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The scientific name of the coffee bean weevil and some additional bibliography (Coleoptera: Anthribidae: Araecerus Schönherr)

Barry D. Valentine Ohio State University, Columbus, Ohio

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Valentine, Barry D., "The scientific name of the coffee bean weevil and some additional bibliography (Coleoptera: Anthribidae: Araecerus Schönherr)" (2005). Insecta Mundi. 98. https://digitalcommons.unl.edu/insectamundi/98

This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 19, No. 4, December, 2005 247

The scientific name of the coffee bean weevil and some additional bibliography (Coleoptera: Anthribidae: Araecerus Schönherr)

Barry D. Valentine Museum of Biological Diversity The Ohio State University Columbus, Ohio 43212 [email protected]

Abstract. The name Araecerus fasciculatus (DeGeer 1775) is resurrected for the coffee bean weevil, with the following synonyms: Bruchus cacao Fabricius 1775, Bruchus peregrinus Herbst 1797, Bruchus capsinicola Fabricius 1798, Anthribus coffeae Fabricius 1801, Amblycerus japonicus Thunberg 1815 (probably a synonym), Anthribus alternans Germar 1824, Phloeobius griseus of Stephens 1831, not Fabricius 1792, Cratoparis parvirostris Thomson 1858, Araecerus seminarius Chevrolat 1871, and Tropideres ( Rhaphitropis) mateui Cobos 1954. Lectotypes are designated for A. fasciculatus, B. capsinicola, and A. alternans. A diagnosis distinguishes Araecerus from all 650 anthribid genera and a diagnosis for the species is as complete as possible with materials studied. Some additional species of Araecerus are discussed: Bruchus crassicornis Fabricius 1798 is distinguished from A. fasciculatus and a lectotype is designated. Araecerus suturalis Boheman 1839 is diagnosed and shown to be dissimilar from the species recently reported as A. suturalis from South Africa. The identity of Araecerus suturalis of Frieser, not Boheman, is not yet clear. Araecerus sambucinus Boisduval 1835 and Trepideres [sic] fragilis Walker 1859 may be synonyms of A. fasciculatus but pertinent data are insufficient. Brachytarsus niveovariegatus Roelofs 1879 (spelled nigrovariegatus by Bovie (1906)), listed as a synonym of A. fasciculatus by Wolfrurn (1929) is removed to Anthribus Forster 1770 with Anthribus lajievorus Chao 1976 as a new synonym. Introduction name fasciculatus to the coffee bean weevil, and restores the name used for this species in over five The name of the coffee bean weevil, a worldwide hundred publications worldwide for over 200 years. pest of cocoa, coffee, and dozens of other dried plant This also ignores the discrepancy between DeGeer’s materials, has recently been changed from Araecerus publication and collection, it assumes that Schön- fasciculatus (DeGeer 1775) to Araecerus coffeae herr (the most outstanding weevil specialist of his (Fabricius 1801) by Zimmerman (1994). Zimmer- time) knew what he was doing when he curated man points out that DeGeer’s original description DeGeer’s collection, it provides nomenclatorial sta- and illustration do not resemble the coffee bean bility for a worldwide pest, and it returns the name weevil, and he concluded that the name Curculio coffeae Fabricius to synonymy. The last point is fasciculatus DeGeer was based on some unknown important because even if fasciculatus is invalidat- beetle. He chose, from several available synonyms, ed, the name coffeae is a junior synonym of several Anthribus coffeae Fabricius 1801, as the replace- older names and should not be used. There has been ment name. These actions are not acceptable for the so much confusion about the identity of C. fascicu- following reasons: the DeGeer collection is pre- latus and its synonyms, I have designated lecto- served in the Naturhistoriska Riksmuseet in Stock- types for all pertinent names. This clarifies both the holm, and was curated by C. J. Schönherr who taxonomy and nomenclature of this economically marked the original DeGeer specimens with small important species. The history of this taxon is de- red (now faded) pin labels. He also discarded some tailed below. specimens and replaced others, but these did not receive the red labels. The collection now contains Araecerus Schönherr three specimens under the name fasciculatus, all with Schönherr’s red pin labels; all are coffee bean See Valentine (1999: 252), for generic synonymy. weevils. I now designate as lectotype of Curculio fasciculatus DeGeer 1775, the male bearing a small The following diagnosis will separate this genus round, British Museum lectotype label (placed by from all 650 genera of World Anthribidae. Antennal me in 1970, but not published). This action binds the insertion dorsal, next to the inner, apical margin of 248 Volume 19, No. 4, December, 2005, INSECTA MUNDI the round eyes