DOCSLIB.ORG

Evolution, Biogeography and Systematics of the Genus Cymbalaria Hill Evolució, Biogeografia I Sistemàtica Del Gènere Cymbalaria Hill Ph.D

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Evolution, Biogeography and Systematics of the Genus Cymbalaria Hill Evolució, Biogeografia I Sistemàtica Del Gènere Cymbalaria Hill Ph.D ADVERTIMENT. Lʼaccés als continguts dʼaquesta tesi queda condicionat a lʼacceptació de les condicions dʼús establertes per la següent llicència Creative Commons: http://cat.creativecommons.org/?page_id=184 ADVERTENCIA. El acceso a los contenidos de esta tesis queda condicionado a la aceptación de las condiciones de uso establecidas por la siguiente licencia Creative Commons: http://es.creativecommons.org/blog/licencias/ WARNING. The access to the contents of this doctoral thesis it is limited to the acceptance of the use conditions set by the following Creative Commons license: https://creativecommons.org/licenses/?lang=en Evolution, biogeography and systematics of the genus Cymbalaria Hill Evolució, biogeografia i sistemàtica del gènere Cymbalaria Hill Ph.D. Thesis Pau Carnicero Campmany Unitat de Botànica Departament de Biologia Animal, Biolo- gia Vegetal i Ecologia Facultat de Biociències Universitat Autònoma de Barcelona Evolution, biogeography and systematics of the genus Cymbalaria Hill Ph.D. Thesis Pau Carnicero Campmany Bellaterra, 2017 Programa de doctorat en Ecologia Terrestre Unitat de Botànica Departament de Biologia Animal, Biolo- gia Vegetal i Ecologia Facultat de Biociències Universitat Autònoma de Barcelona Evolution, biogeography and systematics of the genus Cymbalaria Hill Memòria presentada per: Pau Carnicero Campmany per optar al grau de Doctor amb el vist-i-plau dels directors de tesi: Dra. Mercè Galbany Casals Dr. Llorenç Sáez Gonyalons (Directora i Tutora acadèmica) Unitat de Botànica Unitat de Botànica Departament de Biologia Departament de Biologia Animal, Vegetal i Ecologia Animal, Vegetal i Ecologia Facultat de Biociències Facultat de Biociències Universitat Autònoma de Barcelona Universitat Autònoma de Barcelona Dra. Núria Garcia Jacas Institut Botànic de Barcelona (IBB-CSIC-ICUB) Programa de doctorat en Ecologia Terrestre “When on board of H. M. S ‘Beagle’, I was much struck with certain facts in the distribution of inhabitants of South America, and in the geological relations of the present to the past inhabitants of that continent. These facts seemed to me to throw some light on the origin of the species ̶ that mistery of misterys” C. Darwin, 1859 Als barrancs, que ara sovintejaven, s’hi escampaven gleves denses de silene brodades de floretes rosades i ornades de l’exuberància de les semprevives, mentre que a les concavitats petites treien el cap les últimes anemones, els ranuncles de les geleres, migrats clavells i centenars d’altres flors diminutes de noms estranys i colors vivíssims. W. Bonatti, 1999 (traducció de S. Pohl) “When on board of H. M. S ‘Beagle’, I was much struck with certain facts in the distribution of inhabitants of South America, and in the geological relations of the present to the past inhabitants of that continent. These facts seemed to me to throw some light on the origin of the species ̶ that mistery of misterys” C. Darwin, 1859 Als barrancs, que ara sovintejaven, s’hi escampaven gleves denses de silene brodades de floretes rosades i ornades de l’exuberància de les semprevives, mentre que a les concavitats petites treien el cap les últimes anemones, els ranuncles de les geleres, migrats clavells i centenars d’altres flors diminutes de noms estranys i colors vivíssims. W. Bonatti, 1999 (traducció de S. Pohl) Agraïments Una tesi sol trobar-se envoltada d’una certa aura de transcendència en l’imaginari col·lectiu. Per fer-ne justícia, aquí em trobo, escrivint la part que més persones llegiran en un tren de la línia R3. Abans de res, però, ho dic o rebento: m’ho he passat molt bé fent aquesta tesi. En aquell penúltim any de carrera, vaig ser víctima de l’entusiasme de la Mercè i el Llorenç per la flora i la distribució dels endemismes. Els vostres estils tan diferents em van inspirar en aquelles memorables jornades de camp en què se’m descobria el tresor que tenia sota els peus. Unes primeres col·laboracions estudiant el gènere Helichrysum van relligar la fascinació per la biodiversitat amb una creixent inquietud en l’Evolució. D’aquesta manera vaig conèixer l’Institut