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rgt (886r)6rt S8S:(8)9 ersdnN N i\tsra Vnl.6. N..3 (19881

Methods and presentation

The measurements of the morphological characters and the subsequent descriptions were taken from 650 herbarium specimens. In the descriptions, those character states which occur in fewer than 10% ofthe individuals (ofthe relevant taxon) are enclosed bv parentheses. Parentheses are also used to enclose rarely occurring character states which may be present in an otherwise typical individual specimen.

Generalterminology follows Lawrence (1955),Porter et al. (1973),and Stearn (1973). Terminology for plane shapesfollows Ball et al. (1962).Author and literature abbrevi- ations follow Stafleu & Cowan (1976, 1979, 1981, 1983, 1985).

The distribution summary and the selected citation of specimens examined are grouped according to various regional subdivisions. The subdivisions used fbr the States are:for QueenslandI have fbllowed the pastoral divisions as in Contr. Queensl.Herb. 19 (1975)back end paper, for New South Wales those of Jacobs& Pickard (1981) (which is modified from Anderson 1961),for Victoria those of Cochraneet al. ( 1968),for South those ofLaut et al. (1977a-f),and for those ofBeard (1980).

Morphological characters

Bentham noted that'the species lof Westringial are so closely allied, and run so much into each other as to render it exceedingly difficult to assign to them any tangible characters'(Bentham 18?0,p. 127).This has proved to be particularly true lbr the three taxa discussedin this paper. The type of field observations required for are the same as those needed for all Labiatae. As similar obseryations are required for both Westringia and, Prostanthera, Conn (1980) should be referred to for a discussion of the type of obser- vations required. The lack of useful field observationson most of the herbarium collec- tions of Westrlnglo has made it difficult to evaluate many characters. Only about 30% of the collections examined have information other than the localitv of the collection. Even lesshave information which can be usedto evaluatethe taxonomic valueot certain characterswhich are not readily represenledby herbarium specimens.

Boivin (1949) discussedthe usefulnessof various morphological charactersand his paner should be consulted for further details.

Taxonomic characters currently used

The characterswhich have been most frequently usedto distinguish betweenthe taxa oi this complex are:the number ofleavesper whorl, the sizeand shapeofthe leaves,and the size of the calyx lobes.

The number of Leauesper ri,horl.Although the number of leavesper whorl is a useful character fbr delimiting certain Westringia species(see Boivin 1949), it appears to be of limifed taxonomic value in this complex.For example,some specimensof W. eremicolq have 3 and 4 leaves per whorl (namely Chipstone 25, CopLey 4504, Hunt 233). W. eremicola and W' rlgido usually have 3 leaves per whorl. The Western Australian popu- laiions of !7. dampierl usually have either 3 or 4 (rarely 5) leaves per whorl, whereas the South Australian populations ('W. greaiLtina')appear consistently to have 3 leavesper whorl. So, because of the overlap and the variation even within , it is concluded that the number of leaves per whorl is useful as a secondary feature which mav confirm an initial determination within this complex. 'aJueuodult 'lsaq 'aq 'aloqi\ drBpuorasJo lE o] rBaddEplno,r lr B sBxalduroJ aql JoJjnq 'Dla)lruala urof,!Dpt31J SulqsrnSullsrp JoJ asn auosJo aq {€ru s8urlJ€ur pu€ Jnolo) A4 Z1 'aldJnd 's5ur{JBur BIIoJo) aqJ's8ur{JBur lnorllrd\ )EIII oJ a^nEur alBd alpd Jo u,roJq 'aldrnd ,$ollad JolpuE ,raolla{'poJ q]r,,!\ alrq,\\ raqlra st Jr acurs InJasn f,Ja^ oq lou {eru elloroc aql ?Ja?d?Dp Al uI €rlBrlsny rllnos'qlnos oupuotr\l IE ola)lLuara'll Jo uorlpl -ndod p ur sJa,!\oUalrq,!\ puB a^nBur palou Ii0I uuo, s8ur{J€lu lnoqlr.{\ Jo q}r.r\ sBJIoro) ( p!rr?]qs) alq,,r ^larEr 'aldrnd ro a^neur 'J"H a^ey plo)luara s5ur{rEur 7,6tI 'Ja^a,roH11 {rEl qrrqn sEllorof, a^nBtll puE JBIrIproJar z69z runH pldB00lz Jaup.toc sbur{J€ur u,\\orq aFuEJool a8upJo qll,']a(aFurl a^n€ru qll,$ ualJo) s€lloroc allq^r a^eq Dp!8!J A4 '(Jpr\rurur g g9 iuol rutu '5uol '/1 '5uol B-Z LualduDp 14 :apri\ urul -c-Zt ulur i'l-Z t Dpr8rr :aprd\urur 6 t-9 2 urur B'E-g ZDlnlLuara 1!1flaurEu) uatdutpp l\l pue Dp131J14 r{loq ueq.} JalJ€rrrssr Dlo)rwala '14 rrPd-aqol uBrpalu aql 'rEIn3llJEd uI (8uol urrr 'Jr Buol ulur g Jo IprxBpB 'A4 ZI-B 8-9) uadruDp A4 ueq], selloroJ rallEurs a^erl,{ll€nsn pp?8?r pue Dl@luara Al qtpfI DlloroJ 'aolaq '€xPJ passnrsrpaJB aaJql aql Jo uorlPJturl -ap aql ur lsrss€ 01 readd€ qf,rq,r sral)Er€qc IBuorlrppe asaqJ r{$ual alorlad pup :saqol aql azrs aql snld 'slualuelq ppoulurEls uo urnluaunpul aJuasqs ro IBpouruels Jo 'azrs Jo aJuasaJd:lrqEq :s8ul{J€ur pup rnoloJ €lloJoD :ale ,{aqJ xalduof, srql Jo dulouox€J aql olur slq8lsur raqFnJ raJJo r{rrq,$ sJol)eJEr{J [euorlrppE palpo^ar sEq ^pnls stqJ

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'uuof P.L8ur.asarfi f. ,(rrpU 338 Nultsia Vol. 6, No. 3 (1988)

Calyxlobeto tuberatio

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(Euorullu e o rnoqe 'a'r) alBlorlad,{llloqs ,{raA arE qJrrl,u asoql puE alrssasaJB qcrr{na saAEal uaa,ulaq qsrnBurl 'lnJasn -srp ol llncgJrp sarurlaulos sr Jr ocrlJEJd ur ,{llEJrurouox€l aq ol sJ€addB.r?Jf,BJpqf, '8uol srql q8noqllv (dla^rlradsar ulru 1,0 g 0 pu€ Suol urur 9 0-t 0 alorlad) alelorlad '1,1 ,rl$ual at€ Dlo)17uala A4 pue ua|duDp sBaraq^\ so^€al alssas seq op!8!J Al aplpd '{JBSSaJausr uorl€nle^a ;aq1;ry q8noql ua^a'lnJasn fllB)tulouoxel aq 01 JBaddEsralJer€q, asaqJ (8uol urur -6'OualdluDp pueSuolrulug'O g'ODloJluala '3uo1ruur I ,14 lt g'6-7 6 op?8?J .{[arueu) ex€] aarq] aql uaa^\laq azrsur saJuaraJJrp i!\oqs 14 'dJreq Iprus saqol ppourrllels aqJ aspq aql lp lseal le ale ualdutDp puB Dluu.uara Io ',{rrEr[ A4 M asoql sEaJaq^\ Jo snoJq€lF arE Dp731JM Jo s]uall.rBlrJIBpourur€ls aqI SapoulruDjS '(8rrd'086r) 'J€ raJoqtlv d T {l puE (I89 d'286I) la ru€qFuruunCraJar sarcads srql;o sqde.rSoloqd Jod lualBd uallo aJe saA€olpr8u aqJ pal5uBl LEr{,raurosualJo puB pr8u aJEsaqJuBrq 'qnJqs aqJ adpqs ur lBJuaqdsruraq ualJo pu€ lr€duroD flqBuosEal aleJulur Jo al€c -uB^rp pauuralsrqlnru E.{[Ensn sr rp?t?J dl luauruoJrAua IpJSEoJssalro aJour s]r olpalEl -aIIof,,{lqrssod sr qJrrl,,!\',.{qsau, fl}qErts aq ()1spual. osle sarJadssrq; sapouralur poqs 'lJEdruoJ fla^rlEloJ aql Jo asnEJaq,{llJed sJEaddBqnJr{s aqJ quqs asuap [raA e f,ll€nsn s ualdLuDp (1,(DlhlLuara DuUlloar8DlSutrlr^a[, se pallaqel orn8u] d '186I) AI '086I)L) 'd 69Z su€urratsoCpup (6lg d raJoqllv raJar sarf,adssrql lo sqdBr5oloqdrod 'srxp T O 'Furpuaf,s€ Japro Jaqgrqlxau arll ol "gl uEql ssalal8up ue 1€ ,(lFnsn aJEsaqJu€.rq pu€ sa^pal oq,L ocuBJ€addB uB sBq qJrqn^ qnrqs fqsnq saturlaruosro 'Japuals .lJos, IlB.ra^o e sr plorl?uara 14 paAJasqouaaq a^eq sa?uoJoJJtpa^rlBlupnb Sul,rolloJ aqt',qrl -uEnb (l1 'sarrads (s)ralJ€rer{t llncrJJrp B sr LrqBil qgnorlllv asaql Jo uorlprUrss€lr aql ur pre 01 uorlEurroJur lEuorlrpp€ InJasn lsotu aql apt^oJd ol r{la{rl suraaslrq€q aqJ l?gol{

