The Bristol Dinosaur Project
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M.A. TAYLOR 1 and R.D. CLARK 2 Methodology and Repositories
Ichthyosaurs from the Lower Lias (Lower Jurassic) of Banwell, Somerset ICHTHYOSAURS FROM THE LOWER LIAS (LOWER JURASSIC) OF BANWELL, SOMERSET M.A. TAYLOR 1 AND R.D. CLARK 2 Taylor, M.A. and Clark, R.D. 2016. Ichthyosaurs from the Lower Lias (Lower Jurassic) of Banwell, Somerset. Geoscience in South-West England, 14, 59–71. Two Lower Jurassic ichthyosaurs, lost in a 1940s air raid, are identified from archival records and surviving 19th century plaster casts distributed by their owner, the Bristol Institution for the Advancement of Science, Literature and the Arts. One was the type specimen of Ichthyosaurus latimanus Owen, 1840. The other, initially labelled Ichthyosaurus intermedius Conybeare, 1822, was the first ichthyosaur in which remains of the soft tissue of the tail fin were identified, confirming Richard Owen’s earlier prediction. Unfortunately Owen’s published account conflated the two specimens, apparently by the erroneous transposition of a passage of text. Owen possibly regarded the second specimen as the type of I. intermedius. In 1889 Richard Lydekker referred it to Ichthyosaurus conybeari Lydekker, 1888. For unknown reasons, he caused further confusion by suppressing the fact that both ichthyosaur specimens came from Banwell. Banwell is shown to have been a minor but significant source of Lower Lias fossil vertebrates, and also the type locality of the fish Tetragonolepis monilifer Agassiz, 1837, apparently from quarries in the lowermost Jurassic beds at Knightcott. The Bristol Institution assisted Edward Wilson (1808–1888) to obtain at least some of the West Country marine reptiles which his brother Dr Thomas Bellerby Wilson (1807–1865) donated to the Academy of Natural Sciences of Philadelphia, U.S.A. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Some Unpublished Correspondence of the Rev. W.B. Clarke D.F
Journal & Proceedings of the Royal Society of New South Wales, Vol. 141, p. 1–31, 2008 ISSN 0035-9173/08/02001–31 $4.00/1 Some Unpublished Correspondence of the Rev. W.B. Clarke d.f. branagan and t.g. vallance Abstract: Four previously unpublished letters, with memoranda of Rev. W.B. Clarke to W.S. Macleay, written between 1842 and 1845 clarify the ideas of both about the mode of formation of coal and the age of the stratigraphical succession in the Sydney Basin. Clarke makes the first mention of his discovery of the Lake Macquarie fossil forest, the first identification of the zeolite stilbite in New South Wales and gives details of his study of the volcanic rocks of the Upper Hunter Valley. Keywords: Clarke, Macleay, coal formation, Sydney Basin, stilbite INTRODUCTION essentially self-contained, although, of course, almost no letter can stand alone, but depends When Dr Thomas G. Vallance died in 1993 a on the correspondents. Each letter has some considerable amount of his historical jottings importance in dealing with aspects of Clarke’s and memorabilia on the history of Australian geological work, as will be noted. science, and particularly geology, was passed The first and longest letter, which is ac- on to me (David Branagan) through his wife, companied by a long series of memoranda and Hilary Vallance. For various reasons, only now a labelled sketch fits between two letters from have I been able to delve, even tentatively, into MacLeay (28 June & 4 July 1842) to Clarke. this treasure house. The present note con- Both these MacLeay letters have been repro- cerns four letters of the Reverend W.B. -
Competition Structured a Late Cretaceous Megaherbivorous Dinosaur Assemblage Jordan C
www.nature.com/scientificreports OPEN Competition structured a Late Cretaceous megaherbivorous dinosaur assemblage Jordan C. Mallon 1,2 Modern megaherbivore community richness is limited by bottom-up controls, such as resource limitation and resultant dietary competition. However, the extent to which these same controls impacted the richness of fossil megaherbivore communities is poorly understood. The present study investigates the matter with reference to the megaherbivorous dinosaur assemblage from the middle to upper Campanian Dinosaur Park Formation of Alberta, Canada. Using a meta-analysis of 21 ecomorphological variables measured across 14 genera, contemporaneous taxa are demonstrably well-separated in ecomorphospace at the family/subfamily level. Moreover, this pattern is persistent through the approximately 1.5 Myr timespan of the formation, despite continual species turnover, indicative of underlying structural principles imposed by long-term ecological competition. After considering the implications of ecomorphology for megaherbivorous dinosaur diet, it is concluded that competition structured comparable megaherbivorous dinosaur communities throughout the Late Cretaceous of western North America. Te question of which mechanisms regulate species coexistence is fundamental to understanding the evolution of biodiversity1. Te standing diversity (richness) of extant megaherbivore (herbivores weighing ≥1,000 kg) com- munities appears to be mainly regulated by bottom-up controls2–4 as these animals are virtually invulnerable to top-down down processes (e.g., predation) when fully grown. Tus, while the young may occasionally succumb to predation, fully-grown African elephants (Loxodonta africana), rhinoceroses (Ceratotherium simum and Diceros bicornis), hippopotamuses (Hippopotamus amphibius), and girafes (Girafa camelopardalis) are rarely targeted by predators, and ofen show indiference to their presence in the wild5. -
