A Provisional Atlas of the Coleoptera of Yorkshire (Vice Counties 61 –65)

Part 4 – Suborder Polyphaga: Families Sphaeritidae, Histeridae, Ptiliidae, Leiodidae, Scydmaenidae, Silphidae.

Introduction

Please see the introductory notes in Part 1 of the Atlas. Parts 2, and 3 covering water beetles will be dealt with in due course.

This is Part 4 of the Atlas and covers the above families. The Staphylinidae will be included in part 5.

Previous reviews and reports include the following – Flint (1988), (Denton 1990, 2003, 2008), Marsh (1991, 2002).

Each species in the database is considered and in each case a distribution map representing records on the database (at the end of 2010) is presented. The number of records on the database for each species is given in the account in the form (a,b,c, d,e) where a to e are the number of records from VC61 to VC65 respectively. These figures include undated records (see comment on undated records in the paragraph below on mapping). As a recorder, I shall continue to use the Vice-county recording system, as the county is thereby divided up into manageable areas for recording purposes. For an explanation of the vice-county recording system, under a system devised in Watson (1883) and subsequently documented by Dandy (1969), Britain was divided into convenient recording areas. Thus Yorkshire was divided into “vice-counties”, numbered 61 to 65 inclusive, and notwithstanding fairly recent county boundary reorganisations and changes, the vice-county system remains a convenient one for recording purposes; in the text, reference to “Yorkshire” implies VC61 to VC65 ignoring modern boundary changes.

For some species there are many records, and for others only one or two. In cases where there are six records or less full details of the known records are given. Many common species have quite a high proportion of recent records. This is because the older recorders tended not to record species they regarded as common quite as assiduously as we do today.

“Recent” records are those considered to be later than 1 January 1990 – this is also the break point used in the production of the distribution maps.

Habitat

A brief statement on habitat is included where known and relevant.

Taxon status

The accepted national status of each taxon is given, this being the status as defined in JNCC’s Recorder6 data program. Status is included if Notable B or above. Please bear in mind that many of these statuses are now in need of revision, in the light of recent knowledge of distribution and frequency. The following status criteria are derived from Shirt (1987) and Hyman and Parsons (1994):

RED DATA BOOK CATEGORY 1. RDB1 – ENDANGERED - Species which are known or believed to occur only as a single population within one 10km square of the OS National Grid.

RED DATA BOOK CATEGORY 2. RDB2 – VULNERABLE - Species which are declining throughout their range.

RED DATA BOOK CATEGORY 3. RDB3 – RARE - Species which are estimated to exist in only 15 or fewer 10km squares; this criterion may be relaxed where populations are likely to exist in more than 15 squares but occupy small areas of especially vulnerable habitat.

RED DATA BOOK CATEGORY I. RDB I – INDETERMINATE - Taxa considered to be endangered, Vulnerable or Rare, but where there is not enough information to say which of the three categories (RDB1 to 3) is appropriate.

RED DATA BOOK CATEGORY K. RDBK – INSUFFICIENTLY KNOWN - Taxa recently discovered or recognised in Great Britain, which may prove to be more widespread in the future (although some recent discoveries may be placed in other categories if the group to which they belong is thought to be not under-recorded).

PROVISIONAL RED DATA BOOK. Prdb - The prefix ‘p’ before any red Data Book category implies that the grading is provisional.

Notable A – species which do not fall within the RDB categories but are thought to occur in 30 or fewer 10km squares of the National Grid, or for less well-recorded groups, within seven or fewer vice-counties. Notable B – species which do not fall within the RDB categories but thought to occur in between 31 and 100 10km squares of the National Grid. Nationally Notable - species which are estimated to occur within the range of 16 to 100 10km squares. (subdivision into Notable A and Notable B is not always possible because there may be insufficient information available). No generally superseded by Nationally Scarce, and therefore no longer much used in modern literature.

Phenology Details are given for the range of months in the year for which there are records, with or without a peak month of occurrence, where the number and detail of records are sufficient to give a meaningful conclusion.

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Maps Distribution maps have been produced using Dr Alan Morton’s DMAP software (version 7.0b). It may be worthwhile to note that as DMAP is asked to map pre- and post-1990 records in this application, any undated records will not be mapped.

Nomenclature The YNU database is presently held in a copy of Recorder 6, which uses a new Coleoptera checklist (Duff, 2008) and this replaces that of Pope (1977); the new list incorporates a very large number of taxon name changes. Occasionally I receive records using “noddy names” (see Cooter, (1999) and Key (1999) for an excellent treatise on this subject), e.g. “Snail Hunter”, “Crucifix Beetle” and “Heath-glory ground beetle” (Cychrus caraboides, Panagaeus crux-major and Carabus nitens respectively). Such inventions were designed for use in political documents, and seem to have originated somewhere within Natural England (previously English Nature) or JNCC. For the purposes of serious science these terms are, in my opinion, vague, confusing and unhelpful. I do not accept records using such names only, unless accompanied by specimens or other convincing evidence.

Quality of the records A regrettable aspect of some of the records within the YNU database in particular (and I suspect sometimes within entomological datasets in general) is the lack of attention to detail accompanying the record, especially of accurate grid references, lack of pertinent comments as to the micro-habitat of the capture, its association with plants/animals, and sometimes, in the case of some modern records, a lack of care in determination, and the recording of the sex of the specimen. Old records, although sound, tend not to give much or indeed any of this detail. Old records on the database so rarely give habitat details but sometimes interpretation of the site name can give clues to the habitat type. Grid references could not be applied before 1930 anyway, and grid references do not figure in the paper archive of the YNU Coleoptera database until the late 1970s. Apparently it was not considered necessary to give that level of accuracy. Entering records onto Recorder6 requires a grid reference, and with older records I have added a grid reference where I believe this has been possible. If the site name is a vague one I have used a 10km reference, and where a 1km reference is deemed accurate enough I have included that. There can be little excuse with modern records not to include some accurate comment with the record, and at least a six-figure grid reference.

Taxonomic knowledge and expertise have increased greatly since the days of Walsh, Bayford and Stainforth. Much good literature now exists for the taxonomist in the form of dichotomous keys, and when identifying specimens, especially with critical species pairs or groups, too much reliance on the outcome of keys can be placed on a determination, without apparently considering the habitat in which the specimen was taken, or the host plant/animal. Atlases of the national distributions of various orders of insect are now becoming available and should always be consulted. These often give a guide to the likelihood or otherwise of a determination, especially in scarce taxa. Museum collections containing correctly determined and labelled specimens are invaluable in accurate recording; museum collections are there for our use and should be consulted. If a species is ‘new’ to a vice-county or is very significant in some other respect, a second opinion should always be sought or an acknowledged specialist in the group consulted. A voucher should always be retained by the collector. The recorder can only deal with the information he is given, and often has to beware of identifications made on photographic evidence only, and identifications of ‘difficult’ species by non-specialists. It takes many years to become competent in many areas of entomology, and the recorder often has to rely on the expertise of co-workers, especially when records covering large Orders are being documented.

Although some may disagree, I believe any record of a species afforded Notable B or above, especially if a member of a ‘difficult’ beetle group, should be confirmed by one specialising in that group before the record is entered onto the database. This of course would be difficult to police with large numbers of records and/or large Orders.

Contributors With approaching 200 years of beetle recording in Yorkshire many persons’ names, or simply initials, appear in the records. The initials used of the persons mentioned in the species accounts are listed below alphabetically.

AB - Brindle A ABD – Drane AB AEW - Winter AE AG - Godfrey A AM - Matthews A CJ - Johnson C DB – Bateson D DGH – Hemingway DG DM – Maude D DS – Stenhouse D EAW - Waterhouse EA EFG – Gilmore EF EG – Green E EJS - Smith EJ EWA - Aubrook EW FEK - Kenington FE GBW - Walsh GB HB - Britten H (jr) HMR - Russell HM HP – Pollard H JC - Cooter J

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JF – Jim Flanagan JHF - Flint JH JMB – Brown JM JRB – Beaumont JR JRD - Dibb JR KNAA – Alexander KNA MCB - Barnes MC ME - Eyre M MLD - Denton ML MLT - Thompson ML PB - Buckland P PS - Skidmore P PT – Thorpe P RBA – Angus RB RCW - Welch RC RGB - Booth RG RJM - Marsh RJ RL - Lawson R RRUK - Utthoff-Kaufmann RR RSK – Key RS RW - Wilson R SLM – Mosley SL TB – Bean T TW - Wilkinson T WAE - Ely WA WB – Bunting W WCH - Hey WC WDH - Hincks WD WF – Foggitt W WJF - Fordham WJ WRD - Dolling WR

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HISTERIDAE Gyllenhal, 1808

SPHAERITES Duftschmid, 1805 glabratus (Fabricius, 1792)

Sphaerites glabratus (Fabricius, 1792) 4 WATSONIAN 3 (0,4,0,1,0). 5 records, one YORKSHIRE 2 undated (MLT) therefore not 1 mapped. Recorded from pitfall 0 traps and decaying fungi, in 9 woodland situations. 8 Appletreewick SE0560 in 1938 7 then no records until 1999 6 (Garbutt Wood SE5083) and 2000 5 (Beech Wood SE5982). Very 4 scarce. 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 6/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subfamily ABRAEINAE MacLeay, 1819

Tribe ABRAEINI MacLeay, 1819

ABRAEUS Leach, 1817 granulum Erichson, 1839

Abraeus granulum Erichson, 1839 4 WATSONIAN 3 (0,0,2,0,0) Notable A. Rare in YORKSHIRE 2 Yorkshire. There are only two 1 records on the YNU database and 0 both are recent: Worsborough 9 Country Park (SE ) 1/1997 (JDC 8 det RJM), in rotten wood; Rivelin 7 (SK ) 1/9/1998 (EJS), in rotten 6 wood. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 6/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

4 perpusillus (Marsham, 1802)

Abraeus perpusillus (Marsham, 1802) 4 WATSONIAN 3 (3,7,5,1,1) Widely distributed. YORKSHIRE 2 Recorded from Fagus, Ulmus and 1 possibly Salix, in rotting wood. 0 Also in fungi under the bark of 9 Fagus. Associated with old trees 8 but not necessarily forest 7 situations. Also from subfossil 6 peat deposits. (vi-x) 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 6/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe PLEGADERINI Portevin, 1929

PLEGADERUS Erichson, 1834 dissectus Erichson, 1839 NO RECORDS vulneratus (Panzer, 1796)

Plegaderus vulneratus (Panzer, 1797) 4 WATSONIAN 3 (0,0,2,1,0). Rare in Yorkshire. YORKSHIRE 2 Described as new to Britain (Allen, 1 1968) on specimens from Surrey 0 in 1963. Three Yorkshire records: 9 Cantley SE6202, 28/4/1966 (PS); 8 Sandall Beat SE6103, 5/5/1966 7 (PS); Brayton Barff SE5830, 6 3/7/2002 (RJM). All three records 5 from under bark of fallen conifers. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 6/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

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Tribe ACRITINI Wenzel, 1944

ACRITUS LeConte, 1853 homoeopathicus Wollaston, 1857 NO RECORDS

nigricornis (Hoffmann, J., 1803)

Acritus nigricornis (Hoffmann, J., 1803) 4 WATSONIAN 3 (4,3,6,1,0) Widely distributed in YORKSHIRE 2 the east and south. In compost 1 and manure heaps and in grass 0 and hay piles.Also in 9 archaeological deposits. (iii-x) 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 6/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

AELETES Horn, 1873 atomarius (Aubé, 1842)

Aeletes atomarius (Aubé, 1842) 4 WATSONIAN 3 (0,1,0,0,0). There is a single YORKSHIRE 2 modern record on the database: 1 Duncombe Park SE6082 10/9/1985 0 (RJM) (Marsh, 1988a). The species 9 is also recorded in archaeological 8 deposits of medieval date: 7 Conisbrough SK5198, 2001 (PB). 6 Sometimes associated with Dorcus 5 burrows. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

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HALACRITUS Schmidt, J., 1893

ACRITUS sensu Fowler, 1889 partim non LeConte, 1853 punctum (Aubé, 1843) NO RECORDS

Tribe TERETRIINI Bickhardt, 1916

TERETRIUS Erichson, 1834 fabricii Mazur, 1972 NO RECORDS

Subfamily SAPRININAE Blanchard, 1845

SAPRINUS Erichson, 1834 aeneus (Fabricius, 1775)

Saprinus aeneus (Fabricius, 1775) 4 WATSONIAN 3 (3,2,5,0,0) Most Yorkshire records YORKSHIRE 2 are from old rather dry carcasses 1 of rabbits and birds. Several of our 0 records are old, undated and 9 hence unmapped. (vi-ix) 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

immundus (Gyllenhal, 1827) NO RECORDS

7 planiusculus Motschulsky, 1849

Saprinus planiusculus Motschulsky, 1849 4 WATSONIAN 3 (1,0,0,0,0) Notable B. A single YORKSHIRE 2 record on the YNU database: 1 Spurn TA3910, 23/7/1963 (RCW). 0 Specimens recovered from a 9 stranded, dead porpoise. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

semistriatus (Scriba, 1790)

Saprinus semistriatus (Scriba, 1790) 4 WATSONIAN 3 (9,9,15,7,1) Widespread. Fairly YORKSHIRE 2 frequent in the south, and one of 1 our commonest histerids. In 0 carcasses of cat, fox, rabbit, 9 hedgehog, sheep and various 8 birds, and in baited pitfalls in 7 woodland and grassland. No 6 records from dung. (v-ix) 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

subnitescens Bickhardt, 1909 NO RECORDS

8 virescens (Paykull, 1798)

Saprinus virescens (Paykull, 1798) 4 WATSONIAN 3 (1,0,0,2,0) Very rare in Yorkshire. YORKSHIRE 2 There are three records on the 1 database: Elvington (SE7047), 0 21/8/1941 (MDB); Gateforth 9 (SE5628), 15/5/1945 (JRD); Ox 8 Close (SE3646), 25/5/1953 (HMR) 7 with no habitat details given. 6 Believed to be a predator on 5 larvae of Phaedon (Col., 4 Chrysomelidae) (Halstead, 1963) 3 (Cox, 2007) 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

HYPOCACCUS Thomson, C.G., 1867

SAPRINUS sensu Fowler, 1889 partim non Erichson, 1834

Subgenus HYPOCACCUS Thomson, C.G., 1867 crassipes (Erichson, 1834) NO RECORDS metallicus (Herbst, 1792) NO RECORDS rugiceps (Duftschmid, 1805)

Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805) 4 WATSONIAN 3 (1,0,0,0,0) Notable A. A single YORKSHIRE 2 record of this generally coastal 1 species: Spurn (TA4112), 0 20/6/1990 (MLD), in sheep dung. 9 Nationally, recorded from both 8 dung and carrion. 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

9 rugifrons (Paykull, 1798)

Hypocaccus (Hypocaccus) rugifrons (Paykull, 1798) 4 WATSONIAN 3 (0,1,1,0,0) Notable B. There are YORKSHIRE 2 two records on the YNU database: 1 Eggborough (SE5724), 26/6/1986 0 (RJM), a specimen in a window in 9 lighted building; Forge Valley SE98, 8 undated (and so not mapped), 7 (WCH). Usually in dung and carrion 6 (Halstead, 1963) 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus BAECKMANNIOLUS Reichardt, 1926 dimidiatus (Illiger, 1807)

