Russian Entomol. 1. 2(3-4): 105-118 © RUSSIAN ENTOMOLOGICAL JOURNAL, 1993

New species of (Diptera).

HonMe MALI cemeiticrBa -Lauxaniidae (Diptera).

A.14.111aramcam

Zoological Museum, Moscow State University, Herzen str. 6, Moscow K-9, 103009 Russia. 3ooAorH4ecicrati mysett MTY, yA. repueHa 6, 1Vlocicaa K-9, 103009 Poccmi.

with KEY WORDS:Lauxaniidae, , new species, Palaearctic Region. life- KJIIOLIEBIDIE CJIOBA: Lauxaniidae, TaKCOHOMHA, HOBble ila.meapkuKa. ther ABSTRACT: The following lauxaniid species are eviitbz AayKcaminA: amurensis sp.n., described: Homoneura amurensis sp.n., H.assidua H.assidua sp.n., H.czernyi sp.n., H.ozeroui sp.n., sp.n., H.czernyi sp.n., H.ozerovi sp.n., H.quadrifaria H.quadrifaria sp.n., rohdendorfi DM sp.n., Aulogastromyia rohdendorfi sp.n., Lyciella sp.n., Lyciella bifaria sp.n., L.brachychaeta sp.n., inual bif aria sp.n., L.brachychaeta sp.n., L.oreophila L.oreophila sp.n., L.pectinifera sp.n., sp.n., L.pectinifera sp.n., Sapromyza arctophila arctophila sp.n., S.citrina sp.n., S.fascidula sp.n., sp.n., S.citrina sp.n., Sluscidula sp.n., S.paramerata sp.n., S.pseudopaca sp.n., S.paramerata sp.n., S.pseudopaca sp.n., S.strigillifera sp.n. flavohalterata sp.n., S.strigillifera sp.n. Lauxania flavohalterata sp.n., L.sonora sp.n., S.vitripennis sp.n. ripeACT213.fleHbi L.sonora sp.n., L.vitripennis sp.n. Keys to the Ta6JIHIthl Ai1A onpeAenetam na.neapwrmeckHx utztos Palaearctic species of Lyciella and Lauxania are powa Lyciella a Lauxania. presented. This paper continues the two previous articles of PE3IOME: B pa6o.re 011HCbIBMTCH czeaylotwie the author [Shatalkin, 1992 a,b] and is based on the study of of the family Lauxaniidae. In it 19 new species are described. Besides this, it gives a revised and enlarged keys to the Palaearctic species of the two genera, Lauxania and Lyciella. Types of new species are deposited in the Zoological Museum of Moscow State University and the Zoological Insti- tute (St.Petersbourg),

Genus Homoneura van der Wulp, 1891.

a Among the Palaearctic genera, Homoneura is the most large group, containing about 70 species. The considerable part of them (approximately 50 species) occur in the east area of the Palaearctic Region. While preparing the review of the Palaearctic Homoneura for printing I had examined the compre- hensive material on this genus. As a result of this examination five new species from Russia and China are described. The Chinese material was collected in the last century by the famous geographer and ethnographer O.N.Potanin. Among the others his material contains two specimens, determined by L.Czerny as Homoneura grahami sMalIoch. The Fig. 1. Male pregenital sternites of Homoneura species: examination of these specimens has revealed that a - H.amurensis sp.n.; b - H.ozeroot sp.n. one of them should be reffered to a new species. PHC. 1. Flperemiranbribre c-repiurru camuoa alma Homon- eura: a H.amurensis sp.n.; b H.azeravi sp.n. 106 A.I.Shatalkin

Homoneura amurensis Shatalkin, sp.n. upper margin. Arista black with dense seated black Fig. la. hairs. Its feathering equal to width of third antenna! segment. Frons about 1.2 times wider than its MATERIAL. Holotype (Cr ): Amurskaya Oblast', Zeya, height. Gena low, about 1/5 height of eye. Eyes 6.VII.1982 (Ozerov). - Paratypes: 2 ?7, same locality as with microscopic sparse hairs. Mouthparts and palpi holotype, 4 and 12.VII.1981 (Ozerov); 1 y, same locality yellow. Thorax fulvous. Legs yellow. Fore femur as holotype, 7.VI11.1982 (Shatalkin). DESCRIPTION. MALE. Head yellow. Frons with anteroventral comb of black spinules. All tibiae slightly shining with delicate dark hairs on its with strong preapicals. Wings yellowish with yellow anterior part. Antennae yellow; third antennal veins. ta slightly before middle of discal cell. Ultimate section of about 1.7 times as long as segment about 1.7 times more than its width. Arista M!+s penultimate. Halteres fulvous. Abdomen brownish dark brown with feathering equal to or slightly more yellow. than its thickened basal part. Gena about 1 /3 height of eye. Mouthparts and palpi yellow. Thorax yellow. Chaetotaxy: 1 h, 1 ph, 2 npl, 0+3 dc, ac in 4 rows, prosternum with setulae, 1 weak ppl, 1 mspl, 2 stpl, Mesonotum slightly shining. Legs yellow. Fore femur with anteroventral comb of black spinules. All anterior of them shorter than posterior. Length 2.5 mm. tibiae with strong preapical bristles. Wings yellow- ish with yellow veins and cloudy ta and tp. Scarcely MALE unknown. DIAGNOSIS. In the key of Homoneura by visible clouds at ends of R4.5 and M1+2. ta beyond Malloch [19291 this species runs to H.affinis middle of discal cell. Ultimate section of M1+2 about 1.7 times as long as penultimate. Halteres yellow. Malloch known from Borneo, Malaya, and Philip- Abdomen yellow. Fifth abdominal sternite of male pines [Sasakawa, 19921. It differs from the latter in bilobate posteriorly with stout spines on margin of having four rows of acrostichals (instead of six ones in H.affinis), roundish lobes (Fig. la). Genitalia massive. and denser feathering of the arista. H.assidua sp.n. is also similar to Chaetotaxy: 1 h, 1 ph, 2 npl, I +3 dc, ac in 4 rows H.unguiculata Kert. which differs from the first in having six rows with small setae of same length, prosternum naked, 1 weak ppl, I mspl, 2 stpl with posterior of them of acrostichals, the silvery dusting of parafacials stronger than anterior. discerned by examination from above, and less darkened third antenna! segment. Length 3.1 mm. FEMALE not differs from male. Length 2.8-3.2 mm. Homoneura czernyi Shatalkin, sp.n. DIAGNOSIS. This new species is easily recog- MATERIAL. Holotype (s): China, Sichuan, Ta-xinlun nized by the presence of the presutural dc, thus (Chnyanb, Tagsminy), 12.VII.1893 (Potanin). differing from the majority of species of Homoneura. DESCRIPTION. FEMALE. Head yellow. Frons In general appearance it is similar to H.biumbrata slightly longer than its width, dull, with sparse hairs Lw. The latter differs from H.amurensis sp.n. in a in anterior part. Antennae yellow; third antenna] details of the male genital apparatus and pregenital segment about 1.3 times as long as wide. Arista sternites. In addition it has a two median rows of ac brownish yellow, practically bare. Height of eye much longer than others. On the last character about 1.4 times more than its length. Gena about 1 / H.amurensis sp.n. is separated from those speci- 4 height of eye. Mouthparts and palpi yellow. Thorax mens of H.biumbrata (described by Czerny [19321 yellow. Mesonotum blackish grey in anterior part as H.kowarzi), which have the weak dorsocentrals between humeral calli, with a similar pair of before the suture. postsutural stripes laterad dc, going from presutural darkening to level of third dc. Sternopleuron in Homoneura assidua Shatalkin, sp.n. lower part blackish grey. Legs yellow. Fore femur with anteroventral comb of black spinules. Tibiae MATERIAL. Holotype (7): Primorskiy Kray, Kamen- with preapicals. Wings with costal margin browned ushka (30 km SE from Ussuriysk) , 28. VI.1984 (Shatalkin). - Paratype: 1 7, Primorskiy Kray, Kangaus, 7.VII.1963 and both cross-veins clouded. ta beyond middle of (Nartsbuk). discal cell. Ultimate section of 141 .2 about 1.5 times DESCRIPTION. FEMALE. Head yellow. Frons as long as penultimate. Halteres yellow. Abdomen slightly shining. Antennae yellow. Third antenna! yellow. segment gently concave on dorsal margin, without Chaetotaxy: 1 h, 1 ph, 2 npl, 0+3 dc, ac in 4 rows, apical contraction, about 2.1 times as long as wide. 1 ppl, 1 mspl, 2 stpl with anterior of them shorter It black in apical half beneath and apical third of than posterior. New species of Lauxaniidae 107