which have a small notch at the end internal sac with a group of about 5 small sclerites, of the suprascrobal carina, latter not continued the two largest triangular. posteriad along the inner eye margin; antennomer- Much of the economic literature about this spe- es 5 - 7 symmetrical, not swollen on one side; three cies is listed in the bibliography compiled by Childers club articles small, very loosely articulated; head and Woodruff (1980). not retractile into prothorax, the eyes too wide; prothorax not constricted at base, transverse carina Curculio fasciculatus DeGeer present, basal, lateral angle rectangular, lateral carina not reaching supracoxal suture; short, basi- DeGeer, C. 1775: 276, species #10, pl. 16, fig. 2. lateral carinula present but not or weakly project- ing as an isolated lobe below the humerae; ventro- Type locality: “Surinam”. lateral corners of prosternum pubescent, not gla- brous; scutellum small but visible, not vestigial; Discussion: Lectotype, here designated, a male in elytra with 10 plus scutellar striae; hind coxae the DeGeer collection, with a round British Muse- elongate transverse; all tarsi longer than 1/2 a tibia; um lectotype pin label; paralectotypes: male and male pygidium almost vertical, apex broadly round- female; all three with Schönherr’s red (now pink) ed, female pygidium oblique, apex with a blunt pin labels indicating original DeGeer specimens, point; male metastenum with a weakly-developed these placed below a separate, not original, label median trichobothrium, absent in females; male “10. C. fasciculatus / p. 276”. fore tibiae and tarsi with sexual modifications in- volving length, curvature, rows of asperities, apical Bruchus cacao Fabricius spurs, or areas of dense erect setae. Araecerus has about 75 described species, all in Fabricius, J. C. 1775: 64. that Indo-Pacific area circumscribed by India, Ja- pan, Australia, and Hawaii. One species, Araecerus Type locality: not given; “in Theobromae semini- fasciculatus (DeGeer) is now a worldwide pest, and bus”. has been described as new by several authors. The species is diagnosed below, followed by the syn- Discussion: Holotype female in the Fabrician onyms in chronological order, using the original “Kiel” collection labeled “cacao”. Olivier (1795: 15), name combinations. All the names, except Bruchus who worked closely with Fabricius, says about ca- crassicornis Fabricius are, or appear to be, syn- cao: “Il se trouve á l’Amérique méridionale”. onyms of Araecerus fasciculatus (DeGeer). The species of Araecerus are variable and diffi- Note: I have not been able to determine the priority cult to characterize. Araecerus fasciculatus (De- of the above works and have accepted synonymy Geer) has the following features: dorsum without proposed by Schönherr, 1833: 175. If the name tubercles, tufts, crests, or costae; antennal club very fasciculatus is discarded, the oldest replacement slender, asymmetrical, each article tapering at both name is cacao, not coffeae. ends; prosternal antecoxal strip not linear, rough- ened with fine microsculpture and not coarsely Bruchus peregrinus Herbst punctured; pronotum finely reticulate-punctate; lateral carina not reaching supracoxal suture, fre- Herbst, J.F.W. 1797: 168, pl. 106, fig. 9, F. quently weakly upturned at apex; elytral pubes- cence tessellate dark and light brown and gray plus Type locality: “Amerika”: my translation is: brought a small, paler postscutellar spot; legs with tibiae with seeds from America; in “Mus. Herbst”. relatively long and slender, with four (sometimes very vague) dark annuli or spots often best devel- Note: I have not seen type material, however, syn- oped on hind legs; males with fore tibiae not dentic- onymy with coffeae Fabricius was proposed by Schön- ulate along ventral length, without an apical spur, herr (1833: 175). The Schönherr collection has a and not bent or hooked; male fore tarsus with basal female coffee bean weevil labeled “A. peregrinus / article elongate, narrow, setose ventrally, not bent Hbst. e Cayana. / Chevrolat.” This individual lacks or arched upward in center, about as long as articles a head and is grayer than most, but it has the other 2-5 combined; male mesocoxa without a small tooth features of fasciculatus. The synonymy was accept- mesal to the attachment of the trochanter; male INSECTA MUNDI, Vol. 19, No. 4, December, 2005 249 ed by de la Porte (1840: 288), Sturm (1843: 181), Sehested collection, the Essequibo [British Guiana] Taschenberg (1869: 118), and subsequent authors. specimen is excluded, and the Kiel collection speci- men is probably excluded. I here recognize the male Bruchus capsinicola Fabricius without data in the Sehested and Tonder Lund Collection as the holotype. The specimen is in good Fabricius, J. C. 1798: 159. condition, lacking only the left antenna.