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1.Westringia dampieri R. Br., Prodr.501 (1810);J.D. Hook., Bot. Mag. t.3308 (18i1,1); Benth..Labiat. Gen. Spec.458(1E34); Barll. in Lehmann,Pl. Preiss.1: ll61 (1845); Benth.in DC.,Prodr. 12: 570 (1848); Fl. Austral.5: 129 (1870); F. Muell.,Fragm.9: 163 (18?5);Boivin, Proc. Rov. Soc. Queenslancl 60: 10?(1949 las 19501):Robertson in J.M. Black,Fl. S.Austral. 2nd. edn,,1: 742,fig. 1061(1957); Blackall & Grieve,Western Aus- tra1.Wild1l.3: 577 (1965); J.S. Ileard, Descr. Cat. WeslernAustral. PI.94 (s.dat. IOct. 19651);G. & P. Althofer.Austral. PI. 10:1161 (1980) las'Westr[ngia dampierli']; Grieve (ed.),Blackall & Grieve,Western Austral. Wildfl. 3I}: 430 (1981).Lectotype (herc chosen.i:R. Erorun s.n.lJ.J. Bennett2i381l. anno 1802 5 [8Dec. 1801 1Jan. 1802 (Stearn 1960)].King GeorgesSound, Western Australia (lecto:BM, upper left specimen; isolecto:BM, uppercentre & right specimens). W.clnereo R. Br.,Prodr. 501 (1810);J.D. Hook.' Bot. Mae.t. 330?(183'1); Benth. in DC., Prodr.12: 570 (l8il1). Lectotype (here chosen): -R . Brown s.n.lJ.J. Bennett23831, anno 1802-5[29 Jan. 1802-13Feb. 1802], Bay 3 - 4 5 - 7 [Fowler'sBay, Petrel Bay, Franklin Bav,Waldgrave Island (Stearn 1960)1, (lecto: BM, third specimenfrom left; isolecto:BM other specimenson sheet[excluding type]). W. greuilLinaF. Muell.,Defn. Austral. Pl. 16 ([July (Seberg1986)] 1855); Trans. Phil. Soc.Victoria 1: 49 ([September(Aston 1984)]1E55); J. Bot. Kew. Gard.Misc. 8: 169 (1856);Boivin. Proc.Roy. Soc. Queensland 60: i06 (19,19[as 1950]);G. & P. Althofer, Austral. PL 10: 363 (1980).Lectotype (here chosen):WiLhetmi s.n., s. dat., Cape Donningtonand Tungetta, Port Lincoln,South Australia (lecto: MEL 614327;isolecto: MEL 614328& MEL 614329). W. rigida R. Br. var. dotichophyllaOstenf.. Biol. Meddel.Kc-rngel. Danske Vidensk. Selsk.3: 112(1921): Blackall & Grieve,Western Austral. Wildfl. 3: 577(1965); G. & P. Althoier,Austral. Pl. 10:364(1980): Grieve (ed.), tslackall & Grieve,Western Austral. Wildfl. 38: 429 (1981).Lectotype (here chosen): Ostent'eld 97?, 29.x.1914, Geraldton (lecto:C lowerspecimen; isolecto: C upper specimens,PERTH). W. seniktLiaF. Muell.var. canescensBenth., Fl. Austral.5:130 (1670)..Lectoalpe (here chosen):Mn:m,ell 262, s. dat..Phillips Ranges, Western Australia (lecto: MEL 614549, left specimen;isolecto: MEL 614549,right specimen). lW. senifoLiaauct. non F. Muell. (1855);Blackall & Grieve,Western Austral. Wildfl.3: 5?8(1965).1

Smallshrubs 0.3-1(-1.3) m high.Bronches triangular to quadrangular,or subterete,a smoothor internodeswith raisedridges lrom axil of leaf to next more distal node, denselyhairy (c. 100-230hairs/mm'), hairs appressed, antrorse, simple, c. 0.2mm long. Leauesin whorls of 3 or,1(or 5), spreadinglo recurved,abaxial surface and petiole denselyhairy (150 200hairs/mm'), adaxialsurface densely hairy basally,sparsely hairy to glabrescentdistally; peflole 0.3 0.5 mm long;'Iamina narrowly ovate or narrowly ob- iongto linear,(8-)13 26(-40) mm long,(1 )1.5 3 mm wide(lamina length to width ratio (4-)7.5-15(-19.?),length of maximumwidth from baseto total laminalength ratio up to c.0.1), base cuneate, margin entire and recurved such that onlymidrib of abaxialsuriace visible,apex mucronate (mucro 0.3 0.5mm long);venation not visible,midrib slightly raised on abaxial surface. Inflorescence a frondose racemiform conflorescence, uniflorescencemonadic. PetliceL 0.2 0.8 mm long,densely hairy; prophyLls inserted at baseof calvx,narrowly ovate to nauowly oblong,1.4 1.5mm long,0.2-0.3 mm wide (lengihto width ratio 5 7), denselyhairy, basenarrowly cuneate or prophyllsnot re strictedat base,margin incurved, apex obtuse. (iolyr green, mid-vein of eachsepal thick- ened to form a ridge from baseto apex t.,feach calyx lobe. outer surfacedensely hairv,hairs appressed, antrorse, c.0.2 mm long;tube (2.5-)3.3-4.4(-5.5) mm long,inner suriaceglabrous; lobes depressed to verybroadly triangular, (0.5 )0.8-1.9mm Iong,1.1- 2.3mm wide (lengthto width ratio 0.5-0.8),inner surfacemoderately to denselyhairy, apexobtuse to subacute;(calyx lobes to tuberatio (0.1-)0.20.5( 0.6)\.CarotLa 8-12 mm long,white with purple,red, vellow and/or brown dots mediallyon abaxialsurface of tube and mouth,'or pale purple,pale mauveto lilac with dots absent;outer surface 'sarJads srql ul aDu€JrJru8rsJrurouoxBl alllrl Jo aqollqgnorlls! Iroqi radsaABalloJaq -uInu aqJ 'lroq^\ rad sa^Ealg a^eq ( gI8 ,{aqmaNpu€ a8roaC .{laurEu) suorlralloc ondl 'lJor{.$ 6tI 'JaAa puE rad sa^€ol I, puB t qloq aABqsuorlBlndod rallBl aql Jo suaulJads auros -,{{oH rad sa^pal seq (Brl€Jlsny uJolsa^\ ur EurJJnJJo sp) ualduiDp 'IJoq,{\ IJoqnA t,{llsoul Al Jad sa^€al I a^l?qo},{llualsrsuol parEaddBsuor?Elndod uerleJlsnv qlnoS aq:}paul -uIBXa 'lJoq,r I lEql lpualeur aql lp uI rad sa^l]all SurAEqsaur]auos sE (l,Du1n1nal8M, s€] erlBrlsnv qlnos ualdulDp saquJsap (196l '{J€lg ur) uoslJaqou (Dpuurara '/l Jo) /1 rapun tsaloN, raJar sarcads JaBBI aql Jo uorssnJsrp raqFnJ JoJ) erl€Jitsny qlnos ur rnDco saop Dlo)luala 111l€ql pue ualdruDp 1,1 Jo ur.{uouds B sr Dulnloat8 M iu\11, a^vq Duln1aard sB[\ BrlBJlsnv qlnos ur sB u^\ou{ uoxBl PapnlJuo) I /d ',{lluanbasqnsrr]lor??{aJa/4, 'uorurdo al{l lBt{l srur^rog uroJJ paurnssB (996l) JalqDrg s,ur^rog {Jo^a 'r{r€lg Surssncsrp1noq1ln uorlEJardralur srrallantr\lpa.{\ollol (r96I ur) uosuaqoH luour -ruoJ lnoqlrA BrleJlsnv q:lnos ruo{ Dloc|uala 14 papnlcxa puB DutltoarF 14 pa}B}sur 0r (6t6I) ur^rog ualdLuDp Io -{urouou{s aql ol uox€l srql paDnpar (9lgl) rafiantr{ 'uorsnJuoc l,l '/1 alqeJaprsuoc ul pallnsal sBq Dulll,toal8 Io uorlBlardralur arlJ sa?oN 'Gzt:r*. 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Itu Pl;rrr4sa1l uuoJ I, IrrEg NLiytsiaVol.6. No. ll (1988)