Length. 2.8 mm. Homoneura quadrifaria Shatalkin, sp.n. MALE unknown. DIAGNOSIS. H.czernyi sp.n. is distinct in MATERIAL. Holotype (?): Kurile Isis., Shikotan, (on having the peculiar coloration of mesonotum and Fragmites), 19.VIII.1971 (Nartshuk). sternopleuron. In other characters, first all the DESCRIPTION. FEMALE. Head yellow. Frons costal darkening of wings, it is similar to H.grahami dull with sparse small hairs in its anterior half. Ma!loch, also known from Sichuan. The latter is Antennae yellow. Third antenna! segment about 1.6 characterized by the entirely yellow mesonotum, times as long as wide. Arista brownish; its feathering clear tp, the low gena which about 5.4 times lower equal to thickness of basal part. Frons almost than height of the eye. In addition the feathering of square. Height of eye about 1.3 times more than its the arista is equal to width of its thickened basal part length. Gena about 4.4 times less than height of eye. and the ultimate section of Mi almost 2 times as Mouthparts and palpi yellow. Thorax entirely yel- long as the foregoing in H.grahami. low. Legs yellow. Fore femora with anteroventral This new species is' named for the eminent comb of black spinules. All tibiae with strong dip terologist L.Czerny. preapicals. Middle tibiae with strong and small spurs. Wings with apical and anterior (laterad of Hotnoneura ozerovi Shatalkin, sp.n. mouth of R1) margins fuscous; tp clouded, ta Fig. 1 b. practically not clouded but darker than other veins. ta slightly beyond middle of discal cell. Ultimate MATERIAL. Holotype (Cr ): Ainurslcaya Oblast', Zeya, section of M1.2 about 1.8 times as long as penulti- 6.VII.1982 (Ozerov) . - Paratypes: 2 d'cr same locality mate. Halteres yellow. Abdomen yellow; its margin as holotype, 6.V11.1982 (Ozerov). bristles equal to length of tergites. DESCRIPTION. MALE. Head yellow. Frons dull Chaetotaxy: 1 h, I ph, 2 npl, 0+3 dc, ac in 4 rows with delicate light hairs on anterior part. Antennae with internal setulae about 2 times more than those yellow; third antennal segment about 1.4 times as of external rows, prosternum with black hairs, (ppl long as its width. Arista black; its feathering less ?), 1 mspl, 2 stpl with anterior of them noticebly than thickness of basal part which yellowish. Gena less than posterior, pteropleuron bare. about 1 /2 height of eye. Mouthparts and palpi Length 3 nun. yellow. Thorax yellow. Mesonotum dull. Legs yel- MALE unknown. low. Fore femur with anteroventral comb of black DIAGNOSIS. H.quadrifaria sp.n. is similar to spinules. All tibiae with strong preapical bristles. H.lagena Sasa.et Ike., known from Japan ESasa- Fore tibia with ventral short bristles on tip. Hind kawa, Ikeuchi, 1983], in having the wings with coxae with sparse brush of black spinules. Wings costal cloud and differs from the latter by the yellow hyaline with ta and tp clouded. ta before middle of palpi, practically clear ta, and four-row acrostichals. discal cell. Ultimate section of 114.1.2. about 1.6 times One more the Japanese species with a similar costal as long as penultimate. Halteres yellow. Abdomen cloud, H.hymenophallus Sasa.et Ike., has a larger yellow. Fifth abdominal sternite of male bilobate size and the plumose arista and may be not confused posteriorly with 4 stout spines on each lobe (Fig. with the new species. I b). Genitalia massive. Chaetotaxy: 1 h, 1 ph, 2 rip!, 0+3 dc, ac in 4 rows Genus Aulogastromyia Hendel, I >) \J with setae of inside rows much longer than others of external rows, prosternum with small setulae, I The genus Aulogastromyia shows relationship ppl, 1 mspl, 2 stpl with anterior of them half length to Lyciella Collin in having a presutural pair of de of posterior. and a hairs on the pteropleuron. The distinguishing Length 3.3 rnm. feature of Aulogastromyia is the unique modifica- FEMALE not differs from male. Length 3.2 mm. tion of the male hind tarsus being thickened and DIAGNOSIS. This new species is unique in the shortened with the outer claw of last tarsal segment structure of male pregenital sternites and the hind produced into a long process. coxae with the sparse brush of black spinules. In This genus has been known until now as mono- general appearance H.ozerovi sp.n. is similar to typic with a single European species A.anisodactyla H.tesquae Beck. The last species has a longer Lw. The second species of Aulogastromyia, de- acrostichals and the arista with a hairs which is more scribed below, was collected by B.B.Rohdendorf in than its basal part. Turkmenistan. I have great pleasure in dedicating this new species to my colleague Dr. A.L.Ozerov. 108 A.I.Shatalkin

