Turbinicarpus Heliae (Cactaceae), a New Species from Central Mexico
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Turbinicarpus heliae (Cactaceae), a new species from Central Mexico s 8.2 November ISSN 2285 – 3987 the passion for cacti and other succulents 2015 Turbinicarpus heliae (Cactaceae), a new species from Central Mexico contents This is Xerophilia Special issue No 8.2 - November 2015 and it is not the official first description of Turbinicarpus heliae García-Mor., Díaz-Salim & Gonz.-Bot., sp. nov. that was already published in Xerophilia Special issue No 8.1 - October 2015. 3 · Turbinicarpus heliae García-Mor., Díaz – Salim & Gonz. – Bot., sp. nov. 4 · Taxonomic notes 4 · Aknowledgements 4 · Literature 5 · Figures 1 The graphic structure of this special issue is based on the authors' demand. Founders: Eduart Zimer • Dag Panco • Valentin Posea Nordic representative • Erik Holm Supporter • Mihai Crisbășanu Editor: Eduart Zimer Editorial team's e-Mail: [email protected]. EN edition • Eduart Zimer Graphic layout based on Andrea Cattabriga's pattern. RO edition • Valentin Posea All rights reserved – no part of this publication may be reproduced in any forms SP edition & Field researches • Pedro Nájera Quezada or by any means, without written permission of the Editor. All copyrighted PR & Graphic • Dag Panco photographs have been used under the Creative Commons Attribution 4.0 Cartography • Miguel Angel González Botello International license. Front cover Back cover Juvenile plant - detail of Fig. 1. Flowering plant - detail of Fig. 1. photo by L. J. García-Morales photo by L. J. García-Morales Contents 2 - XEROPHILIA • Special Issue No. 8.2 - November 2015 | ISSN 2285-3987 Turbinicarpus heliae (Cactaceae), unaao new especienouă species specie nueva fromdin del Mexicul Central centro Central. deMexico. México. LECCINUM J. GARCÍA-MORALES1, JOSÉ F. DÍAZ-SALÍM2, MIGUEL A. GONZÁLEZ-BOTELLO3, CRISTIAN PÉREZ BADILLO4 & CLAUDIO A. FLORES LINCE5 1.Herbario, Herbario, Departamento Departamento de de Investigación, Investigación, Museo Museo de de Historia Historia Natural Natural de de Tamaulipas, Tamaulipas, TAMUX, TAMUX, Blvd. Blvd. Fidel Fidel Velázquez Velázquez M-1M-1 L-1L-1 S/N, Col. Hora- contents cio Terán, C.P 87130, Ciudad Victoria, Tamaulipas, Mexico; e-mail: [email protected] 2.Calle Calle Retorno Retorno de de Lomas Lomas Celeste Celeste 602, 602, Lomas Lomas Residencial Residencial Pachuca, Pachuca, C.P. C.P. 42094, 42094, Pachuca, Pachuca, Hidalgo, Hidalgo, Mexico Mexico 3.Privada Privada de de Guacamayas Guacamayas 3046, 3046, El ElQuetzal, Quetzal, C.P. C.P. 67169, 67169, Guadalupe, Guadalupe, Nuevo Nuevo Leon, Leon, Mexico Mexico 4.Calle Calle Azucenas Azucenas 102, 102, Col. Col. Colinas Colinas del del Sur, Sur, C.P. C.P. 76900, 76900, Querétaro, Querétaro, Qro., Qro., Mexico Mexico 5.Monte Monte Sinaí Sinaí 117, 117, Fracc. Fracc. Lomas Lomas de de Vista Vista Hermosa, Hermosa, C.P. C.P. 42026, 42026, Pachuca, Pachuca, Hidalgo, Hidalgo, Mexico Mexico IntroductionIntroducción Turbinicarpus20); que exhibe características heliae morfológcas únicas IntroducereTheEl género genus TurbinicarpusTurbinicarpus (Backeberg)(Backeberg) Buxbaum García-Mor.,cunoscuţi;que lo separan aici delo Díaz-Salim propunem anteriormente ca & peGonz.