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Tiktaalik: Reconstructed at Left Tetrapod Testimony
Fa m i liar Fossils Tiktaalik fossil at right and Fossil site Tiktaalik: reconstructed at left Tetrapod Testimony IMAGINE a creature that looks like a crocodile with a neck but which also has gills like a fish! The stuff of nightmares, did you say? But it is just Tiktaalik roseae – a fish-like fossil that was introduced to the year old rocks. world on April 2006 by a team of researchers led by Neil They had a pretty Shubin, Edward Daeschler and Farish Jenkins, who clear idea about uncovered this bizarre specimen. It has been hailed as what sort of being the species that blurs the boundary between fish animal they were and the four-legged tetrapods. looking for. Tiktaalik was found on Ellesmere Island, Canada, However, success not too far from the North Pole in the Arctic Circle. It has was not been described as, “a cross between a fish and a crocodile.” immediate. It took It lived in the Devonian era lasting from 417m to 354m them five separate years ago. The name Tiktaalik is the Inuktitut word for expeditions to the freshwater fish burbot (Lota lota). The name was Canada before they suggested by Inuit elders of the area where the fossil was raised the discovered. The specific name roseae holds the clue to Tiktaalik. the name of an anonymous donor and is a tribute to the Tiktaalik was dubbed a “fishapod” by Daeschler. The person. media called it a “missing link” between fish and the Tiktaalik was a fish no doubt but a special one. Its four-legged tetrapods. -
Faculdade De Biociências
FACULDADE DE BIOCIÊNCIAS PROGRAMA DE PÓS-GRADUAÇÃO EM ZOOLOGIA ANÁLISE FILOGENÉTICA DE DORADIDAE (PISCES, SILURIFORMES) Maria Angeles Arce Hernández TESE DE DOUTORADO PONTIFÍCIA UNIVERSIDADE CATÓLICA DO RIO GRANDE DO SUL Av. Ipiranga 6681 - Caixa Postal 1429 Fone: (51) 3320-3500 - Fax: (51) 3339-1564 90619-900 Porto Alegre - RS Brasil 2012 PONTIFÍCIA UNIVERSIDADE CATÓLICA DO RIO GRANDE DO SUL FACULDADE DE BIOCIÊNCIAS PROGRAMA DE PÓS-GRADUAÇÃO EM ZOOLOGIA ANÁLISE FILOGENÉTICA DE DORADIDAE (PISCES, SILURIFORMES) Maria Angeles Arce Hernández Orientador: Dr. Roberto E. Reis TESE DE DOUTORADO PORTO ALEGRE - RS - BRASIL 2012 Aviso A presente tese é parte dos requisitos necessários para obtenção do título de Doutor em Zoologia, e como tal, não deve ser vista como uma publicação no senso do Código Internacional de Nomenclatura Zoológica, apesar de disponível publicamente sem restrições. Dessa forma, quaisquer informações inéditas, opiniões, hipóteses e conceitos novos apresentados aqui não estão disponíveis na literatura zoológica. Pessoas interessadas devem estar cientes de que referências públicas ao conteúdo deste estudo somente devem ser feitas com aprovação prévia do autor. Notice This thesis is presented as partial fulfillment of the dissertation requirement for the Ph.D. degree in Zoology and, as such, is not intended as a publication in the sense of the International Code of Zoological Nomenclature, although available without restrictions. Therefore, any new data, opinions, hypothesis and new concepts expressed hererin are not available -
Structure of Tropical River Food Webs Revealed by Stable Isotope Ratios
OIKOS 96: 46–55, 2002 Structure of tropical river food webs revealed by stable isotope ratios David B. Jepsen and Kirk O. Winemiller Jepsen, D. B. and Winemiller, K. O. 2002. Structure of tropical river food webs revealed by stable isotope ratios. – Oikos 96: 46–55. Fish assemblages in tropical river food webs are characterized by high taxonomic diversity, diverse foraging modes, omnivory, and an abundance of detritivores. Feeding links are complex and modified by hydrologic seasonality and system productivity. These properties make it difficult to generalize about feeding relation- ships and to identify dominant linkages of energy flow. We analyzed the stable carbon and nitrogen isotope ratios of 276 fishes and other food web components living in four Venezuelan rivers that differed in basal food resources to determine 1) whether fish trophic guilds integrated food resources in a predictable fashion, thereby providing similar trophic resolution as individual species, 2) whether food chain length differed with system productivity, and 3) how omnivory and detritivory influenced trophic structure within these food webs. Fishes were grouped