Dinosaurs of the District of Columbia
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The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
The Bristol Dinosaur Project
Proceedings of the Geologists’ Association 123 (2012) 210–225 Contents lists available at ScienceDirect Proceedings of the Geologists’ Association jo urnal homepage: www.elsevier.com/locate/pgeola Teaching Perspective The Bristol Dinosaur Project Michael J. Benton *, Remmert Schouten, Edward J.A. Drewitt, Pedro Viegas School of Earth Sciences, University of Bristol, Bristol, BS8 1RJ, UK A R T I C L E I N F O A B S T R A C T Article history: Dinosaurs have been fascinating to the widest public since the 1840s, and that interest has grown step- Received 18 April 2011 wise ever since. Public interest has been harnessed over the years especially by museums in blockbuster Received in revised form 8 July 2011 exhibitions, and in the form of best-selling books and films. Here we describe a major educational Accepted 8 July 2011 initiative, the Bristol Dinosaur Project, which has run for ten years and has reached tens of thousands of Available online 5 August 2011 children and adults, supported by substantial funding. The Bristol Dinosaur Project focuses on the fourth dinosaur ever named in the world, Thecodontosaurus, discovered in Bristol in 1834, and named in 1836. Keywords: The dinosaur is not in itself spectacular, being only 1–2 m long, but its evolutionary role as one of the first Dinosaurs plant-eating dinosaurs in the world justifies our current research, and provides a strong theme for the Education Engagement public presentation. Further, the fact that the dinosaur is found as disarticulated bones in ancient tropical Outreach cave systems, allows us to develop numerous key themes with all age groups: the geological time scale, Geosciences education continental drift, reconstruction of ancient environments, modern landscape analogues, the rock cycle, evolution, biomechanics, and critical assessment of geological and palaeontological evidence. -
Competition Structured a Late Cretaceous Megaherbivorous Dinosaur Assemblage Jordan C
www.nature.com/scientificreports OPEN Competition structured a Late Cretaceous megaherbivorous dinosaur assemblage Jordan C. Mallon 1,2 Modern megaherbivore community richness is limited by bottom-up controls, such as resource limitation and resultant dietary competition. However, the extent to which these same controls impacted the richness of fossil megaherbivore communities is poorly understood. The present study investigates the matter with reference to the megaherbivorous dinosaur assemblage from the middle to upper Campanian Dinosaur Park Formation of Alberta, Canada. Using a meta-analysis of 21 ecomorphological variables measured across 14 genera, contemporaneous taxa are demonstrably well-separated in ecomorphospace at the family/subfamily level. Moreover, this pattern is persistent through the approximately 1.5 Myr timespan of the formation, despite continual species turnover, indicative of underlying structural principles imposed by long-term ecological competition. After considering the implications of ecomorphology for megaherbivorous dinosaur diet, it is concluded that competition structured comparable megaherbivorous dinosaur communities throughout the Late Cretaceous of western North America. Te question of which mechanisms regulate species coexistence is fundamental to understanding the evolution of biodiversity1. Te standing diversity (richness) of extant megaherbivore (herbivores weighing ≥1,000 kg) com- munities appears to be mainly regulated by bottom-up controls2–4 as these animals are virtually invulnerable to top-down down processes (e.g., predation) when fully grown. Tus, while the young may occasionally succumb to predation, fully-grown African elephants (Loxodonta africana), rhinoceroses (Ceratotherium simum and Diceros bicornis), hippopotamuses (Hippopotamus amphibius), and girafes (Girafa camelopardalis) are rarely targeted by predators, and ofen show indiference to their presence in the wild5. -
ABSTRACT Resurrecting Tyrannosaurus Rex Lauren E. Ammerman Director: Jennifer Good, Ph.D. After the First Successful Extraction
ABSTRACT Resurrecting Tyrannosaurus rex Lauren E. Ammerman Director: Jennifer Good, Ph.D. After the first successful extraction of ancient DNA from a fossilized Quagga in 1984, the subsequent development of PCR (Polymerase Chain Reaction) technology opened up a plethora of possibilities in the field of molecular paleontology. Supplied with fragmented ancient genomes, some scientists acted as if the days of resurrecting dinosaurs were a few technical difficulties away. Theories surfaced on the possible applications of ancient DNA technology, and some, such as creating tactical dinosaurs for the U.S. military, were outrageous. A less ridiculous idea surfaced in the form of Michael Crichton’s Jurassic Park, published in 1990. Coupled with Steven Spielberg’s 1993 feature film adaption, the Jurassic Park series created a world in which genetically- engineered