Type locality: “in Cajennae seminibus”. Amblycerus japonicus Thunberg

Discussion: Lectotype, here designated, the more Thunberg, C.P. 1815: 122. complete of two males in the Fabrician “Kiel” collec- tion (lacks right elytron, left elytron adherent to pin; Type locality: “Habitat in Japonia”. Holotype in the paralectotype lacks head and prothorax) labeled the Zoological Museum, Uppsala University, Swe- “capsini / cola”. den (Wallin and Wallin 1989: 16), but I have not seen it. Note: In the same publication, Fabricius (1798: 159) described Bruchus crassicornis which is also a spe- Note: although synonymized with coffeae by Schön- cies of Araecerus, but is clearly distinct from fascic- herr (1839: 273), the identity of this taxon is uncer- ulatus. This species has been widely misidentified, tain. It may be synonymous with Araecerus fascicu- so some brief comments are provided here: latus (DeGeer), or with A. tarsalis Sharp (1891: 323), also described from Japan. The two species Bruchus crassicornis Fabricius share tibiae with dark spots or rings; however, according to Morimoto (1978: 26, 28-29) males of Fabricius, J. C. 1798: 159. fasciculatus have the “fore tibiae simple or at most with a row of minute granules on the underside” Type locality: “in Indiae orientalis seminibus. D. while tarsalis males have “fore tibiae serrate or with Daldorff.” tooth-like granules on the underside.” Also, the internal sac of male fasciculatus has a pair of small Discussion: Lectotype, here designated, one of triangular sclerites plus others which are absent in three damaged females, the first in the series, with tarsalis. the elytra spread by the pin, in the Fabrician “Kiel” collection; paralectotypes: the other two females. Anthribus alternans Germar This distinct species has shorter, non-annulate, red tibiae, antennal club segments short, fat, blackish, Germar, E.F. 1824: 175. almost symmetrical; antecoxal strip of prosternum very short; pronotum with disc more inflated and Type locality: “Habitat in Brasilia.” with three weakly-defined longitudinal pale stripes, the dorsolateral pair much wider than the narrow Discussion: Germar’s Anthribidae are in the Mar- median line. tin-Luther Universitat, Halle, Germany. They are included in the catalogue of weevils by Taschenberg Anthriubus coffeae Fabricius (1869: 115-118), who lists: “A. coffeae F. = peregri- nus Hbn. = alternas [sic] Germ. 7 Expl. Rio de Fabricius, J. C. 1801: 411. Janeiro [sic].” A male and two females have green printed labels with “Rio d. / Pedr.”, and four others Type locality: “In Indiae Coffea. D. Daldorff Mus. on three pins have no labels. Many specimens of D. de Sehested”. other species have very similar green labels with “ Rio d. Jan.” so I believe Taschenberg was careless Discussion: There are two males in the Sehested when he cited Janeiro instead of Pedras. The seven and Tonder Lund Collection, one without data, one individuals are adjacent to a hand-written green labeled “Essequibo. / Smidt. / Mus. S. & T. L. / label: “Coffeae Fabr. / peregrinus Hbr. / alternans fascicula: / tus, DeG. / coffeae. Fabr.” There is also a GM / Bras.” They are coffee bean weevils. I here male in the Kiel collection labeled “coffeae”. Since select as lectotype of Anthribus alternans Germar Fabricius described the species from India in the the male with the green “Rio d. Pedr.” label, and I 250 Volume 19, No. 4, December, 2005, INSECTA MUNDI have placed a round British Museum lectotype label Coleoptera are in the “Mus. Roy. Hist. Nat. Belg., on the pin. The other six specimens are lecto- Brüssel”. When I visited the Institut Royal des paratypes. Sciences Naturelle de Belgique in Brussels in 1989, I saw no recognizable Thomson material, and was Phloeobius griseus Stephens told that, at least for anthribids, Horn and Kahle were incorrect. On the