Hypocaccus (Baeckmanniolus) dimidiatus (Illiger, 1807) 4 WATSONIAN 3 (1,2,0,0,0) Notable B. Very rare YORKSHIRE th 2 and unrecorded since the mid-19 1 century. Three records are 0 mentioned in the database: 9 Sandsend (NZ8613), undated, 8 (WJF); Stockton-on-Tees (NZ42), 7 undated and with recorder not 6 named; Spurn (?TA41), June 1853 5 (WCH). The undated records have 4 not been mapped. Halstead (1963) 3 quotes the species as being coastal 2 on sandhills. In dung and carrion. 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

GNATHONCUS Jacquelin du Val, 1857 buyssoni Auzat, 1917 NO RECORDS communis (Marseul, 1862) NO RECORDS

10 nannetensis (Marseul, 1862)

Gnathoncus nannetensis (Marseul, 1862) 4 WATSONIAN 3 (3,5,1,2,0) Widely distributed but YORKSHIRE 2 scarce, with 11 records. Our records 1 have the species variously recorded 0 from a sparrow nest, in pitfall traps, 9 in poultry manure, on fence palings, 8 in fungi on fallen beech, and in a 7 pile of rotting turnips. (vi-ix) 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

rotundatus (Kugelann, 1792)

Gnathoncus rotundatus (Kugelann, 1792) 4 WATSONIAN 3 (1,0,1,2,0) Very rarely recorded in YORKSHIRE 2 Yorkshire. We have four records: 1 Pannal Ash (SE3052), 1936 and 0 1939 (RRUK), in carrion traps; 9 Margery Wood (SE2506), 2/7/1988 8 (MLD); Welwick (TA3421), 7 10/9/1999 (FEK det RJM), in 6 mealhouse debris in a farm building. 5 Nationally recorded from bird nests, 4 bat roosts, carrion and in granaries 3 (Halstead, 1963). 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

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MYRMETES Marseul, 1862 paykulli Kanaar, 1979

Myrmetes paykulli Kanaar, 1979 4 WATSONIAN 3 (0,5,0,0,0) Notable B. We have five YORKSHIRE 2 records: Harwood Dale (SE9695) 1 4/4/1926 (GBW); Helwath Beck 0 (SE99) 24/10/1936 (HB); Langdale 9 End (SE9492) 1950 (GBW); 8 Scarborough (TA08) 1900 (RL), in 7 ants’ nest; “York”, undated and 6 anon (therefore unmapped). An 5 interesting county distribution. 4 Associated with Formica rufa. 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 7/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subfamily DENDROPHILINAE Reitter, 1909

Tribe DENDROPHILINI Reitter, 1909

DENDROPHILUS Leach, 1817 punctatus (Herbst, 1792)

Dendrophilus punctatus (Herbst, 1792) 4 WATSONIAN 3 (0,1,0,0,0) A single record exists on YORKSHIRE 2 the database: Farndale (SE6697) 1 1935 (GBW), in nest of Formica 0 rufa. A myrmecophilous species. 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

12 pygmaeus (Linnaeus, 1758)

Dendrophilus pygmaeus (Linnaeus, 1758) 4 WATSONIAN 3 (0,0,2,0,0) There are two YNU YORKSHIRE 2 records: Stephen Wood (SE2108) 1 12/4/1963 (EWA), with Formica 0 lugubris (Nat. 1964:21); Ewden 9 (SK2396) 21/2/2003 (EJS), in nest 8 of Formica lugubris. Halstead 7 (1963) quotes Formica rufa as a 6 host. Perhaps F. lugubris is a 5 more common host in the North. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

xavieri Marseul, 1873 NO RECORDS

KISSISTER Marseul, 1862 CARCINOPS sensu Fowler, 1889 partim non Marseul, 1855 minimus (Laporte, 1840)

Kissister minimus (Aubé, 1850) 4 WATSONIAN 3 (0,0,1,0,0) There is a single YORKSHIRE 2 Yorkshire record: Thorne Moor 1 (SE7215) 21/9/2009 (AG det 0 MLD), collected in a mire pitfall 9 trap. There are also nearby 8 records from North Lincolnshire. 7 Usually recorded from ground 6 litter. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

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Tribe PAROMALINI Reitter, 1909

CARCINOPS Marseul, 1855 pumilio (Erichson, 1834)

Carcinops pumilio (Erichson, 1834) 4 WATSONIAN 3 (8,2,14,3,1) With 28 records, this YORKSHIRE 2 species is widely distributed and 1 fairly frequent, except in the far 0 north of the county. Recorded from 9 various habitats including compost 8 and manure heaps, poultry houses, 7 grass and straw heaps, old bones, 6 rotting turnips, from fungi on a 5 dead horse chestnut, and by 4 general sweeping. (v-xi, peak vii). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

PAROMALUS Erichson, 1834

MICROLOMALUS Lewis, 1907 flavicornis (Herbst, 1792)

Paromalus flavicornis (Herbst, 1792) 4 WATSONIAN 3 (0,4,9,3,0) Widely but rather YORKSHIRE 2 centrally distributed. Associated 1 with trees: our records include 0 under bark of beech Fagus and 9 oaks Quercus, and from Polyporus 8 sp. On horse chestnut Aesculus. 7 With 10 of these records recent the 6 species may be on the increase. 5 (i-x, peak vi). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

parallelepipedus (Herbst, 1792) NO RECORDS

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Subfamily ONTHOPHILINAE MacLeay, 1819

ONTHOPHILUS Leach, 1817 punctatus (Müller, O.F., 1776) NO RECORDS striatus (Forster, 1771)

Onthophilus striatus (Forster, 1771) 4 WATSONIAN 3 (18,10,60,10,1) Our commonest YORKSHIRE 2 histerid with nearly 100 records. 1 Widely distributed – most records 0 are from the south and centre of 9 the county. From a wide variety of 8 habitats including rotting 7 vegetation, horse dung, ground 6 litter in gardens and reed beds, in 5 compost and straw heaps. (ii-xi, 4 peak v) 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subfamily TRIBALINAE Bickhardt, 1914 EPIERUS Erichson, 1834 comptus Erichson, 1834 NO RECORDS

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Subfamily HISTERINAE Gyllenhal, 1808

MARGARINOTUS Marseul, 1854 HISTER sensu Fowler, 1889 partim non Linnaeus, 1758

Subgenus PTOMISTER Houlbert & Monnot, 1923 brunneus (Fabricius, 1775)

Margarinotus (Ptomister) brunneus (Fabricius, 1775) 4 WATSONIAN 3 (14,6,23,5,2) Widely distributed YORKSHIRE 2 and frequent. In a wide variety of 1 putrefying dung and carrion 0 situations including the dung of 9 horse, cow and dog, in carcasses 8 of fox, cat, sheep, roe deer, and in 7 baited pitfalls in woodland and on 6 peat moors. Also reported from 5 county archaeological deposits of 4 medieval date. A species better 3 known to coleopterists as Hister 2 impressus or H. cadaverinus. 1 (i-ix, peak vi) 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

merdarius (Hoffmann, J., 1803)

Margarinotus (Ptomister) merdarius (Hoffmann, J., 1803) 4 WATSONIAN 3 (2,0,14,0,0) Widely distributed YORKSHIRE 2 but with a marked distribution in 1 the south of the county. Mainly 0 recorded from putrefying 9 vegetation, with a single record 8 from a carrion habitat. (ii-viii, 7 peak vii). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

16 striola (Sahlberg, C.R., 1819)

Margarinotus (Ptomister) striola (Sahlberg, C.R., 1819) 4 WATSONIAN 3 (2,7,14,7,8) Widely distributed YORKSHIRE 2 with a marked lack of easterly or 1 coastal records. Recorded in 0 Yorkshire from baited pitfalls and 9 carrion traps, putrefying fungi and 8 rotting vegetation, dead sheep and 7 from dry cowdung. (v-ix, peak vi) 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

ssp. succicola (Thomson, C.G., 1862) NO RECORDS

Subgenus STENISTER Reichardt, 1926 obscurus (Kugelann, 1792) NO RECORDS

Subgenus PARALISTER Bickhardt, 1917 neglectus (Germar, 1813)

Margarinotus (Paralister) neglectus (Germar, 1813) 4 WATSONIAN 3 (0,1,0,1,0) Rare in the county. YORKSHIRE 2 There are only two YNU records: 1 Leeds (SE3035) 7/10/1950 (JHF); 0 Rawcliffe Meadows (SE5755) 9 9/5/1997 (MLD), in ground litter. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

17 purpurascens (Herbst, 1792)

Margarinotus (Paralister) purpurascens (Herbst, 1792) 4 WATSONIAN 3 (7,1,2,0,0) Scarce with a south- YORKSHIRE 2 easterly distribution. From rotting 1 vegetation and in compost and 0 straw heaps. Only one recent 9 record. (vi-viii) 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 9/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

ventralis (Marseul, 1854)

Margarinotus (Paralister) ventralis (Marseul, 1854) 4 WATSONIAN 3 (8,6,24,5,2) A widespread and YORKSHIRE 2 frequent histerid. Recorded from 1 dung of horse and cow, in dead 0 rabbit, hedgehog and various 9 birds, baited pitfalls, and in ground 8 litter in woodland. (iv-xii, peak v). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 10/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

18

Subgenus PROMETHISTER Kryzhanovskij, 1966

GRAMMOSTETHUS sensu auctt. partim non Lewis, 1906 marginatus (Erichson, 1834)

Margarinotus (Promethister) marginatus (Erichson, 1834) 4 WATSONIAN 3 (4,2,1,0,0) Notable B. With only 7 YORKSHIRE 2 county records, a very scarce 1 species. No recent records, the 0 latest being North Duffield Carrs 9 (SE6937) 29/5/1987 (MLD). 8 Apparently restricted to moles’ 7 nests, an observation borne out by 6 all the YNU records. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 10/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

HISTER Linnaeus, 1758 bissexstriatus Fabricius, 1801 NO RECORDS illigeri Duftschmid, 1805 NO RECORDS quadrimaculatus Linnaeus, 1758 NO RECORDS quadrinotatus Scriba, 1790 NO RECORDS unicolor Linnaeus, 1758

Hister unicolor Linnaeus, 1758 4 WATSONIAN 3 (5,4,7,0,2) Widespread but with YORKSHIRE 2 few records – most YNU records 1 are post-1990, so the species 0 may be on the increase. 9 Recorded from dead birds and in 8 the dung of horse and cow. 7 (v-viii) 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 10/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

19

ATHOLUS Thomson, C.G., 1859

HISTER sensu Fowler, 1889 partim non Linnaeus, 1758 bimaculatus (Linnaeus, 1758)

Atholus bimaculatus (Linnaeus, 1758) 4 WATSONIAN 3 (2,1,10,2,0) Widely distributed in YORKSHIRE 2 the southern half of the county, 1 with very few recent records. In 0 compost and manure heaps and in 9 rotting vegetation. (vi-xi, peak ix) 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 10/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

duodecimstriatus (Schrank, 1781)

Atholus duodecimstriatus (Schrank, 1781) 4 WATSONIAN 3 (3,4,16,3,1) Fairly frequent and with YORKSHIRE 2 a rather central distribution, the 1 species has been recorded from 0 manure and warm compost heaps, 9 and from horse dung and carrion. 8 (iii-ix) 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 10/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subfamily HETAERIINAE Marseul, 1857

HAETERIUS Dejean, 1833 ferrugineus (Olivier, 1789) NO RECORDS

20

PTILIIDAE Erichson, 1845

Subfamily PTILIINAE Erichson, 1845

Tribe PTENIDIINI Flach, 1889

NOSSIDIUM Erichson, 1845 pilosellum (Marsham, 1802)

Nossidium pilosellum (Marsham, 1802) 4 WATSONIAN 3 (2,0,0,0,0) Nationally Notable. YORKSHIRE 2 Danthorpe (TA2432) 31/7/2001 1 (WRD det CJ), in bracket fungus on 0 tree stump; Burton Constable 9 (TA1737) 17/7/2002 (WRD), in old 8 Polyporus squamosus on ash stump. 7 These are the only county records. 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 10/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

PTENIDIUM Erichson, 1845

Subgenus GRESSNERIUM Ganglbauer, 1899 gressneri Erichson, 1845

Ptenidium (Gressnerium) gressneri Erichson, 1845 4 WATSONIAN 3 (0,0,0,1,0) Nationally Notable. The YORKSHIRE 2 only record we have on the YNU 1 database is: Studley Royal Park 0 (SE26) 1870 (EAW) (Johnson, 9 1990). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 10/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

21

Subgenus MATTHEWSIUM Flach, 1889

laevigatum Erichson, 1845

Ptenidium (Matthewsium) laevigatum Erichson, 1845 4 WATSONIAN 3 (8,8,19,8,1) Widespread and YORKSHIRE 2 frequent. Recorded from grass heaps, 1 straw bales, compost heaps, nests of 0 mole and birds, and in ground litter in 9 reed beds and by lagoons. Often 8 abundant when found. (ii-ix, peak vi) 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 10/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

turgidum Thomson, C.G., 1855

Ptenidium (Matthewsium) turgidum Thomson, C.G., 1855 4 WATSONIAN 3 (0,2,0,1,0) – RDBK. Very rare in YORKSHIRE 2 Yorkshire. There are three records 1 on the database, only two of these 0 being validated: Duncombe Park 9 (SE6082) 26/5/1987 (RJM teste CJ); 8 Studley Royal Park (SE2969) 7 19/6/1989 (RJM teste CJ); 6 Scarborough (TA08) undated and 5 hence unmapped (TW). The latter 4 record is doubtful and would require 3 a voucher to confirm, while the first 2 two are based on specimens from 1 the red-rotted heartwood in oak logs 0 in old parkland sites. 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 13/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus WANKOWIZIUM Flach, 1889 brenskei Flach, 1887 NO RECORDS

22 intermedium Wankowicz, 1869

Ptenidium (Wankowiz ium) intermedium Wankowicz , 1869 4 WATSONIAN 3 (5,1,9,5,1) Mainly distributed in the YORKSHIRE 2 central, more sheltered regions of 1 the county, away from the highland 0 areas. An infrequent ptiliid 9 associated with wetlands, being 8 recorded from decaying vegetation 7 on river banks, lake edges and in 6 reed beds. (iii-ix, peak vi) 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 13/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus PTENIDIUM Erichson, 1845 formicetorum Kraatz, 1851

Ptenidium (Ptenidium) formicetorum Kraatz , 1851 4 WATSONIAN 3 (0,0,1,0,0) There is a single record YORKSHIRE 2 on the YNU database: Fenwick 1 cemetery (SE5815) 20/9/2008 (RJM 0 det CJ), by sweeping around edges 9 of a compost heap in shade of trees. 8 According to Alexander (2002), the 7 main habitat is probably rotting 6 wood, but the species is known from 5 Formica nests (hence its specific 4 name) and decaying vegetable 3 material. 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 13/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

23 fuscicorne Erichson, 1845

Ptenidium (Ptenidium) fuscicorne Erichson, 1845 4 WATSONIAN 3 (13,7,27,6,1) Widespread and fairly YORKSHIRE 2 frequently recorded from the eastern 1 half of the county, with no records 0 from the Pennines or the high 9 ground of the northeast. In 8 vegetable refuse, compost heaps, 7 reedbed detritus, and ground litter 6 at pond and drain edges. 5 (iii-x, peak vi) 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 13/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

longicorne Fuss, 1868 NO RECORDS

punctatum (Gyllenhal, 1827)

Ptenidium (Ptenidium) punctatum (Gyllenhal, 1827) 4 WATSONIAN 3 (4,5,7,0,2) Widely distributed but YORKSHIRE 2 rather scarce, with few recent 1 records. Recorded from rotting 0 straw, grass piles on drain edges 9 and roadsides, and in seaweed and 8 strandline refuse on the Humber 7 shore. (v-xi) 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 13/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

24 pusillum (Gyllenhal, 1808)

