Mark Johnston, Editor-in-Chief University of Colorado-Denver Tracey DePellegrin Connelly, GENETICS Editorial Board Executive Editor

CELLULAR GENETICS Lawrence G. Harshman GENOME AND SYSTEMS Hamish Spencer Orna Cohen-Fix University of Nebraska, Lincoln BIOLOGY University of Otago NIDDK, National Institutes of Health Thomas C. Kaufman Jef Boeke Naoyuki Takahata Susan Dutcher Indiana University Johns Hopkins School of Medicine Graduate University for Washington University School Kenneth J. Kemphues Stanley Fields Advanced Studies of Medicine Cornell University University of Washington Marcy K. Uyenoyama Duke University JoAnne Engebrecht Barbara J. Meyer David Largaespada University of California, Davis University of California, Berkeley University of Minnesota Lindi M. Wahl University of Western Ontario David I. Greenstein Trudi Schüpbach Jeffery F. Miller University of Minnesota Princeton University University of California, Los Angeles John Wakeley Harvard University Michael Nonet Mariana F. Wolfner Peter J. Oefner Washington University School Cornell University Stanford University H. Allen Orr (Perspectives Editor) of Medicine University of Rochester Norbert Perrimon GENE EXPRESSION Adam S. Wilkins (Perspectives Editor) Mark D. Rose Harvard Medical School Princeton University Karen Arndt University of Cambridge Piali Sengupta University of Pittsburgh Patricia J. Pukkila (Genetics Education Linda Siracusa Brandeis University Jefferson Medical College of Thomas James A. Birchler Editor) University of North Carolina Gary Stormo Jefferson University University of Missouri Jasper Rine (Reviews Editor) Washington University School University of California, Berkeley Charles Boone of Medicine COMPLEX TRAITS University of Toronto Michael Turelli (Reviews Editor) Daniel F. Voytas Justin Borevitz University of California, Davis David Jonah Grunwald University of Minnesota University of Chicago University of Utah BOARD OF SENIOR EDITORS Edward Buckler Michael Hampsey EMPIRICAL POPULATION Oliver Hobert Cornell University Robert Wood Johnson Medical GENETICS Columbia University Alain Charcosset School-UMDNJ Montserrat Aguadé Charles H. Langley Institut National de la Recherche Ann Hochschild Universitat de Barcelona University of California, Davis Agronomique Harvard Medical School David Begun Terry R. Magnuson Steve Chenoweth Aaron P. Mitchell University of California, Davis University of North Carolina, Chapel Hill University of Queensland Carnegie Mellon University James J. Bull Lauren M. McIntyre Gary A. Churchill Craig S. Pikaard University of Texas at Austin University of Florida The Jackson Laboratory Indiana University Deborah Charlesworth Rasmus Nielsen Ina Hoeschele Eric Selker University of Edinburgh University of Copenhagen, Centre for Virginia Polytechnic Institute and State University of Oregon Anna Di Rienzo Bioinformatics University John Tamkun University of Chicago Mark Rose Corbin D. Jones University of California, Santa Cruz Jeffrey Lawrence Princeton University The University of North Carolina University of Pittsburgh John C. Schimenti at Chapel Hill GENOME INTEGRITY AND Cornell University Manyuan Long TRANSMISSION Matias Kirst University of Chicago Fred Winston University of Florida Eric E. Alani Harvard Medical School Cornell University Michael Nachman Krista Nichols University of Arizona GENETICS publishes high quality, Purdue University Sharon E. Bickel original research presenting novel Outi Savolainen Dartmouth College findings on a range of topics bearing Fernando F. Pardo Manuel de Villena University of Oulu University of North Carolina Monica Colaiácovo on heredity and variation. GENETICS Harvard Medical School Michel Veuille is a peer-edited journal—all editorial Andrew H. Paterson Ecole Pratique des Hautes Etudes, Gregory Copenhaver decisions are made by the authors’ University of Georgia Paris University of North Carolina peers—with a tradition of rigorous David Threadgill Stephen I. Wright peer review. Publication is open Nancy Hollingsworth University of North Carolina University of Toronto equally to members and non-members Stony Brook University Hongyu Zhao of the Genetics Society of America. Yale University Scott Keeney THEORETICAL POPULATION Submit manuscripts at http://submit. Memorial Sloan-Kettering Fei Zou GENETICS genetics.org. 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