Revision of the Lauraceae I
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719 Revision of the Lauraceae I by A.J.G.H. Kostermans (Utrecht). List of abbreviations: Lam. = Herb. Lamarck (Paris) Herbaria M. = Miinchen B = Brussels Ms. = Meissner Herbar. (New York) Br. = Brit. Museum NY. = New York = Bs. = Herbier Boisskr (Geneve) P. Paris C. = Herb. Cosson (Paris) R. = Rio de Janeiro D. = Berlin-Dahlem St. = Stockholm Dr. = U. Herb. Drake (Paris) = Utrecht DC. = Herb. De Candolle (Geneve) W. = Wien Ws. = Dl. — Herb. Delessert (Geneve) Smithsonian Instit. F. = Field Museum (Chicago) (Washington. D. C.) G. = Gottingen Other abbreviations Gr. = Gray Herb. (Cambridge, Ar. = Arowaccan language (Dutch Mass. U.S.A.) Guiana) H. M. G. Mus. B. W. = = Herb. Goeldi Boschwezen (Forestry Bureau) H. R. = Herb. Rio de Car. Caraibic J. Janeiro = language I. = Koloniaal Instituut (Amsterdam) N. E. = Negro English (Dutch Guiana) K. Sar. Saramaccan = Kjobnhavn = language (Dutch Kw. = Kew Guiana) L. = Leiden S. D. = Surinam Dutch The literature and cited in this — if not synonyms, monograph are, men- tioned otherwise, — verified by myself. All specimens mentioned are identified or verified by myself. I wish best thanks the directors of the above mentioned to express my to herbaria for their great assistance. The specimens from Dutch Guiana have not yet been distributed. ACRODICLIDIUM Nees Acrodiclidium Nees, Laur. Disp. Progr. (1833), p. 13; id., Syst. Laur. Gen. (1836), p. 266; Endl., (1837), p. 319, n. 2042; id., Ench. Pl. II (1841), p. 197; Dietrich, Synops. (1840), p. 1332; Phaner. Steudel. Spach, Hist. nat. Véget., X (1841), p. 471; Nomencl. ed. Gen. 2 (1841), p. 21; Meissn., I (1836—43), p. 326, Diet. univ. II, p. 238; Reichb., Nom., p. 71, n. 2668; Orbigny, *) A proposal to retain this name and to reject Licaria has been made by Koslermans I.c. 720 VII Fl. (1846), p. 259; Lindl., Veg. Kgd. (1846), p. 537; Griseb., Brit. W. I. isl. I in Prodr. (1860), p 280; Meissn. D.C., XV, 1 id. in Fl. Bras. 2 Benth. in Benth. (1864), p. 84; V, (1866), p. 172; et Gen. III Hist. II Hook., (1880), p. 154; Baillon, (1870), p. 474;. Nomencl. Index Pfeiffer, (1873), p. 35; Durand, Gen. (1888), Mez in Bot. Gart. Berl. V p. 349, n. 6190; Jahrb. (1889), p. 81; Pax in Pfl. fam. 2 Dalla Torre Engl.-Prantl, III, (1889), p. 123; Gen. Britton and et Harms, (1900—07), p. 178, n. 2819; Wilson, Porto Rico and isl. Dict. Virg. (1924), p. 316; Lemée, I (1929), Benoist in Arch. V in p. 50; Bot. (1931), p. 65; Kosterm. Pulle, Fl. Surin. II — Licaria Guia. (1936), p. 315; Aubl., I (1775), p. 313; Nees, Syst., p. 344; Endl., Gen, p. 320; id., Ench., p. 197; Steudel, Gen. Spach., l.c.; l.c., p. 41; Meissn., II, p. 238; Lindl., Meissn. in l.c.; D.C., l.c., p. 259; Benth., l.c., p. 150; Baillon, l.c., p. 452; Pfeiffer, l.c., p. 107; Durand, l.c., p. 489; Mez, l.c., p. 220;. dalla et Benoist Torre, l.c., p. 177 585; Lemée, l.c., IV, p. 85; Kosterm. in Meded. Bot. Mus. Utrecht l.c., p. 274; 25 (1936), id. in — p. 34; Pulle, l.c., p. 323; Evonymodaphne Nees, Syst., ed p. 244 et 263; Lindl., Syst. 2 (1836), p. 442; Endl., Gen. p. 319;. id., Ench., p. 