Ptenidium (Ptenidium) pusillum (Gyllenhal, 1808) 4 WATSONIAN 3 (27,5,73,22,7). Frequent and YORKSHIRE 2 widespread, one of our commonest 1 ptiliids. Recorded from a very wide 0 variety of habitats including dung of 9 horse and cow, in piles of rotting 8 mowings, hay and compost, in 7 rotting fungi, in ground litter by 6 pools and drains, in moss and grass 5 tussocks in winter. In gardens, 4 woodland, churchyards, reedbeds 3 and grassland. Few records from 2 high ground. (i-xii, peak vi) 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 13/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus GILLMEISTERIUM Flach, 1889 nitidum (Heer, 1841)

Ptenidium (Gillmeisterium) nitidum (Heer, 1841) 4 WATSONIAN 3 (20,22,87,23,15) Common and YORKSHIRE 2 widespread, more so than pusillum. 1 Habitats as widely varied and similar 0 to that species. 9 (ii-xi, peak vi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 13/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

25

Tribe PTILIINI Erichson, 1845

EURYPTILIUM Matthews, A., 1872 gillmeisteri Flach, 1889

Euryptilium gillmeisteri Flach, 1889 4 WATSONIAN 3 (0,0,0,1,0) There is a single record YORKSHIRE 2 on the YNU database and is the sole 1 British record: Fountains Abbey 0 (SE2768) 3/9/1995 (RJM det CJ), 9 several specimens sieved from 8 deeply shaded oak leaf litter heavily 7 impregnated with bird droppings. Its 6 accepted habitat is old woodland, 5 where it is recorded elsewhere in 4 Europe from decaying organic 3 matter on the woodland floor. See 2 Johnson (2001) for details and 1 differentiation from saxonicum. 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 13/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

saxonicum (Gillmeister, 1845)

Euryptilium saxonicum (Gillmeister, 1845) 4 WATSONIAN 3 (0,0,2,0,0) Nationally Notable. Rare YORKSHIRE 2 in Yorkshire with two records: 1 Farnley Tyas (SE1612) 1975 (CJ); 0 Hey Wood (SE1512) 1975 (CJ). 9 Johnson (1990) also includes a 8 record for Honley. In woodland 7 ground litter (e.g. in grass piles). 6 Known habitats also include carrion. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 13/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

26

PTILIOLA Haldeman, 1848 brevicollis (Matthews, A., 1860) NO RECORDS kunzei (Heer, 1841)

Ptiliola kunz ei (Heer, 1841) 4 WATSONIAN 3 (1,3,9,3,0) With 16 records on the YORKSHIRE 2 database, only one of these is post- 1 1990. In compost heaps, grass piles 0 and old, rather dry horse dung. 9 (iv-x). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 14/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

PTILIOLUM Flach, 1888

Subgenus PTILIOLUM Flach, 1888 fuscum (Erichson, 1845)

Ptiliolum (Ptiliolum) fuscum (Erichson, 1845) 4 WATSONIAN 3 (4,14,11,8,4) Widespread and YORKSHIRE 2 frequent across the county. Mainly 1 from grassland sites and gardens in 0 the following microhabitats: 9 compost heaps, dung of horse and 8 sheep, straw piles, heaps of 7 mowings, in a dead bird, and in 6 ground litter around pond edges and 5 drains. (iv-x, peak vii) 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 14/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

27 marginatum (Aubé, 1850)

Ptiliolum (Ptiliolum) marginatum (Aubé, 1850) 4 WATSONIAN 3 (0,0,1,0,0) There is a single county YORKSHIRE 2 record: Worsborough Reservoir 1 (SE3403) 1975 (CJ). This is noted in 0 Johnson (1990). The usually 9 accepted habitat is woodland, in 8 dung and in putrefying material. 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 14/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

sahlbergi (Flach, 1888)

Ptiliolum (Ptiliolum) sahlbergi (Flach, 1888) 4 WATSONIAN 3 (0,0,0,1,0) A single Yorkshire YORKSHIRE 2 record: Malham Tarn (SD86) 1 Aug/Sept 1967 (AB, CJ, det CJ). 0 Several specimens found during this 9 period in piles of decaying grass. A 8 boreo-alpine species in Europe, 7 there is a record from the Scottish 6 Highlands. See notes in Johnson 5 (1990) and Johnson (1968). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 14/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

28 spencei (Allibert, 1844)

Ptiliolum (Ptiliolum) spencei (Allibert, 1844) 4 WATSONIAN 3 (0,4,13,4,2) A widespread ptiliid YORKSHIRE 2 recorded from compost heaps, piles 1 of grass and old straw, and from old 0 decaying fungi on woodland floor. A 9 few of our records are pre-1967 and 8 are unreliable owing to confusion 7 with other members of the genus. 6 (iii-x) 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 14/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus EUPTILIUM Flach, 1889 caledonicum (Sharp, 1871) NO RECORDS

schwarzi (Flach, 1887)

Ptiliolum (Euptilium) schwarz i (Flach, 1887) 4 WATSONIAN 3 (0,0,2,0,0) Nationally a rare species, YORKSHIRE 2 with only two records on our 1 database: Farnley Tyas (SE1612) 0 12/7/1963 (EWA det CJ); Mollicar 9 Wood (SE1613) 1975 (CJ). A 8 woodland species recorded from 7 decaying vegetation on the 6 woodland floor. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 22/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

29

ACTIDIUM Matthews, A., 1868 aterrimum (Motschulsky, 1845)

Actidium aterrimum (Motschulsky, 1845) 4 WATSONIAN 3 (0,0,0,1,0) There is a single record YORKSHIRE 2 on the database: River Skell, Spa Gill 1 (SE2568 or SE2569) 1871 (EAW). 0 Nationally, recorded from fine 9 alluvium by rivers and streams. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 22/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

coarctatum (Haliday, 1855) NO RECORDS

OLIGELLA Motschulsky, 1869 foveolata (Allibert, 1844)

Oligella foveolata (Allibert, 1844) 4 WATSONIAN 3 (1,0,10,2,0) Nationally Notable. YORKSHIRE 2 There are 13 records on the YNU 1 database, these being mainly due to 0 Colin Johnson in the period 1970- 9 1975 (Johnson, 1990). There is a 8 single recent record : Askham Bog 7 (SE5748) 23/6/1996 (RGB). A 6 grassland species in dungheaps and 5 other putrefying material. (vi-xi). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 22/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

insignis (Matthews, A., 1861) NO RECORDS

30

intermedia Besuchet, 1971

Oligella intermedia Besuchet, 1971 4 WATSONIAN 3 (0,0,1,0,0) There is only one record YORKSHIRE 2 on the YNU database: Melton Wood 1 (SE5103) 2/7/1986 (RJM det CJ), a 0 single specimen sieved from piles of 9 grass cuttings. At the time of 8 capture this was only the second 7 British record. See Marsh (1988b) 6 and Johnson (1990). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 22/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

PTILIUM Gyllenhal, 1827 affine Erichson, 1845 NO RECORDS caesum Erichson, 1845 NO RECORDS exaratum (Allibert, 1844) NO RECORDS

horioni Rosskothen, 1934

Ptilium horioni Rosskothen, 1934 4 WATSONIAN 3 (0,0,4,0,0) Four records: Melton YORKSHIRE 2 Wood (SE5103) 2/7/1986 (RJM det 1 CJ); Sprotbrough (SE5402) 0 6/9/1988 (RJM det CJ); Brodsworth 9 Hall (SE5007) 7/9/1992 (RJM); 8 Pollington Church (SE6119) 7 14/5/1998 (RJM). In rotting piles of 6 grass cuttings and compost heaps. 5 Very scarce. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 22/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

31 myrmecophilum (Allibert, 1844)

Ptilium myrmecophilum (Allibert, 1844) 4 WATSONIAN 3 (0,5,3,0,0) Very scarce with eight YORKSHIRE 2 records on the YNU database. 1 Associated with Formica lugubris 0 nests. Our records are dated 1930 – 9 1970. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 22/8/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

MICRIDIUM Motschulsky, 1869 halidaii (Matthews, A., 1868) NO RECORDS

MILLIDIUM Motschulsky, 1855 minutissimum (Ljungh, 1804)

Millidium minutissimum (Ljungh, 1804) 4 WATSONIAN 3 (0,1,3,0,1) Nationally Notable. YORKSHIRE 2 Another rarely recorded species of 1 grassland situations, in compost and 0 dung heaps. Our five records are: 9 Bolton on Swale (SE2499) 1975 (CJ); 8 Ellerby (NZ79214) 1975 (CJ); Greno 7 Wood (SK3295) 1975 (CJ); 6 Hoylandswaine (SE2604) 1970 (CJ); 5 Langsett (SE2101) 1975 (CJ). There 4 are no recent records. 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 6/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

32

Tribe PTINELLINI Reitter, 1906

MICROPTILIUM Matthews, A., 1872 palustre Kuntzen, 1914 NO RECORDS pulchellum (Allibert, 1844) NO RECORDS

PTINELLA Motschulsky, 1844 aptera (Guérin-Méneville, 1839)

Ptinella aptera (Guérin-Méneville, 1839) 4 WATSONIAN 3 (0,1,2,0,0) The YNU database has YORKSHIRE 2 three records of this saproxylic 1 species which is subcortical on 0 Quercus and Fagus: Strensall 9 Common (SE66) 29/4/1964 (EWA); 8 West Bretton (SE2815) 4/8/1948 7 (EWA); Smith Wood (SE1711) 6 4/9/1962 (EWA). However 5 reference to Johnson (1990) would 4 indicate that these records are 3 referrable to errabunda Johnson. 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 6/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

britannica Matthews, A., 1858

Ptinella britannica Matthews, A., 1858 4 3 WATSONIAN (0,1,0,1,0) There are two records 2 YORKSHIRE on the YNU database: Askham Bog 1 (SE5748) 1970 (CJ); York (SE6153) 0 1970 (PB), in insect remains in 9 archaeological deposits, (Johnson, 8 1990). 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

33 cavelli (Broun, 1893)

Ptinella cavelli (Broun, 1893) 4 3 WATSONIAN (2,3,9,3,0) A woodland species found under bark. Sometimes 2 YORKSHIRE numerous when found. Mainly from 1 broadleaved trees in parkland sites 0 but also from mature trees in other 9 types of location. Also recorded 8 from conifers nationally. (v-xii). 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

denticollis (Fairmaire, 1858)

Ptinella denticollis (Fairmaire, 1858) 4 3 WATSONIAN (0,2,3,2,0) Nationally Notable. 2 YORKSHIRE This rarely recorded species is 1 found in woodland under tight 0 bark of deciduous trees. Yorkshire 9 records quote willow, horse 8 chestnut and beech. (iii-vii). 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

34 errabunda Johnson, 1975

Ptinella errabunda Johnson, 1975 4 3 WATSONIAN (0,2,14,3,1) A fairly common and 2 YORKSHIRE widespread woodland species and 1 undoubtedly under-recorded owing 0 to its small size. Recorded from 9 under the bark of deciduous and 8 coniferous trees, the Yorkshire 7 records often quoting oak Quercus. 6 No post-1990 records. (v-ix). 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

limbata (Heer, 1841) NO RECORDS simsoni (Matthews, A., 1878) NO RECORDS

taylorae Johnson, 1977

Ptinella taylorae Johnson, 1977 4 3 WATSONIAN (0,0,0,0,1) There is a single record 2 YORKSHIRE on the YNU database: Thrintoft 1 (SE3293) 1975 (CJ). Another 0 subcortical woodland species. See 9 Johnson (1990). 8 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

35

PTERYX Matthews, A., 1858 suturalis (Heer, 1841)

Pteryx suturalis (Heer, 1841) 4 3 WATSONIAN (0,3,8,6,2) A widespread woodland 2 YORKSHIRE species with only two recent 1 records. Records indicate that this is 0 a subcortical species, but has been 9 recorded from piles of grass cuttings 8 placed at the bases of trees in 7 woodland situations. (iii-ix). 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subfamily ACROTRICHINAE Reitter, 1909

Tribe NEPHANINI Portevin, 1929

NEPHANES Thomson, C.G., 1859 titan (Newman, 1834)

Nephanes titan (Newman, 1834) 4 3 WATSONIAN (12,5,22,7,7) A widespread and 2 YORKSHIRE common species of dungheaps, with many records from compost piles 1 and other decaying vegetation. 0 (vii-ix). 9 8 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

36

SMICRUS Matthews, A., 1872 filicornis (Fairmaire & Laboulbène, 1855)

Smicrus filicornis (Fairmaire & Laboulbène, 1855) 4 3 WATSONIAN (0,2,0,0,0) There are two records on 2 YORKSHIRE the database, one of which is 1 undated and therefore not mapped: 0 Nunnington (SE67) undated (AM) 9 (Johnson, 1990); Rievaulx Terrace 8 (SE5784) 3/9/1999 (ABD), sieved 7 from moss, the record requiring 6 confirmation. 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

BAEOCRARA Thomson, C.G., 1859 variolosa (Mulsant & Rey, 1861)

Baeocrara variolosa (Mulsant & Rey, 1861) 4 3 WATSONIAN (0,0,0,1,0) A single record exists for 2 YORKSHIRE the county: Malham Tarn (SD8967) 1 1967 (CJ) (Johnson, 1990). A 0 species of woodland, recorded from 9 dung and other putrefying substrates. 8 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

37

Tribe ACROTRICHINI Reitter, 1909

ACTINOPTERYX Matthews, A., 1872 fucicola (Allibert, 1844) NO RECORDS

ACROTRICHIS Motschulsky, 1848 102

Subgenus ACROTRICHIS Motschulsky, 1848 arnoldi Rosskothen, 1935

Acrotrichis (Acrotrichis) arnoldi Rosskothen, 1935 4 3 WATSONIAN (1,0,0,0,0) There is a single 2 YORKSHIRE Yorkshire record: Thornton Ellers 1 (SE7245) 7/9/1989 (RJM det CJ), in 0 decaying ground litter by a pond. 9 Probably a typical habitat for this 8 species. 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

atomaria (De Geer, 1774)

Acrotrichis (Acrotrichis) atomaria (De Geer, 1774) 4 3 WATSONIAN (11,12,50,20,4) A widespread and 2 YORKSHIRE common species of decaying 1 ground litter and mosses where it 0 feeds on fungal spores. Occurring in 9 all types of habitat including 8 wetlands, woodlands, garden compost heaps and grassland, it 7 can occur in large numbers. 6 (iii-xii, peak vi). 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

38 brevipennis (Erichson, 1845) NO RECORDS cephalotes (Allibert, 1844)

Acrotrichis (Acrotrichis) cephalotes (Allibert, 1844) 4 3 WATSONIAN (0,0,1,0,0) This species is the A. 2 YORKSHIRE chevrolati (Allibert) of older lists, and 1 is a rare member of the genus. This 0 very tiny species has only been 9 recorded reliably once in Yorkshire: 8 Greno Wood (SK3295) 1975. The 7 species is recorded from, particularly, 6 old dungheaps by farms (Johnson, 1990). 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

cognata (Matthews, A., 1877)

Acrotrichis (Acrotrichis) cognata (Matthews, A., 1877) 4 3 WATSONIAN (23,15,115,31,18). A very common 2 YORKSHIRE and widespread species recorded in 1 Yorkshire from carrion, rotting fungi 0 and compost heaps, horse dung, 9 leaf litter in woodland, hedgerows, 8 and carr land and from ground litter in Glyceria beds. A distinctive 7 species owing to its overall shape 6 and the Catops-like sheen of the 5 dorsal surface. Many records from 4 the more-sheltered south and centre 3 of the county. (i-xi, peak vi). 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

