no especially interesting con- tributions to genetic knowledge resulted, the results were not putlished. I now believe that this decision was a mistake. -
Phylogeography of Western Mediterranean Cymbalaria
www.nature.com/scientificreports OPEN Phylogeography of western Mediterranean Cymbalaria (Plantaginaceae) reveals two Received: 5 December 2017 Accepted: 19 November 2018 independent long-distance Published: xx xx xxxx dispersals and entails new taxonomic circumscriptions Pau Carnicero 1,2, Peter Schönswetter2, Pere Fraga Arguimbau3, Núria Garcia-Jacas4, Llorenç Sáez1,5 & Mercè Galbany-Casals1 The Balearic Islands, Corsica and Sardinia (BCS) constitute biodiversity hotspots in the western Mediterranean Basin. Oligocene connections and long distance dispersal events have been suggested to cause presence of BCS shared endemic species. One of them is Cymbalaria aequitriloba, which, together with three additional species, constitute a polyploid clade endemic to BCS. Combining amplifed fragment length polymorphism (AFLP) fngerprinting, plastid DNA sequences and morphometrics, we inferred the phylogeography of the group and evaluated the species’ current taxonomic circumscriptions. Based on morphometric and AFLP data we propose a new circumscription for C. fragilis to additionally comprise a group of populations with intermediate morphological characters previously included in C. aequitriloba. Consequently, we suggest to change the IUCN category of C. fragilis from critically endangered (CR) to near threatened (NT). Both morphology and AFLP data support the current taxonomy of the single island endemics C. hepaticifolia and C. muelleri. The four species had a common origin in Corsica-Sardinia, and two long-distance dispersal events to the Balearic Islands were inferred. Finally, plastid DNA data suggest that interspecifc gene fow took place where two species co-occur. Islands in the Mediterranean Basin harbour both high species diversity and endemism. For instance, from the around 5000 islands scattered in the Mediterranean Sea1 all the largest ones (i.e. -
A Phylogenetic Study of the Tribe Antirrhineae: Genome Duplications and Long-Distance Dispersals from the Old World to the New World 1
RESEARCH ARTICLE AMERICAN JOURNAL OF BOTANY A phylogenetic study of the tribe Antirrhineae: Genome duplications and long-distance dispersals from the Old World to the New World 1 Ezgi Ogutcen2 and Jana C. Vamosi PREMISE OF THE STUDY: Antirrhineae is a large tribe within Plantaginaceae. Mostly concentrated in the Mediterranean Basin, the tribe members are present both in the Old World and the New World. Current Antirrhineae phylogenies have diff erent views on taxonomic relationships, and they lack homogeneity in terms of geographic distribution and ploidy levels. This study aims to investigate the changes in the chromosome numbers along with dispersal routes as defi nitive characters identifying clades. METHODS: With the use of multiple DNA regions and taxon sampling enriched with de novo sequences, we provide an extensive phylogeny for Antir- rhineae. The reconstructed phylogeny was then used to investigate changes in ploidy levels and dispersal patterns in the tribe using ChromEvol and RASP, respectively. KEY RESULTS: Antirrhineae is a monophyletic group with six highly supported clades. ChromEvol analysis suggests the ancestral haploid chromosome number for the tribe is six, and that the tribe has experienced several duplications and gain events. The Mediterranean Basin was estimated to be the origin for the tribe with four long-distance dispersals from the Old World to the New World, three of which were associated with genome duplications. CONCLUSIONS: On an updated Antirrhineae phylogeny, we showed that the three out of four dispersals from the Old World to the New World were coupled with changes in ploidy levels. The observed patterns suggest that increases in ploidy levels may facilitate dispersing into new environments. -
Vascular Plants of the Forest River Bi- Ology Station, North Dakota
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln The Prairie Naturalist Great Plains Natural Science Society 6-2015 VASCULAR PLANTS OF THE FOREST RIVER BI- OLOGY STATION, NORTH DAKOTA Alexey Shipunov Kathryn A. Yurkonis John C. La Duke Vera L. Facey Follow this and additional works at: https://digitalcommons.unl.edu/tpn Part of the Biodiversity Commons, Botany Commons, Ecology and Evolutionary Biology Commons, Natural Resources and Conservation Commons, Systems Biology Commons, and the Weed Science Commons This Article is brought to you for free and open access by the Great Plains Natural Science Society at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in The Prairie Naturalist by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. The Prairie Naturalist 47:29–35; 2015 VASCULAR PLANTS OF THE FOREST RIVER BI- known to occur at the site. Despite this effort, 88 species OLOGY STATION, NORTH DAKOTA—During sum- in La Duke et al. (unpublished data) are not yet supported mer 2013 we completed a listing of the plant species of the with collections, but have been included with this list. No- joint University of North Dakota (UND) Forest River Biol- menclature and taxon concepts are given in the accordance ogy Station and North Dakota Game and Fish Department with USDA PLANTS database (United States Department of Wildlife Management Area (FRBS).The FRBS is a 65 ha Agriculture 2013), and the Flora of North America (Flora of tract of land that encompasses the south half of the SW ¼ of North America Editorial Committee 1993). section 11 (acquired by UND in 1952) and the north half of We recorded 498 plant species from 77 families in the the NW ¼ of section 14 (acquired by UND in 1954) in Ink- FRBS (Appendix A), which is greater than the number of ster Township (T154N, R55W). -