197; Dietrich, l.c., p. 1332; Spach, l.c.; Steudel, l.c., Gen. id. p. 621; Meissn., I, p. 326; II, p. 238; Rchb., l.c.; Lindl., Meissn. in id. in Fl. l.c., p. 537; D.C., l.c., p. III; Bras., p. 203; Benth., l.c., p. 158; Baillon, l.c., p. 437; Pfeiffer, l.c., p. 1322; dalla — Durand, l.c., p. 349; Mez, l.c., p. 82; Torre, l.c., p. 177; Fl. Tellur. dalla Triplomeia Rafin., (1838), p. 134; Torre, l.c., p. 178; Mez, l.c. Type species: Acrodiclidium brasiliense Nees. Trees or shrubs in tropical South America and the West Indies.. alternate chartaceous Leaves or opposite, or coriaceous, penniner- ved, usually thickened, along the margin. Panicles or rarely pseudo- and few- racemes axillary sub-terminal, to many-flowered, pro- ducing only few fruit. Flowers hermaphrodite. Involucre wanting. Flower tube small Petals usually distinct, sometimes or wanting. 6, usually nearly equal. The 2 outer rows of stamens transferred into of small, rarely petalous staminodes, rarely abortive. Third row filaments slender stamens fertile, or as broad as anthers, distinct or wanting; base provided outside with 2 glands, rarely glands wanting; anthers 2-celIed, cells introrse, extrorse or extrorse-apically; valves dehiscent from base Fourth of as a rule to top. row stamens usually wanting, rarely staminodial. Ovary ellipsoid, glabrous or sericeous, immersed in the tube, style slender, stigma minute. Fruit at first included in the enlarged tube, at last exserted, base sur- 721 thick- and rounded by the simple- and thinly margined, or double-, rarely triple margined cupule. Cotyledons flat-convex, including and conical radicle. the minute, 2 —4-leaved plumule characterised the 6 staminodes and the fertile inner This genus, by outer 3 ones with basal glands and 2-celled anthers, links Huherodaphne and Phyl- lostemonodaphne with Misanteca and Meiilaurus. Key to the species: ta. Fourth row of staminodes distinct 2 staminodes b. Fourth row of wanting 3 Flower inside. 2a. Leaves cordate. tube glabrous Outer staminodes with fimbriate Fertile sub-orbicular, tops. stamens glabrous with rather large cells. Staminodes of the fourth row short, A. endlicheriaefolium Kosterm. glabrous 1. b. Leaves with rounded-acute base. Tube sericeouls inside. Outer staminodes large, auricled. Fertile stamens sericeous at base, with auricled anthers, cells minute. Staminodes of the fourth row large, densely sericeous. A. 2. A. macrophyllum C. Smith. Cells of the fertile introrse. and leaves 3a. stamens Ovary glabrous 4 b. Cells extrose. and leaves or sericeous Ovary glabrous .. 9 Leaves reticulate both 4a. distinctly, densely, prominently on sides. 1. in sub-racemes. Flowers 4—7 mm A. Mez j. Puchury-major (Mart.) X b. Flowers — I. in \ A. 3 mm panicles 3 Leaves 8. A. Kosterm. 5 a. opposite rigidum b. Leaves alternating 6 the tube. 6a. Tepals patent, as long as Cells distinct 7 b. shorter than the tube. Cells minute 8 Tepals incurved, .. last tube c. Tepals at reflexed, wanting. Cells large. 