39 danica Sundt, 1958

Acrotrichis (Acrotrichis) danica Sundt, 1958 4 3 WATSONIAN (7,3,30,14,3) A widespread and 2 YORKSHIRE common species particularly in 1 damp places, including pond edges, 0 Glyceria and Typha beds, drain sides 9 and moorland edges where it may 8 be found in ground litter, grass piles, straw bales and in Juncus 7 litter. Many records particularly from 6 the south and centre of the 5 recording area. (i-xii, peak vi). 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

dispar (Matthews, A., 1865) NO RECORDS

fascicularis (Herbst, 1793)

Acrotrichis (Acrotrichis) fascicularis (Herbst, 1793) 4 3 WATSONIAN (30,25,122,42,10) Perhaps the most 2 YORKSHIRE commonly encountered Acrotrichis. 1 Recorded mainly from grassland habitats (but also woodland sites) in 0 all kinds of rotting vegetation 9 situations, compost heaps, grass 8 piles, pond edges and under hay 7 bales. Many widely distributed 6 records, but not apparently from 5 very wet situations. (i-xii, peak vi). 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

40 henrici (Matthews, A., 1872)

Acrotrichis (Acrotrichis) henrici (Matthews, A., 1872) 4 3 WATSONIAN (10,0,23,2,0) A wetland species of 2 YORKSHIRE marsh litter. First recognised in 1 Britain at Denaby Ings (SE5000) in 0 1967 (Johnson, 1990), the species 9 has been noted fairly widely in the central south of the county by 8 sieving in reed beds, at pond edges, 7 in beds of Glyceria and Phragmites, 6 and has also been found by 5 sweeping in marshy places, and in 4 hay bales in flooded areas. 3 (ii-xii, peak vii). 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

insularis (Mäklin, 1852)

Acrotrichis (Acrotrichis) insularis (Mäklin, 1852) 4 3 WATSONIAN 6,8,20,4,0) A species of woodland 2 YORKSHIRE and recorded from leaf litter, grass 1 piles and straw bales. A widespread 0 and fairly frequent Acrotrichis. The 9 first British specimens were found in the grounds of Roche Abbey 8 (SK5489) in 1965 (Johnson, 1990). 7 (v-xi, peak vii). 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

41 intermedia (Gillmeister, 1845)

Acrotrichis (Acrotrichis) intermedia (Gillmeister, 1845) 4 3 WATSONIAN (8,12,82,27,3) A widespread and 2 YORKSHIRE common woodland species. May often be recognised in the field in 1 woodland situations owing to its 0 large size (relatively speaking). Often 9 very numerous. Apparently recorded 8 only from broadleaved woodland 7 with no mention of conifers in the 6 Yorkshire records. (i-xii, peak vi). 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

josephi (Matthews, A., 1872)

Acrotrichis (Acrotrichis) josephi (Matthews, A., 1872) 4 3 WATSONIAN (3,0,0,0,0) An interesting species apparently associated with wetlands: the 2 YORKSHIRE A. subcognata Johnson of older lists. 1 Described from New Zealand, the 0 species was first found in Britain (and 9 the Palaearctic) at Aughton Ings 8 (SE6938) 24/6/1987 (RJM det CJ) 7 (Johnson, 1992), in piles of grass mowings and again at a nearby site in 6 1994. The species turned up again at 5 Spurn (TA4215) 30/10/2007 (MLD det 4 RJM teste CJ) in ground litter at the 3 edge of man-made pools. Very similar to 2 the common A. cognata but without the 1 Catops-like sheen, and with very slight 0 differences in the spermatheca. Any specimen of “cognata” from wet sites 9 should be carefully examined. 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

42 lucidula Rosskothen, 1935

Acrotrichis (Acrotrichis) lucidula Rosskothen, 1935 4 3 WATSONIAN (0,1,3,0,0) A rarely recorded wetland species often associated 2 YORKSHIRE with seepages in woodland. Records 1 include one for Forge Valley 0 (SE9886) undated and therefore not 9 mapped (GBW);Ewden Beck 8 (SK2396) 20/6/1987 (EJS det CJ); 7 Salt Springs (SK2496) 21/11/1983 6 (EJS det CJ); Chesterfield Canal 5 (SK5481) 1990 (WAE). 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 19/9/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

montandonii (Allibert, 1844)

Acrotrichis (Acrotrichis) montandonii (Allibert, 1844) 4 3 WATSONIAN (6,12,22,7,0) Widespread and 2 YORKSHIRE common in a wide variety of 1 habitats – in woodland, garden 0 compost heaps, ant nests, piles of 9 grass cuttings, old dungheaps and 8 in ground litter in grassland situations. Few records from higher 7 ground. (ii-x, peak vii). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

43 norvegica Strand, A., 1941

Acrotrichis (Acrotrichis) norvegica Strand, A., 1941 4 3 WATSONIAN (0,0,2,2,0) A rare species of 2 YORKSHIRE grassland sites and recorded from 1 old haystacks. Our records include: 0 Snaith (SE6422) 1970 (CJ);Phippin 9 Parks (SE6519) 1967 (CJ); Copgrove (SE3463) 10/7/1966 (CJ); Studley 8 Park (SE2969) 6/7/1966 (CJ). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

parva Rosskothen, 1935

Acrotrichis (Acrotrichis) parva Rosskothen, 1935 4 3 WATSONIAN (1,0,2,1,0) A rare species of 2 YORKSHIRE woodland situations and occurring in 1 decaying plant refuse. There are 0 four records on the YNU database 9 including: Pocklington (SE84) 1945 8 (WDH); Farnley Tyas (SE1612) 1980 (CJ); Worsbrough (SE3504) 1980 7 (CJ); Malham Tarn (SD86) 1967 6 (CJ), the species first being 5 recognised from Britain from this 4 latter site (Johnson, 1990). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

44 pumila (Erichson, 1845)

Acrotrichis (Acrotrichis) pumila (Erichson, 1845) 4 3 WATSONIAN (1,0,1,0,0) A rare species of 2 YORKSHIRE putrefying substances with only two 1 records on the YNU database: 0 Wheldrake (SE64) 1988 (CJ); 9 Todmorden (SD9424) 1970 (CJ). May also be recorded from old 8 dungheaps and leaf compost. 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

rosskotheni Sundt, 1971

Acrotrichis (Acrotrichis) rosskotheni Sundt, 1971 4 3 WATSONIAN (0,0,17,6,0) Widespread with records 2 YORKSHIRE notably from the western half of the 1 county. Recorded from long- 0 established woodland and parkland 9 sites in leaf litter, grass traps, and 8 there is also a record from under deciduous tree bark. (iii-xi, peak viii). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

45 rugulosa Rosskothen, 1935

Acrotrichis (Acrotrichis) rugulosa Rosskothen, 1935 4 3 WATSONIAN (0,2,2,1,0) Another very scarce 2 YORKSHIRE though widespread species with only 1 five records noted: Mulgrave Woods 0 (NZ81) undated (CJ); Raincliffe Wood (SE9988) undated (GBW); 9 Ewden Beck (SK29) 1970 (CJ); 8 Farnley Tyas (SE1612) 1970 (CJ); 7 Malham Tarn (SD86) 1967 (CJ). The 6 undated records are not mapped. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

sericans (Heer, 1841)

Acrotrichis (Acrotrichis) sericans (Heer, 1841) 4 3 WATSONIAN (11,6,37,11,3) A widespread and 2 YORKSHIRE common ptiliid typically of grassland 1 habitats and is recorded from grass 0 tussocks, old straw piles, compost 9 heaps in gardens, mouldy plant 8 refuse, and occasionally in horse dung. Certainly one of the more 7 frequently encountered Acrotrichis. 6 (i-x, peak vi). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

46 silvatica Rosskothen, 1935

Acrotrichis (Acrotrichis) silvatica Rosskothen, 1935 4 3 WATSONIAN (1,1,8,0,0) Nationally Notable. A 2 YORKSHIRE scarce woodland ptiliid usually 1 occurring in piles of putrefying plant 0 material, e.g. leaf litter and grass piles. Also recorded from woodland 9 pitfall traps. Yorkshire records 8 indicate an association with 7 broadleaved woodland rather than 6 coniferous. There is only one post- 5 1990 record on our database. (ii-x). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

sitkaensis (Motschulsky, 1845)

Acrotrichis (Acrotrichis) sitkaensis (Motschulsky, 1845) 4 3 WATSONIAN (20,2,45,25,4) A common and 2 YORKSHIRE widespread wetland species mainly 1 recorded from central and southern sites in Yorkshire, but with few 0 records from higher ground. Noted 9 from litter in beds of Phragmites, 8 Typha and Glyceria, and at the 7 edges of ponds, drains and 6 reservoirs. Also by sweeping in wet 5 places. (i-xi, peak vi). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

47 strandi Sundt, 1958

Acrotrichis (Acrotrichis) strandi Sundt, 1958 4 3 WATSONIAN (4,1,7,6,1) Nationally Notable. A 2 YORKSHIRE wetland species found in ground 1 litter in marshes, by pools and drains and from river ings. Widely 0 distributed but rather scarce. 9 Askham Bog (SE5748) and 8 Throxenby Mere (produced the first 7 recognised British specimens in 6 1966 (Johnson, 1990). 5 (iii-xi, peak vi). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

thoracica (Waltl, 1838)

Acrotrichis (Acrotrichis) thoracica (Waltl, 1838) 4 3 WATSONIAN (2,7,15,9,3) A widespread species of 2 YORKSHIRE dungheaps and rotting compost. A 1 grassland species with only one 0 recent record. (iii-xi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

48

Subgenus CTENOPTERYX Flach, 1889

grandicollis (Mannerheim, 1844)

Acrotrichis (Ctenopteryx) grandicollis (Mannerheim, 1844) 4 (22,19,59,17,6) A widespread and 3 WATSONIAN very common ptiliid of a wide range 2 YORKSHIRE of habitat types, including horse 1 dung, grass tussocks, old fungi, 0 ground litter in reed beds, in 9 woodland leaf litter, old straw piles, 8 in heaps of road side mowings and in 7 garden compost heaps. Often in very 6 large numbers when discovered. 5 (i-xii, peak vi). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

sanctaehelenae Johnson, 1972

Acrotrichis (Ctenopteryx) sanctaehelenae Johnson, 1972 4 3 WATSONIAN (0,1,1,0,0) Superficially very similar 2 YORKSHIRE to A. grandicollis. New to Britain 1 from Essex specimens in 1985, the 0 species has since been recorded 9 twice in Yorkshire: Rawcliffe 8 Meadows (SE5755) October 1997 7 (RJM det CJ), in grass heap at the 6 edge of a man-made pond; Denaby Ings (SE5000) 16/6/2006 (RJM det 5 CJ), general sweeping alongside a 4 land drain. For comparisons 3 between this and grandicollis see 2 Johnson (1987). 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

49

LEIODIDAE Fleming, 1828

SOGDA Lopatin, 1961

HYDNOBIUS sensu auctt. partim non Schmidt, W.L.E., 1841 TRICHOHYDNOBIUS Vogt, 1961 suturalis (Zetterstedt, 1828)

Sogda suturalis (Zetterstedt, 1828) 4 3 WATSONIAN (0,3,0,0,0) Very rare in Yorkshire. 2 YORKSHIRE Recorded from Saltburn (NZ62) 1 1900 (MLT); Eston (NZ5518) 1912 0 (MLT); Coatham Sands (NZ5725) 9 1995 (ME). Also known from Hartlepool area around 1912, and 8 from Durham, Sussex and Scotland; 7 see Cooter (1996). Associated with 6 sandy soils on the coast. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 7/3/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

HYDNOBIUS Schmidt, W.L.E., 1841 latifrons (Curtis, 1840)

Hydnobius latifrons (Curtis, 1840) 4 3 WATSONIAN (0,0,1,0,0) There is a single record 2 YORKSHIRE on the YNU database: Thorne Moor 1 (SE71) 1978 (PH). Widely scattered 0 in England and Wales but rare. 9 There is an assumed association 8 with subterranean fungi. 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 3/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

50 punctatus (Sturm, 1807)

Hydnobius punctatus (Sturm, 1807) 4 3 WATSONIAN (0,3,4,0,0) Nationally Notable. 2 YORKSHIRE Scarce in Yorkshire. Widely 1 distributed in Britain north to 0 Perthshire (Cooter, 1996) Believed 9 to be associated with subterranean 8 fungi. (vi-xi). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 3/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

spinipes (Gyllenhal, 1813)

Hydnobius spinipes (Gyllenhal, 1813) 4 3 WATSONIAN (0,0,2,0,0) Recorded twice in 2 YORKSHIRE Yorkshire: Deer Hill (SE0711) 1 25/10/1986 (MLD det JC); Withens 0 Clough (SD9823) 24/8/1980 (MLD 9 det JC). Nationally, a rarely recorded 8 autumnal moorland species. 7 Presumably associated with 6 subterranean fungi (Alexander, 5 1995). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 3/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

51

TRIARTHRON Märkel, 1840 maerkelii Märkel, 1840

Triarthron maerkelii Märkel, 1840 4 WATSONIAN 3 (0,2,0,0,0) Nationally Notable. The YORKSHIRE 2 two county records are both from 1 May Moss (SE89) July and 0 September 1996 (LA det MLD). Both 9 records resulted from flight 8 interception trapping. Widely 7 distributed in England and Scotland 6 but rarely recorded. Most records 5 nationally appear to be from 4 woodland. 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 16/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe LEIODINI Fleming, 1821

LEIODES Latreille, 1796 badia (Sturm, 1807)

Leiodes badia (Sturm, 1807) 4 3 WATSONIAN (0,2,2,0,0) The four YNU records 2 YORKSHIRE are: Saltburn (NZ62) 1900 (MLT); 1 Scarborough (TA08) 1880? (MLT); 0 Pot Ridings (SE5200) 29/5/2007 9 (AG det RJM), in pitfall traps in 8 woodland; Thorne Moor (SE7214) 7 14/8/2000 (RJM), in pitfall traps on peat by drain. Widespread and 6 fairly common nationally. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 3/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

52 calcarata (Erichson, 1845)

Leiodes calcarata (Erichson, 1845) 4 3 WATSONIAN (3,8,29,9,3) Widespread and 2 YORKSHIRE common – the polita (Marsham) of 1 earlier checklists. The Leiodes most 0 frequent met with, and at all 9 altitudes. As far as the Yorkshire 8 records are concerned the species 7 has been noted from pitfall and 6 water traps in woodland and by 5 general sweeping in hedgerows and 4 by ponds. (v-ix). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 3/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

ciliaris (Schmidt, W.L.E., 1841)

Leiodes ciliaris (Schmidt, W.L.E., 1841) 4 WATSONIAN 3 (0,1,0,0,0) Nationally Notable. A YORKSHIRE 2 single record: Coatham Sands 1 (NZ5725) 1995 (ME), in area of 0 slack and sand dunes. Associated 9 with hypogeous fungi in sandy 8 situations on the coast. 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 16/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

53 cinnamomea (Panzer, 1793)

Leiodes cinnamomea (Panzer, 1793) 4 3 WATSONIAN (1,2,2,0,0) Nationally Notable. There 2 YORKSHIRE are five records on the YNU 1 database: Hull (?TA02) 1900 0 (Anon); York (?SE65) 1870 9 (Hutchinson); Raincliffe Woods 8 (SE98) 1920 (Anon); North Dean 7 Wood (SE0921) 1984 (MLD); Pot 6 Ridings Wood (SE5200) 21/10/2007 5 (AG det RJM), in pitfalls in woods on 4 Magnesian limestone. Generally 3 easily recognisable because of its 2 large size, and associated with 1 0 truffles in lime-rich soils. The 9 provenance of the Hull and York 8 post-1990 records records cannot be determined. pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 3/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

ferruginea (Fabricius, 1787)