New and Noteworthy Vascular Plant Records for Alabama
Barger, T.W., A. Cressler, B. Finzel, A. Highland, W.M. Knapp, F. Nation, A.R. Schotz, D.D. Spaulding, and C.T. Taylor. New and noteworthy vascular plant records for Alabama. Phytoneuron 2019-16: 1–7. Published 25 April 2019. ISSN 2153 733X NEW AND NOTEWORTHY VASCULAR PLANT RECORDS FOR ALABAMA T. WAYNE BARGER* Alabama Department of Conservation and Natural Resources State Lands Division, Natural Heritage Section Montgomery, Alabama 36130 [email protected] *Corresponding Author ALAN CRESSLER 1790 Pennington Place SE Atlanta, Georgia 30316 [email protected] BRIAN FINZEL St. John Paul II Catholic High School 7301 Old Madison Pike Huntsville, Alabama 35806 [email protected] AMY HIGHLAND Mt. Cuba Center 3120 Barley Mill Road Hockessin, Delaware 19707 [email protected] WESLEY M. KNAPP North Carolina Natural Heritage Program 176 Riceville Road Asheville, North Carolina 28805 [email protected] FRED NATION Weeks Bay Reserve 11300 U. S. Hwy 98 Fairhope, Alabama 36532 [email protected] ALFRED R. SCHOTZ Auburn University Museum of Natural History Alabama Natural Heritage Program Auburn, Alabama 36849 [email protected] DANIEL D. SPAULDING Anniston Museum of Natural History 800 Museum Drive/P.O. Box 1587 Anniston, Alabama 36202 [email protected] CHRIS T. TAYLOR 679 Cauthen Court Auburn, Alabama 36830 [email protected] 2 Barger et al.: Alabama records ABSTRACT Ten non-native species (Achyranthes japonica var. hachijoensis, Atocion armeria, Citrullus amarus, Cuscuta japonica, Cymbalaria muralis, Dioscorea alata, Grindelia ciliata, Odontonema cuspidatum, Paederia foetida, and Parentucellia viscosa), four native species (Clematis versicolor, Heteranthera multiflora, Pellaea glabella, and Quercus xbeadleii), five historic or uncommon species (Asplenium ruta- muraria var. -
Vascular Plant Species of the Comanche National Grassland in United States Department Southeastern Colorado of Agriculture
Vascular Plant Species of the Comanche National Grassland in United States Department Southeastern Colorado of Agriculture Forest Service Donald L. Hazlett Rocky Mountain Research Station General Technical Report RMRS-GTR-130 June 2004 Hazlett, Donald L. 2004. Vascular plant species of the Comanche National Grassland in southeast- ern Colorado. Gen. Tech. Rep. RMRS-GTR-130. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 36 p. Abstract This checklist has 785 species and 801 taxa (for taxa, the varieties and subspecies are included in the count) in 90 plant families. The most common plant families are the grasses (Poaceae) and the sunflower family (Asteraceae). Of this total, 513 taxa are definitely known to occur on the Comanche National Grassland. The remaining 288 taxa occur in nearby areas of southeastern Colorado and may be discovered on the Comanche National Grassland. The Author Dr. Donald L. Hazlett has worked as an ecologist, botanist, ethnobotanist, and teacher in Latin America and in Colorado. He has specialized in the flora of the eastern plains since 1985. His many years in Latin America prompted him to include Spanish common names in this report, names that are seldom reported in floristic pub- lications. He is also compiling plant folklore stories for Great Plains plants. Since Don is a native of Otero county, this project was of special interest. All Photos by the Author Cover: Purgatoire Canyon, Comanche National Grassland You may order additional copies of this publication by sending your mailing information in label form through one of the following media. -
Rare Vascular Plants of the North Slope a Review of the Taxonomy, Distribution, and Ecology of 31 Rare Plant Taxa That Occur in Alaska’S North Slope Region
BLM U. S. Department of the Interior Bureau of Land Management BLM Alaska Technical Report 58 BLM/AK/GI-10/002+6518+F030 December 2009 Rare Vascular Plants of the North Slope A Review of the Taxonomy, Distribution, and Ecology of 31 Rare Plant Taxa That Occur in Alaska’s North Slope Region Helen Cortés-Burns, Matthew L. Carlson, Robert Lipkin, Lindsey Flagstad, and David Yokel Alaska The BLM Mission The Bureau of Land Management sustains the health, diversity and productivity of the Nation’s public lands for the use and enjoyment of present and future generations. Cover Photo Drummond’s bluebells (Mertensii drummondii). © Jo Overholt. This and all other copyrighted material in this report used with permission. Author Helen Cortés-Burns is a botanist at the Alaska Natural Heritage Program (AKNHP) in Anchorage, Alaska. Matthew Carlson is the program botanist at AKNHP and an assistant professor in the Biological Sciences Department, University of Alaska Anchorage. Robert Lipkin worked as a botanist at AKNHP until 2009 and oversaw the botanical information in Alaska’s rare plant database (Biotics). Lindsey Flagstad is a research biologist at AKNHP. David Yokel is a wildlife biologist at the Bureau of Land Management’s Arctic Field Office in Fairbanks. Disclaimer The mention of trade names or commercial products in this report does not constitute endorsement or rec- ommendation for use by the federal government. Technical Reports Technical Reports issued by BLM-Alaska present results of research, studies, investigations, literature searches, testing, or similar endeavors on a variety of scientific and technical subjects. The results pre- sented are final, or a summation and analysis of data at an intermediate point in a long-term research project, and have received objective review by peers in the author’s field.