dinosaurs roamed once again as theme park attractions on a billionaire’s private island, and explored the possible outcomes of a “Jurassic Park” experiment. Jurassic Park ignited scientific debate over the technological feasibility, environmental impact, and ethical questions of a “Jurassic Park” experiment. This thesis continues that conversation by asking, could resurrecting a dinosaur be a productive environmental enterprise, other than a mere display of power over Nature? Focusing on Tyrannosaurus rex, this thesis combines a brief survey the current state of dinosaur genetic research, with analyses of rewilding with large predators, to discuss whether or not scientists should ever attempt to re-create a T. rex in the future. APPROVED BY DIRECTOR OF HONORS THESIS Dr. Jennifer Good, Director of University Scholars APPROVED BY THE HONORS PROGRAM Dr. Elizabeth Corey, Director DATE: RESURRECTING TYRANNOSAURUS REX A Thesis Submitted to the Faculty of Baylor University In Partial Fulfillment of the Requirements for the Honors Program By Lauren E. -
A Jurassic Basal Eusauropod Clade from Iberia Finds Refuge In
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by UCL Discovery Descendants of the Jurassic turiasaurs from Iberia found refuge in the Early Cretaceous of western USA Rafael Royo-Torres, Paul Upchurch, James I. Kirkland, Donald D. DeBlieux, John R. Foster, Alberto Cobos, Luis Alcalá Supplementary Information: I. Abbreviations section II. Geological setting of the Doelling’s Bowl bonebed III. Phylogenetic analyses and list of synapomorphies IV. Holotype of Mierasaurus V. Measurements VI. References I. Abbreviations section BYU (Museum of Paleontology of Brigham Young University, Provo, Utah USA). DBBB Doelling’s Bowl bonebed; DBGI Doelling’s Bowl site; J-K Jurassic- Cretaceous transition; Ma million years ago; My million years; MPT most parsimonious trees; RD name for every dinosaur site from Riodeva in Teruel (Spain); TL Tree length; UMNH the Natural History Museum of Utah (USA); UGS Utah Geological Survey (USA). Anatomical abbreviations: bt basal tubera; bo basioccipital; bpt basipterygoid process; ca crista antotica; ct crista tuberalis; f frontal; fme fenestra metotic; fv fenestra vestivuli (=fenestra ovalis); osc otosphenoidal crest (= crista prootica); p parietal; pa parapophysis; prf prefontal; stf supratemporal fossa; so supraoccipital; eo exoccipital- opisthotic; bo basioccipital; popr paraoccipital process; cranial nerves: I olfactory nerve; II optic nerve; III oculomotor nerve; IV trochlear nerve; V trigeminal nerve (ophthalmic, maxillary, mandibular); VI abductor or abducens nerve; VII facialis nerve; VIII vestibulocochlear nerve. Abbreviations for laminae vertebral: acpl anterior centroparapophyseal lamina; ant. spdl anterior spinodiapophyseal lamina; cprl 1 centroprezygapophyseal lamina; cpol centropostzygapophyseal lamina; pcdl posterior centrodiapophyseal lamina; post. spdl posterior spinodiapophyseal lamina; ppdl paradiapophyseal lamina; prdl prezygodiapophyseal lamina; spol spinopostzygapophyseal lamina. -
The Many Myths, Some Old, Some New, of Dinosaurology
Modern Geology, 1991, Vol. 16, pp. 69-99 © 1991 Gordon and Breach Science Publishers S.A. Reprints available directly from the Publisher Printed in the United Kingdom Photocopying permitted by license only THE MANY MYTHS, SOME OLD, SOME NEW, OF DINOSAUROLOGY GREGORY S. PAUL 3109 N. Calvert St. Side Apt., Baltimore, Maryland 21218, USA (Received November 11, 1990) The study of dinosaurs has always been hindered by a body of unsubstantiated or false beliefs about their biology, and about the biology of other animals that can be applied to dinosaurs. Some of these myths, such as the supposedly horizontal femora of birds, or the importance of heat radiators, are relatively minor. Other myths have greatly distorted our understanding of dinosaur biology, These myths include the persistent selective advantage of energy efficient bradymetabolic physiologies, the impossibility of high speed in large animals, the danger of overheating of large animals, the supposed correlation between brain size and energetics, the great difficulty of flying, especially in large forms, and the inherent vulnerability of the big dinosaurs to extinction. KEY WORDS: Dinosaurs, pterosaurs, archosaurs, anatomy, locomotion, brains, physiology, extinction, myths. INTRODUCTION As soon as dinosaurs were discovered, myths started gathering around them and their archosaur relatives. Among the earliest was the belief that they were overgrown lizards, a myth that continues in modified form today in their classification as reptiles. Some old myths have recently been discredited, such as the theropods' fear of entering the water, and the sauropods' and hadrosaurs' water-loving habits. The polyphyletic split of the Dinosauria into Saurischia and Ornithischia has been displaced by widespread agreement upon the group's monophyly. -
2018 in Review