Leiodes ferruginea (Fabricius, 1787) 4 3 WATSONIAN (2,3,1,1,1) . A widespread but very 2 YORKSHIRE infrequent species recorded from a 1 variety of situations including under 0 carrion, in a light trap and in flood 9 refuse. (vii-x). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 3/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

flavescens (Schmidt, W.L.E., 1841) NO RECORDS

54 furva (Erichson, 1845)

Leiodes furva (Erichson, 1845) 4 WATSONIAN 3 (1,0,0,0,0) Nationally Notable. YORKSHIRE 2 There is a single record on the YNU 1 database: Spurn (TA41) 19/8/1949 0 (AEW). A species associated with 9 hypogeous fungi in coastal 8 sandhills. 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

gallica (Reitter, 1884)

Leiodes gallica (Reitter, 1884) 4 WATSONIAN 3 (0,1,0,0,0) A single record from YORKSHIRE 2 Yorkshire: Scarborough (TA08) c1900 1 (RL teste JC). Nationally extremely 0 scarce with widely distributed records 9 from England. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

55 gyllenhalii Stephens, 1829

Leiodes gyllenhalii Stephens, 1829 4 WATSONIAN 3 (0,2,0,0,0). There are two records YORKSHIRE 2 on the YNU database: Saltburn 1 (NZ62) c1900 (MLT); Scarborough 0 (TA08) c1880 (RL). Cooter (1996) 9 notes a possible woodland 8 association. Another little-known 7 species. 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

litura Stephens, 1832

Leiodes litura Stephens, 1832 4 WATSONIAN 3 (1,3,3,1,0) Widespread but very YORKSHIRE 2 infrequent, and generally associated 1 with wooded sites (Cooter, 1996), 0 and this agrees with our data. Only a 9 single recent record: West Newton 8 (TA1937) 24/8/1999 (WRD). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

longipes (Schmidt, W.L.E., 1841) NO RECORDS

56 lucens (Fairmaire, 1855)

Leiodes lucens (Fairmaire, 1855) 4 3 WATSONIAN (0,0,3,0,0) A species widespread in 2 YORKSHIRE England and Wales with three 1 records on the YNU database: Dean 0 NY NZ NZ OV Wood (SE1213) 13/8/1983 (MLD); SD SE SE TA 9 Sandall Beat (SE6103) 1960 8 (Anon); Wheatley Wood (SE6005) 7 1904 (HHC). This last record would 6 require a voucher specimen for 5 confirmation. L. lucens is one of the 4 less common species. 3 2 1

SD SE SE TA 0 SJ SK SK TF 9

8 post 31/12/1989 pr e 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 20/10/2012 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

lunicollis (Rye, 1872)

Leiodes lunicollis (Rye, 1872) 4 WATSONIAN 3 (1,1,0,0,0) There are two records YORKSHIRE 2 on the YNU database, one recent 1 and one very old: North Ferriby 0 (SE9826) 4/8/1999 (RGB), in 9 seepage at base of low clay cliffs; 8 Scarborough (TA0488) c.1900 (RL 7 teste JC), in flood refuse. 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

macropus (Rye, 1873) NO RECORDS

57 nigrita (Schmidt, W.L.E., 1841)

Leiodes nigrita (Schmidt, W.L.E., 1841) 4 WATSONIAN 3 (0,1,0,0,0) The single record on YORKSHIRE 2 the YNU database is: Scarborough 1 (TA08) c.1900 (RL), in flood refuse 0 in Scalby Beck. A voucher 9 specimen is required to confirm 8 this record. Nationally, recorded 7 from widely scattered localities in 6 England and Scotland. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

obesa (Schmidt, W.L.E., 1841)

Leiodes obesa (Schmidt, W.L.E., 1841) 4 WATSONIAN 3 (1,0,10,5,0) There has been much YORKSHIRE 2 confusion of nomenclature 1 between this species and Leiodes 0 dubia (sensu Joy, 1911 not 9 Kugelann, 1794) See Cooter 8 1996:258 for an explanation. Our 7 records of “dubia” probably refer 6 to L. obesa and so have been 5 included within L. obesa. We have 4 16 records and in the light of the 3 confusion only five of these are 2 considered reliable. 1 0 Widespread and one of the more 9 often recorded Leiodes. 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

oblonga (Erichson, 1845) NO RECORDS

58 picea (Panzer, 1797)

Leiodes picea (Panzer, 1797) 4 WATSONIAN 3 (0,1,0,0,0) The single record we YORKSHIRE 2 have is: Langdale End (SE9492) 1 15/7/1978 (RSK). See Flint, 1986. 0 (The specimen may be L. obesa 9 and would need dissection – 8 J. Cooter pers. comm. to R Marsh). 7 A species of hypogeous fungi, the 6 species is confined to Scotland and 5 northern England (Cooter, 1996) 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

rufipennis (Paykull, 1798)

Leiodes rufipennis (Paykull, 1798) 4 WATSONIAN 3 (0,0,4,0,1) Five records from YORKSHIRE 2 three widely scattered sites: 1 Honley Wood (SE1111) 16/6/1984 0 (MLD); Templeborough (SK4292) 9 2003 (MLD); Low Row (SD9797) 8 11/7/2005 (MLD). 7 Associated with hypogeous fungi. 6 Nationally one of the more 5 widespread and frequently 4 encountered Leiodes. 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

59 rugosa Stephens, 1829

Leiodes rugosa Stephens, 1829 4 3 WATSONIAN (0,2,0,1,0) Nationally Notable. 2 YORKSHIRE There are three records of this 1 widely distributed species, but only 0 one of them is recent: 9 Scarborough (TA08) c.1900 (RL); 8 Raincliffe Woods (SE9988) 10/1920 7 (GBW); Bishop Monkton Railway 6 Cutting YWT Reserve (SE3166) 5 24/10/2010 (coll. RW det RJM teste 4 JC), in grassland pitfall traps. 3 Widespread throughout mainland 2 Britain. Easily recognisable among 1 Leiodes (at least, with the aid of a 0 field lens) owing to its distinctive 9 strigose elytral sculpture. 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 3/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

silesiaca (Kraatz, 1852) NO RECORDS

strigipennis Daffner, 1983

Leiodes strigipennis Daffner, 1983 4 WATSONIAN 3 (0,0,3,2,0) . This species was YORKSHIRE 2 recently separated from L. 1 flavicornis by Daffner in 1983. We 0 have five records on the YNU 9 dadabase: Thorne Moor (SE7415) 8 24/6/1990 (MLD det JC); Pot 7 Ridings (SE5200) (MLD); Pot 6 Ridings (SE5300) (AG det RJM), in 5 water trap; Malham Tarn 4 (SD8967) 16/8/1958 (WDH det 3 JC); Drax (SE6528) 3/6/2009 (PT 2 det DS). 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

60 triepkii (Schmidt, W.L.E., 1841)

Leiodes triepkii (Schmidt, W.L.E., 1841) 4 WATSONIAN 3 YORKSHIRE (0,0,1,0,0) There is a single 2 record on the YNU database: 1 Frickley Park (SE4608) 8/6/1985 0 9 (RJM), by evening sweeping in a 8 woodland ride. JC has seen the 7 specimen and is fairly sure that it 6 is triepkii inspite of its being a 5 female. A little doubt must 4 therefore remain over the 3 determination. Widely distributed 2 but infrequent in England. 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

LIOCYRTUSA Daffner, 1982

CYRTUSA sensu auctt. non Erichson, 1842 minuta (Ahrens, 1812)

Liocyrtusa minuta (Ahrens, 1812) 4 3 WATSONIAN (1,1,0,1,0) Nationally Notable. The 2 YORKSHIRE YNU database has three entries 1 for this species: Spurn (TA41) 0 4/6/1950 WDH) (see Naturalist 9 1952:134); Scarborough (TA08) 8 c.1900 (RL), taken in flood refuse; 7 Ripon Loop, River Ure (SE3173) 6 3/7/2005 (RJM), in pitfall traps on 5 river bank. 4 Nationally a scarce species with a 3 rather northern distributon. 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

61 vittata (Curtis, 1840)

Liocyrtusa vittata (Curtis, 1840) 4 3 WATSONIAN (1,1,2,1,2) With seven records on 2 YORKSHIRE the YNU database, all post-1985, 1 record details include pitfalls in 0 marshy area with Juncus tussocks, 9 in pitfalls in river shingle, roadside 8 sweep-netting, in a mv moth trap 7 and in a light fitting indoors. 6 Widely distributed. Probably 5 overlooked and under-recorded 4 owing to its small size. (vi-vii). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe PSEUDOLIODINI Portevin, 1926

AGARICOPHAGUS Schmidt, W.L.E., 1841

cephalotes Schmidt, W.L.E., 1841

Agaricophagus cephalotes Schmidt, W.L.E., 1841 4 3 WATSONIAN (0,0,1,0,0) There is a single record 2 YORKSHIRE on the YNU database: Potteric Carr 1 (SE5900) 14/8/2002 (DB det EJS), 0 no other details given. Nationally 9 widespread but infrequent 8 throughout England. The species has 7 been recorded from unidentified 6 truffles in other parts of England. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 26/9/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

62

COLENIS Erichson, 1845 immunda (Sturm, 1807)

Colenis immunda (Sturm, 1807) 4 3 WATSONIAN (0,2,6,3,0) With 11 records on the 2 YORKSHIRE database one of the more frequent 1 of the very small leiodids. Noted 0 from woodland rides, canal banks 9 and meadows, by pitfall trapping 8 and general seeping of vegetation. 7 (vi-xi). Association with any 6 particular fungal habitat is not 5 clear from the records. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe SCOTOCRYPTINI Reitter, 1884

AGLYPTINUS Cockerell, 1906 agathidioides Blair, 1930 NO RECORDS

Tribe AGATHIDIINI Westwood, 1838

ANISOTOMA Panzer, 1797 LIODES Erichson, 1845 non von Heyden, 1826 castanea (Herbst, 1792) NO RECORDS

63 glabra (Fabricius, 1792)

Anisotoma glabra (Fabricius, 1792) 4 3 WATSONIAN (0,0,1,0,0) The single record known 2 YORKSHIRE to the YNU recorder is: Hall Dike 1 (SE1112) 7/8/1983 (DM det MLD). 0 Associated with fungal habitats in 9 upland sites, this is a northern 8 species in Britain and our record is 7 at the southern end of its British 6 range. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

humeralis (Fabricius, 1792)

Anisotoma humeralis (Fabricius, 1792) 4 3 WATSONIAN (5,9,66,11,5) A widespread and 2 YORKSHIRE common species at all altitudes and 1 recorded from all five vice-counties. 0 Recorded from various fungi 9 including the larger brackets, from 8 Reticularia and Polyporus species, 7 under the bark of various deciduous 6 and coniferous trees, in pitfalls in 5 woodland and occasionally by 4 sweeping. (v-xi). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

64 orbicularis (Herbst, 1792)

Anisotoma orbicularis (Herbst, 1792) 4 3 WATSONIAN 0,5,8,1,0) Rather a scarce 2 YORKSHIRE Anisotoma, recorded from beneath 1 bark, in debris in old logs and in 0 pitfalls and water traps in deciduous 9 woodland. (iv-ix). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

AMPHICYLLIS Erichson, 1845 globus (Fabricius, 1792)

Amphicyllis globus (Fabricius, 1792) 4 3 WATSONIAN (0,3,10,1,0) Rather scarce with most 2 YORKSHIRE records from the south of the county. 1 This very distinctive leoidid of 0 woodland sites has been recorded from 9 grass traps, pitfalls and leaf litter on 8 the woodland floor where it is 7 associated with various fungi. (v-xii). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

65

AGATHIDIUM Panzer, 1797

Subgenus CYPHOCEBLE Thomson, C.G., 1859 arcticum Thomson, C.G., 1862

Agathidium (Cyphoceble) arcticum Thomson, C.G., 1862 4 3 WATSONIAN (0,0,1,0,0) Our single record on the 2 YORKSHIRE YNU database is the first 1 authenticated record for England: 0 Ramsden Clough (SE1206) 13/6/1981 9 (MLD det JC). A northern species 8 particularly well-known from the 7 Scottish Highlands where it has 6 occurred from v-x. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

nigrinum Sturm, 1807

Agathidium (Cyphoceble) nigrinum Sturm, 1807 4 3 WATSONIAN (2,3,17,4,0) Widespread and fairly 2 YORKSHIRE frequent, and almost exclusively from 1 woodland sites but not recorded from 0 the higher altitudes of the north and 9 west.. Under bark, in pitfalls and in leaf 8 litter. There is also a subfossil record 7 from Thorne Moor. (i-xii). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus NEOCEBLE des Gozis, 1886 confusum Brisout, 1863 NO RECORDS

66 convexum Sharp, 1866

Agathidium (Neoceble) convexum Sharp, 1866 4 3 WATSONIAN (2,0,0,1,0) Very scarce with only three 2 YORKSHIRE records on the YNU database: Allerton 1 Bywater (SE4228) 27/11/1988 (RBA 0 det DGH); Wheldrake Ings (SE64) 9 10/3/1991 (RBA det MLD); Wheldrake 8 Ings (SE74) 13/2/1993 (RBA det MLD). 7 Typical habitats are under fungoid 6 bark, in pitfalls and in moss. An 5 autumn/winter species. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

marginatum Sturm, 1807

Agathidium (Neoceble) marginatum Sturm, 1807 4 3 WATSONIAN (2,1,0,0,0) Nationally Notable. Another 2 YORKSHIRE scarce Agathidium with only three very 1 old records on the database: Spurn 0 (TA41) 7/1952 (JHF), on sea cliffs; 9 Spurn (TA41) 14/7/1952 (SS); Saltburn 8 (NZ62) 1896 (MLT). Nationally very local 7 and mainly recorded from sandy sites, 6 in tussocks, moss and pitfalls. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

67 nigripenne (Fabricius, 1792)

Agathidium (Neoceble) nigripenne (Fabricius, 1792) 4 WATSONIAN 3 (2,5,29,16,3) Widespread and YORKSHIRE 2 frequent, mainly recorded from 1 deciduous woodland sites, in 0 various fungi including Polyporus, 9 under loose fungoid bark and in 8 leaf litter. (iii-xi). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

rotundatum (Gyllenhal, 1827)

Agathidium (Neoceble) rotundatum (Gyllenhal, 1827) 4 WATSONIAN 3 (0,5,4,0,0) A widely distributed but YORKSHIRE 2 infrequent species of rotting fungi 1 and under fungoid bark. Our records 0 are from woodland sites. (iv-vii) 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

68 varians Beck, 1817

Agathidium (Neoceble) varians Beck, 1817 4 WATSONIAN 3 (10,6,32,7,1) Widespread and YORKSHIRE 2 frequent in fungoid habitats 1 including under bark, in faggots, leaf 0 litter, lakeside ground debris, flood 9 refuse and also recorded from a 8 derelict beehive. Records from sites 7 in wooded areas predominate. 6 (ii-xii). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus AGATHIDIUM Panzer, 1797 atrum (Paykull, 1798)

Agathidium (Agathidium) atrum (Paykull, 1798) 4 3 WATSONIAN 1,4,20,3,0) Widespread and fairly 2 YORKSHIRE frequent in grass heaps, leaf litter, 1 in pitfall traps and in moss. The 0 Yorkshire records indicate a 9 preference for woodland situations, 8 although there are records from 7 open moorland and wetland areas. 6 (iv-x). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 26/9/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