Aulogastromyia rohdendorfi Shatalkin, sp.n. Fig. 2. 3(2 4(5 MATERIAL. Holotype (d): Turkmenistan, Ko- pet-Dag, Nuchur, VI.1923 (Rohdendorf). DESCRIPTION. MALE. Head yellow. Antennae 5(4 yellow; third antenna! segment about 1.5 times as long as its width. Arista yellow with feathering equal 6("i to its basal thickness. Width of Irons equal to its height. Gena about 1/5 height of eye. Mouthparts and palpi yellow. Legs yellow. Fore femur without 7" anteroventral comb of spinules. Fore tarsus about 1.3 times more than fore tibia. Hind tarsus about 1.3 8(S times less than hind tibia and comparatively weak thickened. Last tarsomere of hind legs with two sensory hairs apically and outer claw which less than 10 hind metatarsus and spirally twisted (Fig 2). All tibia with strong preapicals. Wings yellowish with yellow 11 veins. to on middle of discal cell. Ultimate section of M1.2 about 1.4 times as long as penultimate. 12 Halteres yellow. Abdomen yellow and shorter than Fig. 2. Male hind leg and last tarsal segment of Aulogastro- thorax. Tergal marginal bristles longer than tergites. myia rohdendorfi sp.n. Chaetotaxy: I h, I ph, 2 npl, 1+3 de, ac in 2 rows 13 PHC. 2. 3a.rourn nora H HOU1eMBH1 gileHiLK aam-teil namut with bristles about 2 times shorter than dc, 1 weak calnia Aulogastromyia rohdendorfi sp.n. ppl, 1 mspl, 2 stpl, pteropleuron with 3 black hairs. 14 With the exception of the Holarctic L.rorida Length 3.4 mm. Fall., the rest yellow species of Lyciella are not FEMALE unknown. DIAGNOSIS. This new species differs from recorded from the Siberia and Far East. It is possible 15 A.anisodactyla Lw. in having wholly yellow anten- that this group has the European origin; the only nae and in possessing the distinct outer claw of hind separate species were able to penetrate into the mountainous area of Middle Asia and West China. tarsi in the male (Fig. 2). It I dedicate this new species to the famous russian Taking into account the lack of clarity in our understanding of the limits of Poecilolycia, zoologist and palaeontologist B.B.Rohdendorf. above I all in respect to the status of a grey Lyciella-like IS Genus Lyciella Collin, 1948. species, in this paper I am confined oneself to the consideration of only yellow species of Lyciella. A In most characters the genus Lyciella agrees with key for their identification is given below. The key 2( Sapromyza Fall. differing from the latter in the does not include the two Becker's species presence of the presutural pair of dorsocentrals. (L.canariensis and L.emarginata), because their 2 Lyciella and created the Shewell [1986] subdivided status is not clear for me. I consider that L.ocellaris 2! Poecilolycia— for species, possessing new genus — Czerny is conspecific with Sapromyza omei Mal- the following combination of characters: pteropleuron loch. The last species therefore belongs to the genus (anepimeron) bare, upper occiput concave, Irons and Lyciella. Thus Lyciella ocellaris Czerny, 1935 is a scutum vittate, thorax grey, usually bluish grey. Out new synonym of Lyciella omei (Mallach, 1929), 2: of the Palaearctic species with the grey coloration of comb. n. body only L.vittata Walk. (quadrivittata Lw.) cor- responds entirely this diagnosis. Other species are 2 distinguished from L. vittata in having the Key to the yellow species of the Palaearctic 2 vertex convexed and in lacking the mesonotal vittae. Lyciella. In possessing the character of a few tiny hairs on pteropleuron the yellow species of Lyciella agree 2 1(10). Fore femora towards tip with anteroventral 2 with Tricholauxania Hendel and Aulogastromyia, comb of black spinules. which are characterized also by the presence of the 2(3). Abdomen yellow with a large blackish mark on presutural dorsocentrals. These three genera form each side of each tergite except for basal one. the monophyletic group which may be opposed to the group of a grey species of Lyciella. New species of Lauxaniidae 109

Wings maculated. - China (Sichuan) 28(27). Tarsi yellow or brownish (if darkened, palpi L.omei Malloch (L,ocellaris Czerny) widely black). 3(2). Abdomen entirely yellow. 29(30). Tip of wing with costal darkening. - Europe 4(5). Antennae yellow or slightly darkened on apex L.illota Lw. of third segment. Palpi. yellow. ac in 4 rows. - 30(29). Wing without costal darkening on tip. Ho!arctic species L.rorida Fall. 31(32). Third antenna! segment black on apical 5(4). At least apical half of third antennal segment third. Genitalia as in Fig. 3e. - Europe black. L.subfasciataZtt. 6(7). Palpi black. Third antenna! segment dark; its 32(31). Third antenna! segment black on apical half. base and basal segments brown. Genitalia as Fig 33(34). Fore tarsi darkened: First segment of hind 3 a,b.- Caucasus L.nigripalpis Czerny tarsi in basic third with ventral black setae. which 7(6). Pa1pi yellow. Base of third segment and basal longer than adjacent ones. Genitalia as in Fig. 31. segment of antennae yellow. - Caucasus L.brachychaeta sp.n. 8(9). ac in 2 rows. Wings with grey clouds on apices 34(33). Fore tarsi not darkened. First segment of of 2-4 veins arid tp. - Europe .. L.compsella hind tarsi without peculiar setae. Genitalia as in 9(8). ac in 4 rows. Wings clear. Genitalia as in Fig. Fig. 3g. - Europe L.mihalyii Papp 3c. - Caucasus L.pecti nifera sp.n. 10(1). Fore femora 'without this comb of black Lyciella bifaria Shatalkin, sp.n. spinules. 11(16). Wings maculated, at least apices of 2-4 veins MATERIAL. Holotype (e): Tadzhikistan, Kondara, with fuscous spots. 30km N from Dushanbe, 1100 in, 29.V1.1965 (Nartshuk). 12(13). Palpi dark. ac in 4 rows. First segment of Paratypes: 3 W, same locality as holotype, 29.V1.1965 male hind tarsi with a dense brush of black (Nartshuk); 3 ??, Tadzhikistan, Gissarskiy Khr., Kwak, spinules. - Europe Ldecempunctata Fall. 1800 m, 6.V111.1969 (Keleinikova); 1 Tadzhikistan, 13(12). Palpi yellow. ac in 2 rows. First segment of Ramit, Kafirnigan River, 25.V11.1939 (Romanov). male hind tarsi without such brush of spinules. DESCRIPTION. MALE. Head yellow. Ocellar 14(15). Antennae yellow. Both cross-veins with triangle brownish yellow. Antennae yellow. Third fuscous patches. Two pairs of large prescutellar antenna! segment about 1.4 times as long as its basal )t. bristles. - Middle Asia (Pamir) .... L.bifaria sp.n. width, narrowing towards on tip. Arista brownish with le 15(14). Third antenna! segment black on apex. Both feathering slightly more than its basal thickness, Frons y cross- veins darkened. Only one pair of large slightly wider than its height. Gena low, about 1/4 e prescutellar bristles. Genitalia as in Fig. 3d. - height of eye. Mouthparts and palpi yellow. Thorax 1. Middle Asia (Pamir) L.oreophila sp.n. yellow. Legs yellow. Fore femur without anteroventral 16(11). Wings clear or slightly darkened on tip. r comb of spinules. All tibiae with strong preapicals. 17(24). Antennae entirely yellow. e Wings yellowish with yellow veins and 5 small 18(21). Palpi entirely yellow. 19(20). Larger species (3.5-4.0 mm). 1 stpl. - brownish spots: on both cross-veins, on apices R2.3, Europe, Transcaucasus .5, M1+2' two latters of them connected fuscous L. () platycephala Lw. border. to at level which crosses discal cell in ratio 20(19). Smaller species (2.5-3.0 mm). 2 stpl. - 1:1.5. Ultimate section of Mi.z about 1.6 times as long Europe, Caucasus L.laeta Ztt. as penultimate. Halteres yellow. Abdomen yellow. 21(18). Palpi black or with black tip. Chaetotaxy: 1 h, I ph, 2 npl, 1+3 dc, ac in 2 rows 22(23). Mesonotum dull with slightly visible greyish with a sparse setulae aranged in second pair of rows stripes. Male tarsi black; female tarsi darkened on posterior part of mesonotum and 2 pairs of large towards tip. - Europe, Caucasus prescutellars, 1 stark ppl, 1 mspl, 2 stpl, pteropleu- L.conjugata Beck. ron with 2-3 black hairs. 23(22). Mesonotum shining without grey stripes. Length 4-.8 mm. Tarsi yellow. - Europe. Transcaucasus FEMALE not differs from mate. Length 5.5-6.0 mm. L.decipiens Lw. DIAGNOSIS. L.bifaria sp.n. is very similar to 24(17). Third antenna] segment black apically. L.decernpunctata Fall. in having a spotty wings and 25(26). Palpi yellow. Tip of wings with costal darkening. ac in two rows. -Europe two pairs of large prescutellar bristles. It differs from L.affinis Ztt. the latter in having wholly yellow palpi and two-row 26(25). At least apex of palpus black. acrostichals, in lacking the brush of black spinules 27(28). Tarsi darkened; fore and hind tarsi almost Of the hind metatarsus. L.bifaria can be confused black. Tip of wings with small costal darkening. also with L.oreophila sp.n. resembling it in a spotty Palpi with very narrow darkening on tip wings, but may be differentiated by characters listed L.affinisZtt. under that species. 110 A.I.Shatalkin