Mueller (1875),and more recently G. & P. Althofer (1980)regarded W. senifoLiavar. conescensfrom Western Australia as a distinct taxon from typical W. senifoLia fuomYic- toria. Boivin (1949) regarded W. senifoliavar. conescensas an indistinct form oftypical W. senifolia. This interpretation is regarded as incorrect since the two taxa are readily distinguishable. W. senifoLiahasleaves which are moderately to densely hairy on all sur iaces (although the abaxial surface is more densely hairy than the adaxial surlace), with spreadinghairs about 0.5-1 mm long. The brancheshave the sametype of indumentum. W. senifoLiavar. canescenshas denselyhairy to glabrescentleaves. The indumentum of the leaves of the latter taxon is sometimes restricted to the abaxial suriace. The hairs ofthe leavesand branchesare appressedand 0,2-0.3mm long. This latter taxon appears to be a small leafed form of W. dampieri. Canseruationsfotus. Considered not at risk.

2. Westringia rigida R. Br., Prodr. 501 (1810);Benth. in DC., Prodr. 12: 570 (1848); Fl. Austral.5: 129 (1870);Rodway, Tasman. F1. 150 (1903);J.M. Black. Fi. S. Austral. 3: 49,1,fig. 200E & F (1926);Ewart, Fl. Victoria 980 (1931 [as 1930]);Boivin, Proc. Roy. Soc.Queensland, 60: 107 (1949 [as 1950]);Itobertson in J. M. Black, Fl. S. Austral. 4:742, figs 1038E& F, i063C (1957);Blackall & Grieve,Western Auslral. Wildfl. 3: 57? (1965); Beard, Desc.Cat. Weslern Austral. PI.94 (s. dat. lOct. 19651);Curtis,StudenL's Fl. Tas- mania,3: 555 (1967);Willis, Handb. Pl. Vicioria, 2: 585 (1973 [as 1972]);G. & P. Althofer, Austral. Pl. 10: 364, & tt. (1980);Grieve (ed.) in Blackall & Grieve, Western Austral. Wildfl. 38: 429 (1981);Haegi in J. Jessop(ed.), Fl. Central. Austral. 311,fig. 413 (1981);Costermans, Native Trees & Shrubs SE Austral.268 & t. (1981);Cunningham et al., Pl. W. New S. Wales 581 & t. (1982 fas 19811).lectofype (herechosen): fr. EroLon 1.1t.lJ.J Bennett2382l, anno 1802-5 [January 1802 (Stearn 1960)],Bay 3 South Coast [GooseIsland Bay, Western Australia (Stearn i960)] (lecto: K -iower left specimen; isolecto:K - other specimenson sheet excluding type).

W. rigida var. breuifolia Benth. in DC., Prodr. 12: 5?0(1848). Lectotlpe (here chosen): Drummond I94, s. dat., Swan River (lecto:K central left specimen;isolecto: K - upper left and right specimens).