into four trophic guilds (herbivores, detritivores/algivores, omnivores, piscivores) based on literature reports and external morphological characteristics. Results of discriminant function analyses showed that isotope data were effective at reclassifying individual fish into their pre-identified trophic category. Nutrient-poor, black-water rivers showed greater compartmentalization in isotope values than more productive rivers, leading to greater reclassification success. In three out of four food webs, omnivores were more often misclassified than other trophic groups, reflecting the diverse food sources they assimilated. When fish d15N values were used to estimate species position in the trophic hierarchy, top piscivores in nutrient-poor rivers had higher trophic positions than those in more productive rivers. -
Amazon Alive: a Decade of Discoveries 1999-2009
Amazon Alive! A decade of discovery 1999-2009 The Amazon is the planet’s largest rainforest and river basin. It supports countless thousands of species, as well as 30 million people. © Brent Stirton / Getty Images / WWF-UK © Brent Stirton / Getty Images The Amazon is the largest rainforest on Earth. It’s famed for its unrivalled biological diversity, with wildlife that includes jaguars, river dolphins, manatees, giant otters, capybaras, harpy eagles, anacondas and piranhas. The many unique habitats in this globally significant region conceal a wealth of hidden species, which scientists continue to discover at an incredible rate. Between 1999 and 2009, at least 1,200 new species of plants and vertebrates have been discovered in the Amazon biome (see page 6 for a map showing the extent of the region that this spans). The new species include 637 plants, 257 fish, 216 amphibians, 55 reptiles, 16 birds and 39 mammals. In addition, thousands of new invertebrate species have been uncovered. Owing to the sheer number of the latter, these are not covered in detail by this report. This report has tried to be comprehensive in its listing of new plants and vertebrates described from the Amazon biome in the last decade. But for the largest groups of life on Earth, such as invertebrates, such lists do not exist – so the number of new species presented here is no doubt an underestimate. Cover image: Ranitomeya benedicta, new poison frog species © Evan Twomey amazon alive! i a decade of discovery 1999-2009 1 Ahmed Djoghlaf, Executive Secretary, Foreword Convention on Biological Diversity The vital importance of the Amazon rainforest is very basic work on the natural history of the well known. -
Zootaxa, Pseudolaguvia Virgulata, a New Sisorid Catfish
Zootaxa 2518: 60–68 (2010) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2010 · Magnolia Press ISSN 1175-5334 (online edition) Pseudolaguvia virgulata, a new sisorid catfish (Teleostei: Sisoridae) from Mizoram, northeastern India HEOK HEE NG1 & LALRAMLIANA2 1Raffles Museum of Biodiversity Research, National University of Singapore, 6 Science Drive 2 #03-01, Singapore 117546. E-mail: [email protected] 2Department of Zoology, Pachhunga University College, Aizawl, Mizoram, India 796001. E-mail: [email protected] Abstract Pseudolaguvia virgulata, a new South Asian sisorid catfish species, is described from the Barak River drainage in Mizoram, India. The new species can be distinguished from congeners in having a brown body with two or three narrow, pale longitudinal stripes and a pale Y-shaped marking on the dorsal surface of the head. Additional distinguishing characters from its congeners are a serrated anterior edge of the dorsal spine, the thoracic adhesive apparatus reaching beyond the base of the last pectoral-fin ray, head width, pectoral-fin length, length of dorsal-fin base, dorsal-spine length, body depth at anus, length of adipose-fin base, caudal peduncle length, caudal peduncle depth, snout length, interorbital distance, and total number of vertebrae. Key words: Siluriformes, Sisoroidea, Barak River, South Asia Introduction Members of the sisorid genus Pseudolaguvia are small catfishes found in rivers draining the sub-Himalayan region and Myanmar. They superficially resemble miniature species of Glyptothorax in overall morphology and in having a thoracic adhesive apparatus with a median depression, but can be distinguished in having prominent postcoracoid processes. Eleven species of Pseudolaguvia are considered valid (Ng 2009): P. -
