Journal of Paleontology, 94(5), 2020, p. 829–851 Copyright © 2020, The Paleontological Society. This is an Open Access article, distributed under the terms of the Creative Commons Attribution licence (http://creativecommons.org/ licenses/by/4.0/), which permits unrestricted re-use, distribution, and reproduction in any medium, provided the original work is properly cited. 0022-3360/20/1937-2337 doi: 10.1017/jpa.2020.25 Ontogenetic analysis of Anisian (Middle Triassic) ptychitid ammonoids from Nevada, USA Eva A. Bischof1* and Jens Lehmann1 1Geowissenschaftliche Sammlung, Fachbereich Geowissenschaften, Universität Bremen, Leobener Strasse 8, 28357 Bremen, Germany <
[email protected]>, <
[email protected]> Abstract.—Ptychites is among the most widely distributed ammonoid genera of the Triassic and is namesake of a family and superfamily. However, representatives of the genus mostly show low-level phenotypic disparity. Furthermore, a large number of taxa are based on only a few poorly preserved specimens, creating challenges to determine ptychitid taxonomy. Consequently, a novel approach is needed to improve ptychitid diversity studies. Here, we investigate Ptychites spp. from the middle and late Anisian of Nevada. The species recorded include Ptychites embreei n. sp., which is distinguished by an average conch diameter that is much smaller and shows a more evolute coiling than most of its relatives. The new species ranges from the Gymnotoceras mimetus to the Gymnotoceras rotelliformis zones, which makes it the longest-ranging species of the genus. For the first time, the ontogenetic development of Pty- chites was obtained from cross sections where possible. Cross-sectioning highlights unique ontogenetic trajectories in ptychitids.