Botànic de Barcelona i la Núria, que des de llavors es va atrevir a pensar que alguna cosa se’n podria fer, de mi. La tesi va anar venint com una seqüència natural, i per una sèrie de caramboles les cimbalàries em van, en certa manera, trobar i convertir-se en les protagonistes d’aquesta tesi. Gràcies, directors, per acollir-me, deixar-me provar coses noves i equivocar-me. Em dec també a tants altres botànics que han anat marcant els meus inicis. Al Ramon Pérez, per dirigir-me a en Llorenç quan et vaig comentar les meves inquietuds. A en Pere Fraga i en Jaume Soler, quin plaer anar al camp amb vosaltres! To all the team from the Evolutionary Systematics Lab in Innsbruck, and specially to Peter, for trusting me and your honesty. The stay in Innsbruck was a milestone in my career. I also thank Pam and Doug Soltis, for being always so positive and open-minded, I learnt a different (and successful!) way of working than we do here. A tot l’equip de la Unitat de Botànica de la UAB i de l’Institut Botànic de Barcelona, entre temps millors i pitjors, m’heu ajudat a anar creixent com a científic. Com oblidar-me de la més gran, cruel i tendra amiga, la Muntanya. Pels grans paisatges regalats, pels petits racons i aquella brisa fresca, per l’equilibrada dança per la vertical. I per la por a l’abisme que atrau, pel fred extrem dels cims a l’hivern, per l’extenuació on l’aire s’enrareix. Aquesta adrenalina no té preu. Gràcies a tots els que m’hi heu acompanyat en un moment o altre. Llorenç, em vas introduir al món de les crestes i les hores de xerrera alpinística em van descobrir fantàstiques muntanyes i rutes, encara estranyes per mi, mai ho oblidaré. A la Gemma per algunes de les escalades més fantàstiques de la meva vida. Óscar i Isma, pel gran aprovisionament del vostre camp base. Al Xavi, mestre, referent, sense paraules. A la Núria i el Nico, molt més que companys d’escalada! A la gent de Chile, Tío Lucho, Henry, grans de les alçades. A tots els escaladors calderins amb qui he fet volar coloms, al Joel, Madu, Peies, Piui... i en especial al Martí, per haver anat complint el somni de les hores de pati d’explorar paradisos verticals, sempre fugint endavant. To Peter, I owe you my (limited) ski touring skills, we made such a good team in the mountain! And to Andy and Alex, Giarda, Moritz... just strong people in Innsbruck! Hi ha tota una corrua de gent fantàstica que no vull oblidar: Uri, Dídac, Cesc, Martí, Peies... Ben tronats! Als companys de militància, la Sandra i l’Anna especialment, quina feinada esteu fent! Als companys de carrera i màster, Marta, Núria, Judit, María... per haver anat compartint penes i alegries (amb la certesa que les alegries han guanyat). A Noe, por los titánicos esfuerzos en provar todas las cervezas artesanas de Gainesville y hacer más llevadera la vida yankee. No oblido la família, pel mèrit que té recolzar-me sense entendre quin sentit té el què faig! Mama, pel teu recolzament i confiança sincera, no saps quant t’ho agraeixo. Papa, per transmetre’m que sempre es pot anar una mica més enllà. Als avis, exemple admiradíssim de perseverança i il·lusió. Avi, sembla que finalment hi haurà un doctor Carnicero a la família! A la Iaia i la Conxita, per haver-vos enamorat també de les cimbalàries. I des de la sorpresa i la pena de no haver coincidit prou, llegint-te, “Tio” Manel, crec que podríem haver passat hores parlant de les plantes que poblen els volts d’Albinyana tot prenent la fresca al sagrat... Mar, gràcies per haver-me acompanyat tot aquest temps, per comprendre les meves excentricitats, calmar-me quan m’esverava i escoltar, críticament, els meus projectes i idees. Pels grandíssims moments envoltats de natura i les petites “descobertes”, sovint mundanes, que per nosaltres eren com un nou continent. I sobretot, perquè sempre he sentit que creus en mi, i això dóna una embranzida molt bèstia. Table of contents Abstract ...................................................................................................................... 11 Resum ......................................................................................................................... 13 General introduction .................................................................................................. 15 Plant systematics ................................................................................................. 