69 laevigatum Erichson, 1845

Agathidium (Agathidium) laevigatum Erichson, 1845 4 WATSONIAN 3 17,3,23,4,1) Widespread and fairly YORKSHIRE 2 frequent. Yorkshire records come 1 from a wide variety of habitats, 0 many of which are damp to quite 9 wet, ranging from peat and upland 8 moors, drains and strand lines on 7 beaches, piles of cut reeds and 6 straw bales, to damp woodland. 5 (i-x). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

pisanum Brisout, 1872

Agathidium (Agathidium) pisanum Brisout, 1872 4 WATSONIAN 3 (0,0,0,1,0) There is a single record YORKSHIRE 2 on the YNU database: Birks Wood 1 (SD9376) 25/5/1993 (KNAA det JC), 0 see Cooter (2004). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

70 seminulum (Linnaeus, 1758), 1821

Agathidium (Agathidium) seminulum (Linnaeus, 1758) 4 WATSONIAN 3 (0,2,3,5,0) A scarce species in YORKSHIRE 2 Yorkshire, and all our records are from 1 wooded sites. There is a subfossil 0 record from the peat strata on Thorne 9 Moor. (i-xi). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subfamily COLONINAE Horn, 1880

COLON Herbst, 1797 KOLON Herbst, 1797 115

Subgenus EURYCOLON Ganglbauer, 1899 latum Kraatz, 1850

Colon (Eurycolon) latum Kraatz, 1850 4 WATSONIAN 3 (0,2,1,0,0) Nationally Notable. YORKSHIRE 2 A scarcely recorded species in the 1 county. The three records we have 0 are: Scarborough (TA08) c.1900 9 (RL); Wykeham Forest (SE9588) 8 9/7/1995 (LA); Pot Ridings Wood 7 (SE5200) 12/8/2002 (EG det MLD), 6 in pitfall traps in woodland. Both the 5 Scarborough and Wykeham records 4 require vouchers for confirmation. 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

rufescens Kraatz, 1850 NO RECORDS

71

Subgenus MYLOECHUS Latreille, 1807 angulare Erichson, 1837 NO RECORDS

appendiculatum (Sahlberg, C.R., 1822)

Colon (Myloechus) appendiculatum (Sahlberg, C.R., 1822) 4 WATSONIAN 3 (0,1,0,0,0) There is a single record YORKSHIRE 2 on the YNU database: Forge Valley 1 (SE98) 1925 (GBW). A voucher 0 specimen is required for confirmation 9 of this record. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

brunneum (Latreille, 1807)

Colon (Myloechus) brunneum (Latreille, 1807) 4 WATSONIAN 3 (1,2,3,3,0) Widespread but rarely YORKSHIRE 2 recorded. Recent records are from 1 pitfall traps. Noted from a wide 0 variety of habitat types. (iv-vi). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

72 dentipes (Sahlberg, C.R., 1822)

Colon (Myloechus) dentipes (Sahlberg, C.R., 1822) 4 WATSONIAN 3 (0,1,0,1,0) Very rare with only two YORKSHIRE 2 records known from the county and 1 both would require vouchers to 0 confirm their identity: Forge Valley 9 (SE98) pre-1950 (GBW); “Ripon” 8 (possibly Studley) (SE26) 1871 7 (EAW), mentioned in EMM 1871:38. 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

zebei Kraatz, 1854 NO RECORDS

Subgenus COLON Herbst, 1797

serripes (Sahlberg, C.R., 1822)

Colon (Colon) serripes (Sahlberg, C.R., 1822) 4 3 WATSONIAN (0,2,1,1,0) We have four records on 2 YORKSHIRE the YNU database: Scarborough 1 (TA08) c.1880 (WCH); Scalby Beck 0 (TA09) 1936 (GBW); Potteric Carr 9 (SE5900) 20/5/2002 (RJM det JC), in 8 pitfalls around pond; Healey Mills 7 Ossett (SE2619) 25/6/2011 (MLD). 6 The very old Scarborough record has 5 to be regarded as doubtful without a 4 voucher specimen. 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 29/8/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

73 viennense Herbst, 1797

Colon (Colon) viennense Herbst, 1797 4 WATSONIAN 3 (0,0,0,0,1) There is a single record YORKSHIRE 2 for the county: Reeth (SE0398) 1 29/7/2005 (AG det MLD), in pitfall 0 traps in riverside shingle. 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subfamily CHOLEVINAE Kirby, 1837

Tribe ANEMADINI Hatch, 1928

NEMADUS Thomson, C.G., 1867 colonoides (Kraatz, 1851)

Nemadus colonoides (Kraatz, 1851) 4 WATSONIAN 3 (1,1,0,1,0) Very scarce with only YORKSHIRE 2 three records known for Yorkshire: 1 Skipwith Common (SE6637) 0 7/9/2000 (DGH det MLD); 9 Beningbrough Park (SE5157) 8 4/9/1999 (ABD), in leaf 7 litter;Studley Royal (SE2969) 6 5/1871 (EAW). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

74

Tribe CHOLEVINI Kirby, 1837

NARGUS Thomson, C.G., 1867

Subgenus NARGUS Thomson, C.G., 1867 velox (Spence, 1813)

Nargus (Nargus) velox (Spence, 1813) 4 WATSONIAN 3 (21,6,95,18,6) Widespread and very YORKSHIRE 2 common in leaf litter in woodland, 1 by reservoirs and other wet places, 0 in grass piles and in compost heaps 9 in gardens, in pitfalls and flight 8 interception traps. (i-xii). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

wilkinii (Spence, 1813)

Nargus (Nargus) wilkinii (Spence, 1813) 4 WATSONIAN 3 (3,6,95,18,6) As widespread but a YORKSHIRE 2 little less common than N. velox. A 1 similar range of habitats to that 0 species has been noted from the 9 records, but apparently more often 8 associated with woodland sites. 7 (i-xii). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

75

Subgenus DEMOCHRUS Thomson, C.G., 1867 anisotomoides (Spence, 1813)

Nargus (Demochrus) anisotomoides (Spence, 1813) 4 WATSONIAN 3 1,4,16,0,0) Rather uncommon and YORKSHIRE 2 mainly recorded from sites in the 1 far south of the county. Mainly 0 restricted to woodland sites in 9 damp situations, with a few from 8 wetland margins in carr land. 7 Found mainly by pitfall trapping 6 and sieving leaf litter on the 5 woodland floor. (i-xii). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

CHOLEVA Latreille, 1796

Subgenus CHOLEVOPSIS Jeannel, 1922 spadicea (Sturm, 1839)

Choleva (Cholevopsis) spadicea (Sturm, 1839) 4 WATSONIAN 3 (0,2,7,0,0) In keeping with all our YORKSHIRE 2 Choleva species, rather scarce 1 especially in modern times. This 0 species is widespread and recorded 9 mainly from woodland sites and best 8 found by pitfall trapping on the 7 woodland floor. There is a record 6 from a window in a power station 5 building. Apparently a spring and 4 early summer species. Choleva 3 species generally are believed to be 2 associated with the nests and runs 1 of small mammals, although the 0 Yorkshire records make little 9 mention of this. (iii-vi). 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

76

Subgenus CHOLEVA Latreille, 1796 agilis (Illiger, 1798)

Choleva (Choleva) agilis (Illiger, 1798) 4 WATSONIAN 3 (2,3,9,0,0) Widespread but rarely YORKSHIRE 2 recorded. A variety of habitats is 1 recorded from both woodland and 0 wetland sites. Again very few recent 9 records. (iv-xii). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

angustata (Fabricius, 1781)

Choleva (Choleva) angustata (Fabricius, 1781) 4 WATSONIAN 3 (7,2,12,5,0) One of the more YORKSHIRE 2 commonly recorded Choleva 1 species. Widespread in the county 0 except in the far north and west. 9 Mainly recorded from damp open 8 areas with one or two woodland 7 sites mentioned. (i-x). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

77 cisteloides (Frölich, 1799)

Choleva (Choleva) cisteloides (Frölich, 1799) 4 WATSONIAN 3 (0,0,1,0,0) There is a single record YORKSHIRE 2 on the YNU database: Wath-on- 1 Dearne (SE4300) c.1900 (EGB). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

elongata (Paykull, 1798) NO RECORDS

fagniezi Jeannel, 1922

Choleva (Choleva) fagniezi Jeannel, 1922 4 WATSONIAN 3 (0,0,15,5,0) Widespread and rather YORKSHIRE 2 scarce, mainly in the south- western 1 quarter of the county. (ii-xii). 0 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

78 glauca Britten, 1918

Choleva (Choleva) glauca Britten, 1918 4 WATSONIAN 3 (1,0,2,4,0) Nationally Notable. YORKSHIRE 2 Widely distributed but very scarce. 1 Taken in pitfall traps and in ground 0 litter. The records do not associate 9 the species with any particular 8 habitat type. (ii-ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

jeanneli Britten, 1922

Choleva (Choleva) jeanneli Britten, 1922 4 WATSONIAN 3 (7,2,10,7,0) Widely distributed with YORKSHIRE 2 only four recent records. Recorded 1 from bogs, wet meadows, damp 0 woodland, but also from drier 9 situations such as quarry floors and 8 limestone woodland. Several of our 7 very old records undated and 6 therefore not mapped. (ii-xi). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

lederiana Reitter, 1902 NO RECORDS

79 oblonga Latreille, 1807

Choleva (Choleva) oblonga Latreille, 1807 4 WATSONIAN 3 (1,1,6,4,0) Scarce but widely YORKSHIRE 2 distributed. Recorded mainly from 1 damp habitats – wet meadows, 0 ings, reservoir margins – usually in 9 pitfall traps, but also under stones 8 and in ground litter. Few recent 7 records. (i-xii). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

SCIODREPOIDES Hatch, 1933

SCIODREPA sensu auctt. non Thomson, C.G., 1859 fumatus (Spence, 1813)

Sciodrepoides fumatus (Spence, 1813) 4 WATSONIAN 3 (3,6,18,13,2) Very widespread and YORKSHIRE 2 fairly frequent. Mainly recorded 1 from carrion including hedgehog, 0 various birds, and dead sheep, but 9 occasionally from fungi on 8 deciduous trees. (iv-x). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

80 watsoni (Spence, 1813)

Sciodrepoides watsoni (Spence, 1813) 4 WATSONIAN 3 (17,14,70,20,4) A widespread and YORKSHIRE 2 common species recorded from a 1 wide variety of habitats including 0 carrion (particularly birds), in 9 pitfalls on heaths and in woodland 8 and marshy areas, in straw heaps 7 and bales, in fungi on trees, and in 6 water traps in woodland rides. 5 Often very numerous in woodland 4 pitfalls traps. 3 (ii-xii, with a marked peak in vi). 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

CATOPS Paykull, 1798 chrysomeloides (Panzer, 1798)

Catops chrysomeloides (Panzer, 1798) 4 WATSONIAN 3 9,4,10,8,0) Widespread, and a YORKSHIRE 2 rather uncommon Catops. From a 1 wide variety of site types, usually 0 recorded from carrion and baited 9 pitfall traps, but also occasionally 8 from fungi of the gill type and from 7 fungoid growths under bark. (i-xi). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

81 coracinus Kellner, 1846

Catops coracinus Kellner, 1846 4 WATSONIAN 3 (5,5,15,6,3) Widespread in the county. YORKSHIRE 2 Abundance and habitats as noted in C. 1 chrysomeloides, but also recorded 0 from water traps in woodland rides. 9 (ii-ix). 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

fuliginosus Erichson, 1837

Catops fuliginosus Erichson, 1837 4 WATSONIAN 3 (15,4,66,11,0) The most often YORKSHIRE 2 recorded Catops species in the county, 1 but not, so far, from VC65. Noted from 0 carrion (birds and rabbits), pitfall traps 9 (baited and otherwise) in woodland, in 8 leaf litter, from Phragmites litter and 7 from piles of cut grass. Often 6 abundant when found. (i-xii). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

82 fuscus (Panzer, 1794)

Catops fuscus (Panzer, 1794) 4 WATSONIAN 3 (3,1,4,1,0) One of the rarer Catops YORKSHIRE 2 with only nine records on the 1 database. Habitat details attached to 0 the Yorkshire records are sparse – the 9 remains of dead birds are mentioned 8 as a habitat. 7 (vi-xii). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

grandicollis Erichson, 1837

Catops grandicollis Erichson, 1837 4 WATSONIAN 3 (11,4,33,8,4) A common and YORKSHIRE 2 widespread species of carrion of 1 sheep, fox, rabbit, various birds, in 0 compost heaps and in piles of old 9 straw, in pitfalls (baited and 8 unbaited) in woodland, in water traps 7 in woodland rides and in animal dung 6 in a garden. (ii-xi, peak ix). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

83 kirbii (Spence, 1813)

Catops kirbii (Spence, 1813) 4 WATSONIAN 3 (6,7,21,7,2) Widespread and fairly YORKSHIRE 2 common in Yorkshire. Noted from a 1 similar range of habitats to Catops 0 grandicollis, but records include more 9 records from fungi and badger carrion 8 is also included. (ii-x, peak ix). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

longulus Kellner, 1846

Catops longulus Kellner, 1846 4 WATSONIAN 3 (0,5,7,3,1) Nationally Notable. Rather YORKSHIRE 2 scarce. Recorded from fungi of the 1 bracket type on trees and also the gill 0 type on the woodland floor. Also noted 9 from bird nests, and from grass 8 tussocks in winter. (ix-xii). 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

84 morio (Fabricius, 1787)

Catops morio (Fabricius, 1787) 4 WATSONIAN 3 (20,8,32,11,2). Widespread and fairly YORKSHIRE 2 common in dead sheep and birds, with 1 many records being from pitfall traps in 0 a wide range of habitat types ranging 9 from dunes on the coast to woodland 8 at all altitudes. Records from rotting 7 fungi are also included. A mainly 6 autumnal species. (i-xii, peak ix). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

nigricans (Spence, 1813)

Catops nigricans (Spence, 1813) 4 WATSONIAN 3 (17,3,35,6,1). Common and YORKSHIRE 2 widespread except in the far north 1 and far west of our recording area. 0 Recorded from pitfalls in habitats 9 ranging from dunes on the coast to 8 woodland at all altitudes, from fungi, 7 compost heaps in gardens and 6 churchyards, and from carrion of 5 sheep and rabbits. (i-xii, peak vi). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

nigriclavis Gerhardt, 1900 NO RECORDS

85 nigrita Erichson, 1837

Catops nigrita Erichson, 1837 4 WATSONIAN 3 (0,5,43,2,2) There are plenty of YORKSHIRE 2 records for Yorkshire but the species 1 seems confined mainly to the south of 0 the region. Habitats include pitfall 9 traps, water traps and in fungi. Most of 8 our records come from woodland sites 7 but the species is also recorded from 6 the peat moors and river margins. 5 (iii-xi, peak vi). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

tristis (Panzer, 1793)

Catops tristis (Panzer, 1793) 4 WATSONIAN 3 (11,12,97,17,2) Widespread and YORKSHIRE 2 common, especially in the south. In a 1 wide variety of habits from coastal 0 sites to woodland to wet places, and 9 recorded from leaf litter, woodland and 8 marshland pitfalls, carrion and fungi, 7 and in any putrefying animal or 6 vegetable material. (i-xii, peaks in vi 5 and x). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

86

CATOPIDIUS Jeannel, 1922 depressus (Murray, 1856)

Catopidius depressus (Murray, 1856) 4 WATSONIAN 3 (1,0,1,0,0) Nationally Notable. Very YORKSHIRE 2 scarce – there are only two confirmed 1 records for Yorkshire: Hatfield Moor 0 (SE7006) 9/9/1991 (PS), in pitfall traps 9 on moorland; Kelsey Hill (TA2326) 8 19/7/1998 (WRD det RJM), a single 7 specimen beaten from sallows. This 6 species is usually associated with the 5 runs and burrows of mammals such as 4 badger and rabbit. Recorded from 3 pitfalls in rabbit burrows in 2 neighbouring North Nottinghamshire 1 (RJM). 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