-Bot., mencionado o specie nouă y el &Genul Backeberg Turbinicarpus (1937: 27)(Backeberg) ises aun small grupo Buxbaum group pequeño of &20-30 Ba-de sp.pentruresto nov. de ştiinţă. los (Fig. taxones 1, pag. relativos 5). conocidos; aquí pro- speciesckebergentre 20 and (1937: y 30several especies27) esteinfraspecific uncon grup muchos taxamic deoftaxones small20-30 cacti inde- ponemos como una especie nueva para la ciencia. endemicspeciifraespecíficos şi maito the multi yChihuahuan endémicos taxoni infraspecifici Desert del desierto and adjoiningde chihuacactuşi- TurbinicarpusPlanta affinis T. pseudomacrochele heliae subsp. minimus areashuensemici, endemici of yCentral regiones ai Mexico Deşertului áridas deserts adyacentes Chihuahua (Bravo & delşi Sánchez- ai centro zone- TurbinicarpusGarcía-Mor.,(G. Frank) Luthy etDíaz-Salim heliae A. Hofer , caule & Gonz.-Bot., simplici, spinis iu- Mejoradalorde Méxicoadiacente 1991,(Bravo ale Donatideşerturilor & Sánchez-Mejorada & Zanovello din Mexicul 2005, 1991, Central Hunt Do- sp.García-Mor.,venilibus nov. maturisque (Fig. 1. Díaz-Salim page numerosioribus 5). & Gonz.-Bot., longioribusque, 2006).(Bravonati & ZanovelloOn& Sánchez- a study 2005, developed Mejorada Hunt 2006.)1991,by Doweld DonatiEn una (1998), & investi Zano he- sp.caule nov. globoso (Fig. usque , pag. ad 5) brevicylindrico. atque radice upgradedvellogación 2005, realizada the Hunt taxonomic por2006). Doweld Într-un status (1998), studiu of elevóTurbinicarpus efectuat el rango de Plantanapiformi affinis longiore, a T. pseudomacrochelemaioribus tuberculis, subsp altioribus. mini- Doweldtaxonómicopseudomacrochele (1998), de acestaTurbinicarpus (Backeberg) a actualizat Buxbaumpseudomacrochele statutul & Backe taxo- musPlantalatioribusque, (G. affinisFrank) maioribus LüthyT. pseudomacrochele & A. latioribusque Hofer, cum caulis(insubsp. diametrum) sim-plex, mini- bergnomic(Backeberg) (1937: al Turbinicarpus 27)Buxbaum and two & pseudomacrocheleadditional Backeberg varieties (1937: (Backe of 27) this y- spinaemusfloribus (G. juvenilis albisFrank) fructibusque et Luthy maturus et A.numerosibusovatis Hofer circiter, caule et5 longibus,mmsimplici, longis, species,berg)dos variantes Buxbaum creating de & éstathe Backeberg genusepecie Kadenicarpus (1937:adicionalmente, 27) adăugând Doweld cre- caulisspinisseminibus globosusiuvenilibus pyriformis vel maturisque brevicylindricus nigris, circiter numerosioribus 1 mmet ra-dice longis. napi lon- (1998:şiando două el 22), varietăţigénero on the Kadenicarpus suplimentarebasis of the globoseDoweld ale acestei to(1998: cylindric specii, 22), formegioribusque, longibus, caule tuberculae globoso majoribususque ad brevicylindricoalti et lati, flo- body,creândbasándose 5–8 genul straighten laKadenicarpus forma to slightly cilíndrico-globosa Doweld curved spines,(1998: del tallo, 22),dry risatqueType albus radice: MEXICO.majoribus napiformi Hidalgo diametens longiore, State: et maioribus fructusActopan, ovalis tubercu2200 ad m 5- fruitbazându-se5-8 espinas at maturity, rectaspe corpul ovoid o ligeramente globularto pyriform până curveadas, seeds la cilindric, with fruto a mmlis,elev., altioribus longus, 13 February semina latioribusque, pyriformis2014, L.maioribus nigerGarcía-Morales ad 1latioribusque mm longus. 