17 Systematics tools ................................................................................................. 20 The Meditarranean Basin .................................................................................... 27 Cymbalaria Hill (Plantaginaceae) ......................................................................... 28 Structure of the Thesis ......................................................................................... 31 Objectives ............................................................................................................ 33 Chapter I: Different speciation types meet in a Mediterranean genus: the biogeographic history of Cymbalaria (Plantaginaceae) ........................................... 41 Chapter II: Phylogeography of western Mediterranean Cymbalaria (Plantaginaceae) reveals two independent long-distance dispersals and entails new taxonomic circumscriptions ................................................................... 83 Chapter III: Cymbalaria muelleri subsp. villosa subsp. nov., a new morphologically and genetically divergent
Load more

Recommended publications
  • Proceedings Amurga Co
    PROCEEDINGS OF THE AMURGA INTERNATIONAL CONFERENCES ON ISLAND BIODIVERSITY 2011 PROCEEDINGS OF THE AMURGA INTERNATIONAL CONFERENCES ON ISLAND BIODIVERSITY 2011 Coordination: Juli Caujapé-Castells Funded and edited by: Fundación Canaria Amurga Maspalomas Colaboration: Faro Media Cover design & layout: Estudio Creativo Javier Ojeda © Fundación Canaria Amurga Maspalomas Gran Canaria, December 2013 ISBN: 978-84-616-7394-0 How to cite this volume: Caujapé-Castells J, Nieto Feliner G, Fernández Palacios JM (eds.) (2013) Proceedings of the Amurga international conferences on island biodiversity 2011. Fundación Canaria Amurga-Maspalomas, Las Palmas de Gran Canaria, Spain. All rights reserved. Any unauthorized reprint or use of this material is prohibited. No part of this book may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopying, recording, or by any information storage and retrieval system without express written permission from the author / publisher. SCIENTIFIC EDITORS Juli Caujapé-Castells Jardín Botánico Canario “Viera y Clavijo” - Unidad Asociada CSIC Consejería de Medio Ambiente y Emergencias, Cabildo de Gran Canaria Gonzalo Nieto Feliner Real Jardín Botánico de Madrid-CSIC José María Fernández Palacios Universidad de La Laguna SCIENTIFIC COMMITTEE Juli Caujapé-Castells, Gonzalo Nieto Feliner, David Bramwell, Águedo Marrero Rodríguez, Julia Pérez de Paz, Bernardo Navarro-Valdivielso, Ruth Jaén-Molina, Rosa Febles Hernández, Pablo Vargas. Isabel Sanmartín. ORGANIZING COMMITTEE Pedro
    [Show full text]
  • The Phenology of an Urban Street Flora: a Transect Study C.D
    British & Irish Botany 2(1): 1–26, 2020 The phenology of an urban street flora: a transect study C.D. Preston Cambridge, England Corresponding author: [email protected] This pdf constitutes the Version of Record published on 26th February 2020 Abstract Vascular plants in flower along a fixed 3.8 km route in eight streets in a primarily residential area of urban Cambridge, U.K., were recorded at monthly intervals between January 2016 and December 2019. There was a consistent annual pattern over the four years; the number of flowering species was greatest in June or July but there were still appreciable numbers of species flowering when totals were at their lowest in February or March. Five annuals (Capsella bursa-pastoris, Euphorbia peplus, Poa annua, Senecio vulgaris, Stellaria media) and one perennial (Parietaria judaica) were very frequent and flowered from January to December. Perennial species showed greater variation through the year than annual species. In most months the number of flowering British native species exceeded the combined number of archaeophytes and neophytes, but the native total peaked earlier in the summer and then declined more rapidly than that of the introductions. The transect method appeared to be effective in identifying the main annual phenological trends and also revealed the effects of extreme weather on the patterns in some seasons. Keywords: annual; archaeophyte; Cambridge; native; neophyte; perennial; weather; weedkiller. Introduction In January 2016 I took part in the BSBI New Year Plant Hunt, in which participants were invited to spend up to three hours in the field listing the vascular plants they could find in flower (Marsh, 2016).