W. r[gidavar. brachl'phyllnOstenf., Biol. Meddel. Kongel. Danske Vidensk. Seisk.3: 112 ( 1921);Blackall & Grieve,Western Austral. Wildll.3:577 (1965);G. & P. Althofer, Aus- tral. Pl. 10: 364 (1980);Grieve (ed.) in Blackall & Grieve, Western Austral. Wildfl.38: 429 (1981).Lectotype (here chosen):OstenfeLd 982,7.x.1914, Kalgoorlie (lecto: C lelt specimen;isolecto: AD, C - centre and dght specimens,MEL, PERTH). Small shrubs,0.3-0.6(-1)m high. Bronclzessubterete with two slightly raised lateral ridges, sparsely to densely hairy distally, glabrous basally; hairs appressed, simple, an- trorse, c. 0.3 mm long. leaues in whorls of 3 (or 4), spreadingto recurved,sessile; lamlnc ovate to nauowly ovate, 1.9 5.2(-9.8)mm long, 1-1.7('2.2)mm wide (lamina length to width ratio (1.5-)2-7(-8.2),length of maximum widlh trom baseto total lamina lensth ratio 0.1-0.6),rigid, sparselyto moderatelyhairy, hairs usuallypersistent on abaxial sur- face, but adaxial surlace becoming sparsely hairy or rarely glabrous, base rounded to broadly obtuse, margin enlire and recurved such that abaxial surface almost concealed, apex subrnucronate to subpungent (rigid point c. 0.8 mm long); venation not visible, midrib slightly raised on abaxial surface. InfLorescence a frondose racemilbrm conflorescence,uniflorescence monadic. Pedicel 0.2-0.'/(-7) mm long, densely hairy; prophylls inserted near baseofcalyx, narrow, ovate to suboblong,1 1.5mm 1ong,0.3-0.5 mm wide (length to width ratio 2.7-5.6),moderately to densely hairy, base narrowly- cuneate, margin often slightly incurved, apex obtuse. Caly;r green, mid-vein ofeach sepal thickened to form a ridge irom base to apex of each calyx lobe, outer surface densely hairy, hairs appressed,antrorse,less than 0.2 mm long;tube 2.6 3.6 mm long, inner sur- face glabrous;Iobes depressedtriangular to very broadly triangular, (0.6 )0.9 1.6 mm long, 1-1.? mm wide at base (length to width ratio 0.7-1),inner surface moderately to denselyhairy, apex subacute;(calyx lobes to tube ratio (0.2-)0.3-0.5).Corolla 6-7 mm '.L-96I '7,0tI rapdo) '196l xr8 'us radooJ :(EI€ll€I :(OY) {aIU }urod ^ II taldo) I) '1,96I'xItZ'Z()g :(sure14roog) :(Oy) BtuooI/{JogSS ur{ ryoftlg t(PllnlaalA)surPld 'tS6I ' 9', JInC:(OV) euotJlA trod l!l^ 6 u s puDPlJ :(B_ruErn)elnsuruad a{ro^ uraqlnos :(qy) q-ouy e11e{q16'1161'xrI ''u's puDPlC:(BUPfqllA):(OV) BIIe{q^AJo l uI{ il 'i SS'6f'*gtfOi uo3lrl4:(ooqal{cng) sureld [l€[I\l uraqloN:(OY) Sutprseqqruoox ]o salul gI '696I ' u's uo1.u,4s:(EInrrIl!\) :(OV) EuuIpnI\ Jo l!\ salur I 996I lI!^ 6I N''\r's 4 0t 'r!\ 'ZqGI "u uaIlH :(€llnru€{X) :(Oy) oolE{oogJo salur f lll^ 64 s raryDg:(OV) pue pllBdI{A\uaaralaq'896I'x '9t2,{alg?ql4:(aa8prt\) :(aturTlr.\\elurd) :(QV) 11a,ro3 ') 'Zg6I't\\^'LZI '8916I {eg ouiy go trN ur{ ZI ra\11r1g:(a^alC) :(OV) u€rra^ }tr l.Jo M rrr{ c !196I x'62 'tr69]L :(s{)urH) saun(I pu€ sur"ld aalFtr'\JIBrluaC :((IY) uloD t '696I'llxWQIV '8gQ4lqads -ul.I ld Jo N ur{ 0zI r 8 :(aluBidPddEx)lsBoC lsal!\ :sElnsuruad a1.ro1pue ar.{g (I09tI9 .ISl,{) (ral,$€Clo gN lul tI J) IaarJ rlES'8t8Il:62 - ''u 'ri'silfzlog qtil ir irqag:1eisoreg.l :1gyf elplFunJ ro S ur{ I J It6I'rIA s radoo, i1qy1 p1euoqce1,n.,.tt".q 'gg6i'tll't 'II9 33Di :(a^orBrapues)spueldn :g.ropriqefi1 'Z96I Blnsuruad :>lcolg dgo.I (OY) sPuelraqlns ru{8 ) Ir^ tI lI I Jo .US.{\ '06I '69guqaog :lipuBlraqlns)l(OV) ra^\og ur{ g I '896l'111 prDL{r.lO:([.IEIN Jo'I16IIAS 't998 6Z 1]/\I):(q1y) rauio3 puPIra^OJo N lul ZI c x!92 falqlqlA :(PloorrBd):(CY) pg rriag-ue8ro111uo'dossolC rBau'I96I xrg rUIII porr"qs :(rlrEuuau):((IV) aua{rP^\ '7gV j" ur1 c 'tr961'xl91 n7aaqt14:(raploH) :(OV) ue5rotrl Jo gNN uI{ 6I ) '9/6I.ttS.tt '9LIL9f x 9I talq!11,M:(uiolaqruBlg) :(ov) unuu€t\l Jo gNg ru{ 0I |196I IlI^ l 'roII spuE'I {errntr^lraddn :(oV) puog uIa[EJ salrqr rauuo1 :(upqtund) Jo N ''u 'A96I 'ZO1:IpDlrDrls :(.{€rrnl^I ra,\\oT) :(OV) aFplra ^BuntrNr396I xrl s 0I ll^ tZ '89II 3?r?s1:(llrH poolA) :(OV) unlBN'1961 4 n pDrrpUS:(spu€lrool,\l) :(OY) ooraurE'I Jo g url g c'116Ixgz'us raqrsu T iioJvlnns :(PpuoorP)4)spuBlql€aH aallBIN ts€a-qtnoS:((V)u,{dlEuooJ'ZgOI x gZ' u'ss1uotll1.A4:(u^dleuooJ) :((V) {oopuPruooJ '[6171po.ttotlg :(PuElu€J)suteld pue sagptUa]ruarPJIPJ ;o g-a1rur i-'igOt'*tZf 1816l r6ql9 u.raq1ro51:(uorl)allof, paxtu ov) saut€ld sa{ooc o.l JJournl x0z raauoC' :(EuupuBxalvarle'I) sa{B'I ,{errntrN:aallBw .{Prrntr{ :VITYUJSnY '696I'x HJnOS'116 '('ISI,{ 'OV) 1o sallur r rlnw loqs8eg N'{rPrJ 9 '896I06 'VZ9,8Z :surBlduraqFoN :(.IgI I) {rBd IBuorl€NplaJrad,tl[ >lD€1lno x I aplpFnDag il,rasaq 3rg) :(19tr{) dnad1er11'6161llt,t'08'IZE uolrotrI'(Igtr\l) BarESJJTIJ pau 'a^rasagEunBd prold '186I [r^'I '9181?)IrroJ :aollB] :vIuoJJIA T lrBd\als'((v) '296I'M{'I8'IttIIl uolsng uro4 sallul6I sd?t|l11d:sur€ld ural.salA r€d qlnos :(zI9rI9'IsI I) pleuErlEgrEau €;uBI a{E-I 3A.]-'09BI 'u's 'IgW '099tI9 'IgI lE xl gI Dl4)ag:sur"ld uralsa^\ qlnos :(itrgl9 I) ra^tE SulTr€or?ddn 'gLgI ouuB ''u s fDC' :sur€ld uralsaj^ r{lroN :SSTY,^\ HJnOS ,&\-SNW '186I 'Gy) arols plard atPC puPtlP^\o^ uaoalaq pPor Jo gSS uI{ q-n x! qZ 'gggp I€dO {cBIg n4.tog:oiaher( :qNylgNgg 1b'(paulruDxa 08{:)paul11nxa suaLurlads papaPS '8uol urln I.r')'dur|nourdre^oqo,{I,\\oJJEupdua}}ell 'paJS:d[,{lsJodseqpuo^Jqpapual\Jlutu uru{g Jal^ls i 6'111e;srpaucil tuur gi1 g:1td.iu,rJ"7,13ur-'1 urtuL0-i0sdqol eut8rl':5uo1 jsr6' 'q5-rq 'l€rupurT{r 's?g 3uo1utut g g-tr'g *;"i;; i 1 g ,t,,enoIxo"1 .utu 9 c Jr urur i 0 'ayq^l i8uo1^ruur saqr4Topou??tlD?s :sno.IqBl5 ,tilenin 6'0-l'0 sluauottl !'q"'i1?]€ :snoJqEI:';uol.gI?! unir-7 6-1 0 uaulnlP lEseqalnurul E q1r,\\aqol'8uol uIuI 8 0-1 0 s!a{luo xadB papunor -* ; ; i i sluaturt1i11ou'rt o1g qln

'uuoJ Pl.iurJ:tsa14 I irrPg NuytsiaVol. 6, No. 3 (1988)

(AD); (Barung): Co_opers.n., 23.viii.1966, Hummock Mi (AD); 9l4i"g.(Glendellar: lg.Bowans(hinnorh 1433.25.\ii.l9;4. Mr GraingerrADr. FlindersRange.: Nontern l.omple\ l Y ('hinnoch udnam ulana\.. 12l ..20.v1g7J.2 km N trrMudlamurana Well {AD). W_esternPastoral: Gawle-rUp!ands (Gaqler): Eotes s.n., 10.x.1976,Scrubbypeak 'Centrai iAOi; {1(uolcutta): Donncr 3184, 23.ix.1969,Cariewerloo Homestead (MEL); Saii (Acraman): f,gkes.ary!Plateayx. Hi.lton s.n., 13.vii.19b4,top of Uio ntuif llly; Creat Victoria Desert (Yellabina):Moa.,]lng 36, 2?.ix.1976,15 km W of Barton (AD). Nj*n"r" Arid: Western Sandplains (Mt Sir Thomas): Ilelms s.n., 23.vi.1g91,ijO t i" S ot Mi Lindsav (AD). WESTERN AUSTRALIA: Eremaean (Helms): Ceorge3?54, 19.viii.1962,12 miles E "Ij1r:11l9 NewberryTPERTHT: rEur'lar:Apiin 1690.-1.ix.1962,6 miles N of Eucla {PEH I H); (Austin): Smrth 66/513, 15.ix.1966,? miles S of Broad Arrow on Menzies_ Kalgoo_rlij,Road(MEL). Souih-West(Roe): Ioremon 789, 19.ix.1984,,1 km E of Laie (M-EL); King (Avon): Snzirh 261_,10.ix.1983, Amery (MEL); 1?Darling): Ciin-s.i., anno l889.sources ofBlackwood River IMEL 614625). Di.stribution. Queensland, New South Wales, Victoria, , South Australia, Western Australia.