Lyciella brachychaeta Shatalkin, sp.n. arista evidently less than its basal thickness. Frons Fig. 3f. about 1.2 times wider than its height. Gena about 2.7 times less than height of eye. Mouthparts and MATERIAL Holotype (e): Caucasus, North Osetiya, palpi yellow. Thorax yellow. Legs yellow. Fore Alagir, Lapparerak, 1700 m, 29.VI.1990 (Shatalkin). femur without anteroventral comb of spinules. All Paratypes: 11 do', 4 W, same locality as holotype, 28- 29.V1.1990 (Shatalkin);1 ?, North Osetiya, Tsey gorge, tibiae with yellow preapicals. Wings hyaline with 3 2500 m, 6.V1.1989 (Ozerov); 3 dd, same locality, brown spots on apices R2.3, R4.5, and MI+, (both 16.111989 (Shatalkin). cross-veins not clouded but darker than all other DESCRIPTION. MALE. Head yellow. Ocellar veins). ta slightly beyond middle of discal cell. triangle black. Antennae yellow. Third antenna' Ultimate section of M1.2 about 1.5 times as long as segment about 1.7 times as long as its width; it black penultimate. Halteres yellow. Abdomen yellow. apical half and more. Arista dark brown with Genitalia as in Fig. 3d. a microscopic hairs. Frons about 1.3 times wider than Chaetotaxy: 1 h, 1 ph, 2 npl, 1+3 dc, ac in 2 rows height. Gena about 3.2 times less than height of eye. with only one pair of large prescutellar bristles, 1 Mouthparts yellow. Palpi black. Thorax yellow. weak ppl, 1 mspl, 2 stpl, pteropleuron with 1-2 black Legs yellow. Tarsi darkened. Fore femur without hairs. anteroventral comb of spinules. All tibiae with Length 3.9 mm. preapicals. First segment of hind tarsus in basic FEMALE not differs from male. Length 3.5-4.2 third with black setae ventrally which visibly strong- mm. er than those in the neighbourhood of them. Wings DIAGNOSIS. L.oreophila sp.n. is closely relat- yellowish with yellow veins. ta beyond middle of ed to L.decempunctata Fall. and L.bifaria sp.n. It discal cell. Ultimate section of M1.2 about 1.5 times is separated from the both by transparent ta and tp, as long as penultimate. Halteres yellow. Abdomen and only a pair of large prescutellar bristles. In yellow. Genitalia (Fig. 3f) with massive and asym- addition it differs from the former in having yellow metrical parameres. palpi, two-row ac, in lacking the brush of black Chaetotaxy: 1 h, I ph, 2 npl, 1+3 dc, ac in 2 rows spinules on the hind metatarsus, from the second in with comparatively massive setulae which about 2 having the third antenna' segment with the black times shorter than dc, 1 weak ppl, 1 mspl, 2 stpl, apex. pteropleuron with 3-4 hairs. Length 3.6 mm. Lyciella pectinifera Shatalkin, sp.n. FEMALE not differs male. Length 3.2-3.8 mm. Fig. 3c. DIAGNOSIS. L.brachychaeta sp.n. may be confused with L.nahalyii and L.subfasciata. The MATERIAL. Holotype (d'): Caucasus, North exterior characters, indicated in the key, is not Osetiya, Alagir, 22.IX.1989 (Shatalkin). - Para- reliable for their exact identification. The new types: 10 Cr Cr, 7 W, same locality as holotype, 22.IX.1989 (Shatalkin); 6 W, same locality 18.V.1989 species therefore can be safely distinguished only by ztt the structure of male genitalia. The latter in (Ozerov). L.brachychaeta sp.n. shows most resemblance to DESCRIPTION. MALE. Head yellow. Ocellar L.s that of L.mihalyii. triangle dark. Antennae yellow. Third antenna! segment about 1.6 times as long as its width; it from _Lyciella oreophila Shatalkin, sp.n. dark brown to black above level of insertion of arista. Fig. 3d. Arista brownish with feathering equal to thickness of its basal part. Frons near square. Gena about 3.3 (si MATERIAL. Holotype (cf ): Tativhikistan, Gissarskiy times less than height of eye. Mouthparts and palpi Khr., Kondara, Kwak, 1800 m, 14.VIII.1969 (Keleiniko- yellow. Thorax yellow. Legs yellow. Fore femur with L. va). Paratypes: 2 crc , 8 ?, same locality as holotype, anteroventral comb of 7-8 black spinules. All tibiae in 11-15.V111.1969 (Keleinikova); I cf, Tadzhikistan, 15 km with strong preapical bristles. Wings yellowish with th from DzhayPgan, Kizylsu River; 2400 m, 3.VI.1983 yellow veins. ta slightly beyond middle of discal cell. aE (Pavlinov). Ultimate section of MI.2 about 1.6 times as long as L DESCRIPTION. MALE. Head yellow. Antennae penultimate. Halteres yellow. Abdomen yellow. yellow. Third antennal segment not narrowing on le Genitalia as in Fig. 3c. SE tip, straight dorsally and convex ventrally, about 1.7 Chaetotaxy: 1 h, 1 ph, 2 npl, I +3 dc, ac in 4 rows times as long as its basal width; it black in apical with internal setulae longer than those of external third. Arista darkish with yellow base; feathering of rows, prosternum with black hairs, 1 weak ppl, 1 h