Phylogenetic Relationships of the South American Doradoidea (Ostariophysi: Siluriformes)
Neotropical Ichthyology, 12(3): 451-564, 2014 Copyright © 2014 Sociedade Brasileira de Ictiologia DOI: 10.1590/1982-0224-20120027 Phylogenetic relationships of the South American Doradoidea (Ostariophysi: Siluriformes) José L. O. Birindelli A phylogenetic analysis based on 311 morphological characters is presented for most species of the Doradidae, all genera of the Auchenipteridae, and representatives of 16 other catfish families. The hypothesis that was derived from the six most parsimonious trees support the monophyly of the South American Doradoidea (Doradidae plus Auchenipteridae), as well as the monophyly of the clade Doradoidea plus the African Mochokidae. In addition, the clade with Sisoroidea plus Aspredinidae was considered sister to Doradoidea plus Mochokidae. Within the Auchenipteridae, the results support the monophyly of the Centromochlinae and Auchenipterinae. The latter is composed of Tocantinsia, and four monophyletic units, two small with Asterophysus and Liosomadoras, and Pseudotatia and Pseudauchenipterus, respectively, and two large ones with the remaining genera. Within the Doradidae, parsimony analysis recovered Wertheimeria as sister to Kalyptodoras, composing a clade sister to all remaining doradids, which include Franciscodoras and two monophyletic groups: Astrodoradinae (plus Acanthodoras and Agamyxis) and Doradinae (new arrangement). Wertheimerinae, new subfamily, is described for Kalyptodoras and Wertheimeria. Doradinae is corroborated as monophyletic and composed of four groups, one including Centrochir and Platydoras, the other with the large-size species of doradids (except Oxydoras), another with Orinocodoras, Rhinodoras, and Rhynchodoras, and another with Oxydoras plus all the fimbriate-barbel doradids. Based on the results, the species of Opsodoras are included in Hemidoras; and Tenellus, new genus, is described to include Nemadoras trimaculatus, N. -
Homoplasies, Consistency Index and the Complexity of Morphological Evolution: Catfishes As a Case Study for General Discussions on Phylogeny and Macroevolution
Int. J. Morphol., 25(4):831-837, 2007. Homoplasies, Consistency Index and the Complexity of Morphological Evolution: Catfishes as a Case Study for General Discussions on Phylogeny and Macroevolution Homoplasias, Índice de Consistencia y la Complejidad de la Evolución Morfológica: Peces Gato como un Estudio de Caso para Discusiones Generales en Filogenia y Macroevolución *,** Rui Diogo DIOGO, R. Homoplasies, consistency index and the complexity of morphological evolution: Catfishes as a case study for general discussions on phylogeny and macroevolution. Int. J. Morphol., 25(4):831-837, 2007. SUMMARY: Catfishes constitute a highly diversified, cosmopolitan group that represents about one third of all freshwater fishes and is one of the most diverse Vertebrate taxa. The detailed study of the Siluriformes can, thus, provide useful data, and illustrative examples, for broader discussions on general phylogeny and macroevolution. In this short note I briefly expose how the study of this remarkably diverse group of fishes reveals an example of highly homoplasic, complex 'mosaic' morphological evolution. KEY WORDS: Catfishes; Homoplasies; Morphological macroevolution; Phylogeny; Siluriformes; Teleostei. INTRODUCTION The catfishes, or Siluriformes, found in North, Cen- and diversity surely resulting from several homoplasic tral and South America, Africa, Europe, Asia and Australia, events. This was precisely the main reason to choose this with fossils inclusively found in Antarctica, constitute a amazing group of fishes as a case study for discussing gene- highly diversified, cosmopolitan group, which, with more ral topics on phylogeny and macroevolution. But the exam than 2700 species, represents about one third of all freshwater of more and more morphological phylogenetic characters in fishes and is one of the most diverse Vertebrate taxa (e.g. -
Summary Report of Freshwater Nonindigenous Aquatic Species in U.S