Ecologl'.It usually occursin Eacolyptrrsmallee or upen Eacolypt us woodland communi_ lies.asiocjaled wir h. Erirolypr us erythronema.E. ctetandii.'E.trillithsli. E. "toosi,E. rrons(anttnpntotts.IWpratpucct un.tnola..A(ariq spp.. Errmophils.pp.. Trlndic spp., fipyeyto lpslllpa6ulrii. Dodr,napobursaritlutia and,Hahoo *ittil;r"oio. The .oils'aie usually sandy,shallow or deep.frequently overlying limestone or sandston",oft;;i;;s_ socralronwlttr tatpnttcgravel. rarelv clavey.

Notes._W.-rigida is an extremely variable species which is often diUicult to distinguish ftom W. dampierii and W. eremicctlo.The distinguishing 'Key characters are summarizied in the to species" White & Francis (1922) believed that the Queensland material referred to W. rigid.a by Bailey ( 1901) actually belonged t o W. cheelli. A detailed reappraisal of these two iaxa is required._Arecent collection (Barker 4856)from the Warrego district of Queensland appears to be a typical specimen ol W. rigida. The calyx of S.A. PastorcrlBoard s.n (AD 97628557)is verv similar to W. eremico\a. It has calyx lobes of about 2.8 mm long and a calyx lobe to calyx tube ratio of about 0.8. However, the vegetative characters are typical of W. rigida.' Canseruqtionstatus. Although this speciesis often locally rare, it is not consideredto be AI ITSK. VernqcuLernamei Stiff Westringia(Willis i973, Cunninghamet al. 1982);Stiff West- ern RosemaryrCunningham et ,1. I982r.

13.Westringia eremicola A. Cunn. ex Benth., Labiat. Gen Spec.,159 (1834); J.D. Hook.,Boi. Mag. t.3,138 (1835); Benth. in DC.,Prodr. 12:5?1 (18,18);Fl. Austral.5: 130 (1870);F. Muell.,Fragm. 9: 163(18i5) las'W. longif,rliap.p.l; Ewart, Fl. Victoria980 (19_31{as19301N Boivin, Proc. Roy. S,,c. Queensland60: r0? i1949 [as 1'9b0]); Robedson in J M Black,Fl. S. Austral.2nd.edn.4: T{2, tig. 1063A& B (r9b?);Beadie, Evans & Carolin,Fl. Sy4!,eyREion 519(19631; Burbidge & M. Cray,Fl. Austral.Cap. Terr. 316, fig _318(197-0)lWlllis, Handb. PI. Victoria2: 58i t19?3las i972l); G. Cunninghamet al., Pl. W. New S.Wales 581 (1982 1981]). (here 'Arid [as lecrotlpe chosen):A . Cunniighams.n-, anno 1817, wasteson the LachlanIt. fRiver]' New SouthWales (lecto: K - left specrmen). lW. angustifoliaauct. non R. Br. (1810):J.M. Black,Fl. S. Austral.494 (1926).1 lW. dampieriiauct. non R. Br. (1810):G.& P. Althofer,Austral. Pl. 10:361(1980).1 [W. greuiLLinaauct. non F. Muell. (1855):H. Eichler,Suppl. J.M. Black'sFL S.Austral. 270 (1965);Costermans, Native Trees& ShrubsSE Austral.268(1981) (p.p.)1. '(-IgI 'OV);uorEt\ I 'ZL6I'|'6I 't6I?, url Jo S ru{ 0I qxlrtDD:surEld uralsal\ qlnos :((Y) uqBrreN Jo IAS gg'qurs E8rrlrd'tl6I 'ggL uuDLul?.tiS:suleld ura'tsalAq]roN :(OY) Ilx ZI 'lt0t [,{roluraJ I€tr -rdaC uBrlPrtsnyl p€oU Bl[qulqpIJ uo ralloJ r!ro.I]sallul g'ig6I llx t aroo.lttl:puPI 'lI6I ' -a1qu; ulaqlnog :(8ilSl,S 'IUnl) qercurey x trI u's ntlDg :sadols u.IalsalA,l€r1l 'prurer.{.1gn1g 'A96I 'tll6 "'ab LfSI,\l) s1141a13unqurn.r.re6 .\lolaq xI 08 aayrl,tr' :sadols ura1sar11qliop :5i{.IV.{ HJnOS IA4N (pauluDxa 8II) pau|LuDxasuauDads papaPs