ABSTRACT Resurrecting Tyrannosaurus Rex Lauren E. Ammerman Director: Jennifer Good, Ph.D. After the First Successful Extraction
ABSTRACT Resurrecting Tyrannosaurus rex Lauren E. Ammerman Director: Jennifer Good, Ph.D. After the first successful extraction of ancient DNA from a fossilized Quagga in 1984, the subsequent development of PCR (Polymerase Chain Reaction) technology opened up a plethora of possibilities in the field of molecular paleontology. Supplied with fragmented ancient genomes, some scientists acted as if the days of resurrecting dinosaurs were a few technical difficulties away. Theories surfaced on the possible applications of ancient DNA technology, and some, such as creating tactical dinosaurs for the U.S. military, were outrageous. A less ridiculous idea surfaced in the form of Michael Crichton’s Jurassic Park, published in 1990. Coupled with Steven Spielberg’s 1993 feature film adaption, the Jurassic Park series created a world in which genetically- engineered dinosaurs roamed once again as theme park attractions on a billionaire’s private island, and explored the possible outcomes of a “Jurassic Park” experiment. Jurassic Park ignited scientific debate over the technological feasibility, environmental impact, and ethical questions of a “Jurassic Park” experiment. This thesis continues that conversation by asking, could resurrecting a dinosaur be a productive environmental enterprise, other than a mere display of power over Nature? Focusing on Tyrannosaurus rex, this thesis combines a brief survey the current state of dinosaur genetic research, with analyses of rewilding with large predators, to discuss whether or not scientists should ever attempt to re-create a T. rex in the future. APPROVED BY DIRECTOR OF HONORS THESIS Dr. Jennifer Good, Director of University Scholars APPROVED BY THE HONORS PROGRAM Dr. Elizabeth Corey, Director DATE: RESURRECTING TYRANNOSAURUS REX A Thesis Submitted to the Faculty of Baylor University In Partial Fulfillment of the Requirements for the Honors Program By Lauren E. -
The Many Myths, Some Old, Some New, of Dinosaurology
Modern Geology, 1991, Vol. 16, pp. 69-99 © 1991 Gordon and Breach Science Publishers S.A. Reprints available directly from the Publisher Printed in the United Kingdom Photocopying permitted by license only THE MANY MYTHS, SOME OLD, SOME NEW, OF DINOSAUROLOGY GREGORY S. PAUL 3109 N. Calvert St. Side Apt., Baltimore, Maryland 21218, USA (Received November 11, 1990) The study of dinosaurs has always been hindered by a body of unsubstantiated or false beliefs about their biology, and about the biology of other animals that can be applied to dinosaurs. Some of these myths, such as the supposedly horizontal femora of birds, or the importance of heat radiators, are relatively minor. Other myths have greatly distorted our understanding of dinosaur biology, These myths include the persistent selective advantage of energy efficient bradymetabolic physiologies, the impossibility of high speed in large animals, the danger of overheating of large animals, the supposed correlation between brain size and energetics, the great difficulty of flying, especially in large forms, and the inherent vulnerability of the big dinosaurs to extinction. KEY WORDS: Dinosaurs, pterosaurs, archosaurs, anatomy, locomotion, brains, physiology, extinction, myths. INTRODUCTION As soon as dinosaurs were discovered, myths started gathering around them and their archosaur relatives. Among the earliest was the belief that they were overgrown lizards, a myth that continues in modified form today in their classification as reptiles. Some old myths have recently been discredited, such as the theropods' fear of entering the water, and the sauropods' and hadrosaurs' water-loving habits. The polyphyletic split of the Dinosauria into Saurischia and Ornithischia has been displaced by widespread agreement upon the group's monophyly. -
Advocates for Cold-Blooded Dinosaurs: the New Generation of Heretics
