CHECKLIST of PENNSYLVANIA BUTTERFLIES based on the Butterflies and Moths of North America (BAMONA) project http://www.butterfliesandmoths.org/ 153 species identified in PA

Common Name Scientific Name

SPREAD-WING SKIPPERS Family: Hesperiidae Subfamily: Pyrginae

Silver-spotted Skipper Epargyreus clarus Long-tailed Skipper Urbanus proteus Golden-banded Skipper Autochton cellus Hoary Edge Achalarus lyciades Northern Cloudywing Thorybes pylades Southern Cloudywing Thorybes bathyllus Confused Cloudywing Thorybes confusis Hayhurst's Scallopwing Staphylus hayhurstii Dreamy Duskywing Erynnis icelus Sleepy Duskywing Erynnis brizo Juvenal's Duskywing Erynnis juvenalis Horace's Duskywing Erynnis horatius Mottled Duskywing Erynnis martialis Zarucco Duskywing Erynnis zarucco Funereal Duskywing Erynnis funeralis Wild Indigo Duskywing Erynnis baptisiae Columbine Duskywing Erynnis lucilius Persius Duskywing Erynnis persius Grizzled Skipper Pyrgus centaureae Common Checkered-Skipper Pyrgus communis Common Sootywing Pholisora catullus

GRASS SKIPPERS Family: Hesperiidae Subfamily: Hesperiinae

Arctic Skipper Carterocephalus palaemon Swarthy Skipper Nastra lherminier Clouded Skipper Lerema accius Least Skipper Ancyloxypha numitor European Skipper Thymelicus lineola Fiery Skipper Hylephila phyleus Leonard's Skipper Hesperia leonardus Cobweb Skipper Hesperia metea Dotted Skipper Hesperia attalus Indian Skipper Hesperia sassacus Sachem Atalopedes campestris Peck's Skipper Polites peckius Tawny-edged Skipper Polites themistocles Crossline Skipper Polites origenes Long Dash Polites mystic Whirlabout Polites vibex Northern Broken-Dash Wallengrenia egeremet Little Glassywing Pompeius verna Arogos Skipper Atrytone arogos Delaware Skipper Anatrytone logan Hobomok Skipper Poanes hobomok Zabulon Skipper Poanes zabulon Mulberry Wing Poanes massasoit Broad-winged Skipper Poanes viator Black Dash Euphyes conspicua Dion Skipper Euphyes dion Two-spotted Skipper Euphyes bimacula Dun Skipper Euphyes vestris Dusted Skipper Atrytonopsis hianna Pepper and Salt Skipper Amblyscirtes hegon Common Roadside-Skipper Amblyscirtes vialis Twin-spot Skipper Oligoria maculata Brazilian Skipper Calpodes ethlius Salt Marsh Skipper Panoquina panoquin Ocola Skipper Panoquina ocola

SWALLOWTAILS Family: Papilionidae Subfamily: Papilioninae

Pipevine Swallowtail Battus philenor Zebra Swallowtail Eurytides marcellus Black Swallowtail Papilio polyxenes Canadian Tiger Swallowtail Papilio canadensis Appalachian Tiger Swallowtail Papilio appalachiensis Eastern Tiger Swallowtail Papilio glaucus Spicebush Swallowtail Papilio troilus Palamedes Swallowtail Papilio palamedes Giant Swallowtail Papilio cresphontes

WHITES Family: Pieridae Subfamily: Pierinae

Falcate Orangetip Anthocharis midea Olympia Marble Euchloe olympia West Virginia White Pieris virginiensis Cabbage White Pieris rapae Checkered White Pontia protodice

SULPHURS Family: Pieridae Subfamily: Coliadinae

Clouded Sulphur Colias philodice Orange Sulphur Colias eurytheme Pink-edged Sulphur Colias interior Southern Dogface Zerene cesonia Cloudless Sulphur Phoebis sennae Orange-barred Sulphur Phoebis philea Little Yellow Pyrisitia lisa Sleepy Orange Abaeis nicippe Dainty Sulphur Nathalis iole