New species of Lauxaniidae 111

Fig. 3. Male fermi nalia otLyciella species: a,b - Lragripalpis Czerny; c -L.pectinifera sp.n.; d -Loreophila sp.n.; e- L.subfasciata Ztt.; I - L.brachychaeta sp.n.; g - L.mihalyii Papp (after Papp, 1978, Figs. 24-25). Scale = 0.1 mm. Pric. 3. TepmnHanint camnos 1114,11013 Lyciella: a,b L.nigripalpis Czerny; c - Lpectinifera sp.n.; d - Lareophila sp.n.; e - L.subfasciata Ztt.; I - L.brachychaeta sp.n.; g L.mihalgii Papp (Ha Papp, 1978, pnc. 24-25). Macuvra6 - 0.1 MM.

mspl, 2 stpl, pteropleuron with 2-3 black hairs. This genus is predominantly Holarctic. Out of Length 4.8 mm. more than 60 Palaearctic species [Papp, 1984; FEMALE resembles male. Length 3.5-4.1 mm Shatalkin, 1992; Yarom, 19901 the majority falls on (spring forms) and 4.5-5.2 mm (autumnal forms). to the west and central areas of the Region. As DIAGNOSIS. This new species is very similar to distinct from Homoneura, Sapromyza is not nu- L.compsella Hendel and L.rorida Fall. It differs merous in the Far East. There are only 16 species from the latter by the shorter hairs on the arista, the known from here. Eight of these species occur also third antennal segment black apically, the longer in Europe. Such is S.albifacies Czerny, S.amabilis aedeagus, the stark bristles on epandrium. Frey, S.atripes Mg., S.basalis Ztt., S.opaca Beck., L.compsella have the two rows of acrostichals, the S.sexpunctata Mg., S.viciespunctata Czerny, less dimension of the black on the third antennal S.zetterstedti Hendel. The male of S.albtfacies was segment. recorded for Primorskiy Kray [Shatalkin, 19921. I have before me the three females of this species Genus Sapromyza Fallen, 1820. (known also as S.decaspila Lw.= S.octopunctata v.Ros.) from Kurile Isis. (Kunashir, 4Mendeleyeva* 112 A.I.Shatalkin

Volcano, 26 and 28.VII.1985, leg. Czurkin; DESCRIPTION. MALE. Head lemon-yellow. 1.V11.1971, leg. Nartshuk). S.viciespunctata is a ()cellar triangle brown. Antennae lemon-yellow. rare species with the bluish-grey thorax and the Third antenna! segment about 1.3 times as long as system of black spots on the tergites of abdomen. A its width. Arista brown with yellow base and single female exemplar of this species was taking by feathering less than thickness of its basal part. Frons me 3.VII.1988 in Kamenushka (Primorskiy Kray). about 1.4 times wider than its height. Gena about In this paper 6 new species of the genus 4.5 times less than height of eye. Mouthparts and Sapromyza are described. palpi yellow. Thorax lemon-yellow. Legs yellow. All tibiae with preapicals. Wings hyaline with yellow Sapromyza arctophila Shatalkin, sp.n. veins. ta before middle of discal cell. Ultimate section of M1.2 about 1.7 times as long as penulti- MATERIAL. Holotype (s): Magadanskaya oblast', mate. Halteres yellow. Abdomen yellow. Genitalia Arkagala, 23.VIIL1978 (Kovalev). (Fig. 4): surstylus broad with apical dent-like DESCRIPTION. FEMALE. Head yellow. Anten- projection sclerotized (black) and curved on middle nae yellow. Third antenna! segment black in apical line; parameres asymmetrical and sclerotized (black) half. Arista practically bare. Frons about 1.3 times on tip, one of them longer than aedeagus and curved wider than its height. Eyes roundish. Gena high, on dorsal side apically; there is a pair of small narrow only 2.4 times less than height of eye. Mouthparts sclerites above base of aedeagus. brown. Palpi yellow, slightly darkened on tip. Chaetotaxy: I h, 1 ph, 2 npl, 0+3 dc, ac in 2 rows Thorax yellow; mesonotum with a pair of narrow of 4-5 pairs of large bristles, which equal to black stripes between de and middle rows of ac, prescutellars, I ppl, 1 mspl, 2 stpl with anterior of going from its anterior margin to 2nd dc, and a them half length of posterior. similar but wider stripes laterad dc, divided into Length 2.5 mm. presutural and postsutural patches. Upper of them FEMALE not differs from male. Length 2.5-3.0. roundish, located mesad ph; postsutural patch DIAGNOSIS. Sapromyza citrina sp.n. may be extended, reached 3rd dc, and practically mixed with recognised by the following combination of characters: median stripe. Legs yellow; tarsi darkened. All tibiae antennae and palpi yellow, large acrostichal bristles with preapicals. Wings clear, slightly greyish. ta arranged in 4-5 pairs, abdomen without black spots. wit before middle of discal cell. Ultimate section of MI., stp about 1.9 times as Iong as penultimate. Costa with Sapromyza fuscidula Shatalkin, sp.n. 2nd, 3rd, and 4th sections in proportions of 20:12:45. ta at a level which divides 3rd costal section about MATERIAL. Holotype (): Kurile Isis, Kunashir, 13:11 of latter's length from its beginning. Halteres *Mendeleyevas. Volcano, 28.V1.1985 (Czurkin). - Para- dist yellowish white. Abdomen brownish yellow. types: 2 ??, same locality as holotype, 8. and 26.VII.1985 col, Chaetotaxy: 1 h, 1 ph, 2 npl, 0+3 dc, ac in 4 rows (Czurkin). fou with internal setulae slightly more than those of DESCRIPTION. FEMALE. Head yellow. Frons external rows, 1 ppl, 1 mspl, 2 stpl with anterior of with a pair of brownish vittae placed in insertion them visibly less than posterior, pteropleuron bare. orbitals and with black minute hairs on its anterior Length 3.8 mm. part. ()cellar triangle widely black. Antennae brown, MALE unknown. its first segment (sometimes second and base of DIAGNOSIS. S.arctophila sp.n. is an easily rec- third ones) yellowish. Arista bare with microscopic *M ognized species due to the presence of the black hairs on apical half. Occiput sloping. Eyes roundish. mesonotal stripes. The other peculiarity is the compar- Gena about 3.5 times less than height of eye. trig atively long third costal segment and as the result ta Mouthparts brownish yellow. Palpi black. Thorax nal is situated far before the level of the end of R1. Perhaps yellow. Mesonotum brown with median yellowish Ari this new species is related to SJuscidula sp.n. stripe between acrostichals, a pair yellowish stripes ba$. on line of dorsocentrals, and a similar but shorter hei Sapromyza citrina Shatalkin, sp.n. stripes laterad ph, embracing humeri and notopleu- Oe Fig. 4. rae. Scutellum brownish with yellow margin and a 0Cf median yellow stripe, appearing continuation of bel MATERIAL. Holotype (cf ): Primorskiy Kray, Kamen- middle mesonotal stripe. Pleurae yellow; sternop- br( ushka (30 km SE from liuriysk), 5.VIII.1984 (Shatalkin). leurae brownish. Legs yellow.All tibiae with preapi- Le - Paratypes: di g, 31.VII. and 10.V111.1983; 4 CO', 2W, cals. Wings greyish with tp clouded. Halteres ye 15.V11-5.V111.1984; 4 d c /1, 3 W, 15-20.IX.1987 (Shatalkin), yellow. Abdomen yellow with dark fasciae (narrow- of all from same locality as holotype. ing in the middle) on hind margin of tergites. New species of Lauxaniidae 113