    [Show full text]
  • Different Speciation Types Meet in a Mediterranean Genus: The
    This is an Accepted Manuscript of an article published in Taxon on 4 May 2017, available online: https://doi.org/10.12705/662.7 Different speciation types meet in a Mediterranean genus: the biogeographic history of Cymbalaria (Plantaginaceae). Running head: Phylogeny and biogeographic history of Cymbalaria Pau Carnicero1, Llorenç Sáez1, 2, Núria Garcia-Jacas3 and Mercè Galbany- Casals1 1 Departament de Biologia Animal, Biologia Vegetal i Ecologia, Facultat de Biociències, Universitat Autònoma de Barcelona, 08193 Bellaterra, Spain. 2 Societat d’Història Natural de les Balears (SHNB), C/ Margarida Xirgu 16, E-07011 Palma de Mallorca, Balearic Islands, Spain. 3 Institut Botànic de Barcelona (IBB-CSIC-ICUB), Pg. del Migdia s/n, ES-08038 Barcelona, Spain Author for correspondence: Pau Carnicero, [email protected] ORCID: P.C., http://orcid.org/0000-0002-8345-3309 Abstract Cymbalaria comprises ten species and six subspecies growing in rocky habitats in the Mediterranean Basin. Several features, such as the genus’ highly fragmented distribution as well as noticeable ecological differentiation between partially sympatric species and presence of ploidy barriers between species suggest the involvement of different speciation types in its evolution. The aims of this study were to test the monophyly of Cymbalaria and to reconstruct infrageneric phylogenetic relationships, to infer the genus’ biogeographic history by estimating divergence times and ancestral distribution areas of lineages, and to disentangle the role of different speciation types. To address these issues, we constructed a phylogeny with a complete taxon sampling based on ITS, 3'ETS, ndhF and rpl32-trnL sequences. We used the nuclear ribosomal DNA data to produce a time-calibrated phylogeny, which served as basis for estimating ploidy level evolution and biogeographic history.
    [Show full text]
  • 2011 Biodiversity Snapshot. Guernsey Appendices
    UK Overseas Territories and Crown Dependencies: 2011 Biodiversity snapshot. Guernsey: Appendices. Author: Dr Charles David Guernsey Biological Records Centre, States of Guernsey Environment Department & La Societe Guernesiaise. More information available at: www.biologicalrecordscentre.gov.gg This section includes a series of appendices that provide additional information relating to that provided in the Guernsey chapter of the publication: UK Overseas Territories and Crown Dependencies: 2011 Biodiversity snapshot. All information relating to Guernsey is available at http://jncc.defra.gov.uk/page-5743 The entire publication is available for download at http://jncc.defra.gov.uk/page-5821 Commissioned by the States of Guernsey Environment Department for the Joint Nature Conservation Committee Prepared by Dr C T David Guernsey Biological Records Centre August 2010 1 Contents Appendix 1: Bailiwick of Guernsey – Location and Introduction ............................. 3 Location, Area, Number of Islands, Population 3 Topography 4 Main economic sectors 4 Constitutional Position 4 Appendix 2: Multilateral Environmental Agreements. ............................................... 5 Appendix 3: National Legislation ................................................................................ 8 Planning 8 Ancient Monuments 8 Coast and beaches 8 Land 8 Fauna 8 Flora 9 Trees 9 Import/export 9 Marine environment 9 Waste 9 Water 9 Appendix 4: National Strategies ................................................................................ 11 Appendix
    [Show full text]
  • Vascular Plants and a Brief History of the Kiowa and Rita Blanca National Grasslands