2018 in Review By: Sajal Shukla 2018 was a very influential year in the world. There were many bad and good events that took place, but they all contributed to a very unique year. As the year 2018 has come to an end, let’s take a look at some of the major events that happened this year. One of the first major events that happened in 2018 were the winter Olympics in Pyeongchang, South Korea. Ninety-six coun- tries came together to compete in various winter sports such as ski- ing, curling, figure skating, and ice hockey. One of our very own alumni, Kendall Coyne, competed with the U.S. women’s ice hockey team, which one gold at the competition. Overall, the U.S. won nine gold, eight silver, and six bronze metals. Another event that took place this year was the tragic school shooting in Parkland, Florida. Seventeen people died in the shooting, making it the largest school shooting in U.S. history. Though this event was catastrophic, it brought up important discussions about gun control in the United States. It also brought people together to mourn the loss of those seventeen lives and to see how we can bring about positive change in the world. Something else that happened this year was the royal wedding that took place in England. Meghan Markle made his- tory as the first divorced, biracial American woman to marry someone in the royal family. The marriage of Prince Harry and Meghan Markle was a beautiful moment people all over the world were able to celebrate. -
APS Bulletin June 1986
ALBERTA • PALAEONTOLOG I CAL • SOC I ETY ULUULLETINLLLLLEEIETININ VOLUMEBB 11 NUMBER 2 JUNE 1996 1 ALBERTA PALAEONTOLOGICAL SOCIETY OFFICERS President Wayne Braunberger 278-5154 Program Coordinator Kris Vasudevan 288-7955 Vice-President Peter Meyer 289-4135 Curator Harvey Negrich 249-4497 Treasurer Joe LeBlanc 246-7601 Librarian Dr. Gerry Morgan 241-0963 Secretary Don Sabo 289-0138 Field Trip Coordinator Les Fazekas 248-7245 Past-President Les Adler 289-9972 Director at Large Dr. David Mundy 281-3668 DIRECTORS Editor Howard Allen 274-1858 APAC† Representative Don Sabo 278-8045 Membership Vaclav Marsovsky 547-0182 †APAC is the Alberta Palaeontological Advisory Committee The Society was incorporated in 1986, as a non-profit organization formed to: a. Promote the science of palaeontology through study and education. b. Make contributions to the science by: 1) discovery 4) education of the general public 2) collection 5) preservation of material for study and the future 3) description c. Provide information and expertise to other collectors. d. Work with professionals at museums and universities to add to the palaeontological collections of the province (preserve Alberta’s heritage). MEMBERSHIP: Any person with a sincere interest in palaeontology is eligible to present their application for membership in the Society. (Please enclose membership dues with your request for application.) Single membership $15.00 annually Family or Institution $20.00 annually THE BULLETIN WILL BE PUBLISHED QUARTERLY: March, June, September and December. Deadline for submitting material for publication is the 15th of the month prior to publication. Society Mailing Address: Material for Bulletin: Alberta Palaeontological Society Howard Allen, Editor, APS P.O. -
Advocates for Cold-Blooded Dinosaurs: the New Generation of Heretics
AL SOCIET IC Y G O O F L A O M E E G R I E C H A T GROUNDWORK Furthering the Influence GROUNDWORK: of Earth Science Furthering the Influence of Earth Science Advocates for cold-blooded dinosaurs: The new generation of heretics Timothy L. Clarey, Delta College, Dept. of Geology, 1961 sion of highly vascularized bone merely means the animal was Delta Road, University Center, Michigan 48710-0002, USA, active and not necessarily endothermic. Owerkowicz raised [email protected] two savannah monitor lizards and exercised one daily while letting the other remain inactive. After a few years, he killed INTRODUCTION the pair and examined their respective bone microstructure. Robert Bakker (1986) branded himself and a small group He found that the active lizard had a bone microstructure that of groundbreaking and upstart paleontologists in the 1960s was highly vascularized, mimicking an endothermic animal and 1970s “heretics” because they challenged the mainstream and containing numerous Haversian canals. The inactive lizard view of dinosaurs as slow, sluggish, and cold-blooded reptiles. had a poorly vascularized microstructure, typical of modern This view, that dinosaurs were warm-blooded, active, bird-like ectothermic animals. His study only “proved” that dinosaurs creatures, much different than extant reptiles of today, is now were active. widely accepted by scientists and the public alike. Yet, this Ruben and Jones (2000) believe that the presence of respira- viewpoint has been challenged in recent years by a small but tory turbinates makes a better “proof” of endothermy. Ninety- active minority group of researchers, herein described as the nine percent of endothermic animals have turbinates or coils new generation of heretics. -
Activities in Historical Geology with IGO's
