Flora Von Kamtschatka – 31
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A Medical-Historical Examination of the Death of Alexander the Great
A MEDICAL-HISTORICAL EXAMINATION OF THE DEATH OF ALEXANDER THE GREAT Abstract: Alexander the Great’s cause of death has been contentious since antiquity. Historians and physicians alike have proposed a multitude of hypotheses. However, neither party is without their analytical flaws. The historians often neglect obvious medical refutations. Meanwhile, the physicians often err by forsaking disciplined historical methodology. Therefore, the authors of this paper subject these prior hypotheses to both Nathan Gamble medical and historical criticism, in order to provide a multidisciplinary University of Toronto, Toronto, Canada approach to a longstanding mystery. Some hypotheses have more weight St. Mary’s University, Twickenham, London than others, as is discussed. The most probable of the poisoning hypotheses, [email protected] which aligns with the Vulgate tradition of Alexander’s death, cites the use of Veratrum album, a plant derived bane. When the Court tradition is considered, Edmund F. Bloedow i.e. that no foul play occurred, acute pancreatitis induced by alcohol abuse holds greatest credence as a hypothesis. It is hoped that the approach used Augustine College, Ottawa, Ontario will not only increase clarity regarding Alexander’s death and challenge weak [email protected] ideas but also provide an approach by which speculation about other medical diagnoses in history may be tempered and critiqued. Keywords: Alexander the Great, Retrospective Diagnosis, Regicide, Murder, Death DOI: 10.14795/j.v4i3.269 ISSN 2360 – 266X ISSN–L 2360 – 266X ACKNOWLEDGMENTS: The authors would like to thank Joel Gamble and Jonathan Madany for the kind help in reviewing and preparing the manuscript for publication. -
"National List of Vascular Plant Species That Occur in Wetlands: 1996 National Summary."
Intro 1996 National List of Vascular Plant Species That Occur in Wetlands The Fish and Wildlife Service has prepared a National List of Vascular Plant Species That Occur in Wetlands: 1996 National Summary (1996 National List). The 1996 National List is a draft revision of the National List of Plant Species That Occur in Wetlands: 1988 National Summary (Reed 1988) (1988 National List). The 1996 National List is provided to encourage additional public review and comments on the draft regional wetland indicator assignments. The 1996 National List reflects a significant amount of new information that has become available since 1988 on the wetland affinity of vascular plants. This new information has resulted from the extensive use of the 1988 National List in the field by individuals involved in wetland and other resource inventories, wetland identification and delineation, and wetland research. Interim Regional Interagency Review Panel (Regional Panel) changes in indicator status as well as additions and deletions to the 1988 National List were documented in Regional supplements. The National List was originally developed as an appendix to the Classification of Wetlands and Deepwater Habitats of the United States (Cowardin et al.1979) to aid in the consistent application of this classification system for wetlands in the field.. The 1996 National List also was developed to aid in determining the presence of hydrophytic vegetation in the Clean Water Act Section 404 wetland regulatory program and in the implementation of the swampbuster provisions of the Food Security Act. While not required by law or regulation, the Fish and Wildlife Service is making the 1996 National List available for review and comment. -
National List of Vascular Plant Species That Occur in Wetlands 1996
National List of Vascular Plant Species that Occur in Wetlands: 1996 National Summary Indicator by Region and Subregion Scientific Name/ North North Central South Inter- National Subregion Northeast Southeast Central Plains Plains Plains Southwest mountain Northwest California Alaska Caribbean Hawaii Indicator Range Abies amabilis (Dougl. ex Loud.) Dougl. ex Forbes FACU FACU UPL UPL,FACU Abies balsamea (L.) P. Mill. FAC FACW FAC,FACW Abies concolor (Gord. & Glend.) Lindl. ex Hildebr. NI NI NI NI NI UPL UPL Abies fraseri (Pursh) Poir. FACU FACU FACU Abies grandis (Dougl. ex D. Don) Lindl. FACU-* NI FACU-* Abies lasiocarpa (Hook.) Nutt. NI NI FACU+ FACU- FACU FAC