PARABATHYSCIA Jeannel, 1908

BATHYSCIA sensu auctt. partim non Schiødte, 1849 wollastoni (Janson, E.W., 1857) NO RECORDS

Tribe PTOMAPHAGINI Jeannel, 1911

PTOMAPHAGUS Illiger, 1798 medius Rey, 1889

Ptomaphagus medius Rey, 1889 4 WATSONIAN 3 (3,3,10,5,1) Rather scarce in Yorkshire, YORKSHIRE 2 but widely distributed and with few 1 recent records. Noted from ground 0 litter and carrion traps. (iii-ix). 9 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

87 subvillosus (Goeze, 1777)

Ptomaphagus subvillosus (Goeze, 1777) 4 WATSONIAN 3 (14,19,108,30,3) A widespread and very YORKSHIRE 2 common species, possibly the most 1 common leoidid of the woodland floor. 0 Besides woodland, the insect is found in 9 pitfall traps in wetland, in roadside 8 mowings and any fungoid and 7 putrefying plant debris, in compost 6 heaps and in flight interception and 5 water traps. Adults are found in all 4 months. (i-xii, peak vi). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

varicornis (Rosenhauer, 1847)

Ptomaphagus varicornis (Rosenhauer, 1847) 4 WATSONIAN 3 (1,0,0,0,0). A rare insect with only one YORKSHIRE 2 Yorkshire record: Millington Wood NR 1 (SE8353) 24/4/2009 (WRD), by 0 sweeping Mercuralis perennis by a 9 footpath. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

88

Subfamily PLATYPSYLLINAE Ritsema, 1869

LEPTINUS Müller, P.W.J., 1817 testaceus Müller, P.W.J., 1817

Leptinus testaceus Müller, P.W.J., 1817 4 WATSONIAN 3 (1,5,8,3,0). Scarce though widely YORKSHIRE 2 distributed. The few recent records 1 received are from the south of the 0 county. Recorded from the nests of 9 mole and field mouse, and one 8 specimen from the pelage of a dead 7 unidentified rodent away from the 6 nest environment. (ii-ix). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 18. Family SCYDMAENIDAE Leach, 1815 119

Subfamily SCYDMAENINAE Leach, 1815

Tribe EUTHEIINI Casey, 1897

EUTHEIA Stephens, 1830

EUTHIA auctt. (misspelling)

formicetorum Reitter, 1881 NO RECORDS linearis Mulsant, 1861 NO RECORDS plicata (Gyllenhal, 1813) NO RECORDS

89 schaumii Kiesenwetter, 1858

Eutheia schaumii Kiesenwetter, 1858 4 WATSONIAN 3 (0,2,1,0,0). Nationally Notable. Very YORKSHIRE 2 scarcely recorded. As with 1 scydmaenids generally, these tiny 0 beetles with cryptic habits may be 9 more widespread and numerous than 8 our records suggest. There are three 7 records on the YNU database: 6 Scarborough (TA08) 1919 (GBW), 5 taken by evening sweeping (see 4 Naturalist 1920:201); Scarborough 3 Mere (TA0386) 7/1921 (GBW), over 2 300(!) specimens collected by 1 sweeping in a restricted area; Haw 0 Park (SE3615) 1980 (EWA). The 9 species would be expected in compost 8 post-1990 records heaps and similar situations. pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

scydmaenoides Stephens, 1830

Eutheia scydmaenoides Stephens, 1830 4 WATSONIAN 3 (0,2,2,1,0) Nationally Notable. YORKSHIRE 2 Widespread but very scarce with only 1 five records: Raincliffe Woods 0 (SE9988) 1870 (RL); Harton (SE7162) 9 v/1969 (EWA); Hatfield Lings (SE6507) 8 13/10/1973 (CJ); Hatfield Lings 7 (SE6507) 1975 (PS); Copgrove 6 (SE3463) 12/7/1967 (CJ). We have 5 received no recent records. Recorded 4 from decaying material in ground litter. 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

90

EUTHICONUS Reitter, 1881/2 conicicollis (Fairmaire & Laboulbène, 1855) NO RECORDS Tribe CEPHENNIINI Reitter, 1882

CEPHENNIUM Müller, P.W.J. & Kunze, 1822 gallicum Ganglbauer, 1899

Cephennium gallicum Ganglbauer, 1899 4 3 WATSONIAN (2,0,3,0,0) Widely distributed in the 2 YORKSHIRE south of our recording area but scarce. 1 There are five records on the YNU 0 NY NZ NZ OV database: Brantingham (SE9430) SD SE SE TA 9 20/5/2000 (WRD; Hull – Louis Street 8 LNR (TA0829) 30/9/2001 (WRD); 7 Frickley Park (SE4608) 8/6/1985 6 (RJM); Honley Wood (SE1211) 5 21/7/1985 (JRB det MLD); Park Wood 4 (SE1122) 21/9/1985 (EJS). Recorded 3 by general sweeping, in moss and leaf 2 litter and in vegetable refuse. (v-ix). 1

SD SE SE TA 0 SJ SK SK TF 9

8 post 31/12/1989 pr e 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 20/10/2012 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe CYRTOSCYDMINI Schaufuss, 1889

EUCONNUS Thomson, C.G., 1859

Subgenus CLADOCONNUS Reitter, 1909 denticornis (Müller, P.W.J. & Kunze, 1822)

Euconnus (Cladoconnus) denticornis (Müller & Kunze,1822) 4 3 WATSONIAN (0,2,0,0,0) There are two, both very 2 YORKSHIRE old, records for Yorkshire: Marton in 1 Cleveland (NZ51) 1832 (YNU), noted as 0 recorded by GT Rudd with no other 9 details known; Saltburn (NZ62) 7/1897 8 (MLT). Both these records would 7 require vouchers for confirmation, but 6 in the meantime they will remain on 5 the database. Typical habitat would be 4 compost heaps, ground litter in 3 woodland, and similar situations. 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

91

Subgenus EUCONNUS Thomson, C.G., 1859 fimetarius (Chaudoir, 1845)

Euconnus (Euconnus) fimetarius (Chaudoir, 1845) 4 WATSONIAN 3 (0,1,1,0,0) Very scarce with only two YORKSHIRE 2 records: Scarborough (TA08) c.1890 1 (RL); Green Wood (SE2108) 4/1/2009 0 (JF det RJM). The recent record was 9 noted from a sample of litter that was 8 part of a nest of the wood ant Formica 7 lugubris. The 1890 record would 6 require a voucher for confirmation, but 5 remains on the database. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

hirticollis (Illiger, 1798)

Euconnus (Euconnus) hirticollis (Illiger, 1798) 4 WATSONIAN 3 (1,2,1,8,0) The commonest of the YORKSHIRE 2 genus in our region with 12 records, 1 although nonetheless a scarce species. 0 Recorded from leaf litter, in moss and 9 in ground detritus in a willow bog, the 8 species has been consistently and 7 often recorded at Askham Bog from 6 the 1960s to date, with very few 5 records from anywhere else. (iii-ix). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

rutilipennis (Müller, P.W.J. & Kunze, 1822) NO RECORDS

92

Subgenus NAPOCHUS Thomson, C.G., 1859 duboisi Méquignon, 1929

Euconnus (Napochus) duboisi Méquignon, 1929 4 WATSONIAN 3 (2,0,1,0,0) So far, rare in Yorkshire, YORKSHIRE 2 with three very recent records from 1 only two sites: North Cave Wetlands 0 (SE8833) 21/8/2007 (WRD det RJM), 9 sieved from piles of woodchips; North 8 Cave Wetlands (SE8833) 9/6/2008 7 (RJM), in the same habitat as above; 6 Green Wood Denby Dale (SE2108) 5 4/1/2009 (JF det RJM), in a sample of 4 litter that was part of a nest of the 3 wood ant Formica lugubris. The 2 question arises: is this very distinctive 1 species a newcomer to the county or 0 has it been overlooked in the past? 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

pragensis (Machulka, 1923) NO RECORDS

Subgenus NEONAPOCHUS Machulka, 1929 maeklinii (Mannerheim, 1844) NO RECORDS

MICROSCYDMUS Saulcy & Croissandeau, 1893 EUCONNUS sensu auctt. partim non Thomson, C.G., 1859 minimus (Chaudoir, 1845) NO RECORDS nanus (Schaum, 1844)

Microscydmus nanus (Schaum, 1844) 4 WATSONIAN 3 (0,1,0,1,0) Nationally Notable. Very rare YORKSHIRE 2 in Yorkshire with only two records: 1 Scarborough (TA08) c.1890 (RL); 0 Askham Bog (SE5748) 2/9/1994 (RGB), 9 in ground litter. 8 7 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

93

NEURAPHES Thomson, C.G., 1859 121

Subgenus NEURAPHES Thomson, C.G., 1859 angulatus (Müller, P.W.J. & Kunze, 1822)

Neuraphes angulatus (Müller & Kunze, 1822) 4 3 WATSONIAN (0,3,1,0,0) Widespread but very scarce 2 YORKSHIRE with only four records known: Saltburn 1 (NZ62) 7/1899 (MLT); Sandsend 0 (NZ8613) 1934 (HB), see Naturalist 9 1934:276; Rosedale (SE7395) 8 14/5/1980 (RSK); Toftwood, Rivelin 7 (SK3287) 17/10/1998 (EJS). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5 carinatus (Mulsant & Rey, 1861) NO RECORDS

elongatulus (Müller, P.W.J. & Kunze, 1822)

Neuraphes elongatulus (Müller & Kunze, 1822) 4 3 WATSONIAN (0,9,27,7,1) Widespread, and one of 2 YORKSHIRE our more common Scydmaenidae. This 1 Neuraphes is recorded from a wide 0 range of ground-based habitats: leaf 9 litter, moss, pitfalls in woodland, at the 8 bases of compost heaps and old straw 7 bales, in grass traps, and in ground 6 detritus in wetland situations, the 5 species often occurring in large 4 numbers when found. (i-xi, peak ix). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

94 talparum Lokay, 1921

Neuraphes (Neuraphes) talparum Lokay, 1921 4 3 WATSONIAN (0,3,0,1,0) A rare species with only 2 YORKSHIRE four Yorkshire records: Raincliffe 1 Woods (SE9988) 1871 (RL), under 0 bark; Scarborough (TA08) 1880 9 (WCH); Beck Hole (NZ8202) 8 13/2/1937 (HB), in haystack refuse; 7 Studley Royal Park (SE2969) 1871 6 (EAW), in an old nest in a rotting 5 beech tree. 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 12/9/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subgenus PARARAPHES Reitter, 1891 plicicollis Reitter, 1879

Neuraphes plicicollis Reitter, 1879 4 3 WATSONIAN (0,0,0,3,0) Nationally Notable. There 2 YORKSHIRE are three recent records, and only from 1 a single site: Askham Bog (SE5748) 0 2/9/1994 (RGB); Askham Bog (SE5648) 9 22/6/1996 (JAO); Askham Bog 8 (SE5748) 23/6/1996 (JAO). The two 7 JAO records may either refer to two 6 findings on consecutive days or may 5 refer to the same record. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced for Yorkshire Naturali sts' Union by DMAP 5/4/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

praeteritus Rye, 1872 NO RECORDS

95

SCYDMORAPHES Reitter, 1891 NEURAPHES sensu auctt. partim non Thomson, C.G., 1859 helvolus (Schaum, 1844)

Scydmoraphes helvolus (Schaum, 1844) 4 WATSONIAN 3 (4,0,5,1,0) Nationally Notable. YORKSHIRE 2 Widespread in the south-eastern 1 quarter of the county with ten records. 0 The species has been reported 9 recently, and is recorded from a 8 decayed bracket fungus, ground litter 7 in woodland and at the base of a 6 hedgerow, from grass piles and heaps 5 of dry twigs in a garden. (ii-x). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

sparshalli (Denny, 1825)

Scydmoraphes sparshalli (Denny, 1825) 4 WATSONIAN 3 (0,2,0,1,0) A rare species with only YORKSHIRE 2 three records from Yorkshire: 1 Scarborough (TA08) c.1890 (RL); Beck 0 Hole (NZ8202) 9/5/1936 (HB); Studley 9 Royal Park (SE2969) 1870 (EAW). 8 There are no habitat details given with 7 any of these records, but may be 6 expected in a range of situations similar 5 to the previous species. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

96

STENICHNUS Thomson, C.G., 1859

CYRTOSCYDMUS Motschulsky, 1870

SCYDMAENUS sensu Fowler, 1888 non Latreille, 1802 bicolor (Denny, 1825)

Stenichnus bicolor (Denny, 1825) 4 WATSONIAN 3 (0,2,8,3,0) Widespread but scarce, with YORKSHIRE 2 only three recent records. Generally 1 associated with rotten wood and under 0 bark, both deciduous, but two of our 9 recent captures came from nests of 8 ants Formica spp.. Considered by 7 Alexander (2002) to be saproxylic, and 6 with national records mainly from 5 ancient wood pastures. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

collaris (Müller, P.W.J. & Kunze, 1822)

Stenichnus collaris (Müller, P.W.J. & Kunz e, 1822) 4 WATSONIAN 3 (8,23,42,20,2) A widespread and YORKSHIRE 2 common species of the woodland floor, 1 in moss, plant litter and pitfall traps, 0 from grassland sites, in decaying fungi, 9 grass traps, compost heaps, at the 8 edges of reservoirs and canals 7 wherever decaying plant litter occurs. 6 A specimen has also been reported 5 from a dead jackdaw. 4 (ii-xii, peak vi and ix). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

godarti (Latreille, 1806) NO RECORDS poweri (Fowler, 1884) NO RECORDS pusillus (Müller, P.W.J. & Kunze, 1822) NO RECORDS

97 scutellaris (Müller, P.W.J. & Kunze, 1822)

Stenichnus scutellaris (Müller, P.W.J. & Kunz e, 1822) 4 WATSONIAN 3 (8,4,9,1,0) Widespread, but mainly YORKSHIRE 2 from the south-eastern quarter, and 1 never very common. In Yorkshire 0 recorded from compost heaps, leaf 9 litter, piles of lawn mowings in a 8 garden, in old decaying bracket fungi, 7 in wetlands in ground litter under 6 Carex paniculata and in ground debris 5 at the water’s edge. (i-xi, peak ix). 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Tribe SCYDMAENINI Leach, 1815

SCYDMAENUS Latreille, 1802 EUMICRUS Laporte, 1840

Subgenus CHOLERUS Thomson, C.G., 1859 rufus Müller, P.W.J. & Kunze, 1822 NO RECORDS

Subgenus SCYDMAENUS Latreille, 1802 tarsatus Müller, P.W.J. & Kunze, 1822

Scydmaenus tarsatus Müller, P.W.J. & Kunz e, 1822 4 3 WATSONIAN (19,10,88,19,2) This widespread 2 YORKSHIRE species of grass piles, compost heaps 1 and silage, and occasionally from 0 carrion, is the most commonly recorded 9 scydmaenid in Yorkshire. 8 (iii-xii, peak vi). 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 19/9/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

98

Subfamily SILPHINAE Latreille, 1807

NECRODES Leach, 1815 littoralis (Linnaeus, 1758)

Necrodes littoralis (Linnaeus, 1758) 4 WATSONIAN 3 (6,5,19,7,30) A widespread species of YORKSHIRE 2 carrion. The high number of records 1 from VC65 reflects collecting effort – 0 many of those records were obtained 9 during a light trapping survey, and 8 indeed the species often comes to 7 moth traps, and occasionally house and 6 street lights. (iii-x, peak viii). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