Summary Report of Freshwater Nonindigenous Aquatic Species in U.S. Fish and Wildlife Service Region 4—An Update April 2013 Prepared by: Pam L. Fuller, Amy J. Benson, and Matthew J. Cannister U.S. Geological Survey Southeast Ecological Science Center Gainesville, Florida Prepared for: U.S. Fish and Wildlife Service Southeast Region Atlanta, Georgia Cover Photos: Silver Carp, Hypophthalmichthys molitrix – Auburn University Giant Applesnail, Pomacea maculata – David Knott Straightedge Crayfish, Procambarus hayi – U.S. Forest Service i Table of Contents Table of Contents ...................................................................................................................................... ii List of Figures ............................................................................................................................................ v List of Tables ............................................................................................................................................ vi INTRODUCTION ............................................................................................................................................. 1 Overview of Region 4 Introductions Since 2000 ....................................................................................... 1 Format of Species Accounts ...................................................................................................................... 2 Explanation of Maps ................................................................................................................................ -
Amazon Alive!
Amazon Alive! A decade of discovery 1999-2009 The Amazon is the planet’s largest rainforest and river basin. It supports countless thousands of species, as well as 30 million people. © Brent Stirton / Getty Images / WWF-UK © Brent Stirton / Getty Images The Amazon is the largest rainforest on Earth. It’s famed for its unrivalled biological diversity, with wildlife that includes jaguars, river dolphins, manatees, giant otters, capybaras, harpy eagles, anacondas and piranhas. The many unique habitats in this globally significant region conceal a wealth of hidden species, which scientists continue to discover at an incredible rate. Between 1999 and 2009, at least 1,200 new species of plants and vertebrates have been discovered in the Amazon biome (see page 6 for a map showing the extent of the region that this spans). The new species include 637 plants, 257 fish, 216 amphibians, 55 reptiles, 16 birds and 39 mammals. In addition, thousands of new invertebrate species have been uncovered. Owing to the sheer number of the latter, these are not covered in detail by this report. This report has tried to be comprehensive in its listing of new plants and vertebrates described from the Amazon biome in the last decade. But for the largest groups of life on Earth, such as invertebrates, such lists do not exist – so the number of new species presented here is no doubt an underestimate. Cover image: Ranitomeya benedicta, new poison frog species © Evan Twomey amazon alive! i a decade of discovery 1999-2009 1 Ahmed Djoghlaf, Executive Secretary, Foreword Convention on Biological Diversity The vital importance of the Amazon rainforest is very basic work on the natural history of the well known. -
Phylogeny of Basal Tetrapoda
Stuart S. Sumida Biology 342 Phylogeny of Basal Tetrapoda The group of bony fishes that gave rise to land-dwelling vertebrates and their descendants (Tetrapoda, or colloquially, “tetrapods”) was the lobe-finned fishes, or Sarcopterygii. Sarcoptrygii includes coelacanths (which retain one living form, Latimeria), lungfish, and crossopterygians. The transition from sarcopterygian fishes to stem tetrapods proceeded through a series of groups – not all of which are included here. There was no sharp and distinct transition, rather it was a continuum from very tetrapod-like fishes to very fish-like tetrapods. SARCOPTERYGII – THE LOBE-FINNED FISHES Includes •Actinista (including Coelacanths) •Dipnoi (lungfishes) •Crossopterygii Crossopterygians include “tetrapods” – 4- legged land-dwelling vertebrates. The Actinista date back to the Devonian. They have very well developed lobed-fins. There remains one livnig representative of the group, the coelacanth, Latimeria chalumnae. A lungfish The Crossopterygii include numerous representatives, the best known of which include Eusthenopteron (pictured here) and Panderichthyes. Panderichthyids were the most tetrapod-like of the sarcopterygian fishes. Panderichthyes – note the lack of dorsal fine, but retention of tail fin. Coelacanths Lungfish Rhizodontids Eusthenopteron