Russian Entomol. 1. 2(3-4): 105-118 © RUSSIAN ENTOMOLOGICAL JOURNAL, 1993 New species of Lauxaniidae (Diptera). HonMe MALI cemeiticrBa -Lauxaniidae (Diptera). A.14.111aramcam Zoological Museum, Moscow State University, Herzen str. 6, Moscow K-9, 103009 Russia. 3ooAorH4ecicrati mysett MTY, yA. repueHa 6, 1Vlocicaa K-9, 103009 Poccmi. with KEY WORDS:Lauxaniidae, taxonomy, new species, Palaearctic Region. life- KJIIOLIEBIDIE CJIOBA: Lauxaniidae, TaKCOHOMHA, HOBble ila.meapkuKa. ther ABSTRACT: The following lauxaniid species are eviitbz AayKcaminA: Homoneura amurensis sp.n., described: Homoneura amurensis sp.n., H.assidua H.assidua sp.n., H.czernyi sp.n., H.ozeroui sp.n., sp.n., H.czernyi sp.n., H.ozerovi sp.n., H.quadrifaria H.quadrifaria sp.n., Aulogastromyia rohdendorfi DM sp.n., Aulogastromyia rohdendorfi sp.n., Lyciella sp.n., Lyciella bifaria sp.n., L.brachychaeta sp.n., inual bif aria sp.n., L.brachychaeta sp.n., L.oreophila L.oreophila sp.n., L.pectinifera sp.n., Sapromyza sp.n., L.pectinifera sp.n., Sapromyza arctophila arctophila sp.n., S.citrina sp.n., S.fascidula sp.n., sp.n., S.citrina sp.n., Sluscidula sp.n., S.paramerata sp.n., S.pseudopaca sp.n., S.paramerata sp.n., S.pseudopaca sp.n., S.strigillifera sp.n. Lauxania flavohalterata sp.n., S.strigillifera sp.n. Lauxania flavohalterata sp.n., L.sonora sp.n., S.vitripennis sp.n. ripeACT213.fleHbi L.sonora sp.n., L.vitripennis sp.n. Keys to the Ta6JIHIthl Ai1A onpeAenetam na.neapwrmeckHx utztos Palaearctic species of Lyciella and Lauxania are powa Lyciella a Lauxania. presented. This paper continues the two previous articles of PE3IOME: B pa6o.re 011HCbIBMTCH czeaylotwie the author [Shatalkin, 1992 a,b] and is based on the study of flies of the family Lauxaniidae. In it 19 new species are described. Besides this, it gives a revised and enlarged keys to the Palaearctic species of the two genera, Lauxania and Lyciella. Types of new species are deposited in the Zoological Museum of Moscow State University and the Zoological Insti- tute (St.Petersbourg), Genus Homoneura van der Wulp, 1891. a Among the Palaearctic genera, Homoneura is the most large group, containing about 70 species. The considerable part of them (approximately 50 species) occur in the east area of the Palaearctic Region. While preparing the review of the Palaearctic Homoneura for printing I had examined the compre- hensive material on this genus. As a result of this examination five new species from Russia and China are described. The Chinese material was collected in the last century by the famous geographer and ethnographer O.N.Potanin. Among the others his material contains two specimens, determined by L.Czerny as Homoneura grahami sMalIoch. The Fig. 1. Male pregenital sternites of Homoneura species: examination of these specimens has revealed that a - H.amurensis sp.n.; b - H.ozeroot sp.n. one of them should be reffered to a new species. PHC. 1. Flperemiranbribre c-repiurru camuoa alma Homon- eura: a H.amurensis sp.n.; b H.azeravi sp.n. 106 A.I.Shatalkin Homoneura amurensis Shatalkin, sp.n. upper margin. Arista black with dense seated black Fig. la. hairs. Its feathering equal to width of third antenna! segment. Frons about 1.2 times wider than its MATERIAL. Holotype (Cr ): Amurskaya Oblast', Zeya, height. Gena low, about 1/5 height of eye. Eyes 6.VII.1982 (Ozerov). - Paratypes: 2 ?7, same locality as with microscopic sparse hairs. Mouthparts and palpi holotype, 4 and 12.VII.1981 (Ozerov); 1 y, same locality yellow. Thorax fulvous. Legs yellow. Fore femur as holotype, 7.VI11.1982 (Shatalkin). DESCRIPTION. MALE. Head yellow. Frons with anteroventral comb of black spinules. All tibiae slightly shining with delicate dark hairs on its with strong preapicals. Wings yellowish with yellow anterior part. Antennae yellow; third antennal veins. ta slightly before middle of discal cell. Ultimate section of about 1.7 times as long as segment about 1.7 times more than its width. Arista M!+s penultimate. Halteres fulvous. Abdomen brownish dark brown with feathering equal to or slightly more yellow. than its thickened basal part. Gena about 1 /3 height of eye. Mouthparts and palpi yellow. Thorax yellow. Chaetotaxy: 1 h, 1 ph, 2 npl, 0+3 dc, ac in 4 rows, prosternum with setulae, 1 weak ppl, 1 mspl, 2 stpl, Mesonotum slightly shining. Legs yellow. Fore femur with anteroventral comb of black spinules. All anterior of them shorter than posterior. Length 2.5 mm. tibiae with strong preapical bristles. Wings yellow- ish with yellow veins and cloudy ta and tp. Scarcely MALE unknown. DIAGNOSIS. In the key of Homoneura by visible clouds at ends of R4.5 and M1+2. ta beyond Malloch [19291 this species runs to H.affinis middle of discal cell. Ultimate section of M1+2 about 1.7 times as long as penultimate. Halteres