'5uol 'ouqlno 'pau€118u ulul 8I q'I , uI ale^oqo fl^\o Bu + sPaas '5uo1 sdrDuralN EuoI :al,{'l.sJo aseq puo.{aq popualia *- 8 0-lt0 ,(lptsrp urul 9I 8-I ur,.t c saqol eru8ili :3uo1uru l-g g al{ls :8uol luru I-1, 0 ,{JP^o:8uol ruru 8-L lpsld 16 :r-'1el.rpuq.{r rallq,\\ (lrel 'q8rq urur g g rirg Suol ruul 8 0-9 0 saqolppQuluDls :(asuq}E '8uol nrryed) ,'{.rreq,{lasrpds uIuI t I q 0 ,^luaLuDllJlDpoulrurls uaurn)? {PSEqlnoqll^\ BIIo aqL'l';uof urur t I-t 0 sraqluD :snorqPlS'jluoluIuI z-t'I sruauDll| lDur?rD?s|qlnoru lbj or pjpes,,, -nirJo.rpu y xadupdpunorP qlr.{\qf,Ba pu€ (e I I orler t{}pl.t!01q}Eual) '(Euol '(a a1e,ro'flpeorq rl€d aqol rlJBa ulur g I 8 I snuls) paqollq I-6 0 orlPr Jo JIBq ';uo1 '.re1r'Bar qfpr,tl oi q$riali ap,ar tir'ur ruru 8 t-9 2 rlDd aqol uDlpau 1-DlrDpD '(I i:8 9 i 'xuol -.ri + pue papunor xodE Z-9 1 orlEr qlpr^\ o1 ql-Fual) apt,$ tuul a-9 I rqru t t 6 Z 'aie,tciqo.t11q:qs o1 8uo1qosdqal pratpl:1Fuo1 uur snuts) paqolq pu€ rBlnEaur l 0-g 0 ';uol 'a]B^oqo 'papunoJ xadE '(Z otlBr qlpr.{\ (]1ql8ual) apr,|Aullu g 8-6 2 ulur 6 8-6 2 I-I '{llBtslp 01 Suolqo + aqoluDlpaLu :\uur J ql.noul lP Jalauerp ulroJlouunJ sJBad lDlxDqD '.{rPAoZ -ae aqnj leql qcns ltio.rql ui pa1u11p aloq€ dlalplpauwt pal:luJsuot-'f,rPAo P,unorB '.re1nqn'i ':uby 'Surpeards ,llpseq palipp "i* s g c aqni ::uo1 urur i 0-t 0( z 0) pue '.^,11e1irp 'saqol ']Eorql ljira + s.r,er1 snorqelE uaUo saqol ro lr€d IESBquo ,{leq ^lasJeds riuur :8uol ruru 'asrorluu pue passarddPqns ol l,ara ur ,{.rreq .tladuaii alblrns i 0-T 0 'flleS€q sjreq'siqol uo ftreq ilasuap'aqn17o-ped 1e1srpuo frteq {lalerapo!u 01,{lasr€ds snorqelS are;rns Jalno :luasqB {lluaJ€dd€ slop Jo'saqol IEtxeqBpuE IeialPI uo oslB clop qlnourpue;qnl d 'elJns u" (llelpJtuslop u\\oJqol ddurJoqlr^\'dllq^\ ''a1d.rnrl JL, 'Fuollel\eqE ,{1aier ro aAnEtu rElll ruru q 8-q g DIIoroC (.I'I-)t'I t 0(-t 0) orreJ-aqnl n\ aqol *k1er1:alnreqns xadelkrteq [lasuap a]EJrnsrauur'(9'A-Z I oller qlpr.lr ol qr]Fual) "p'^ -*; I auoltuul rd uiril | {Pd\JnJdJ.{lluo.tl'utitBtu dcne\aq JelrriuPlJl llq cnoJqelbJu'qlnnuI .{i^\orJeu;urredddP,{llsn"nrrelniuetrI-dq,|:Inoqin('JqI urc'rnJqDIn '.{.re,ro 'osPq Surpuno.r.rns1ed uo ialellaarpad spuBIS)J€lnpu€lt ,'{lasuap01 ,'{la}EJapoul IP jcul.rns 'Fuoi 'osrorluB 'passardde snorqelE "flasuaprauut ruut (g:t-)t g Z aqnl :Euol-ruul l 0-!10 s.neq' '.ileq J:relrds .ra1no 'aqn1 i,ilir uo saipy uroJ 01 paua{r-rq] {l}qFIIS 'uaar8 xad€ 'palJnJur sl)dds go suie,l leur8reru lue ural-prur rf,?rJ alnJtqns o1 asnlqo uego ur8.reut'aseq palJlJlsar sll,{qdord Jo alEaunc,{l^\oJIEu asEq'[rtBq {jrqirti lB }ou '8uol '(gt g'orler q1pr.l ,,1q$ua1; apr\\ tuur luul 9 Z I ,tiaduJpo1 i1a1d.,apo- t 9 0-2 0 '(llnq "reeuriol alria.o,{1i,ror.reu 'xflP) asEqrP)u pa}Josuts1l,\qduJd:,{Jr€q,'{lasuap ut Jo 'aJu_aJsaJolluo:) Suol lrIIlI A ol dn) Suol luul (l, t)a I-9 0/a;tPad JIPeuou af,qaJsaroulun urJolruraJer asopuor; P auatsa.tollu1 aJPlJns IElx€q€ uo pastsr {11qFqs qllpltu 'alqrsl^ ]ou uorl.Bua^:8uol ruul t 0'c o'1.dn orrnur q:ll^\ alEuoJ)nurqnsxadE'pal€aruo) tcruLuleaJEJJns IBlxeqe leqIqJnsudu0 PJ\JnJaJpuedJlludurElBulaseq le Pd-lJ-lJlsJJlual eururelJo dsnlqoaspq qupttu iunle frlPq tldsuapdJelJns [EIxEqe'{'noJqPIE bulJxaodP 'aruaq) '.{JIeq sa.realainleur uo lualsrs;ad saseq lEq q]!.^\ {luo uaUU ,{lasuop^o1dla}BJa '(g'0 -pour , otrdn orlBJ qlSual €urulPl o1 asPqulo4 qlpl.\\ unrutxBlrr Jo tll-6ual (9t-) lBl.ot '3uo1 ortBr qlpl1^ o1 ql5ual) eprMruur (9'I I-8 0( 9 91 urlu (fA-)0U-8(-t) 0Z'9(-f) 'F)t l€a -ur1o1 ci1cli11a,tprorreu nirr ri.,oflFireq {1asu ep uo1utu.t 1 0 g 0 srlo?lad :pa^rn)ar .{llqFITs '(n 'asrorlue .{llEnsnpue Surp€ards ro)t Jo slroqr ut saoDa? Suol uu (1 0 )tl 0-4 0 'aldurrs passa.rdde+ srtBq :(zurur/sJlPqqZI ,{Jlerl,{lasuap ol fla]srapou'olaralqns 0I) 'sqnrrls o{ reln8uerpenb ro rElnEuBuJ sarpuDrg rlSlq ul (Z )S I S 0(-S 0) IIPUTS '(suaunrads lJal - Iotll9'IUI :olralosr :uaur lpJluar T 'qroJuaIIJ.rPau I -rcads1q8r.r - IoitIg.IgI I :olJal) salBlAr{lnoc .\\aN soITInrJua^EqlPoqs 'q98I - 'Bt araot[ :(uasoqr areq) Jd,{]r-,1Ja7(10-q6I sBl 6i6I) g0I :69 puelsuean$ It 'ur^rogi] 'rlluag eog ,{og cord :(ofgl) otl :g lErlsnY Id DuraJDnb'ru^ Dlouuara M

!rhorrr\i_ll uu,'.) f,i.r.reH Nuyl.siaVol.6. No. ll (1988)

VICTORIA: Eastern Highlands: Victorian Alps: lTolier s.n.,-.xi.1891, Pine Mountain (MEL 1515914);East Gippsland: tr4/illlss.n., 12.xi.1968,Little River Gorge (MEL 614355);East Gippsland Plains: BeauglehoLe(& Finch) 32354.13.xii.1969,between The Narrows and Totem Poinl, Mallacoota Inlet National Park (MEL). - Northern Plains: PhiLLipsCBG 24052,8.xi.1965. Whipstick Scrub, N of Bendigo (AD). - Mallee: Big Des- ert BeaugLehoLe(& -Finch)29023, 7.x.1968, Rudd's Rocks,W side of Wyperteld National Park (AD, MEL); Wimmera: BeauglehoLe19005, 3.ix.1962, 8 miles N of Mt Arapiles (MEL); Liitle Deserl: BeauglehoLe42930, -.x.1948, Woraigworm (MEL). SOUTH AUSTRALIA: Murray Mallee: Murray Lakes (Lake Alexandrina): Hergstrom s.n., .ix.1961,Cooke Plains (AD); Northern Calcarenite Ridges and Plains (Cantana): WiLson7472,3.viii..1960, c.8 km from Coonalpyn (AD); (Coonalpyn):Symon 12776,9.ix.1980, between Kiki & Coonalpyn (AD). South-East Mallee Heathlands (Bordertown): Herb. Ising s.n.,16.x.1925,Bordertown (AD); (Cannawigara):Bcxtmsma 281, 26.ix.1977, 20 km NE of Bordertown (AD); (The Big Desert): Symon 8622, 21.x.19?3,Scorpion Springs ConservationPark (AD); (Karoonda): Phllllps CBG 23844, 20.ix.1961r,7 miles S of Lock (AD); (Moorlands): Donner 6757,20.x.19?8,turnoff to Cooke Plains (AD). Mt Lofty Block: Kangaroo Island (Amberley\: Coctperin herb. Sctuthcott B176, 8-14.viii.1961, near Muston (AD); (MacGillivray): Tepper 7257, 13.iii.1884,near Lashmar's Lagoon (MEL 61,1650);Peninsula Uplands (Sandergrove): Smith 311.,22.ix.1967,c. 3 km S of Monarto South (AD); (Claredon):Gill 251, s. dat., Milang scrub (MEL); (Mt Terrible): CLipstone25, 6.xi.1980,near Athelstone (AD): (Hahndorf): ?.Erschers.n., s. dat., near Mt Barker (MEL 614651). Eyre and Yorke Peninsulas: Southern Highlands and Plains (Lincoln): Alcoch 763, 3.x.1965, Cape Donnington (AD); West Coast (Kappawanla\: Jackson 1131, 4.x.1967,c. 9 km S of BascombeWeil Homestead (AD); (Polda): lsing s.n., 16.ix.1938,Venus Bay (AD); Cen- tral Mallee Plains and Dunes (Hincks): WheelerI040,11.x.1968, Hincks National Park (AD); (Lock): Philllps CBG 2384,1,20.ix.1965, 7 miles S of Lock (AD); (Hambidge): ALcoch1.02'i, 28.viii.1966. SW corner of Hambidge Flora and Fauna Reserve (AD); Southern Yorke Peninsula (Innes): Coplel 4504,30.viii.197.1, Warrenben National Park (AD). DistribuLion. ?Queensland,New South Wales (incl. Australian Capital Territory), Vic- toria and South Australia.