    United States Department of Agriculture Vascular Plants and a Brief Forest Service Rocky Mountain History of the Kiowa and Rita Research Station General Technical Report Blanca National Grasslands RMRS-GTR-233 December 2009 Donald L. Hazlett, Michael H. Schiebout, and Paulette L. Ford Hazlett, Donald L.; Schiebout, Michael H.; and Ford, Paulette L. 2009. Vascular plants and a brief history of the Kiowa and Rita Blanca National Grasslands. Gen. Tech. Rep. RMRS- GTR-233. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 44 p. Abstract Administered by the USDA Forest Service, the Kiowa and Rita Blanca National Grasslands occupy 230,000 acres of public land extending from northeastern New Mexico into the panhandles of Oklahoma and Texas. A mosaic of topographic features including canyons, plateaus, rolling grasslands and outcrops supports a diverse flora. Eight hundred twenty six (826) species of vascular plant species representing 81 plant families are known to occur on or near these public lands. This report includes a history of the area; ethnobotanical information; an introductory overview of the area including its climate, geology, vegetation, habitats, fauna, and ecological history; and a plant survey and information about the rare, poisonous, and exotic species from the area. A vascular plant checklist of 816 vascular plant taxa in the appendix includes scientific and common names, habitat types, and general distribution data for each species. This list is based on extensive plant collections and available herbarium collections. Authors Donald L. Hazlett is an ethnobotanist, Director of New World Plants and People consulting, and a research associate at the Denver Botanic Gardens, Denver, CO.
    [Show full text]
  • Alien Flora of Europe: Species Diversity, Temporal Trends, Geographical Patterns and Research Needs
    Preslia 80: 101–149, 2008 101 Alien flora of Europe: species diversity, temporal trends, geographical patterns and research needs Zavlečená flóra Evropy: druhová diverzita, časové trendy, zákonitosti geografického rozšíření a oblasti budoucího výzkumu Philip W. L a m b d o n1,2#, Petr P y š e k3,4*, Corina B a s n o u5, Martin H e j d a3,4, Margari- taArianoutsou6, Franz E s s l7, Vojtěch J a r o š í k4,3, Jan P e r g l3, Marten W i n t e r8, Paulina A n a s t a s i u9, Pavlos A n d r i opoulos6, Ioannis B a z o s6, Giuseppe Brundu10, Laura C e l e s t i - G r a p o w11, Philippe C h a s s o t12, Pinelopi D e l i p e t - rou13, Melanie J o s e f s s o n14, Salit K a r k15, Stefan K l o t z8, Yannis K o k k o r i s6, Ingolf K ü h n8, Hélia M a r c h a n t e16, Irena P e r g l o v á3, Joan P i n o5, Montserrat Vilà17, Andreas Z i k o s6, David R o y1 & Philip E. H u l m e18 1Centre for Ecology and Hydrology, Hill of Brathens, Banchory, Aberdeenshire AB31 4BW, Scotland, e-mail; [email protected], [email protected]; 2Kew Herbarium, Royal Botanic Gardens Kew, Richmond, Surrey, TW9 3AB, United Kingdom; 3Institute of Bot- any, Academy of Sciences of the Czech Republic, CZ-252 43 Průhonice, Czech Republic, e-mail: [email protected], [email protected], [email protected], [email protected]; 4Department of Ecology, Faculty of Science, Charles University, Viničná 7, CZ-128 01 Praha 2, Czech Republic; e-mail: [email protected]; 5Center for Ecological Research and Forestry Applications, Universitat Autònoma de Barcelona, 08193 Bellaterra, Spain, e-mail: [email protected], [email protected]; 6University of Athens, Faculty of Biology, Department of Ecology & Systematics, 15784 Athens, Greece, e-mail: [email protected], [email protected], [email protected], [email protected], [email protected]; 7Federal Environment Agency, Department of Nature Conservation, Spittelauer Lände 5, 1090 Vienna, Austria, e-mail: [email protected]; 8Helmholtz Centre for Environmental Research – UFZ, Department of Community Ecology, Theodor-Lieser- Str.