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0. Fig. 4. Male terminalia of Sapromyza citrina sp.n.: a - dorsal view; b - ventral view. Scale = 0.1 mm. be Prtc. 4. Tepr,uimam4H camtia Sapromyza citrina sp.n.: a - BHA caepxy; b - BHA unto. Macarra6 - 0.1 MM. •s: es Chaetotaxy: 1 h, I ph, 2 npl, 0+3 dc, ac in 4 rows times as long as penultimate. Hateres yellow. S. with a small setae of same length, 1 ppl, 1 mspl, 2 Abdomen yellow; tergites V and VI with black spots stpl with anterior of them half length of posterior. laterally as in S.zetterstedti. Genitalia as in Fig. 5 Length 3.5 mm. with massive parameres, similar to those of MALE unknown. S.(Sapromyzosoma) bipunctata Mg. ir, DIAGNOSIS. S.fuscidula sp.n. can easily be Chaetotaxy: I h, 1 ph, 2 npl, 0+3 dc, ac in 4 rows, distinguished from all other species by the peculiar prosternum with hairs, 1 ppl, I mspl, 2 stpl, 85 coloration of body. Until the male of this species is p teropleuron bare. found it is difficult to suggest it affinities. Length 4.4 mm its FEMALE unknown. )11 Sapromyza paramerata Shatalkin, sp.n. DIAGNOSIS. This new species is similar to or Fig. 5. S.zetterstedti in having the two pairs of the abdominal n, spots. It can be distinguished from the latter in its large of MATERIAL. Holotype (d): Kurile Isis., Kunashir, size and by the structure of the male genitalia. The other is *Mendeleyeva.> Volcano, 29.VII.1985 (Czurkin). species which shows the essential resemblance to h. DESCRIPTION. MALE. Head yellow. Ocellar S.paramerata sp.n. is S.pseudopaca sp.n. described e.. triangle darkish grey. Antennae yellow; third anten- bellow. S.pseudopaca sp.n. differs from the first in ax nal segment about 1.2 times as long as its width. possessing the three pairs of the abdominal spots. sh Arista dark with feathering slightly more than its es basal thickness. Frans about 1.4 times wider than Sapromyza pseudopaca Shatalkin, sp.n. er height. Gena about 4.7 times less than height of eye. Fig. 6. u.- Ocellar bristles inserted within triangle formed by a ocelli. Mouthparts yellow. Palpi yellow, darkened MATERIAL. Holotype (cr ): Primorskiy Kray, Kamen- of below. Thorax yellow. Mesonotum with a pair of ushka (30km SE from Ussuriysk) 23.V1.1983 (Shatalkin). P. brownish stripes placed on lines of dorsocentrals. - Paratypes: 2 d d , 5W, same locality as holotype, 16 and )i- Legs yellow. All tibiae with strong preapicals. Wings 711V1.1984, 13.VIII.1984, 9.1X.1987 (Shatalkin), 23 and 27.VI11.1987 (Antropov). es yellowish with yellow veins. to slightly before middle DESCRIPTION. MALE. Head yellow. Frons of discal cell. Ultimate section of Mi2 about 1.8 with a weak brownish stripe. Ocellar triangle 114 AIShatalkin

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Th ab, na ye rrif wi he ap H Fig. 5. Sapromyza paramerata sm.: a - caudal segments of male abdomen; b - parameres (ventral view). hi Pic. 5. Sapromyza paramerata sp.n.: a - molten 6pioarKa camua; h - napamepu (arui, ciniay). VE m brownish. Gena with a weak brownish mark below preapical bristles. Wings yellowish with brownish eye. Antennae yellow. Third antennal segment veins. to strikingly before middle of discal cell. A bi about 1.8 times as long as its width. Arista brown Ultimate section of M12 about 1.5 times as long as except for its base which yellow. Feathering of arista penultimate. Halteres yellow. Abdomen yellow. equal to thickness of its basal part. Frons nearly Tergites IV, V and VI each with a pair of lateral black p' square. Gena about 62 times less than height of eye. spots. Genitalia as in Fig. 6. Parameres asymmet- er Mouthparts and palpi yellow. Thorax yellow. Mes- rical: left paramere visibly less than right paramere onotum with a pair of weak brownish stripes going and with distinct configuration; parameral tips b on line of dc from mesonotal anterior margin to hind almost black. rt dorsocentrals. Legs yellow. All tibiae with strong Chaetotaxy: 1 h, 1 ph, 2 npl, one of male paratypes (13.VIII.1984, leg. Shatalkin) with intra- alar bristles, 0+3 dc, ac in 4 rows, 1 weak ppl, 1 mspl, 2 stpl, pteropleuron bare. Length 3.5 mm. FEMALE not differs from male. Length 3.1-3.8 mm. DIAGNOSIS. S.pseudopaca sp.n. is similar to S.opaca Beck. in having the black spots on the IV- VI tergites of abdomen and the dull mesonotum. However this new species may be recognized by the presence of the mesonotal stripes and the massive parameres (Fig. 6). As distinct from S.pseudopaca sp.n. the males of S.opaca have the long and narrow left paramere. The male paratype of S.pseudopaca sp.n., having the intra-alar bristle, can be separated from most species of by the °cellar bristles placed within triangle, formed by ocelli.