Panderichthyes Tiktaalik Ventastega Acanthostega Ichthyostega Tulerpeton Whatcheeria Pederpes More advanced amphibians Tiktaalik roseae – a lobe-finned fish intermediate between typical sarcopterygians and basal tetrapods. Mid to Late Devonian; 375 million years old. The back end of Tiktaalik’s skull is intermediate between fishes and tetrapods. Tiktaalik is a fish with wrist bones, yet still retaining fin rays. The posture of Tiktaalik’s fin/limb is intermediate between that of fishes an tetrapods. Coelacanths Lungfish Rhizodontids Eusthenopteron Panderichthyes Tiktaalik Ventastega Acanthostega Ichthyostega Tulerpeton Whatcheeria Pederpes More advanced amphibians Reconstructions of the basal tetrapod Ventastega. -
Teacher's Guide: Tiktaalik: a Fish out of Water
Teacher’s Guide: Tiktaalik: A Fish Out of Water Recommended Grade Level: 5–8 (also applicable to grades 9–12 for students requiring significant support in learning) Suggested Time: About 50–60 minutes spread over one or more class periods, plus additional time to complete a writing assignment Goals Vocabulary Following are the big ideas that students • fossil should take away after completing this lesson: • characteristic • Transitional fossils help scientists establish • transition how living things are related • tetrapod • Physical features and behaviors may • species change over time to help living things sur- vive where they live • amphibian • evolution Key Literacy Strategies Following are the primary literacy strategies students will use to complete this activity: • Categorizing basic facts and ideas (screen 10) • Making inferences (screens 4 and 7, writing assignment 2) • Identifying and using text features (screens 4, 6, and 9) • Determining important information (screen 7, writing assignment 1) • Sequencing events (screen 9) Note: In addition to using the key literacy strategies listed above, students will use each of the strategies below to complete this lesson: • Monitoring comprehension • Synthesizing • Making predictions • Developing vocabulary • Connecting prior knowledge to new learning • Developing a topic in writing • Identifying and using text features (photographs, captions, diagrams, and/or maps) Overview Tiktaalik: A Fish Out of Water is a student-directed learning experience. However, while students are expected to work through the lesson on their own, teachers should be available to keep the lesson on track, organize groupings, facilitate discussions, answer questions, and ensure that all learning goals are met. Teacher’s Guide: Tiktaalik: A Fish Out of Water 1 The following is a summary of the lesson screens: Screen 1: Students learn that they will explore evolutionary relationships between animal groups that do not appear to be related. -
Peces De La Cuenca Del Pastaza, Ecuador Peces De La Cuenca Del Pastaza, Ecuador 4 5
Peces de la Cuenca del Pastaza ECUADOR Ri'o Bobonaza.. cerca deI pobla::lo de Canelo Peces de la Cuenca del Pastaza, Ecuador Peces de la Cuenca del Pastaza, Ecuador 4 5 Contenido: Juan Francisco Rivadeneira, Consultor - Fundación Natura, Gerencia de Proyectos, Quito, Ecuador Elizabeth Anderson, Global Water for Sustainability (GLOWS) program, Florida International University, Miami, USA CONTENIDO Sistema de Información Geográfica (SIG) y Seguimiento: Agradecimientos .................................................................................................... 7 Sara Dávila Fundación Natura, Gerencia de Proyectos, Quito, Ecuador La Cuenca del Pastaza ............................................................................................... 11 Supervisión: Jorge Rivas, Ruth Elena Ruíz Peces del Pastaza ....................................................................................................... 19 Fundación Natura, Gerencia de Proyectos, Quito, Ecuador • Órdenes y Familias ................................................................ 23 • Characiformes ................................................................ 23 Fotografías: • Siluriformes .................................................................... 34 Byron Freeman Peter Frey • Gymnotiformes ............................................................... 42 Juan Francisco Rivadeneira • Perciformes ........................................................................ 45 • Synbranchiformes .........................................................