AL SOCIET IC Y G O O F L A O M E E G R I E C H A T GROUNDWORK Furthering the Influence GROUNDWORK: of Earth Science Furthering the Influence of Earth Science Advocates for cold-blooded dinosaurs: The new generation of heretics Timothy L. Clarey, Delta College, Dept. of Geology, 1961 sion of highly vascularized bone merely means the animal was Delta Road, University Center, Michigan 48710-0002, USA, active and not necessarily endothermic. Owerkowicz raised [email protected] two savannah monitor lizards and exercised one daily while letting the other remain inactive. After a few years, he killed INTRODUCTION the pair and examined their respective bone microstructure. Robert Bakker (1986) branded himself and a small group He found that the active lizard had a bone microstructure that of groundbreaking and upstart paleontologists in the 1960s was highly vascularized, mimicking an endothermic animal and 1970s “heretics” because they challenged the mainstream and containing numerous Haversian canals. The inactive lizard view of dinosaurs as slow, sluggish, and cold-blooded reptiles. had a poorly vascularized microstructure, typical of modern This view, that dinosaurs were warm-blooded, active, bird-like ectothermic animals. His study only “proved” that dinosaurs creatures, much different than extant reptiles of today, is now were active. widely accepted by scientists and the public alike. Yet, this Ruben and Jones (2000) believe that the presence of respira- viewpoint has been challenged in recent years by a small but tory turbinates makes a better “proof” of endothermy. Ninety- active minority group of researchers, herein described as the nine percent of endothermic animals have turbinates or coils new generation of heretics. -
Activities in Historical Geology with IGO's
Activities in Historical Geology With IGO's 2001 Historical Geology West Virginia Instructional Goals and Objectives Historical geology is an integral portion of science instruction in the state of West Virginia as evidenced in the list of Instructional Goals and Objectives listed below. Every attempt was made to incorporate these WVIGOs into the activities which follow. These topic- specific WVIGOs, as well as others, are identified for each of the activities that appear in this document. Fourth Grade 4.27 engage in active inquiries, investigations, and hands-on activities for a minimum of 50% of the instructional time to develop conceptual understanding and laboratory skills 4.62 4,5 research evidence to discover the age of the earth – systems 4.63 10 associate fossils with the periods in which they were formed – systems 4.66 5 compare and explain the relative time differences to erode materials (e.g., a sand pile, mud pile, rock pile) – changes Fifth Grade 5.27 3,4,6,8,9,10,11 classify living and non- living things according to properties (e.g., structures and functions, mass, volume, density, solubility, conductivity, magnetism, weight, shape, color, freezing point, boiling point, evaporation, rocks and minerals) – systems 5.30 3,4,6,7,8,9,10,11 recognize and explore methods for investigating physical changes (e.g., evaporating, condensing, boiling, freezing, melting, salinity, density, shape and size) – changes 5.32 2,3,4,7,8,9,10 explain how the different characteristics of plants and animals help them to survive in different niches -
Evidence for Endothermic Ancestors of Crocodiles at the Stem of Archosaur Evolution Author(S): Roger S
Division of Comparative Physiology and Biochemistry, Society for Integrative and Comparative Biology Evidence for Endothermic Ancestors of Crocodiles at the Stem of Archosaur Evolution Author(s): Roger S. Seymour, Christina L. Bennett‐Stamper, Sonya D. Johnston, David R. Carrier, and Gordon C. Grigg Source: Physiological and Biochemical Zoology, Vol. 77, No. 6, Sixth International Congress of Comparative Physiology and Biochemistry Symposium Papers: Evolution and Advantages of Endothermy (November/December 2004), pp. 1051-1067 Published by: The University of Chicago Press. Sponsored by the Division of Comparative Physiology and Biochemistry, Society for Integrative and Comparative Biology Stable URL: http://www.jstor.org/stable/10.1086/422766 . Accessed: 04/11/2015 23:31 Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at . http://www.jstor.org/page/info/about/policies/terms.jsp . JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. The University of Chicago Press and Division of Comparative Physiology and Biochemistry, Society for Integrative and Comparative Biology are collaborating with JSTOR to digitize, preserve and extend access to Physiological and Biochemical Zoology. http://www.jstor.org This content downloaded from 23.235.32.0 on Wed, 4 Nov 2015 23:31:05 PM All use subject to JSTOR Terms and Conditions 1051 Evidence for Endothermic Ancestors of Crocodiles at the Stem of Archosaur Evolution Roger S. -
Hadrosaurs As Ungulate Parallels: Lost Lifestyles and Deficient Data