Sapromyza slrig1111 f era, Shatalkin sp.n. Fig. 6. Male terminalia of Sapromyza pseudopaca sp.n.: a Fig. 7. - lateral view; b parameres and aedeagus (ventral view). Scale = 0.1 mm. MATERIAL. Holotype(d ): Primorskiy Kray, Kamen- Pric. 6. TephiprElarrpim camna Sapromyzapseudopaca sp.n.: a BHA c6oKy; b - riapamepu x aKearyc (aM cnnay). Macurra6 - 0.1 MM. ushka (30 km SE from Ussuriysk), 20.IX.87 (Shatalkin). New species of Lauxaniidae 115

Paratypes: 12 C1' cr, 8 ??, 6-8.V111.1984, 1-20.IX.1987 posterior, pteropleuron bare. (Shatalkin), 1 , 21.111987 (Antropov), all from same Length 4.2 mm. locality as holotype; paratypes from Gornotayezhnoe (25 FEMALE. No apical brush of black spinules on km SE from Ussuriysk): 3 crcr , 1-2.VI11.1963 (Nartshuk); ventral surface of hind tibiae. In all other external 1 , 1 ?, 26.IX.1980 (Shatalkin); paratypes from AKe- characters it resembles male. Length 3.5-4.2 mm. drovaya Pad► Reserve (Primorskiy Kray, Khasan Distr.): 3 W,19.X.1968 (Gorocilcov);4W,21.1X1980and20.V111.1984 DIAGNOSIS. S.strigillifera sp.n. is very similar (Shatalkin); 1 Cr, 2., 5.IX.1984 (Czurkin). to S.speciosa Remm, Elberg [1980], which was DESCRIPTION. MALE. Head yellow. Ocellar described from Mongolia. The both have a close type triangle grey. Frons subshining. Antennae yellow. of the genitalia; their distinctions may be found in Third antennal segment black on apical half and the structure of surstyles internal corners of which about 2.2 times as long as its width; it without apical rounded in S.speciasa and cuted out in the new narrowing, slightly concave dorsally and convex species. In addition the aedeagus with upper lobus ventrally. Arista brown with feathering slightly which half of lower one and the male hind tibia more than its baSal thickness. Frans about 1,3 times without an apical brush of spinules in S.speciosa. wider than its height. Gena about 5 times less than height of eye. Mouthparts yellow. Palpi black in Genus Lauxania Latreille, 1804. apical half. Thorax yellow, subshining. Legs yellow. Hind preapical bristle equal to tibial thickness. Male The genus Lauxania was known hitherto from hind tibia with dense apical brush of black spinules the three Palaearctic species. Among them ventrally. Wings yellowish with yellow veins. to on L.cylindricarnis F. is the Holarctic species. The middle of discal cell. Ultimate section of M1 .2 about second species L.kerzhneri Remm, Elberg [1980] is 1.8 times as long as penultimate. Halteres yellow. known from Mongolia. The third species L.minor Abdomen yellow. Genitalia as in Fig. 7. Aedeagus Martinek [1974] was described from Czechoslova- bilobate and wrinkly; both lobes plate-like apically. kia. It occur also in Mongolia. I have this species Parameres asymmetrical: right one bilobate, left taken by me in the Far Eait (Primorskiy Kray, paramere broad and short. There is plate-like sclerite Kamenushka, 3 d'd', I 2-20.VI.1984; 1 d', 6.VIII.1988; covering up aedeagus from above (dorsally). 5 d`d, 2 ??, 29.V-20.VI.1989). Chaetotaxy: 1 h, 1 ph, 2 npl, 0+3 dc, ac in 6 rows In addition I have the three more species of Lauxania before suture and 4 rows posteriorly, 1 weak ppl, 1 which are described as a new below. A key to all known mspl, 2 stpl with anterior of them half length of Palaearctic species of Lauxania is also given.

C

Fig. 7. Male terminalia of Saprontyza strigiltifera sp.n.: a - lateral view; b - dorsal view; c - aedeagus and parameres (ventral view). Scale = 0.1 mm. Pm. 7. TepnwHaAHH caMu2 Saprarnyza strigillifera sp.n.: a - BHA c6oHy; b - BHA ceepxy; c - aaearyc H napaMepI (BHA cHaray). Macurra6 - 0.1 MM. 116 A.I.Shatalkin

C

Fig. 8. Lateral view of heads of Lauxania species: a L.minor Martinek; b L.vitripennis sp.n.; c L.flavohalterata sp.n. Pic. 8. rozoaa (c6oxy) Biwa Lauxania: a - L.minor Martinek; b - L.vitripennis sp.n.; c - L. flavohalterata sp.n.

Key to the Palaearctic species of Lauxania Lauxania flavohalterata Shatalkin, sp.n. Fig. 8c. Wing 1(4). infuscated at base. MATERIAL. Holotype (?): Amurskaya Oblast', Zeya, 2(3). Head with projecting Pronto-facial angle; face 10.V1.1978 (Shatalkin). without a swelling on its upper part (Fig. 8a). DESCRIPTION. FEMALE. Head (Fig. 8c) black First antennal segment shorter than second one. and shining. Parafacials white dusted; postgena - Europe, Mongolia, Primorskiy Kray L.minor Martinek brownish dusted. Antennae widely arranged: dis- 3(2). Head without projecting fronto-facial angle; tance between them about 1.2 times less than that Face with a moderate swelling on upper part. between antennae and eyes. Ratio of antennal First antennal segment about 2 times longer segments equal to I (for first segment): 1.5:3.8. than second one. - Holarctic species Antennae yellow; its third segment brown in dorsal L.cylindricornis F. half and about 3 times as long as width. Arista white 4(1). Wing entirely clear; no dark spot at its base. with yellowish base and whitish hairs. Feathering of 5(6). ac in 2 rows. At least femora yellow. - arista slightly more than its basal thickness. Frons -Mongolia L.kerzhneri Remm & Elberg visible wider than its height. Face glossy. Gena 6(5). ac in 4-6 rows. All femora black. about 3.7 times less than height of eye. Mouthparts 7(8). Larger species (4.7-5.7 mm). Apical part of and palpi black. Thorax black, shining. Legs dark hind tibiae of male and female with dense brush brown except for middle and hind tibiae, and also all of black spinules ventrally. Genitalia as in Fig. 9 tarsi which yellowish brown. All tibiae with a weak - Primorskiy Kray L.sonora sp.n. preapicals. Wings yellowish; no black spot on its 8(7). Smaller species (3.1-3.5 mm) No such brush root. to on middle of discal cell. Ultimate section of of spinules on hind tibiae. Mt .2 about 1.4 times as long as penultimate. 9(10). Halteres with black knob. Head as in Fig. 8b. - Kurile Isis L.vitripennissp.n. Halteres yellow. Abdomen black and shining. 10(9). Halteres yellow. Head as in Fig. 8c.- Amur- Chaetotaxy: 1 h, 1 ph, 0+3 dc, ac in 4 rows, 1 skaya Oblast' (Zeya) L.fiavohaiterata sp.n. strong ppl, 1 mspl, 2 stpl with anterior of them half of posterior, pteropleuron bare. Length 3.5 mm. New species of Lauxaniidae 117

Fig. 9. Male terrninalia of Lauxania sonora sp.n.; a - lateral view; b - ventral view. Scale = 0.1 mm. Pm. 9. Tepsimianmia onus Lauxania sonora sp.n.: a - axu cOoKy; b - orkri outlay. Macarra6 - 0.1 sm.