UPL UPL,FAC Abies magnifica A. Murr. NI UPL NI FACU UPL,FACU Abildgaardia ovata (Burm. f.) Kral FACW+ FAC+ FAC+,FACW+ Abutilon theophrasti Medik. UPL FACU- FACU- UPL UPL UPL UPL UPL NI NI UPL,FACU- Acacia choriophylla Benth. FAC* FAC* Acacia farnesiana (L.) Willd. FACU NI NI* NI NI FACU Acacia greggii Gray UPL UPL FACU FACU UPL,FACU Acacia macracantha Humb. & Bonpl. ex Willd. NI FAC FAC Acacia minuta ssp. minuta (M.E. Jones) Beauchamp FACU FACU Acaena exigua Gray OBL OBL Acalypha bisetosa Bertol. ex Spreng. FACW FACW Acalypha virginica L. FACU- FACU- FAC- FACU- FACU- FACU* FACU-,FAC- Acalypha virginica var. rhomboidea (Raf.) Cooperrider FACU- FAC- FACU FACU- FACU- FACU* FACU-,FAC- Acanthocereus tetragonus (L.) Humm. FAC* NI NI FAC* Acanthomintha ilicifolia (Gray) Gray FAC* FAC* Acanthus ebracteatus Vahl OBL OBL Acer circinatum Pursh FAC- FAC NI FAC-,FAC Acer glabrum Torr. FAC FAC FAC FACU FACU* FAC FACU FACU*,FAC Acer grandidentatum Nutt. -
Veratrum Steroidal Alkaloid Toxicity Following Ingestion of Foraged
Veratrum Steroidal Alkaloid Toxicity Following Ingestion of Foraged Veratrum Parviflorum M Anwar 1, MW Turner 2, N Farrell 3, R Kleiman 4, WB Zomlefer 5, OM McDougal 2, BW Morgan 1 1Emory University School of Medicine, Atlanta, GA; 5Boise State University, Boise, ID; 3Rhode Island Hospital, Providence, RI; 4Wellstar Kennestone Hospital, Marietta, GA; 5University of Georgia, Athens, GA BACKGROUND RESULTS DISCUSSION • Steroidal alkaloids are found in the Veratrum genus of • The specimen was identified as V. parviflorum by botanists • Steroidal alkaloids have previously been isolated and toxicity plants. at the University of Georgia . has been reported from many species of Veratrum plants. ¡ ¢ £ ¡ ¤ ¥ ¢ ¦ ¨ § • Their toxicity manifests as GI illness followed by a x107 1.0 • This is the first reported case of Veratrum toxicity from V. Bezold-Jarisch reflex: hypopnea, hypotension and 0.5 ©¨ parviflorum with identified steroidal alkaloids. bradycardia. x107 1.0 • As far as we know, there is no previous study to characterize • Some Veratrum steroidal alkaloids are also teratogens 0.5 ¨ x107 the steroidal alkaloids in V. parviflorum . interfering with the hedgehog-2 signaling pathway which 1.0 causes cyclopsia and holoprosencephaly. 0.5 ¨ • A prior study shows some cross reactivity between Veratrum x107 1.0 steroidal alkaloids and the digoxin assay but no digoxin CASE PRESENTATION 0.5 ¨ immune fab binding. x107 • A 27 year old man (patient 1) and his 25 year old wife 1.0 0.5 (patient 2) presented to the ED with nausea and 0 x107 f) vomiting after foraging and ingesting what they believed 1.0 to be wild leeks from the Appalachian Trail in Georgia, 0.5 10 USA. -
South American Cacti in Time and Space: Studies on the Diversification of the Tribe Cereeae, with Particular Focus on Subtribe Trichocereinae (Cactaceae)
Zurich Open Repository and Archive University of Zurich Main Library Strickhofstrasse 39 CH-8057 Zurich www.zora.uzh.ch Year: 2013 South American Cacti in time and space: studies on the diversification of the tribe Cereeae, with particular focus on subtribe Trichocereinae (Cactaceae) Lendel, Anita Posted at the Zurich Open Repository and Archive, University of Zurich ZORA URL: https://doi.org/10.5167/uzh-93287 Dissertation Published Version Originally published at: Lendel, Anita. South American Cacti in time and space: studies on the diversification of the tribe Cereeae, with particular focus on subtribe Trichocereinae (Cactaceae). 2013, University of Zurich, Faculty of Science. South American Cacti in Time and Space: Studies on the Diversification of the Tribe Cereeae, with Particular Focus on Subtribe Trichocereinae (Cactaceae) _________________________________________________________________________________ Dissertation zur Erlangung der naturwissenschaftlichen Doktorwürde (Dr.sc.nat.) vorgelegt der Mathematisch-naturwissenschaftlichen Fakultät der Universität Zürich von Anita Lendel aus Kroatien Promotionskomitee: Prof. Dr. H. Peter Linder (Vorsitz) PD. Dr. Reto Nyffeler Prof. Dr. Elena Conti Zürich, 2013 Table of Contents Acknowledgments 1 Introduction 3 Chapter 1. Phylogenetics and taxonomy of the tribe Cereeae s.l., with particular focus 15 on the subtribe Trichocereinae (Cactaceae – Cactoideae) Chapter 2. Floral evolution in the South American tribe Cereeae s.l. (Cactaceae: 53 Cactoideae): Pollination syndromes in a comparative phylogenetic context Chapter 3. Contemporaneous and recent radiations of the world’s major succulent 86 plant lineages Chapter 4. Tackling the molecular dating paradox: underestimated pitfalls and best 121 strategies when fossils are scarce Outlook and Future Research 207 Curriculum Vitae 209 Summary 211 Zusammenfassung 213 Acknowledgments I really believe that no one can go through the process of doing a PhD and come out without being changed at a very profound level. -