    [Show full text]
  • Flora Mediterranea 26
    FLORA MEDITERRANEA 26 Published under the auspices of OPTIMA by the Herbarium Mediterraneum Panormitanum Palermo – 2016 FLORA MEDITERRANEA Edited on behalf of the International Foundation pro Herbario Mediterraneo by Francesco M. Raimondo, Werner Greuter & Gianniantonio Domina Editorial board G. Domina (Palermo), F. Garbari (Pisa), W. Greuter (Berlin), S. L. Jury (Reading), G. Kamari (Patras), P. Mazzola (Palermo), S. Pignatti (Roma), F. M. Raimondo (Palermo), C. Salmeri (Palermo), B. Valdés (Sevilla), G. Venturella (Palermo). Advisory Committee P. V. Arrigoni (Firenze) P. Küpfer (Neuchatel) H. M. Burdet (Genève) J. Mathez (Montpellier) A. Carapezza (Palermo) G. Moggi (Firenze) C. D. K. Cook (Zurich) E. Nardi (Firenze) R. Courtecuisse (Lille) P. L. Nimis (Trieste) V. Demoulin (Liège) D. Phitos (Patras) F. Ehrendorfer (Wien) L. Poldini (Trieste) M. Erben (Munchen) R. M. Ros Espín (Murcia) G. Giaccone (Catania) A. Strid (Copenhagen) V. H. Heywood (Reading) B. Zimmer (Berlin) Editorial Office Editorial assistance: A. M. Mannino Editorial secretariat: V. Spadaro & P. Campisi Layout & Tecnical editing: E. Di Gristina & F. La Sorte Design: V. Magro & L. C. Raimondo Redazione di "Flora Mediterranea" Herbarium Mediterraneum Panormitanum, Università di Palermo Via Lincoln, 2 I-90133 Palermo, Italy [email protected] Printed by Luxograph s.r.l., Piazza Bartolomeo da Messina, 2/E - Palermo Registration at Tribunale di Palermo, no. 27 of 12 July 1991 ISSN: 1120-4052 printed, 2240-4538 online DOI: 10.7320/FlMedit26.001 Copyright © by International Foundation pro Herbario Mediterraneo, Palermo Contents V. Hugonnot & L. Chavoutier: A modern record of one of the rarest European mosses, Ptychomitrium incurvum (Ptychomitriaceae), in Eastern Pyrenees, France . 5 P. Chène, M.
    [Show full text]
  • (Linaria Vulgaris) and Dalmatian Toadflax (Linaria
    DISSERTATION VIABILITY AND INVASIVE POTENTIAL OF HYBRIDS BETWEEN YELLOW TOADFLAX (LINARIA VULGARIS) AND DALMATIAN TOADFLAX (LINARIA DALMATICA) Submitted by Marie F.S. Turner Department of Soil and Crop Sciences In partial fulfillment of the requirements For the Degree of Doctor of Philosophy Colorado State University Fort Collins, Colorado Fall 2012 Doctoral Committee: Advisor: Sarah Ward Christopher Richards David Steingraeber George Beck Sharlene Sing Copyright by Marie Frances Sundem Turner 2012 All Rights Reserved ABSTRACT VIABILITY AND INVASIVE POTENTIAL OF HYBRIDS BETWEEN YELLOW TOADFLAX (LINARIA VULGARIS) AND DALMATIAN TOADFLAX (LINARIA DALMATICA) Although outcomes of hybridization are highly variable, it is now considered to play an important role in evolution, speciation, and invasion. Hybridization has recently been confirmed between populations of yellow (or common) toadflax (Linaria vulgaris) and Dalmatian toadflax (Linaria dalmatica) in the Rocky Mountain region of the United States. The presence of hybrid toadflax populations on public lands is of concern, as both parents are aggressive invaders already listed as noxious weeds in multiple western states. A common garden experiment was designed to measure differences in quantitative (shoot length, biomass, flowering stems, seed capsule production) phenological (time of emergence, first flowering and seed maturity) and ecophysiological (photosynthesis, transpiration and water use efficiency (WUE)) traits for yellow and Dalmatian toadflax, F1 and BC1 hybrids, as well as natural field-collected hybrids from two sites. Genotypes were cloned to produce true replicates and the entire common garden was also replicated at two locations (Colorado and Montana); physiological data were collected only in Colorado. All genotypes grew larger and were more reproductively active in Colorado than in Montana, and hybrids outperformed parent taxa across vegetative and reproductive traits indicating heterosis.
