Viruses 2013, 5, 2129-2152; doi:10.3390/v5092129 OPEN ACCESS viruses ISSN 1999-4915 www.mdpi.com/journal/viruses Article Movements of Wild Ruddy Shelducks in the Central Asian Flyway and Their Spatial Relationship to Outbreaks of Highly Pathogenic Avian Influenza H5N1 John Y. Takekawa 1,*, Diann J. Prosser 2, Bridget M. Collins 2, David C. Douglas 3, William M. Perry 4, Baoping Yan 5, Luo Ze 5, Yuansheng Hou 6, Fumin Lei 7, Tianxian Li 8, Yongdong Li 8 and Scott H. Newman 9,† 1 San Francisco Bay Estuary Field Station, Western Ecological Research Center, U.S. Geological Survey, 505 Azuar Drive, Vallejo, CA 94592, USA 2 Patuxent Wildlife Research Center, U.S. Geological Survey, Beltsville, MD 20705, USA; E-Mails:
[email protected] (D.J.P.);
[email protected] (B.M.C.) 3 Alaska Science Center, U.S. Geological Survey, Juneau, AK 99801, USA; E-Mail:
[email protected] 4 Dixon Field Station, Western Ecological Research Center, U.S. Geological Survey, 800 Business Park Drive, Suite D, Dixon, CA 95620, USA; E-Mail:
[email protected] 5 Computer Network Information Center (CNIC), Chinese Academy of Sciences, Beijing 100080, China; E-Mails:
[email protected] (B.Y.);
[email protected] (L.Z.) 6 Qinghai State Forestry Administration, Qinghai Lake National Nature Reserve (QLNNR), Xining 25700, Qinghai, China; E-Mail:
[email protected] 7 Institute of Zoology (IOZ), Chinese Academy of Sciences, Beijing 100101, China; E-Mail:
[email protected] 8 Institute of Virology (WIV), Chinese Academy of Sciences, Wuhan 430071, China; E-Mails:
[email protected] (T.L.);
[email protected] (Y.L.) 9 EMPRES Wildlife Health and Ecology Unit, Animal Health Service, Animal Production and Health Division, Food and Agriculture Organization of the United Nations, Rome 00153, Italy; E-Mail:
[email protected] † Current address: Emergency Center for Transboundary Animal Diseases (ECTAD)-Vietnam, Food and Agriculture Organization of the United Nations (FAO), No.