Helleborus Niger Helleborus Niger, Commonly Called Christmas Rose Or Black Hellebore, Is an Evergreen Perennial Flowering Plant in the Buttercup Family, Ranunculaceae
Helleborus niger Helleborus niger, commonly called Christmas rose or black hellebore, is an evergreen perennial flowering plant in the buttercup family, Ranunculaceae. It is poisonous. Although the flowers resemble wild roses (and despite its common name), Christmas rose does not belong to the rose family (Rosaceae). Taxonomy The black hellebore was described by Carl Linnaeus in volume one of his Species Plantarum in 1753.The Latin specific name niger (black) may refer to the colour of the roots. There are two subspecies: H. niger niger and H. niger macranthus, which has larger flowers (up to 3.75 in/9 cm across). In the wild, H. niger niger is generally found in mountainous areas in Switzerland, southern Germany, Austria, Slovenia, Croatia and northern Italy. Helleborus niger macranthus is found only in northern Italy and possibly adjoining parts of Slovenia. Description Helleborus niger is an evergreen plant with dark leathery pedate leaves carried on stems 9–12 in (23–30 cm) tall. The large flat flowers, borne on short stems from midwinter to early spring, are generally white, but occasionally purple or pink. The tips of the petals may be flushed pink or green, and there is a prominent central boss of yellow. Horticulture The plant is a traditional cottage garden favourite because it flowers in the depths of winter. Large- flowered cultivars are available, as are pink-flowered and double-floweredselections. It can be difficult to grow well; acid soil is unsuitable, as are poor, dry conditions and full sun. Moist, humus-rich, alkaline soil in dappled shade is preferable. Leaf-mould can be dug in to improve heavy clay or light sandy soils; lime can be added to 'sweeten' acid soils. -
Investigation of Spatial and Temporal Distribution Pattern of Major
1 1 A taxonomically harmonized and temporally standardized fossil pollen dataset 2 from Siberia covering the last 40 ka 3 Xianyong Cao1*, Fang Tian1, Andrei Andreev1,2,, Patricia M. Anderson3, Anatoly V. Lozhkin4, 4 Elena Bezrukova5,6, Jian Ni7, Natalia Rudaya1,6,, Astrid Stobbe8, Mareike Wieczorek1, Ulrike 5 Herzschuh1,9,10 6 1 Alfred Wegener Institute for Polar and Marine Research, Research Unit Potsdam, Telegrafenberg A43, 14473 7 Potsdam, Germany 8 2 Institute of Geology and Petroleum Technologies, Kazan Federal University, Kremlevskaya 18, 420008 Kazan, 9 Russia 10 3 Earth and Space Sciences and Quaternary Research Center, University of Washington, Seattle, WA 98185, USA 11 4 North East Interdisciplinary Science Research Institute, Far East Branch Russian Academy of Sciences, 685000 12 Magadan, Russia 13 5 Vinogradov Institute of Geochemistry, Siberian Branch, Russian Academy of Sciences, ul. Favorskogo 1a, 14 664033 Irkutsk, Russia 15 6 Institute of Archeology and Ethnography, Siberian Branch, Russian Academy of Sciences, pr. Akad. Lavrentieva 16 17, 630090 Novosibirsk, Russia 17 7 College of Chemistry and Life Sciences, Zhejiang Normal University, Yingbin Road 688, 321004 Jinhua, China 18 8 Goethe University, Norbert-Wollheim-Platz 1, Frankfurt am Main 60629, Germany 19 9 Institute of Environmental Sciences and Geography, University of Potsdam, Karl-Liebknecht-Str. 24, 14476 20 Potsdam, Germany 21 10 Institute of Biochemistry and Biology, University of Potsdam, Karl-Liebknecht-Str. 24, Potsdam 14476, 22 Germany 23 Correspondence: Ulrike Herzschuh ([email protected]) and Xianyong Cao ([email protected]; 24 [email protected]) 2 25 * Present address: Key Laboratory of Alpine Ecology, CAS Center for Excellence in Tibetan Plateau Earth ciences, 26 Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing, 100101, China 27 Abstract 28 Pollen records from Siberia are mostly absent in global or Northern Hemisphere 29 synthesis works. -
The Claytonia Arctica Complex in Alaska