Hadrosaurs as ungulate parallels: Lost lifestyles and deficient data MATTHEW T. CARRANO, CHRISTINE M. JANIS, and J. JOHN SEPKOSKI,JR.? Carrano, M.T, Janis, C.M., & Sepkoski, J.J., Jr. 1999. Hadrosaurs as ungulate parallels: Lost Lifestyles and deficient data. -Acta Palaeontologica Polonica 44,3,237-261. Hadrosaur dinosaurs (Ornithischia: Hadrosauridae) were abundant in Late Cretaceous terrestrial environments of North America and Asia. Their derived dental and postcranial specializations for herbivory have led to the suggestion that these dinosaurs were analo- gous to modem ungulates in lifestyle. Ungulates display a suite of morphological fea- tures that are correlated with diet, habitat preference, and sexual dimorphism. In this pa- per we examine several of these same features in hadrosaurs in order to determine whether they display similar patterns under multivariate analysis. Initial results confirm the resemblances between hadrosaurs and ungulates but suggest that missing data may considerably affect the outcomes of statistical analyses. Using the hadrosaur dataset as a template, we artificially degrade the (previously complete) ungulate datasets and per- form the same analyses. Results are consistent with earlier results and provide an oppor- tunity to assess the impact of missing data on the original multivariate structure. Our re- sults support the hypothesis that hadrosaurs were similar to ungulates in patterns of ecomorphology. Key words : Dinosauria, Ornithischia, Ornithopoda, multivariate statistics, paleoeco- logy, Marnmalia, NMDS. Matthew T. Carrano [[email protected]],Department of Anatomical Sci- ences, Health Sciences Center T-8, State University of New York at Stony Brook, Stony Brook, New York, 11 794 USA. Christine M. Janis [[email protected]], Department of Ecology and Evolution- ary Biology, Division of Biology and Medicine, Box G-B209, Brown University, Provi- dence, Rhode Island, 02912 USA. -
Physical Evidence of Predatory Behavior in Tyrannosaurus Rex
Physical evidence of predatory behavior in Tyrannosaurus rex Robert A. DePalma IIa, David A. Burnhamb,1, Larry D. Martinb,2, Bruce M. Rothschildb, and Peter L. Larsonc aDepartment of Paleontology, Palm Beach Museum of Natural History, Ft. Lauderdale, FL 33306; bDivision of Vertebrate Paleontology, University of Kansas Biodiversity Institute, Lawrence, KS 66045; and cBlack Hills Institute of Geological Research, Inc., Hill City, SD 57745 Edited by Mark A. Norell, American Museum of Natural History, New York, NY, and accepted by the Editorial Board May 21, 2013 (received for review September 27, 2012) Feeding strategies of the large theropod, Tyrannosaurus rex, ei- scavenger (26). Horner (25) argued that T. rex was too slow to ther as a predator or a scavenger, have been a topic of debate pursue and capture prey items (14) and that large theropods previously compromised by lack of definitive physical evidence. procured food solely through scavenging, rather than hunting Tooth drag and bone puncture marks have been documented on (11, 25). Horner also suggested that the enlarged olfactory lobes suggested prey items, but are often difficult to attribute to a spe- in T.rex were characteristic of scavengers (25). More recent cific theropod. Further, postmortem damage cannot be distin- studies (28, 29) determined the olfactory lobes of modern birds guished from intravital occurrences, unless evidence of healing is are “poorly developed,” inferring that enlarged olfactory lobes in present. Here we report definitive evidence of predation by T. rex: T. rex are actually a secondary adaptation for predation navigation a tooth crown embedded in a hadrosaurid caudal centrum, sur- “to track mobile, dispersed prey” (30). -
Dinosaurs! Pdf, Epub, Ebook
DINOSAURS! PDF, EPUB, EBOOK Gail Gibbons | 32 pages | 05 Mar 2009 | Holiday House | 9780823421978 | English | New York, NY, United States Dinosaurs! PDF Book Comment and Reply. There is general agreement that some behaviors that are common in birds, as well as in crocodiles birds' closest living relatives , were also common among extinct dinosaur groups. Anchiornis - A four-winged dino-bird that resembled Microraptor. Let us know if you have suggestions to improve this article requires login. Dinosaur legs extend directly beneath the body, whereas the legs of lizards and crocodilians sprawl out to either side. November—December It has been suggested that because small mammals, squamata and birds occupied the ecological niches suited for small body size, non-avian dinosaurs never evolved a diverse fauna of small-bodied species, which led to their downfall when large-bodied terrestrial tetrapods were hit by the mass extinction event. They were wrong. New York: W. This dinosaur was likely resting in a burrow before it was abruptly buried alive. The earliest part of this time saw the spread of ankylosaurians, iguanodontians , and brachiosaurids through Europe , North America, and northern Africa. Edward Drinker Cope. Sinoceratops - A rare ceratopsian from late Cretaceous China. The battle of the sourpusses! Denisovans are an extinct species of hominid and a close relative to modern humans. Dutton , ; London : Evans Brothers Ltd , ]. Anatosaurus - This dinosaur is now known as either Anatotitan or Edmontosaurus. One of the best examples of soft-tissue impressions in a fossil dinosaur was discovered in the Pietraroia Plattenkalk in southern Italy. Neanderthals are an extinct species of hominids that were the closest relatives to modern human beings.