9. A. armeniacum (Nees) Mez staminodes Basal the 7a. Outer rectangular. glands as long as sessile Cells A. Mez istamens. large 3. amarum b. Outer staminodes clavate or spathulate. Cells small. Basal half the filaments broad glands as long ais stamens; as as the anthers A. Rob. 4. Camara Schomb. reticulate 8 a. Leaves coriaceous, elliptic, 8—12 cm 1., on both Basal distinct. sides. Panicles 3 —12 cm 1. glands Cupule verrucous, triple-margined, outer one of 6 large lobes. 6. A. Canella (Meissn.) Mez b. Leaves veins incons- rigid-coriaceous, elliptic, 10—14 cm 1., Panicles 1. Basal picuous. 3 cm glands wanting. Cupule 722 smooth with shortly Idbed outer margin. 7. A. cayennense (Meissn.) Mez 9a. Ovary glabrous 10 b. Ovary sericeous 22 toa. Leaves opposite 11 b. Leaves alternate, rarely sub-oppo(site 14 11a. Leaves densely aureo-sericeous beneath 12 b. sericeouls Leaves sparsely, minutely or glabrescent beneath 13 12a. Leaves sericeous beneath. Flowers sulb-globose. Stamens in- cluded. Outer staminodes distinct. Basal glands aS long as filaments. Cells Hub. minute ij. A. aureum b. Leaves sericeous-tomentellous beneath. Flowers cylindrical. Stamens exserted. Outer istaminodes as a rule abortive. Bas il glands minute or wanting. Cells large. 16. A. oppositifolium Nees Leaves 6 —8 — Flowers 13a. rigid-coriaceous, X 1Y2 2 cm. unknown. Cupule thick, 28 mm h., 35 mm diam., warty; margin triple: inner one 1—2 mm h., outer one 8 mm thick, with lobes in double patent, large irregular a row. 8. A. rigidum Kosterm. b. Flowers Leaves chartaceous, 10—13 X 2 —3 cm. narrowly urceolate. Outer staminodes slender, spathulate, 1 mm 1. Stamens exserted. h., Cupule thin, smooth, 7 mm 15 mm diam.; margin simple 14. A. Meissneri Mez c. Leaves thin-chartaceous, 10—12 X 3—4Y2 cm. Flowers Outer staminodes broadly funnel-shaped. ovate or broadly 1. included. elliptic, X mm Stamens Cupule thin, 7 mm h., A. debile Mez 15 mm diam.; margin simple 13. Flowers 14a. 3 mm diam.; tube nearly wanting; petals reflexed after flowering. Outer staminodes i}/2 mm I. Basal glands tepaloid nearly as long as the stamens. 9. A. armeniacum (Nees) Mez b. Flowers tube up to 2 mm diam.; distinct; tepals erect or Outer staminodes to 1. Basal erect-patent. up 1 mm glands shorter than the stamens 15: Flowers 1. 15a. cylindrical-urceolate, mm 10. A. guianense Nees b. Flowers subglobose or broadly infundibuliform, up to 2 Y2 mm 1., rarely sub-urceolate, but then only mm 1. 16 .. 16a. Leaves reticulate above distinctly prominulously .... 17 b. Leaves smooth above 18 Leaves 6 Flowers 17a. lanceolate, —9 X 1^/2—2 cm. sub-globose; petals incurved. Outer istaminodes Y2 mm !•> filaments of 723 fertile stamens not distinct. West Indian species. A. salicifolium Gr. 17. (Sw.) b. Leaves —16 —6 Flowers infundibuli- elliptical, 12 X 3 cm. form; tepals erect-patent; outer staminodes X mm 1. Fila- ments of fertile stamens slender, distinct. Pucheri 12. A. (R. et P.) Kosterm. areolate 18a. Leaves elliptical, 7—10 X iVi —3X2 cm, minutely above. Flowers sub-urceolate, rather slender. n. A. multiflorum Kcxsterm. b. Leaves lanceolate, 6—9 X ij'a— 3 cm or elliptical, 12—19 smooth above. Flowers X 4—9 cm, globose 19 19a. Leaves lanceolate, 6—9 X 1Y2 —3 cm., soon glabrous beneath. West Indian 18. A. sericeum species .... Griueb. Leaves X tomentellous beneath b.