Ecologl. This speciesusually occursin sanclysoils (rarely in clays),overlying or derived from sandstones,shales or granitic rocks. Often associatedrvith buckshot gravel.In the drier areas it is usually associatedwith mallee-eucalyptcommunities wiih an under- storey typically ol laeuigatum (.L coriaceum\, brachybotrya, CaLytrixtetragona, Deuiesia spp.,Leucctpogort spp. and ?rlodla sp. hr central Victoria it is associatedrvith Box Ironbark EucaL',ptuslbrests. rvhereasin easternVictoria it is associatedrvith coastalto foothills ripariair clitl communities ol E. botrToides,Banhsia integrifolia, and Acoclq klbeanensis, EumLyptus glaucescens, Leptospermum scopar[um (respectively).plus l)rlloblum erborescens,Pomqderris andromedifr.tlia and P.aurea.In the Australian Capitai l'erritorl, it. has been recordedas occurring in -Euco h'ptus cordieri E. diues E. uininalis dominalecl ibrests. Nofes. Black (1926) included W. angusLit'oLiaR. Br. in the flora of South Australia. Boivin (1949)correctly restricted this taxon to'l'asmania, 'lf. ' but he appearsnot to have in- cluded cngustlollo sensuBlack in his rreatment oi the genus.Robertson (in Black 1957) realized that the taxon referred to W. angusti.t'oLi.a(by Black 1926) was W 'W. eremicola.Eichler ( i965) assumedthat Boivir.rregarded angustit'olia'sensu Black as synonymous with W. greuiLLi.na.lt appears likelv that Boivin did not examine South Australian material ol W. eremicoLa. I have applied a relatively broad speciesconcept to this species.The typical form of this speciesoccurs in central New South Wales and central Victoria. This speciesis often exlremely diilicult to distinguish from other closelyrelated taxa. Althuugh popu lations from the drier parts of New South Wales and Victoria are usuallv readilvldentifi able,the South Australian populations arevery similar to somepopulations of W. rigido. '^ '(I88I '[(0I8I) u qcrrlar0) 68I raqals :ad,{J I0g 'rpord 'rg 'rads U uou](tg8I) gqt uat lBIq€rI qluag xa raqarsslrpau|I DraUJuDisard '(r98rr9'IgI '^'u satE^\ '1epI ''u t{g) qlnos 1daN'[.ra.ug] asorp '[t08I raquaca0 - lsr$nv .{lqPqord] s s unorg lraJorlllv ?l :adfi (086I)Agt :0I Id IErlsny d T C :(L0S6IsEl 6nOI) 60I :09pu€lsuaan(J ')oS doE 'cord 'urArog:(868I) gSt sal€l iAaN qpuPH 'aqctog g arooj S Id T' "CO I 'C:(Sr8I) :6'lu8Erd'llanl ;g IIq'rpord ur 89I'rads ' I',{:(0t8I) IgI 'rpordlBrlsnv r'rg Id:(818I) 'U :(tg8l) 09t uaC l€rqE'I qtuag :(0I8I) I09 BrloJISuoIEl8uulsadl

'l\oTa0 papr.ro.rdore,{Jeururns uotlnqrJlsrp pu€ uot1€llJ arnlBra}ITIInJ E'ssaualalclurocroj '0I8I ur paqsrlqnd seL\DlloJlSuol '/1 Jo an:ololord aqtrsEaraq^\'(I88I t{4r}alo) tzSI rJquaJoG Mun aunf luo{ €rlBrlsnv ur palJJlloJJdqJrq JSneJdq IerJalEur Jd^l lL,u'{rsltlJ (00{l d'188I l.lJIJldrcpue 99hfI9 fCiW ""Fr) roirrlEJrqcl€rtod,{} P aq ot paulnsardsr srq; 991 raga?Sse [(8t8I allopueC uO ureqtuag dq pailrJ flsnor^ard] 88I raga?S paltJ ,{llrarro)ur (018I) ur€qluag 'sarJads srql Jo uorssnJsrp lEqorllppP f,oI DloJlLuala 14 rapun .sal.oN, aas 'sarJads Dlo)luara lllt ol stql Jo leualBur adlil '(I98tI9 'IgI 'u .U aql Jo UEd sr q3lqld I) s 1lo?or8' parraJar^llcoJlorur (6t6I) urArog 'urnasntr\l qsrlug aql ur s.unAoJBpaulur€xa 'arolaraqJl€l.talEur a^pq I llun addlolDalE Jo Sursooqcaql pa^€lap aA€I{I ur^org ^qpa}ElouuP uaaq atq qcrq^,\ unasntr l qsrlrJg aql ur uaunrods I€npL\IpuI alaldruoJ ?soru aql uro{ uasoqc aq plnirqs saDads ueruuorg E Jo ad.{lolral aql l€ql pa}sa88ns (096I) urEals 'JI{ 'E ollol18uol Dt8ult$aM uo satoN

'(286I }a ueq;uruunC) ArPlllosou ura]sol\ 'tl,6i lB rapuals :(2861 1e1a rueq8uluun3 slllllA) er3urr1sa11.rapuelS sauDu lDln)DulaA '{srl LBl.ou paJAplSuOJr^nJD:JS uolJDaJaSuaJ 'uorlBgrlsa^ur polelap sarlnbal sJol -c?rBqcasaqlJo aJueJgru8rscrulouoxpl ar{J uoxpl Jollel arll uPq}'x^lPc aqllo acPJrns 'saqruerq 'acuaq) ralno aql uo oslB pu€ aql Jo s]J€d (8uno[ IElsrp aql uo ,{IBq ,{losuap arour sr raurJoJ oql:1"q1sl BxEl od\l aq:I uaa^\laq aDuaJaJJIpsnouqo ]sour oqL plo)lruara 'r14 ;o suorlelndod uEuolcr^ uJalsea aql ol paleloJ ,{lasolc ,{ra^ sr nluldoutuatt ,17 'puBlsuaantl DUoJlSuo!1!1 .{lqlssod sr Dlorlluara Al oltparoJar st qrlq,'!'! Jo suor8ar rall.a,!\ aql uorJ aql l{rntr\l s€lloJoJ JB[[ o] aldJnd s€rl ,{llBnsn 1)loJ?uaJa1l sPaJar{,{\ 'se11o.rocIBrral€ur Jo 'arouuaqlrnd -1rq.u sBq {llBnsn DqollSuol ll palBacuot IIaBTPI areJrns l€rxe -qe aq1qlru'apr,u urur g I-I aJBsa^eal arll puP,{rIBq {lasuap o1.{lalPJapolusr x{lPJ aql Jo oJ€JJnsralno a:]ilD{))lruala rl uI pasodxa dlaFrel ale;rns IBIXeqeaql ql!^\'ap!,lA urru g -q aJBsa^€ol aq1 puB snorqBl5(s^BulE lou fllEnsn sr x,{lP) aI{]Jo acBJJnsralno U I Jr) 'puPISuaantJ aqt,DUoll1uol l,1 uI lDurlsrp se paurPl.urBurar€ o,{\1aqJ, (sal€l\Aqlnos .^aN i7'nt1'oir1uof jl q1r,rl sarlrleyuns Suor]s .$orls suor]Elndod raslrol aqJ suorl€lndod uBrJolDr^ raqlo aql uBql saABalJa8uol q)nur aAEqsuorlelndod uEuolcl^ uJalsea aqJ 'oAlsnlJxa 'ilalPurlJoJuil fllBnlnu oJ€ salpJsralf,preqJ aqt ,salrads o1 ,{aX, aq} uI PazI lou 'd1 -ieufluns are opr,?] rqo.rl sarcadsstql SutqslnFullslp roJ sJal)erBql Inlasn lsolu aI{J 'Dto)luata '1!1 Jo suorlBlndod salBlA qlnos i aN puB u€rrolJr^ oql JoI sr lr sE )rlsou8€tp, flJBalJ del6u sr a.rnlee; srql 'suorlBlndod u€q€rlsnv qlnos aql ur'aroJaraqJ uorlBlndod u€tlerl -snv qlnos aqi u€ql sanl€^ ra3.re1a,teq (Buolcr^ puB salEl qlnos d\aNJo) suorlPlndod uJalseaaql lPq} qans'orlEJ aqnl c)]aqol x^lBJ aql ul uorlJnpal B sl aJdql'JIdurEXorod

P.rnurrlsall uuoJ f .\rru8 348 Nultsia Vol. 6, No. 3 (1988)

lW. eremi.colaauct. non A. Cunn.ex Benth.:F.M. Bailey,Queensl. Fl.5: 1206(1902) p.p.l. Distribution. ?Queenslandand New South Wales.