other hand, the account of Stephens, J.F. 1831: 211, pl. XXI, fig. 2. This is not this museum in Arnett, Samuelson, and Nishida Phloeobius griseus (Fabricius, 1792: 377), see discus- (1993: 24) mentions holdings from the collections of sion below. Thomson and Oberthur. Based on these items, it appears that some beetle groups from Thomson are Type locality: “I have taken a specimen in London; in Brussels, but not the Anthribidae. and others have occurred in Suffolk and in Devon- I consider this taxon a probable junior synonym shire”. ofAraecerus fasciculatus (DeGeer) because, although the genus is not native to Africa, A. fasciculatus Discussion: Stephens appears to have had coffee occurs continent-wide. I have seen dozens of speci- bean weevils that he misidentified as Anthribus mens from Gabon, Cameroon, Zaire, Ghana, Togo, griseus Fabricius (1792: 377), a species transferred Nigeria, Angola, South Africa, and Somalia, and it to Phloeobius by Schönherr (1823: 1135). I agree has been reported from Kenya (Jordan 1914: 344), with the synonymy first proposed by Schönherr Tanzania (Kolbe 1897: 291), Rwanda (Jordan 1955: (1839: 273). 347), and Malawi (Lee 1971: 2). Identification of this species in Africa is compli- Cratoparis? parvirostris Thomson cated by the recent report (Frieser 1993:41) of a second species, Araecerus suturalis Boheman (1839: Thomson, J. 1858: 113. 273), type locality: Calcutta, India, which was reared from dry male cycad cones in South Africa. A male Type locality: By inference and present designa- and female from this series, determined by Frieser, tion: Gabon, West Africa, based on Thomson’s pub- have been received in exchange from the National lication of this species in his “Voyage au Gabon”. Collection of Insects, Pretoria, South Afica. They Holotype unknown, see following discussion. differ from A. fasciculatus as follows: tibiae dull reddish-brown, non-annulate; pronotum more con- Discussion: A male coffee bean weevil labeled vex; elytral pubescence without a post-scutellar “Gabon” is in the R. Oberthur collection in the pale spot; male fore tibiae with a ventral row of Museum Nationale d’Histoire Naturelle in Paris. It denticles; male fore tarsi weakly depressed, the bears the above name and fits Thomson’s short basal article weakly arched and bent (more convex description. There is no indication it is a type, but on dorsal and exterior surfaces) with a few long this is the only individual I have seen with this name ventral setae. They differ from the male holotype of in the many collections examined. A. suturalis Boheman, in Stockholm, because the Horn and Kahle (1936: 279) and von Hayek latter has broadly depressed, densely setose fore (1989: 83) reported that in 1865 James Thomson tarsi, and longitudinally striped elytra, interstriae began to sell parts of his huge beetle collection of 1-3 and 7-11 are dark, 4-6 are pale, interstriae 3, 7, over 35,000 species. Some families went to René 8, 9 with 2 to 6 pale spots each, interstriae 1, 2, 10, Oberthur, who in turn sent material to Neervoort 11 uniformly dark; male fore tibiae unarmed. Males van de Poll and Maurice Sedillot. The van de Poll of all three species lack mid-coxal tubercles. The collection was purchased by the Tring Museum in prosternal antecoxal area of A. fasciculatus and the England and was amalgamated with Karl Jordan’s South African specimens lack coarse punctures, anthribids and then combined with the British this condition is unknown for A. suturalis. It is clear Museum (Natural History) to form the most impor- that two introduced species of Araecerus occur in tant collection of Anthribidae in the World: 180 South Africa: fasciculatus (DeGeer) and suturalis of drawers, housed in London. The Sedillot Anthri- Frieser, not Boheman. bidae (which include those from the Chevrolat col- lection) is in the Paris Museum. I have not recog- Araecerus seminarius Chevrolat nized Thomson types in either collection. Horn and Kahle also state that Thomson’s Voyage au Gabon Chevrolat, L.A.A. 1871: 7, pl. 1, fig. 5. INSECTA MUNDI, Vol. 19, No. 4, December, 2005 251