Activities in Historical Geology With IGO's 2001 Historical Geology West Virginia Instructional Goals and Objectives Historical geology is an integral portion of science instruction in the state of West Virginia as evidenced in the list of Instructional Goals and Objectives listed below. Every attempt was made to incorporate these WVIGOs into the activities which follow. These topic- specific WVIGOs, as well as others, are identified for each of the activities that appear in this document. Fourth Grade 4.27 engage in active inquiries, investigations, and hands-on activities for a minimum of 50% of the instructional time to develop conceptual understanding and laboratory skills 4.62 4,5 research evidence to discover the age of the earth – systems 4.63 10 associate fossils with the periods in which they were formed – systems 4.66 5 compare and explain the relative time differences to erode materials (e.g., a sand pile, mud pile, rock pile) – changes Fifth Grade 5.27 3,4,6,8,9,10,11 classify living and non- living things according to properties (e.g., structures and functions, mass, volume, density, solubility, conductivity, magnetism, weight, shape, color, freezing point, boiling point, evaporation, rocks and minerals) – systems 5.30 3,4,6,7,8,9,10,11 recognize and explore methods for investigating physical changes (e.g., evaporating, condensing, boiling, freezing, melting, salinity, density, shape and size) – changes 5.32 2,3,4,7,8,9,10 explain how the different characteristics of plants and animals help them to survive in different niches -
Evidence for Endothermic Ancestors of Crocodiles at the Stem of Archosaur Evolution Author(S): Roger S
Division of Comparative Physiology and Biochemistry, Society for Integrative and Comparative Biology Evidence for Endothermic Ancestors of Crocodiles at the Stem of Archosaur Evolution Author(s): Roger S. Seymour, Christina L. Bennett‐Stamper, Sonya D. Johnston, David R. Carrier, and Gordon C. Grigg Source: Physiological and Biochemical Zoology, Vol. 77, No. 6, Sixth International Congress of Comparative Physiology and Biochemistry Symposium Papers: Evolution and Advantages of Endothermy (November/December 2004), pp. 1051-1067 Published by: The University of Chicago Press. Sponsored by the Division of Comparative Physiology and Biochemistry, Society for Integrative and Comparative Biology Stable URL: http://www.jstor.org/stable/10.1086/422766 . Accessed: 04/11/2015 23:31 Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at . http://www.jstor.org/page/info/about/policies/terms.jsp . JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. The University of Chicago Press and Division of Comparative Physiology and Biochemistry, Society for Integrative and Comparative Biology are collaborating with JSTOR to digitize, preserve and extend access to Physiological and Biochemical Zoology. http://www.jstor.org This content downloaded from 23.235.32.0 on Wed, 4 Nov 2015 23:31:05 PM All use subject to JSTOR Terms and Conditions 1051 Evidence for Endothermic Ancestors of Crocodiles at the Stem of Archosaur Evolution Roger S. -
Hadrosaurs As Ungulate Parallels: Lost Lifestyles and Deficient Data
Hadrosaurs as ungulate parallels: Lost lifestyles and deficient data MATTHEW T. CARRANO, CHRISTINE M. JANIS, and J. JOHN SEPKOSKI,JR.? Carrano, M.T, Janis, C.M., & Sepkoski, J.J., Jr. 1999. Hadrosaurs as ungulate parallels: Lost Lifestyles and deficient data. -Acta Palaeontologica Polonica 44,3,237-261. Hadrosaur dinosaurs (Ornithischia: Hadrosauridae) were abundant in Late Cretaceous terrestrial environments of North America and Asia. Their derived dental and postcranial specializations for herbivory have led to the suggestion that these dinosaurs were analo- gous to modem ungulates in lifestyle. Ungulates display a suite of morphological fea- tures that are correlated with diet, habitat preference, and sexual dimorphism. In this pa- per we examine several of these same features in hadrosaurs in order to determine whether they display similar patterns under multivariate analysis. Initial results confirm the resemblances between hadrosaurs and ungulates but suggest that missing data may considerably affect the outcomes of statistical analyses. Using the hadrosaur dataset as a template, we artificially degrade the (previously complete) ungulate datasets and per- form the same analyses. Results are consistent with earlier results and provide an oppor- tunity to assess the impact of missing data on the original multivariate structure. Our re- sults support the hypothesis that hadrosaurs were similar to ungulates in patterns of ecomorphology. Key words : Dinosauria, Ornithischia, Ornithopoda, multivariate statistics, paleoeco- logy, Marnmalia, NMDS. Matthew T. Carrano [[email protected]],Department of Anatomical Sci- ences, Health Sciences Center T-8, State University of New York at Stony Brook, Stony Brook, New York, 11 794 USA. Christine M. Janis [[email protected]], Department of Ecology and Evolution- ary Biology, Division of Biology and Medicine, Box G-B209, Brown University, Provi- dence, Rhode Island, 02912 USA.