3398 papillose5-8secos spini al drepţimadurar,testa pânăbut semillas withoutla uşor ovoidescurbaţi,a cuticle fructea ornament,piriformes uscate (in(Holotype diametrum) ITCV ,floribus Isotype GBHalbis). fructibusque ovatis cir- andlacon maturitate, unaa big testa and seminţe papilosasucculent ovoide pero embryo sin până ornamento without la piriforme perisp en cula- citerTip 5: mmMEXICO longis,. Hidalgo seminibus State: pyriformis Actopan, nigris, 2200 circiter m al- erm.ocutícula testă Donati papiloasă y un (2003) embrión , dardowngraded grande fără ornament y suculento the status cuticular, carente of Kad şi- titudine,1 mmYoung longis. 13stems February simple, 2014, elongated, L. García-Morales 40–60 mm ×3398 5–9 undeenicarpus perisperma.embrion to maresubgenus Donati şi suculent, (2003) level bajó underfără el perisperm. rangoTurbinicarpus de Kad Do- (Holotipmm, claviform, ITCV, Isotip with 16–20 GBH). white pectinate short ra- remarkingnatienicarpus (2003) aathe declasatsubgénero infertility statutul bajoof the KadenicarpusTurbinicarpus back–crosses ladesta nivel be- dialTipo spines: MEXICO to 1 mm. Estadolong, appressed, de Hidalgo: areoles Actopan, ovoid, tweendecando subgen bothla infertilidad subgeneraîn Turbinicarpus de due las to copias vicariance,remarcând de los recogniz infertili cruces- 22000.8–1Tulpinile msnm.,× 0.2–0.3 tinere 13 mm. Febrerosunt Mature simple, 2014, elongate,plant L. García-Morales stems 40–60 simple, mm ingtateaentre two încrucişărilorambos sections subgéneros with în ambelethree debido species subgenuri a andla vicarianza, dintwo cauza sub- ×339830–60 5–9 (Holotipo mm,× 12–20 claviforme, ITCVmm,, Isotipoclearly cu 16-20 GBHseparated spini). pectinaţi, by a neck ra- species.vicarianţei,reconociendo This recunoscând narrow así dos group secciones însă of doua poorly con secţiuni, tres known especies cu taxa trei diali,from albi,the scurţi,roots. deTubercles până la 1conical-pyramidal, mm lungime, presaţi, 2–3 asspeciiy dos proposed şisubespecies. două by subspecii. Bravo Este & AcestSánchez-Mejorada pequeño grup degrupo taxoni de (1991), puţin tax- areolemmTallos long ovoidale,jóvenes and wide simples, 0,8–1 at base, × elongados, 0,2–0,3 dark green mm. 40-60 with Tulpinile mm glau ×- Glasscunoscuţi,ones mal(1997), conocidos propus Doweld de según (1998), Bravo lo Lüthy &propuesto Sánchez-Mejorada (2002), por Guzmán Bravo plantelor5-9mm,cous tinged claviformes, mature areas, sunt arranged con simple, 16-20 in 30–60 9–10 espinas ×spiral 12–20 radiales series. mm, et(1991),y Sánchez-Mejorada al. (2003), Glass Donati(1997), &Doweld(1991), Zanovello Glass(1998), (2005), (1997), Lüthy and Doweld (2002), Hunt separateblancasMain root pectinadas detuberose rădăcini or de tapîn hasta mod root, 1 clar40–70 mm de demm un largo, longgât. andadTu- (2006),Guzmán(1998), comprisesLüthy et al. (2002), (2003), 1 to Guzmán Donati3 species & et Zanovelloand al. (2003),several (2005) Donatiuncer şi- berculipresas,15–20 mm conici-piramidali,areolas wide, ovoideswith secondary 0.8-1 2-3 mm× shot-roots0.2-0.3 lungi mm.şi appearlargi Plan la-