THE CLAYTONIA ARCTICA Stefanie M. Ickert-Bond,2* David Murray,2 COMPLEX IN ALASKA— Margaret G. Oliver,2,3 Hazel K. Berrios,2 and 2 ANALYZING A BERINGIAN Campbell O. Webb TAXONOMIC PUZZLE USING TAXONOMIC CONCEPTS1 ABSTRACT Trans-Beringia taxa often present complex puzzles for taxonomists, a reflection of differing traditions and opinions, taxonomic approaches, and access to material from both sides of the Bering Strait. There is wide biological variation in perceived or circumscribed taxa whose populations are widespread within the regions and yet biogeographically isolated in Asia and/or America. The Claytonia arctica complex is one such example; it illustrates these issues well and has been dealt with by North American and Russian botanists in decidedly different ways. We reviewed specimens and examined the various taxonomic concepts of C. arctica through time and source publications. The relationships (alignments) among taxonomic concepts are presented in a graphical format. We found that much of the confusion related to C. arctica in Beringia stems from overlooking C. scammaniana Hult´en sensu Hult´en (1939), and placing too much emphasis on the woody caudex and perennation structures, during the creation of two taxonomic concepts: C. arctica Adams sensu Porsild and C. porsildii Jurtzev sensu Yurtsev. The C. arctica complex (in our current sense) is an evolutionary work in progress, resulting in partially differentiated races with much overlapping variability and intergradation of characters (particularly in C. scammaniana according to our current sense) that have not reached the level of stability (i.e., individuals may still intergrade freely) usually associated with the concept of species in other arctic lineages. -
Alaska Natural Heritage Program Environment and Natural Resources Institute University of Alaska Anchorage 707 a Street AK Anchorage, Alaska 99501 NHP
STEWART RIVER TRAINING AREA RARE PLANT SURVEY 2006 Matthew L. Carlson, Robert Lipkin, Helen Cortes-Burns, Irina V. Lapina Alaska Natural Heritage Program Environment and Natural Resources Institute University of Alaska Anchorage 707 A Street AK Anchorage, Alaska 99501 NHP prepared for Alaska Army National Guard State of Alaska Department of Military and Veteran’s Affairs Environmental Section Building 57040, Camp Carrol P.O. Box 5-549 Fort Richardson, Alaska 99505-0549 November 2006 Abstract Stewart River Training Area was surveyed for the presence of rare vascular plants in July 2006. A crew of four botanists surveyed all major habitats and substrates in the course of five days. A total of 155 specimens were collected, including a number of species that are rare in the state or globally. The rare species were arctic wormwood, (Artemisia senjavinensis; Alaska Natural Heritage Program Rank: G3-S2S3), Blaisdell’s bittercress (Cardamine blaisdellii; G4T3T4-S2S3), mountain stitchwort (Minuartia biflora; G5-S3S4), Walpole’s poppy (Papaver walpolei; G3-S3), Anvil Mountain primrose (Primula anvilensis; G3G4-S3S4), Greenland buttercup (Ranunculus auricomus; G5-S2), and matted starwort (Stellaria dicranoides; G3-S3). An additional species of candytuft (Smelowskia) may be a rare species, but determination by taxonomic experts is required. Four of these species (A. senjavinensis, P. walpolei, Smelowskia sp., and Stellaria dicranoides) were associated with barren carbonate outcrops in multiple sites in the training area. Blaisdell’s bittercress (C. blaisdellii) and Anvil Mountain primrose (Primula anvilensis) were encountered frequently in wet herbaceous meadows with some ground flow of water. We collected Greenland buttercup (R. auricomus) in only one location, composed of few individuals, in a shrub-meadow mosaic. -
Germinating Seeds Or Bulbils in 87 of 113 Tested Arctic Species Indicate Potential for Ex Situ Seed Bank Storage