Type locality: Paris, France, apparently imported Wolfrum (1929: 107) lists another synonym: the from Brasil. species described as Brachytarsus niveovariegatus Roelofs 1879:1v, from Japan (spelled “nigrovariega- Discussion: The female holotype in Paris is la- tus” by Bovie (1906: 313)). The species is not in beled: “_ _ [illegible] / vivant a / Paris” written on a Araecerus, it belongs in Anthribus Forster 1770, green label (green was a widely used code for the occurring in Japan, China, and Korea. Chinese Neotropics). There is also a separate box label: specimens were redescribed as a new species: An- “Araecerus / seminarius Chvt / Brasilia D. D. Huet” thribus lajievorus Chao 1976: 339, new synonymy. I written on green paper. Chevrolat states “Acun have seen a topotypic paratype, and specimens from renseignement bien précis ne m’a été fourni sur la all three countries. mode de vie de cet insecte”. The holotype is a normal coffee bean weevil. References cited

Tropideres (Rhaphitropis) mateui Cobos Arnett, R.H. Jr., G.A. Samuelson, and G.M. Nishida. 1993. The Insect and Spider Collec- Cobos, A. 1954: 41, fig. 1. tions of the World. Flora & Fauna Handbook No. 11. Sandhill Crane Press, Gainesville, FL. vi+ Type locality: Spain: “Tiana (Prov. de Barcelona); 310 p. J. Mateu, coll.” Boheman, C.H. 1839. [Descriptions]. In C.J. Schön- herr, 1839, which is cited below. Discussion: Holotype female in the Instituto de Boisduval, J.B.A. 1835. Faune entomologique de Aclimatación, Almeria, Spain. Cobos states the l’Océan Pacifique, avec l’illustration des insect- unique type is a male, but his illustration clearly es nouveaux recueillis pendant le voyage. Duex- shows the pointed pygidium characteristic of fe- ième partie, coléoptères et autres orders. Voy- males. Synonymy by Valentine (1999: 252). ages de découvertes de l’Astrolabe. Exécuté or- It is difficult to understand why Cobos placed dre du Roi, pendant les années 1826-1827-1828- this taxon in the wrong subfamily, aligned with two 1829, sous le commandement de M.J. Dumont genus-group names that are unrelated and fall into D’Urville. Paris, 716 p. very different tribes. Although the type has not been Bovie, A. 1906. Catalogue des Anthribides. An- examined, the description and figure convince me nales de la Société Entomologique de Belgique, that this is a coffee bean weevil. 49(11): 218-334. Chao, Y-C. 1976. A new species of Anthribus For- The ten synonyms discussed above do not com- ster predaceous upon the Chinese wax scale plete the picture. Zimmerman (1994: 205,208) lists (Coleoptera: Anthribidae). Acta Entomologica Araecerus sambucinus Boisduval 1835: 299, and Sinica, 19(3): 339-341, fig. 1. [In Chinese and Trepideres [sic] fragilis Walker 1859: 220, as syn- English]. onyms of A. coffeae (Fabricius), for the former citing Chevrolat, L.A.A. 1871. Description de six synonymy by Dejean (1837: 259 ), [actually 1836] Coléoptères exotiques. Annales de la Société and for the latter, synonymy by Pascoe (1859: 438) Entomologique de Belgique, 14: 5-8, pl. 1, fig. 3a, [actually p. 439]. I have not seen Boisduval’s origi- b. nal description, and the location of his type, if it still Childers, C.C. and R.E. Woodruff. 