Acknowledgements I wish to sincerelythank Drs D.B. Foreman,P.S. Short (Both MEL) and J.G. West (CANB) for their constructive comments on the manuscript.

References

Althofer, G.W. & Althofer, P. McD. (1980).The genus tv€stringia Sm. Austml. Pl. 10:343 351,359-866. Anderson,R.H. (1961).Introduction. Contr. New S. Wales Natl Herb. Nos 1-18: 1-15. Aston, H.I. (1984).Publication dates of esrly sci€ntificjournals in Victoria. Muelle a 5:281-288. "Queenstand Bailey, F.M. (1901). Labiatae. In Flora." Vol.4, pp. 1205 & 1206. (Govemment Printer: Brisbane.) Ball, H.W. et al. (eds)(1962). IL Terminology of simplesymmetrical plane shapes(chart 1). Taxon 11: 14b- 156,fis. 19. Beard,J.S. (1980).A new phytogeographicmap of Westem Australia. WesternAustral. Herb. Res.Notes B: 3? 58. "Flora Bentham, G. (1870).Labiatae. In Austraiiensis."Vol.5, pp. ?0 137. (Reeve:London.) "Flora Black, J.M. (1957).Labiatae- In of South Australia." Patt 4, pp.126-'142.(Government Printer: Adelaide.) "How Blackall,W.E. & Grieve,B.J. (1965)..In to Know WesternAustralian Wildflowers." PartIII. pp. 573- 595. (UniveNity of Western Australia Press: [Nedlands].) Boivin, B. ( 1949).W€srlingio, an Australiangenus of Labiatae.Proc. Roy. Soc.Queenstand 60: 99 110,i. 9. Candolle,A.P. de (1848).Labiatae. In "ProdromusSystematis Naturalis RegniVegetabilis..." yoi.12,pp.27 - 603. (Treuttel & Wultz: Pa s.) Cochrane,G.R., Fuhrer, 8.A., Rotherham,E.R. & Willis, J.H. 53(1968)."Australian Flora in Colour.Flowers and Plants of Victoria." lA.H. & A.W. Reed:Sydney.) Conn, B.J. (1980).The lack of field obseNationsand collectingtechniques with referenceto Prostanther&. Ausiral. Syst. Bot. Soc. Newsletter 25: 6 9. Conn, B.J. (1984). A taxonomic revision of Prostanthera Labill. section Klonderio (F.v.Muell.) Benth. (Labiatae).J. AdelaideBot. Gard. 6t 20'7-438. "Flora Conn, B.J. (1986).l4l€srriDgia. In Jessop,J.P. & Toelken,H.R. (eds). of South Australia.',palt B.pD. 1230- 1232.(Government Printer: Adelaide.) "Native Costermans,L. (1981). Trees and Shrubs of South easternAustralia." (Rigby: Adelaide.) Cunn_ingham,^G.M.,Mulhan, W.E., Milthorpe. P.L. & Leigh, J.H. 11982). Plants of Western New South Wales.'(Governmpnr Prinier: Sydney.) "The Cuftis, W.M. (1967).Labiatae. In Student's Fbra of Tasmania." Part 3, pp. 542-55?.(Government P nter: lHobart] Tasmania.) Dietrich, F.C. (1881).Franz Wilhelm Sieber,ein Beitrag zur Geschichteder Botanik vor sechzigJahren. Jahrb. bot. Gart. Mus. Berlin 1: 278-306. "Supplement Eichler, H. (1962).Lamiaceae. In to J.M. Black's Flora of South Australia."pp. 267-270.(Gov ernment P nter: Adelaide.) Holmgren,P.K., Keuken,W. & Schofield,E.K. (1981).Index herbadorum.Pat t. The herbariaofthe world, ed.?. Reg.Veg. 106: 1-452. "Plants Jacobs,S.W.L. & Pickard, J. (1981). of New South Wales."(Govemment P nter Sydney.) "Environments Laut, P.C-etal. (19?7a). ofSouth Australia.Province 2 Murray Mallee."(Land UseResearch. C.S.l.R.O.:Canberra.) "Envircnments Laut, P.C. et al. (1977b). of South Australia- Province 3 Mt. Loftv Block." (Land Use Re- .earch.C.S.l.R.O.: canberra.t "Envircnments Laut, P.C. et al. (197?c). of South Australia. Province4 Eyre and Yorke Peninsulas.,,(Land Use Research.C.S.I.R.O.: Canberra.) "Environments Laut, P.C. et al. (1977d). of South Australia. Province 6 FlindeN Ranges.,,(Land Use Re search.C.S.I.R.O.: Canberra.) ( uollleC :ssard^lrsra^ '999 'sln,{4. -run aurnoqler{) {' 9Bg dd'Z IoA,. Buotcr^ ur s?ueldot looqpue}IV,,uj DlSul4sail (ef6t) Hf 'C9t-69t:tC puelsurenbJoS 'IoU rsr'uerd 'e?rq^r\ cord Erold puelsueonoeq? ot suoltnqFluoC ul rus ,?811?J$aA\ZZ6t) O 1\| T J O 'noqqv 'ureals ('urElug lBero ro$raN :salrEqC? phe0) upe puz ., urle.I IE uetog,, (gl,6l) J A\ ( ulraqutelA:suueulaSug) upe elufl6cBd rl ^ dd,, (0e8I) runu4rd runlqaualddns pue (0IBI) uoluar( UBA aelnsq ta aelpuelloH a€^oN aElold snurorpord,, 'u 's,$orE .(096I) .J.l .uJ€els uI .aErpuelloHa€^oN eBJoldsnworpoJd, s,u.rorg u 01 uolpnporlul '990I-I 'u€^loC 'nauels :ZIT 8aA 8aE olsles :A ToA upa puz aJntpJaII rllrlouoxEl (S86I) S U T' v d ',IZI-1 'uB^loC 'naUE?S :0II se^ 8aE rIBS-d :AI IoA upa puz aJnterall rlurouox€I (eS6I) S U T v J '086-T '(186I) 'ue^1oC 'naUE?S :90T 8eA 8aU O-,{.I :III IoA upa puz alnteJa}lrrlulottoxeJ S E T y J 'T66-I 'u€^1oC 'nagE?S :86 sa^ seu e.I-H :II IoA upa puz aJnteratlrrrruouoxeJ (616I) S U T, V J '9811-1 {@^\oC 'na[Bts :t6 8eA 8eU C-y :I IoA upa puz aJnleralrtrruouoxEJ (91,6I) S U T' y J 'IrZ-ZgZ :98uoxEJ siadBdcluouox€t dlrBos.rellenl^l H fpueurpreduo uortBurroJrrrAraN (9861) O'Sraqos old)uorrcru,sep pun "o-I3$5'l,"Ki$"p'l?t:Tlil.:"1,!":liffl 'etFawY...cpaFuorsr relrqeq ,n,,ro,ooo,o-',3, iuBrlBuow rra8rx'tr{ qFoNJo erolJ aql 01srolnquluoC roJ aprno V,, (tf6l) g f T' A|U O'ralJod 'ggl'Igt 're :6 IrrBBrJ aBteqe.I (qlgl) uo^ I H fl d anl,^J locuar^Brl ( Irotr ,raN :uEllrurcElD ,,truouoxpJ luEld ol uortrnporlul uV,, (996I) I\I.H.C 'qcrBasau tenequeJ:oulsc asnpue.I),! puv uieql.IoN 8 ecur^ord eqertsnv qlnosJo stueuuol^u9,, (1116I).Ista.C.d,lnE.I (.ErrequeS:.o.u qcrBas .(a/r6l) I.s'P c .C.d .lnE.I -aU esn pue-I) .,]€ro?spduralsar!\ , arul^ord erlerlsnv qlnos Jo stuouuol^ug,J ta

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