THANATOPHILUS Leach, 1815 dispar (Herbst, 1793)

Thanatophilus dispar (Herbst, 1793) 4 3 WATSONIAN (0,1,0,2,0) A very scarce species of 2 YORKSHIRE carrion, unrecorded since 1926, with 1 only three Yorkshire records: Coxwold 0 (SE5377) 1880 (WF); Ripon (SE37) 9 1880 (EAW); Leeds (SE33) 1926 (JRD). 8 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 13/9/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

99 rugosus (Linnaeus, 1758)

Thanatophilus rugosus (Linnaeus, 1758) 4 3 WATSONIAN (27,6,48,21,10) Widespread and 2 YORKSHIRE common on a variety of dead animals, 1 including birds, foxes, rabbits, sheep, 0 rats, roe deer and badgers, in 9 woodland, grassland, wetlands and on 8 coastal dunes, and at all altitudes. Also 7 reported from dry animal dung. 6 (i-ix, peak vi). 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 13/9/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

sinuatus (Fabricius, 1775)

Thanatophilus sinuatus (Fabricius, 1775) 4 3 WATSONIAN (9,4,21,6,1) A widespread species of 2 YORKSHIRE carrion of fox, rabbit, sheep and birds. 1 Also recorded from pitfall traps on peat 0 moorland and at edges of water bodies. 9 Most of our records are from sites that 8 are well inland but at all altitudes 7 except the very highest ground. 6 (v-ix, peak vii). 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 13/9/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

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OICEOPTOMA Leach, 1815 thoracicum (Linnaeus, 1758)

Oiceoptoma thoracicum (Linnaeus, 1758) 4 WATSONIAN 3 (9,25,32,8,0) In woodland, grassland YORKSHIRE 2 and wetland sites and recorded from the 1 carrion of fox, sheep, rabbit, various 0 birds, and once in a dead fish by a drain 9 on peat moorland. Widespread but not 8 reported from the higher altitudes in the 7 west of the county. (iii-x, peak viii). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

ACLYPEA Reitter, 1884 opaca (Linnaeus, 1758)

Aclypea opaca (Linnaeus, 1758) 4 WATSONIAN 3 (5,7,6,10,0) Nationally Notable A. YORKSHIRE 2 Widespread, not common and with 1 very few recent records. This species 0 is vegetarian and is known to damage 9 turnip and beet crops. Few of our 8 records go into habitat details but the 7 species in Yorkshire has been recorded 6 from turnips, rotting mangolds, from 5 pitfalls by a land drain on peat 4 moorland and from under pine bark 3 (the last probably an overwintering 2 site). A much declined species in our 1 region, possibly due to change in 0 agricultural practices and use of 9 insecticides. (iv-xii, peak v). 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

101 undata (Müller, O.F., 1776)

Aclypea undata (Müller, O.F., 1776) 4 WATSONIAN 3 (0,1,0,0,0) There is a single record on YORKSHIRE 2 the YNU database: Buttercrambe 1 (SE7358) 1936 (GBW), in carrion (see 0 Naturalist 1936:166). Another 9 phytophage, so it is strange that the 8 record should mention carrion. I hope it 7 is not a misidentification. On mainland 6 Europe, the insect is associated with 5 beet and carrot crops. 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

DENDROXENA Motschulsky, 1858

XYLODREPA Thomson, C.G., 1859 quadrimaculata (Scopoli, 1772)

Dendroxena quadrimaculata (Scopoli, 1772) 4 WATSONIAN 3 (3,2,22,2,0) Nationally Notable B. YORKSHIRE 2 Recorded from the southern half of the 1 county. Very local with few recent 0 records. All our records seem to be 9 from woodland sites. The adults are 8 known to run about on foliage in search 7 of the larvae of Lepidoptera. Apparently 6 absent from locations at higher 5 altitudes. (Most records show only 4 years, in a few May and June are 3 mentioned.) 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

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SILPHA Linnaeus, 1758 atrata Linnaeus, 1758

Silpha atrata Linnaeus, 1758 4 WATSONIAN 3 (12,27,49,13,9) Widespread and fairly YORKSHIRE 2 common. There are plenty of recent 1 records. An easily recognisable species 0 with mouthparts adapted to feed on 9 land molluscs. Recorded from loose 8 bark and moss, pitfall traps and in plant 7 litter where their prey items occur. 6 Noted from sites that are woodland, 5 damp grassland, moorland and 4 wetlands. (i-xi, peaks vi and ix). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

carinata Herbst, 1783 NO RECORDS

laevigata Fabricius, 1775

Silpha laevigata Fabricius, 1775 4 WATSONIAN 3 (0,2,3,0,0) There are five records on YORKSHIRE 2 the YNU database, three of which are 1 undated so therefore not mapped. 0 One of these is from archaeological 9 deposits of early medieval age: 8 Scarborough (TA08) date unknown 7 (TB); Whitby (NZ81) date unknown 6 (HP); Conisbrough (SK5198) AD 650 5 approx (PB), in ditch excavation near 4 church; Wath (SE4300) 1893 (EGB); 3 2 Barnsley (SE30) 1907 (EGB). 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

103 obscura Linnaeus, 1758

Silpha obscura Linnaeus, 1758 4 WATSONIAN 3 (0,0,1,0,0) A very local insect YORKSHIRE 2 generally in Britain, we have only a 1 single record on the YNU database: 0 Skelmanthorpe (SE2311) 18/4/1920 9 (Anon), these data on a specimen in 8 the Tolson Museum Huddersfield. 7 Associated with a wide variety of 6 carrion. 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

tristis Illiger, 1798

Silpha tristis Illiger, 1798 4 WATSONIAN 3 (0,1,42,9,1) A carrion beetle with an YORKSHIRE 2 odd distribution in Yorkshire. The high 1 number of VC63 records results from 0 captures of large numbers of the insect 9 in equally large numbers of pitfall traps 8 during two major surveys of Calluna 7 heath and peat moorland in recent 6 years. It could be that the southern 5 corner of VC63 is the major centre of 4 population of the insect in Yorkshire. 3 (iii-xii, peak ix). 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

104 tyrolensis Laicharting, 1781

Silpha tyrolensis Laicharting, 1781 4 3 WATSONIAN (0,3,5,2,3) Nationally Notable B. 2 YORKSHIRE Widespread but scarce with no records 1 since Semerdale (SD9186) 1935 (JMB). 0 The species may now be long-extinct in 9 the county. Associated with a wide 8 variety of carrion. 7 6 5 4 3 2 1 0 9 8 post 31/12/1989 pre 31/12/1989 7 Map produced for Yorkshire Naturali sts' Union by DMAP 13/9/2011 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Subfamily NICROPHORINAE Kirby, 1837

NICROPHORUS Fabricius, 1775 germanicus (Linnaeus, 1758) NO RECORDS

humator (Gleditsch, 1767)

Nicrophorus humator (Gleditsch, 1767) 4 WATSONIAN 3 (22,3,87,31,135) Widespread and YORKSHIRE 2 common in Yorkshire, this species is 1 recorded from carrion of fox, badger, 0 rabbit and various birds. Also widely 9 pitfall trapped and with many dozens of 8 recent light-trap records from a small 7 area of VC64/65. (i-xi, peaks v and ix). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

105 interruptus Stephens, 1830

Nicrophorus interruptus Stephens, 1830 4 WATSONIAN 3 (0,1,3,0,0) Nationally Notable B. YORKSHIRE 2 Widespread but very rarely recorded, 1 with only four records on the YNU 0 database. One of these is undated 9 and therefore unmapped: Coxwold 8 (SE5377) 1880 (WF); Skelmanthorpe 7 (SE21) date unknown (SLM); 6 Ravensknowle Park (SE1616) 5 1/7/1946 (EWA); Thorne (SE6813) 4 28/8/1972 (WB). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

investigator Zetterstedt, 1824

Nicrophorus investigator Zetterstedt, 1824 4 WATSONIAN 3 (12,4,33,24,107) Widespread with large YORKSHIRE 2 numbers attracted to light in VC64/65. 1 Distribution indicates that higher 0 altitudes are not detrimental to the 9 species. Noted from carrion of mole, 8 rabbit and sheep, and from pitfalls on 7 peat moorland and Calluna heath (see 6 comments under Silpha tristis), pitfalls 5 in woodland and in gardens. 4 (iv-x, peak ix). 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

106 vespillo (Linnaeus, 1758)

Nicrophorus vespillo (Linnaeus, 1758) 4 WATSONIAN 3 (10,7,36,15,6) Widespread and fairly YORKSHIRE 2 common with plenty of recent records. 1 Recorded from pitfalls in moorland 0 woodland and grassland in dead rats, 9 sheep and rabbits and various bird 8 species including pheasant. 7 (i-xii, peak ix). 6 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

vespilloides Herbst, 1783

Nicrophorus vespilloides Herbst, 1783 4 WATSONIAN 3 (6,20,122,6,3) Another widespread and YORKSHIRE 2 common Nicrophorus species, recorded 1 from moorland, woodland and reservoir 0 margins, where carrion may be found. 9 Noted from dead sheep and rabbits, 8 and birds including gulls and crows. 7 Often in large numbers in baited pitfall 6 traps. (i-xii, peak ix). 5 4 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

107 vestigator Herschel, 1807

Nicrophorus vestigator Herschel, 1807 4 WATSONIAN 3 (0,4,0,0,0) Nationally Notable A. Very YORKSHIRE 2 rarely reported with only four, none 1 recent, records on the YNU database: 0 Raincliffe Woods (SE9988) 1933 9 (GBW); Seamer Moor (TA0183) 1933 8 (GBW); Scarborough (TA08) 8/1937 7 (Anon); Scarborough (TA0488) 8/1951 6 (EFG). Precise habitat details are 5 lacking in all these records but carrion 4 is assumed. 3 2 1 0 9 8 post-1990 records pre-1990 records 7 Map produced by DMAP for Yorkshire Naturali sts' Union 20/9/2010 5 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 5

Acknowledgements

The recording system could not function effectively without the input from Yorkshire coleopterists (and other workers further afield) past and present. I am very grateful to a ‘hard core’ of present members of the YNU Entomological Section for passing their records to me for inclusion in the database, in order to keep the database up to date. In this respect I thank Bill Dolling, the late Frank Kenington, Peter Kendall, Mike Denton and Roy Crossley. I am also grateful to Colin Johnson and the late Dr. Peter Skidmore for giving patiently of their time and expertise in identifying/checking many specimens (some in very battered condition) presented to them over many years. Their help and encouragement in my study of Coleoptera has been outstanding. I also thank Derek Whitely and Eric Smith of the Sorby Natural History Society, Sheffield, for passing on records to me regularly; to Dmitri Logunov and Philip Rispin of Manchester University Museum for use of bench and microscope facilities, access to Coleoptera collections and searching for specimens and literature references for me; to Martin Limbert for continued access to the Coleoptera collection at Doncaster Museum; to Mike Denton for providing advice and sharing of his expertise over many years. Any omissions or mistakes in the text or in the mapping are my responsibility. Any errors which come to the notice of readers will I hope be fed back to me.

For the section on Ptiliidae, I have drawn heavily for information on the work by Colin Johnson in Naturalist 115 (1990) (see reference below). This paper gathered together all the known authenticated records for Watsonian Yorkshire up to that date. For the section on Leiodidae I have referred to papers by Jonathan Cooter, particularly Cooter (1996).

References

Alexander, K.N.A., (1995) A new locality for Hydnobius spinipes (Gyllenhal) (Col: Leiodidae). Coleopterist 3 (3) (1995):80-81.

Alexander, K.N.A., (2002) The invertebrates of living & decaying timber in Britain and Ireland – a provisional annotated checklist. English Nature Research Reports, no. 467. English Nature, Peterborough.

Allen, A.A., (1968) Plegaderus vulneratus Panz., a histerid beetle new to Britain. Entomologist’s mon. Mag. 104 :110-112.

Cooter, J., (1996) Annotated keys to the British Leiodinae (Col., Leiodidae) Entomologist’s mon. Mag. 132 (1996):205-?

Cooter, J., (1999) On “English‟ names for British Coleoptera. Coleopterist 8 (1): 43-45.

Cooter, J., (2004) Agathidium pisanum Brisout (Col., Leiodidae) in Britain. Entomologist’s mon. Mag. 140 (2004):59-64.

Cox, M.L., (2007) Atlas of the Seed and Leaf Beetles of Britain and Ireland. Pisces Publications.

Dandy J.E. 1969. Watsonian Vice-counties of Great Britain. The Ray Society, London

Denton, M.L. 1990. Entomogical Reports for 1988-1989. Coleoptera: Staphylinidae (Aleocharinae). Naturalist 115 (1990): 97- 102.

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Denton, M.L. 2003. The Aleocharinae (Coleoptera: Staphylinidae) of Yorkshire. Naturalist 128 (2003): 103-125.

Denton, M.L. 2008. Entomological records for 2003-2007. (Coleoptera:Staphylinidae (Aleocharinae). Naturalist 133 (2008): 96-100 .

Duff, A.G. (2008). Coleoptera Checklist of the British Isles. 2008 Edition. Coleopterist website www.coleopterist.org.uk

Flint, J.H., (1986) Entomological Reports for 1973-1983, Coleoptera:part 2, Haliplidae – Scolytidae. Naturalist 111 (1986):26.

Flint, J.H. 1988. Entomological reports for 1984-1986: Coleoptera. Naturalist 113 (1988): 69-72.

Halstead, D.G.H., (1963) Handbooks Ident. Br. Insects. Coleoptera: Histeroidea. Royal Entomological Society, London.

JNCC, (1994). Hyman, P.S., and Parsons, M.S. A Review of the scarce and threatened Coleoptera of Great Britain. Part 2. Peterborough.

Johnson, C., (1968) Two new British species of Coleoptera from Yorkshire. Entomologist 101 :64-66.

Johnson, C., (1987) Additions and corrections to the British list of Ptiliidae (Coleoptera). Ent. Gazette 38 (1987):117-122.

Johnson, C., (1990) The Feather-wing beetles of Yorkshire (Coleoptera: Ptiliidae). Naturalist 115 : 57-71.

Johnson, C., (1992) Additions and corrections to the British list of Coleoptera. Ent.Rec.J.Var. 104:305-310.

Johnson, C., (2001) Euryptilium gillmeisteri Flach (Coleoptera: Ptiliidae) new to Britain. Entomologists Gazette 52 : 181-182.

Key, R.S., (1999). More on “noddy names‟ for British beetles. Coleopterist 8 (2): 94.

Marsh, R.J., (1988a) Aeletes atomarius (Aube) (Col., Histeridae) in Yorkshire. Entomologist’s mon. Mag. 124 :166.

Marsh, R.J., (1988b) Oligella intermedia Besuchet (Col., Ptiliidae) – a second British record. Entomologist’s mon. Mag. 124:242

Marsh, R.J. 1991. Coleoptera records for Yorkshire: A review and update. Naturalist 116 (1991): 101-106.

Marsh, R.J. 2002. Coleoptera report for 1995-2001. Naturalist 127 (2002): 69-80.

Pope, R.D. (1977). Kloet & Hincks. A Check List of British Insects. Part 3: Coleoptera and Strepsiptera. Second revised edition. Handbks. Ident. Brit. Insects 11(3), pp. xiv+105. Royal Entomological Society, London.

Shirt, D.B., ed. (1987). British Red Data Books: 2. Insects. Nature Conservancy Council, Peterborough.

Watson, H.C., (1883). 2nd. ed. Topographical Botany. Quaritch, London.

R. J.Marsh FRES YNU Coleoptera recorder 20 October 2012

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