1980. A bibli- exists, is unknown. The type of Walker’s fragilis is ography of the Coffee Bean Weevil Araecerus a male from “Ceylon” in the British Museum, Lon- fasciculatus (Coleoptera: Anthribidae). Bulle- don. When I examined it, I did not know enough tin of the Entomological Society of America, about the genus to form a strong opinion about its 26(3): 384-394. identity, except that it was clearly in the group of Cobos, A. 1954. Dos especies nuevas de Tropideres species that includes fasciculatus, lutatus (Fair- Schönh. (Col. Anthribidae) de España. Archivos maire) (=vieillardi Montrouzier 1860, of older au- del Instituto de Aclimatación, 3: 41-44, fig. 1, 2. thors), and others. A male Ceylon topotype looks DeGeer, C. 1 775 . Me mo ir e s p o u r s e rv ir a l’Histoire like fasciculatus with simple fore tibiae and non- des Insectes. Vol. 5. Stockholm, Pierre Hessel- punctured antecoxal strip of the prosternum; but it berg, viii+448 p., 16 pl. has tuberculate mid coxae. The identity of fragilis Dejean, P.F.M.A. 1834. Catalogue des Coléoptères Walker remains unsettled. de la Collection de M. le Compte Dejean. Paris, 252 Volume 19, No. 4, December, 2005, INSECTA MUNDI

Méquignon-Marvis Père et Fils. Livraison 3: Jordan, H.E.K. 1914. Insectes Colopteres. X: An- 177-256 [includes fascicules 23-32]. The other thribidae. Voyage de Ch. Alluaud et R. Jeannel livraisons span 1833-1836, p. 1-443. This, the en Afrique Orientale (1911-1912). Résultats Sci- third of Dejean’s catalogues, was mostly de- entifiques, Coleoptera, X: 327-346, 1 photo. stroyed by fire before distribution; anthribids Jordan, H.E.K. 1955. Contributions à l’étude de la on p. 234-237. fauna entomologique du Ruanda-Urundi (Mis- D ej e an , P . F . M . A. 1836. Catalogue des Coléoptères sion P. Basilewsky 1953). LXVIII. Coleoptera de la Collection de M. le Compte Dejean. Paris, Anthribidae. Annales du Musée Royal du Congo Troisième Èdition, Revue, Corrigée et Aug- Belge. Sciences Zoologiques, 40(2): 338-347, fig. mentée. Livraison 1-4, 1836: 1-384; Livraison 5, 1-11. 1837: 385-503. This is the replacement “third” Kolbe, H.J. 1897. Coleopteren. Die Käfer Deutsch- edition; anthribids on p. 256-259. Ost-Afrikas. Berlin, Verlag von Dietrich Re- Fabricius, J.C. 1775. Systema Entomologiae, sis- imer, IV: 1-367, pl. I-IV. tens Insectorum Classes, Ordines, Genera, Spe- Lee, R.F. 1971. A preliminary annotated list of cies ... etc. Flensburgi et Lipsiae, Libraria Ko- Malawi forest insects. Malawi Forest Research rtii, [i-xxxii], 1-832. Anthribids on p. 63, 64, 151. Institute Research Record, 40: i-viii, 1-132, 1 Fabricius, J.C. 1792. Entomologia Systematica folded map. Emendata et Aucta.... etc. Hafniae, Christ. Got- Morimoto, K. 1978. The family Anthribidae of tl. Proft. 1(2): 1-538. Bruchus, p. 369-374, An- Japan (Coleoptera). Part 1. Esakia, (12):17-47, thribus, p. 375-378. fig. 1-11, photos A-R. Fabricius, J.C. 1798. Supplementum Entomologi- Pascoe, F.P. 1859. On some new Anthribidae. ae Systematicae. Hafniae, Proft et Storch, p. i- Annals and Magazine of Natural History iv, 1-572. Bruchus, p. 158-160, Anthribus, p. [3]4(24): 431-439 [reprint p. 7-15]. This is the 160-161. second part of an article in three consecutive Fabricius, J.C. 1801. Systema Eleutheratorum issues of the Ann. Mag. Nat. Hist.; two plates secundum Ordines, Genera, Species ... etc. Kil- appeared, with part 3 (1860). iae, Bibliopolii Academici Novi, p. 1-687. Bru- de la Porte, F.L.N. de C. [also cited as Comte de chus, p. 396-403, Anthribus, p. 404-412. Castelnau] 1840. Histoire Naturelle des Insect- Frieser, R. 1993. New species and new records of es Coléoptères. Vol. 2. Paris, P. Duménil, p. [i- Anthribidae from South Africa and the Comoro- iv], 1-563, [1]. Anthribidae on p. 283-288, pl. 21, Islands (Coleoptera, Anthribidae). Acta Co- index p. 530. leopterologica, 9(2): 31-50, fig. 1-4. Roelofs, W. 1879. Diagnoses de nouvelles especes Germar, E.F. 1824. Insectorum Species novae aut de Curculionides, Brenthides, Anthribides et minus cognitae, descriptionibus illustratae. Bruchides du Japon. Annales de la Société En- Halae, J. C. Hendelii et Filii. Volume 1, Co- tomologique de Belgique, 22: Bulletin, p. LV. leoptera, p. i-xxiv, 