Polar Biol (2013) 36:819–830 DOI 10.1007/s00300-013-1307-7 ORIGINAL PAPER Germinating seeds or bulbils in 87 of 113 tested Arctic species indicate potential for ex situ seed bank storage Inger Greve Alsos • Eike Mu¨ller • Pernille Bronken Eidesen Received: 14 October 2012 / Revised: 8 February 2013 / Accepted: 13 February 2013 / Published online: 26 March 2013 Ó The Author(s) 2013. This article is published with open access at Springerlink.com Abstract Arctic plant species are expected to lose range in 1969 (n = 51) and 0.7 times that observed in 2008 due to climate change. One approach to preserve the (n = 22), indicating that recent climate warming improves genetic and species diversity for the future is to store germination in the Arctic. While in situ conservation is of propagules in seed vaults. However, germinability of seeds vital importance, ex situ conservation in seed banks is a is assumed to be low for Arctic species. We evaluated ex potential complementary conservation strategy for the situ storage potential of 113 of the 161 native angiosperms majority of Arctic vascular plant species. For species that of Svalbard by studying seed ripening and germination. did not germinate, other methods for ex situ conservation Seeds or bulbils were collected, and germinability was should be sought, for example, growing in botanical tested after one winter of storage in the Svalbard Global gardens. Seed Vault. Twenty-six of the species did not produce ripe propagules, 8 produced bulbils, and 79 produced seeds. Keywords Arctic Á Bulbils Á Conservation Á Bulbils sprouted to high percentages. -
Literature Cited
Literature Cited Robert W. Kiger, Editor This is a consolidated list of all works cited in volume 8, whether as selected references, in text, or in nomenclatural contexts. In citations of articles, both here and in the taxonomic treat- ments, and also in nomenclatural citations, the titles of serials are rendered in the forms recom- mended in G. D. R. Bridson and E. R. Smith (1991). When those forms are abbreviated, as most are, cross references to the corresponding full serial titles are interpolated here alphabetically by abbreviated form. In nomenclatural citations (only), book titles are rendered in the abbreviated forms recommended in F. A. Stafleu and R. S. Cowan (1976–1988) and F. A. Stafleu et al. (1992– 2009). Here, those abbreviated forms are indicated parenthetically following the full citations of the corresponding works, and cross references to the full citations are interpolated in the list alpha- betically by abbreviated form. Two or more works published in the same year by the same author or group of coauthors will be distinguished uniquely and consistently throughout all volumes of Flora of North America by lower-case letters (b, c, d, ...) suffixed to the date for the second and subsequent works in the set. The suffixes are assigned in order of editorial encounter and do not reflect chronological sequence of publication. The first work by any particular author or group from any given year carries the implicit date suffix “a”; thus, the sequence of explicit suffixes begins with “b”. There may be citations in this list that have dates suffixed “b” but that are not preceded by citations of “[a]” works for the same year, or that have dates suffixed “c,” “d,” or “e” but that are not preceded by citations of “[a],” “b,” “c,” and/or “d” works for that year. -
Arctic Biodiversity Assessment
310 Arctic Biodiversity Assessment Purple saxifrage Saxifraga oppositifolia is a very common plant in poorly vegetated areas all over the high Arctic. It even grows on Kaffeklubben Island in N Greenland, at 83°40’ N, the most northerly plant locality in the world. It is one of the first plants to flower in spring and serves as the territorial flower of Nunavut in Canada. Zackenberg 2003. Photo: Erik Thomsen. 311 Chapter 9 Plants Lead Authors Fred J.A. Daniëls, Lynn J. Gillespie and Michel Poulin Contributing Authors Olga M. Afonina, Inger Greve Alsos, Mora Aronsson, Helga Bültmann, Stefanie Ickert-Bond, Nadya A. Konstantinova, Connie Lovejoy, Henry Väre and Kristine Bakke Westergaard Contents Summary ..............................................................312 9.4. Algae ..............................................................339 9.1. Introduction ......................................................313 9.4.1. Major algal groups ..........................................341 9.4.2. Arctic algal taxonomic diversity and regionality ..............342 9.2. Vascular plants ....................................................314 9.4.2.1. Russia ...............................................343 9.2.1. Taxonomic categories and species groups ....................314 9.4.2.2. Svalbard ............................................344 9.2.2. The Arctic territory and its subdivision .......................315 9.4.2.3. Greenland ...........................................344 9.2.3. The flora of the Arctic ........................................316