HARVESTERS Family: Lycaenidae Subfamily: Miletinae

Harvester Feniseca tarquinius

COPPERS Family: Lycaenidae Subfamily: Lycaeninae

American Copper Lycaena phlaeas Bronze Copper Lycaena hyllus Bog Copper Lycaena epixanthe

HAIRSTREAKS Family: Lycaenidae Subfamily: Theclinae

Great Purple Hairstreak Atlides halesus Juniper Hairstreak Callophrys gryneus Brown Elfin Callophrys augustinus Hoary Elfin Callophrys polios Frosted Elfin Callophrys irus Henry's Elfin Callophrys henrici Eastern Pine Elfin Callophrys niphon Oak Hairstreak Satyrium favonius Coral Hairstreak Satyrium titus Acadian Hairstreak Satyrium acadica Hickory Hairstreak Satyrium caryaevorus Edwards' Hairstreak Satyrium edwardsii Banded Hairstreak Satryium calanus Striped Hairstreak Satyrium liparops Red-banded Hairstreak Calycopis cecrops Gray Hairstreak Strymon melinus White M Hairstreak Parrhasius m-album Early Hairstreak Erora laeta

BLUES Family: Lycaenidae Subfamily: Polyommatinae

Marine Blue Leptotes marina Eastern Tailed-Blue Cupido comyntas Spring Azure Celastrina 'ladon' Northern Azure Celastrina lucia Summer Azure Celastrina neglecta Dusky Azure Celastrina nigra Appalachian Azure Celastrina neglecta-major Cherry Gall Azure Celastrina serotina Silvery Blue Glaucopsyche lygdamus Melissa Blue Plebejus melissa

TRUE METALMARKS Family: Riodinidae Subfamily: Riodininae

Northern Metalmark Calephelis borealis

SNOUTS Family: Nymphalidae Subfamily: Libytheinae

American Snout Libytheana carinenta

MILKWEED BUTTERFLIES Family: Nymphalidae Subfamily: Danainae

Monarch Danaus plexippus Queen Danaus gilippus

LONGWINGS Family: Nymphalidae Subfamily: Heliconiinae

Gulf Fritillary Agraulis vanillae Variegated Fritillary Euptoieta claudia Great Spangled Fritillary Speyeria cybele Aphrodite Fritillary Speyeria aphrodite Regal Fritillary Speyeria idalia Atlantis Fritillary Speyeria atlantis Silver-bordered Fritillary Boloria selene Meadow Fritillary Boloria bellona

ADMIRALS AND RELATIVES Family: Nymphalidae Subfamily: Limenitidinae

Red-spotted Purple Limenitis arthemis Viceroy Limenitis archippus

EMPERORS Family: Nymphalidae Subfamily: Apaturinae

Hackberry Emperor Asterocampa celtis Tawny Emperor Asterocampa clyton

TRUE BRUSH-FOOTS Family: Nymphalidae Subfamily: Nymphalinae

Silvery Checkerspot Chlosyne nycteis Gorgone Checkerspot Chlosyne gorgone Harris' Checkerspot Chlosyne harrisii Pearl Crescent Phyciodes tharos Northern Crescent Phyciodes cocyta Tawny Crescent Phyciodes batesii Baltimore Checkerspot Euphydryas phaeton Common Buckeye Junonia coenia Question Mark Polygonia interrogationis Eastern Comma Polygonia comma Green Comma Polygonia faunus Gray Comma Polygonia progne Milbert's Tortoiseshell Aglais milberti Compton Tortoiseshell Nymphalis vaualbum Mourning Cloak Nymphalis antiopa California Tortoiseshell Nymphalis californica Red Admiral Vanessa atalanta Painted Lady Vanessa cardui American Lady Vanessa virginiensis

SATYRS Family: Nymphalidae Subfamily: Satyrinae

Northern Pearly-eye Enodia anthedon Eyed Brown Satyrodes eurydice Appalachian Brown Satyrodes appalachia Common Ringlet Coenonympha tullia Little Wood Satyr Megisto cymela Carolina Satyr Hermeuptychia sosybius Common Wood Nymph Cercyonis pegala