    [Show full text]
  • ABSTRACTS 117 Systematics Section, BSA / ASPT / IOPB
    Systematics Section, BSA / ASPT / IOPB 466 HARDY, CHRISTOPHER R.1,2*, JERROLD I DAVIS1, breeding system. This effectively reproductively isolates the species. ROBERT B. FADEN3, AND DENNIS W. STEVENSON1,2 Previous studies have provided extensive genetic, phylogenetic and 1Bailey Hortorium, Cornell University, Ithaca, NY 14853; 2New York natural selection data which allow for a rare opportunity to now Botanical Garden, Bronx, NY 10458; 3Dept. of Botany, National study and interpret ontogenetic changes as sources of evolutionary Museum of Natural History, Smithsonian Institution, Washington, novelties in floral form. Three populations of M. cardinalis and four DC 20560 populations of M. lewisii (representing both described races) were studied from initiation of floral apex to anthesis using SEM and light Phylogenetics of Cochliostema, Geogenanthus, and microscopy. Allometric analyses were conducted on data derived an undescribed genus (Commelinaceae) using from floral organs. Sympatric populations of the species from morphology and DNA sequence data from 26S, 5S- Yosemite National Park were compared. Calyces of M. lewisii initi- NTS, rbcL, and trnL-F loci ate later than those of M. cardinalis relative to the inner whorls, and sepals are taller and more acute. Relative times of initiation of phylogenetic study was conducted on a group of three small petals, sepals and pistil are similar in both species. Petal shapes dif- genera of neotropical Commelinaceae that exhibit a variety fer between species throughout development. Corolla aperture of unusual floral morphologies and habits. Morphological A shape becomes dorso-ventrally narrow during development of M. characters and DNA sequence data from plastid (rbcL, trnL-F) and lewisii, and laterally narrow in M.
    [Show full text]
  • Sibthorpia Peregrina L
    Nuevas citas y observaciones Sibthorpia peregrina L. (Plantaginaceae) novedad para España (Parque Natural de los Alcornocales, Cádiz). ¿Neófita o relicta macaronésica? Enrique Sánchez Gullón1 & Íñigo Sánchez García2 1 Paraje Natural Marismas del Odiel, Ctra. del Dique Juan Carlos I, Apdo. 720, 21071, Huelva. 2 ZooBotánico Jerez. C/ Madreselva s/n. 11404, Jerez de la Frontera. Recibido: 2 de noviembre de 2019. Aceptado (versión revisada): 6 de noviembre de 2020. Publicado en línea: 10 de noviembre de 2020. Sibthorpia peregrina L. (Plantaginaceae) novelty for Spain (Parque Natural de los Alcornocales, Cádiz). Neophyte or Macaronesian relict? Palabras claves: Sibthorpia peregrina; Corología; España; Andalucía; Cádiz. Keywords: Sibthorpia peregrina; Chorology; Spain; Andalusia; Cadiz. Resumen Abstract Se registra la presencia de Sibthorpia peregrina L. en el Parque The presence of Sibthorpia peregrina L. is registered in the Los Natural de los Alcornocales (Cádiz) por primera vez en España. Se Alcornocales Natural Park (Cádiz) for the first ime in Spain. It is discute si esta especie, considerada hasta el momento endémica de debated whether this species, considered endemic to Madeira unil Madeira, se encuentra aquí como un neófito de llegada reciente o now, is found here as a recently arrived neophyte or, on the contrary, por el contrario se trata de una especie relicta de distribución más it is a relict species with a wider distribuion not detected to date. amplia no detectada hasta la fecha. El género Sibthorpia L. (Plantaginaceae) (etimológicamente Esta población aparece localizada sobre sustratos de areniscas dedicado a Humphrey Waldo Sibthorp -1713-1797-, médico y oligocenas con rankers aluvial, en suelos muy higroturbosos botánico inglés, profesor de botánica en Oxford), comprende dentro de una aliseda con Rhododendrum ponticum, 5 especies: S.