MALE unknown. Thorax black and shining with green reflection. Legs DIAGNOSIS. L.flavottatterata sp.n. differs from black. Anterior knees, middle and hind tibiae, and other species of Lauxania in the following combina- also tarsi brownish yellow. All tibiae with strong tion of characters: head without projecting fronto- preapicals. Hind tibiae with dense brush of black facial angle, face glossy, halteres yellow, wing clear spinules apically on its ventral surface. Wings without black spot in its base, four-row acrostichals. yellowish with yellow veins; no black spot on wing root. to slightly beyond middle of discal cell. Ultimate section of MI , about 1.5 times as long as ya, Lauxania sonora Shatalkin, sp.n. Fig. 9. penultimate. Halteres yellow. Abdomen black, shin- ing with green reflection. Genitalia as Fig. 9. ick !na MATERIAL. Holotype (cr ): Primorskiy Kray, Kamen- Chaetotaxy: 1 h, 1 ph, 0+3 dc, ac in 6 rows, I uslika (30 km SE from LIssuriysk) 5.V1II.1984 (Shatalkin). fis- strong ppl, I mspl, 2 stpl with anterior of them - Paratypes: 5 do' , 13 ??, 14.VII-1.VIII.1983, 13.VII- visibly less than posterior, pteropleuron bare. t 6.VIII.1984, 15.VIII.1987 (Shatalkin); 1 ?, 28.VII.1983 na I Length 5.5 mm. (Ozerov); 2 W, 19 and 24.VIII.1986 (Czurkin) — all FEMALE. In all exterior characters it resembles .8. from same locality as bolotype; 2 d'd, 1 Primorskiy to male. Length 5.2-5.7 mm. sal Kray, Kiyevka (Lazo Preserve), 4-6.IX. 1980 (Shatalkin). ite DESCRIPTION. MALE. Head black and shining, DIAGNOSIS. L.sonora sp.n. differs from all Lauxania ; of with 2 narrow yellowish marks along margin eye on other known Palaearctic species of in its ins level of antennae and postgena. Frons slightly large size and tibial brush of black spinules. na greyish dusted on anterior part and middle line. its Paralacials white dusted. Antennae widely arranged: irk distance between them equal to that between all antennae and eyes. Ratio of antenna! segments Lauxania vitripennis Shatalkin, sp.n. ak equal to 1:1.2:4.0. Basal segments of antennae Fig. 8b. MATERIAL. Holotype (Cr ): Kurile Isls., Kunashir, its yellow or brownish yellow. Third antennal segment Tretyakovo, 10.V11.1985 (Czurkin). of brown dorsally and brownish yellow ventrally, and DESCRIPTION. MALE. Shape of head (Fig. 8b) e. about 4 times as long as its width. Arista white with as in L.minor Martinek. Frons picked up in anterior yellowish base and whitish hairs. Feathering of third, nearly straight over antennae. Face black, 1 arista slightly more than its basal thickness. Frons shining, with bronze reflection. Parafacials brown- alf visibly wider than high, diverging a little posteriorly. ish with a silvery reflection. Gena brown, postgena Face moderately glossy. Gena about 3.3 times less black. Frons black, lightly shining, dark brown in its than height of eye. Mouthparts and palpi black. anterior third. Vertex and occiput black, lightly 118 A.I.Shatalkin Russi' shining on margin of eye. Antennae widely ar- References ranged: distance between therm equal to that from Czerny L. 1932. 50. Lauxaniidae (Sapromyzidae) // Fllegen antennae to eyes. Ratio of antenna! segments equal Palaearktischen Region. 5.1-76_ to 1:1.6:4.1. Basal antenna! segments brownish; Malloch J.R. 1929. Notes on some Oriental Sapromyzid flies third segment dark brown with yellowish base. (Diptera), with particular reference to the Philippine species Arista white with yellowish base, its hairs white, // Proc.U.S.Nat.Mus. Vol.74, Art.6. No.2751. P.1-97. Martinek V. 1974. New European species Lauxania minor sp.n. and dense seated, short (feathering of arista slightly redescription of species Lauxania cytindricornis F. (Diptera, more than thickness of its basal part). Gena about Lauxaniidae) // Biologia (Bratislava). Roe.29. P.609-617. 3.5 times less than height of eye. Mouthparts and Papp L. 1978. Contribution to the revision of the Palaearctic palpi black. Thorax black, shining. Humeri with Lai maniidae (Diptera) // Arin.liist.-nat.Mus.Nation.Hung. 1.70. P.213-231. bronze reflection. Legs black. Middle and hind tarsi Papp L. 1984. Family Lauxaniidae (Sapromyzidae) // Catalogue whitish with more darkish apical tarsomeres. All of Palaearctic Diptera. Akademiai !Gado, Budapest. Vol.9. Schmi tibiae with preapical bristles. Wings yellowish, P.193-217. Hycrt without darkened spot in its base. Halteres black. Remm E., Elberg K. 1980.1 On the Mongolian fauna of Lauxaniidae (Diptera)] // of Mongolia. No.7. K Abdomen black, shining. P.423-436 fin Russian]. Chaetotaxy: 1 h, 1 ph, 0+3 de, ac in 4 rows with Sasakawa M. 1992. Lauxaniidae (Dip tera) of Malaysia (Part 2): K median setulae noticebly more than those of exter- A revision of Homoneura van der Wulp // Insects CI4CT Maisumurana. N.Ser. Vol.46. P.133-210. nal rows, 1 ppl, 1 msel, 2 stpl, pteropleuron bare. Sasakawa M., Ikeuchi Sh. 1983. A revision of the Japanese species Length 3.1 mm. of Homoneura (Homoneura)(Diptera, Lauxaniidae) Part 2 FEMALE unknown. / / Kontyu. Vol.51. No.2. P.289-297. reco DIAGNOSIS. This new species shows most Shatalkin A.I. I 992a. [ New Lauxaniid flies (Diptera, Lauxaniidae) It is resemblance to L.minor Martinek in the structure of from the Amur region and the Far East] // Zool.Zhurn. rich Vol.71. No.9. P.79-87 [in Russian]. beet head, the coloration of body and halteres, but it differs Shatalkin A.I. 1992b. New and little-known Palaearctic Diptera of from the latter in the clear wing without the cloud in the families , , and Lauxaniidae // H.bL its base and the entirely black tibiae (the middle and Russian Entomol.J. Vol.!. No.2. P.59-74, H.p hind tibiae is yellow on the apical third in L.minor). Shewell G.E. 1986. New American genera of Lauxaniidae, based yna on species of earlier authors, and a note on Lyciella rorida Dirt (Fallen) in North America (Diptera) // Canad.Entomol. ACKNOWLEDGEMENTS. I wish to express my cordial Vol.118. No.6. P.537-547. syn thanks to the following persons for the material and for Shewell G.E. 1987. Lauxaniidae // Manual of Nearctic Diptera. Che providing the recent papers: DrE.P.Nartshuk (Zoological Ottawa: Minister Supply and Services Canada. Vol.2. lect P.951-964. sp.; Institute, S tPetersburg), DrALOzerov(Zoological Museum, Yarom I. 1990. A review:of the genus Sapromyza Fallen in Israel Moscow), Dr.LPapp (Hungarian Nat.Hist.Museurn, Buda- with remarcs on S.(Sapromyzosoma) quadripunctata Kyr pest), Dr.M.Sasakawa(Kyoto University, Japan), Dr.G.E.Shewell (Linnaeus) (Diptera: Lauxaniidae) // Entomologica Scan- (Bicsysternatics Research Institute, Canada). dinavica. Vol.21, P.289-304.

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