1-624, pl. 1. Roelofs, W. 1880. Additions a la Faune du Japon. von Hayek, C.M.F. 1989. A short biography of the Nouvelles espèces de Curculionides et familles entomologist James Thomson and the dates of voisines. Observations sur les espèces déja pub- publication of the Archives Entomologiques, liées. Annales de la Société Entomologique de Arcana Naturae, Monographie des Cicindéil- Belgique, 24: 29. ides, Musée Scientifique and Physis. Archives of Roelofs, W. ?1888 or later. De Japansche Curcu- Natural History 16(l): 81-99. lionidum -Fauna vergeleken met die van andre Herbst, J.F.W. 1797. Sechs und sechzigste Gat- landen. Tijdschrift voor Entomologie, 32: 20-28. tung der Käferartigen Insecten. Anthribus. Na- Schönherr, C.J. 1823. Curculionides. Isis von Oken, tursyste aller bekannten in- und auslandischen 1823(10): 1132-1146. Insecten, als eine Fortsetzung der von Büf- Schönherr, C.J. 1833. Genera et Species Curcu- fonchen Naturgeschichte, 7: xi+346, col. pl. lionidum, cum synonymia. hujus familiae. Pa- XCVI-CXVI, Q-U. Anthribus list, index, plate risiis; Roret, 1(1): i-xv, 1-381. On p. 175, the numbers on p. vi, text on p. 156-169, plate species fasciculatusDeGeer,peregrinus Herbst, CVI[106]. cacao Fabricius (and incorrectly, crassicornis Horn, W. and I. Kahle. 1936. Über entomologische Fabricius) are combined in the synonymy of Sammlungen. Teil 11. Entomologische Beihefte coffeae Fabricius. aus Berlin-Dahlem, 3: 161-296, pl. XVII-XXVI. INSECTA MUNDI, Vol. 19, No. 4, December, 2005 253

Schönherr, C. J. 1839. Genera et Species Curcu- 1856 et en 1857 par M. Henry C. Deyrolle sous lionidum cum synonymia. hujus familiae. Pa- les aspices de MM. Le Comte de Mniszech et risiis; Roret, 5(l): i-viii, 1-456. On p. 273, the James Thomson précédée de l’histoire du voy- species sambucinus Boisduval, japonicus Thun- age. Insectes. 1. Ordre Coléoptères. Archives berg, and griseus Stephens are added to the Entomologiques, 2:29-239, pl. i-viii. synonymy of coffeae Fabricius. Thunberg, C.P. 1815. De coleopteris rostratis. Sharp, D. 1891. The Rhynchophorous Coleoptera Nova Acta Regiae Societatis Scientiarum Upsa- of Japan. Part II. Apionidae and Anthribidae. liensis, 7: 104-125. Transactions of the Entomological Society of Valentine, B.D. 1998 (1999). A review of Nearctic London, 1891(2): 293-328. and some related Anthribidae (Coleoptera). In- Stephens. J.F. 1831. Illustrations of British Ento- secta Mundi, 12(3+4): 251-296, fig. 1-23, tab. 1. mology; or, a Synopsis of Endigenous Insects. . Walker, F. 1859. Characters of some apparently . etc. Mandibulata. Vol. IV. London; Baldwin undescribed Ceylon Insects. Annals and Maga- and Cradock, p. 1-413, Col. pl. xx-xxiii. zine of Natural History [3]4(21): 217-224. Sturm, J. 1843. Catalog der Kaefer-Sammlung von Wallin, L. and H. Wallin. 1989. Catalogue of type J. Sturm. Nürnberg; Rosten des Verfassers, 386 specimens. I. C. P. Thunberg (1743-1828), In- pp., 6 pls. Anthribids on p. 179-182. secta. Uppsala University Zoological Museum, Taschenberg, E.L. 1869. Verzeichniss der im zool- 83 pp. ogischen Museum der Universitat Halle-Wit- Wolfrum, P. 1929. Anthribidae. In Junk, W. and S. tenberg aufgestellten Russelkäfer. Zeitschrift Schenkling. Coleopterorum Catalogus, Pars 102: für die Gesammten Naturwissenschaften Halle, 1-145. 33: 1-248. Anthribidae p. 115-118. This journal Zimmerman, E.C. 1994. Family Anthribidae Bill- citation may be incorrect, but the anthribid berg, In Australian Weevils (Coleoptera: Curcu- page numbers are accurate. lionoidea). Vol 1. Orthoceri. Anthribidae to At- Thomson, J. 1858. Voyage au Gabon. Histoire telabidae. The Primitive Weevils, p. 9-11, 13-23, Naturelle des Insectes et des Arachnides re- 35-239, fig. 1- 162. cueillis pendant un voyage fait au Gabon in 254 Volume 19, No. 4, December, 2005, INSECTA MUNDI