yellow. Malloch known from Borneo, Malaya, and Philip- Abdomen yellow. Fifth abdominal sternite of male pines [Sasakawa, 19921. It differs from the latter in bilobate posteriorly with stout spines on margin of having four rows of acrostichals (instead of six ones in H.affinis), roundish lobes (Fig. la). Genitalia massive. and denser feathering of the arista. H.assidua sp.n. is also similar to Chaetotaxy: 1 h, 1 ph, 2 npl, I +3 dc, ac in 4 rows H.unguiculata Kert. which differs from the first in having six rows with small setae of same length, prosternum naked, 1 weak ppl, I mspl, 2 stpl with posterior of them of acrostichals, the silvery dusting of parafacials stronger than anterior. discerned by examination from above, and less darkened third antenna! segment. Length 3.1 mm. FEMALE not differs from male. Length 2.8-3.2 mm. Homoneura czernyi Shatalkin, sp.n. DIAGNOSIS. This new species is easily recog- MATERIAL. Holotype (s): China, Sichuan, Ta-xinlun nized by the presence of the presutural dc, thus (Chnyanb, Tagsminy), 12.VII.1893 (Potanin). differing from the majority of species of Homoneura. DESCRIPTION. FEMALE. Head yellow. Frons In general appearance it is similar to H.biumbrata slightly longer than its width, dull, with sparse hairs Lw. The latter differs from H.amurensis sp.n. in a in anterior part. Antennae yellow; third antenna] details of the male genital apparatus and pregenital segment about 1.3 times as long as wide. Arista sternites. In addition it has a two median rows of ac brownish yellow, practically bare. Height of eye much longer than others. On the last character about 1.4 times more than its length. Gena about 1 / H.amurensis sp.n. is separated from those speci- 4 height of eye. Mouthparts and palpi yellow. Thorax mens of H.biumbrata (described by Czerny [19321 yellow. Mesonotum blackish grey in anterior part as H.kowarzi), which have the weak dorsocentrals between humeral calli, with a similar pair of before the suture. postsutural stripes laterad dc, going from presutural darkening to level of third dc. Sternopleuron in Homoneura assidua Shatalkin, sp.n. lower part blackish grey. Legs yellow. Fore femur with anteroventral comb of black spinules. Tibiae MATERIAL. Holotype (7): Primorskiy Kray, Kamen- with preapicals. Wings with costal margin browned ushka (30 km SE from Ussuriysk) , 28. VI.1984 (Shatalkin). - Paratype: 1 7, Primorskiy Kray, Kangaus, 7.VII.1963 and both cross-veins clouded. ta beyond middle of (Nartsbuk). discal cell. Ultimate section of 141 .2 about 1.5 times DESCRIPTION. FEMALE. Head yellow. Frons as long as penultimate. Halteres yellow. Abdomen slightly shining. Antennae yellow. Third antenna! yellow. segment gently concave on dorsal margin, without Chaetotaxy: 1 h, 1 ph, 2 npl, 0+3 dc, ac in 4 rows, apical contraction, about 2.1 times as long as wide. 1 ppl, 1 mspl, 2 stpl with anterior of them shorter It black in apical half beneath and apical third of than posterior. New species of Lauxaniidae 107 Length. 2.8 mm. Homoneura quadrifaria Shatalkin, sp.n. MALE unknown. DIAGNOSIS. H.czernyi sp.n. is distinct in MATERIAL. Holotype (?): Kurile Isis., Shikotan, (on having the peculiar coloration of mesonotum and Fragmites), 19.VIII.1971 (Nartshuk). sternopleuron. In other characters, first all the DESCRIPTION. FEMALE. Head yellow. Frons costal darkening of wings, it is similar to H.grahami dull with sparse small hairs in its anterior half. Ma!loch, also known from Sichuan. The latter is Antennae yellow. Third antenna! segment about 1.6 characterized by the entirely yellow mesonotum, times as long as wide. Arista brownish; its feathering clear tp, the low gena which about 5.4 times lower equal to thickness of basal part. Frons almost than height of the eye. In addition the feathering of square. Height of eye about 1.3 times more than its the arista is equal to width of its thickened basal part length. Gena about 4.4 times less than height of eye. and the ultimate section of Mi almost 2 times as Mouthparts and palpi yellow. Thorax entirely yel- long as the foregoing in H.grahami. low. Legs yellow. Fore femora with anteroventral This new species is' named for the eminent comb of black spinules. All tibiae with strong dip terologist L.Czerny. preapicals. Middle tibiae with strong and small spurs. Wings with apical and anterior (laterad of Hotnoneura ozerovi Shatalkin, sp.n. mouth of R1) margins fuscous; tp clouded, ta Fig. 1 b. practically not clouded but darker than other veins. ta slightly beyond middle of discal cell. Ultimate MATERIAL. Holotype (Cr ): Ainurslcaya Oblast', Zeya, section of M1.2 about 1.8 times as long as penulti- 6.VII.1982 (Ozerov) . - Paratypes: 2 d'cr same locality mate. Halteres yellow. Abdomen yellow; its margin as holotype, 6.V11.1982 (Ozerov).