    [Show full text]
  • Development of Distribution Maps of Grassland Habitats of EUNIS Habitat Classification
    Development of distribution maps of grassland habitats of EUNIS habitat classification Joop H.J. Schaminée Milan Chytrý Jürgen Dengler Stephan M. Hennekens John A.M. Janssen Borja Jiménez-Alfaro Ilona Knollová Flavia Landucci Corrado Marcenò John S. Rodwell Lubomír Tichý and data-providers Report EEA/NSS/16/005 1 Alterra, Institute within the legal entity Stichting Dienst Landbouwkundig Onderzoek Professor Joop Schaminée Stephan Hennekens Partners Professor John Rodwell, Ecologist, Lancaster, UK Professor Milan Chytrý, Masaryk University, Brno, Czech Republic Doctor Ilona Knollová, Masaryk University, Brno, Czech Republic Doctor Lubomír Tichý, Masaryk University, Brno, Czech Republic Date: 07 December 2016 Alterra Postbus 47 6700 AA Wageningen (NL) Telephone: 0317 – 48 07 00 Fax: 0317 – 41 90 00 In 2003 Alterra has implemented a certified quality management system, according to the standard ISO 9001:2008. Since 2006 Alterra works with a certified environmental care system according to the standard ISO 14001:2004. © 2014 Stichting Dienst Landbouwkundig Onderzoek All rights reserved. No part of this document may be reproduced, stored in a retrieval system, or transmitted in any form or by any means - electronic, mechanical, photocopying, recording, or otherwise - without the prior permission in writing of Stichting Dienst Landbouwkundig Onderzoek. 2 TABLE OF CONTENTS 1 Introduction 2 Scope of the project 2.1 Background 2.2 Review of the EUNIS grassland habitat types 3 Indicator species of the revised EUNIS grassland habitat types 3.1 Background
    [Show full text]
  • Palinotaxonomía De Scrophulariaceae Sensu Lato
    Bol. Soc. Argent. Bot. 51 (2) 2016 M. M. Sosa y C. R. Salgado - Palinotaxonomía de ScrophulariaceaeISSN sensu 0373-580 lato X Bol. Soc. Argent. Bot. 51 (2): 299-321. 2016 VALOR TAXONÓMICO DEL POLEN EN SCROPHULARIACEAE SENSU LATO MARÍA DE LAS MERCEDES SOSA1,3 y CRISTINA R. SALGADO2,3 Resumen: Este trabajo es el resultado del estudio palinomorfológico de 32 especies pertenecientes a 19 géneros de Scrophulariaceae sensu lato. Además se realizó un registro exhaustivo de la información bibliográfica existente sobre la morfología polínica con el objetivo de analizar las relaciones entre las variables palinológicas y la clasificación sistemática actual. Los granos de polen analizados son: mónades, radiosimétricos e isopolares, de tamaño variable desde pequeños (11 µm) hasta medianos (51 µm), predominantemente esferoidales (P/E= 1), a veces suboblatos (P/E= 0,76), oblato-esferoidales (P/E= 0,95), prolatos (P/E=1,63) y perprolatos (P/E= 2,63). Inaperturados, 3-(4-5) colpados, 3-(4-5) colporados, espiraperturados, 2-8 sincolpados, 3-diploporados y pantoporados. La exina puede ser semitectada (retipilada, reticulada o estriada perforada) y tectada (psilada, foveolada, perforada, escábrida, rugulada, verrugosa, granulada, estriada o equinulada). Se realizó un análisis estadístico en base a una matriz de datos, compuesta por los caracteres palinológicos de 32 especies estudiadas sumado a la recopilación bibliográfica de 140 géneros y 218 especies pertenecientes a Scrophulariaceae sensu lato. El análisis de agrupamiento (UPGMA), confirmó que las diferencias palinomorfológicas apoyan la clasificación sistemática actual. Palabras clave: Morfología del polen, sistemática, euripolínico, Scrophulariaceae. Summary: Taxonomic value of pollen in Scrophulariaceae sensu lato. This paper is result of a study of pollen morphology of 32 species belonging to 19 genera of Scrophulariaceae sensu lato.
    [Show full text]