FOLIA ENTOMOLOGICA HUNGARICA ROVARTANI KOZLEMENYEK Volume64 2003 pp.69-84.

Checklist of the Hungarianmayfly fauna (Ephemeroptera)

T. KoVAcSand E. BAUERNFEIND

Abstract:A checklistof 9l Ephemeropteraspecies from Hungary is given.

Key words: Ephemeroptera,checklist, Hungary

INTRODUCTION

The first paper surnmarizingfaunistic data of was written by Mo- csdry (1899),who recorded25 speciesfrom the presentarea of Hungary. Ujhelyi (1959, 1966)listed 46 and 5l ,respectively. The preliminary list by T6th (1992b) gaveatotal of 70 speciesfor Hungary.Recent changes in taxonomicopin- ions (e.g. Kluge & Novikova 1992,Belfiore et al. 1999,Thomas et a|.2000) and new resultsfrom faunistic researchas well as the needto investigatesome earlier doubtful recordsmake it necessaryto compile a new checklistof Hungarianmay- flies. Unfortunatelythe attributionof generic(and/or subgeneric) names to several taxa within the families Baetidae, and Ephemerellidae is still rather controversial.To facilitate comparisonwith recentkeys the nomenclaturein the present checklist follows the (provisional) use in Bauernfeind and Humpesch (2002).Fol a more detaileddiscussion of alternativeopinions see Haybach ( 1998), Bauernfeind& Moog (2001) and Bauernfeind& Humpesch(1.c.).

MATERIAL AND METHODS

The presentpaper is basedon a careful evaluationof previously published faunistic records.Literature sourcespublished prior to Mocsdry (1899) have not beenincluded in the presentstudy. Records based on previouslypublished data are

Foliae|t hun8.64,2003 70 T. Kovics& E. Bruernfeind also omitted. So, only thoserecords afe taken into accountfrom Ujhelyi (1959) which were new as comparedto the former literature. In caseof taxonomic or nomenclaturalchanges the name originally usedin the refered publication is given in parentheses. Omitted arealso all recordsbased on specimensonly doubtfully identified by the respectiveauth or hirnseTf'.Ecdyonurus hebeticus (B,aton,1885): Sdtori 1939a; Ecdyonurusfluminum (Pictet, 1843) [= ? E. dispar (Curtis, 1834)]: Andrikovics 1991a, Csbrgits 2000:,Ecdyonurus subalpinus Klaprilek, 1905: Csijrgits 2000; Rhithrogenagermanica Earon, 1885:T6th 1992a. Caenis lactect,without author and year of description,was mertioned by Bdba& Ferencz( 1971),Andlikovics & Kiss (2000)and Szit6 (2002).It is unclear whether this name refers to Caenis lactea (B:urmerster,1839) or Caenis lactea Pictet,1843. The latteris presently(Malzacher 1986 et auct.sequ.) considered to representa synonymof Caenishoraria (Linnaeus,1758). Also not included in the presentcheck list ale the following species,which havebeen listed by T6th (19920),but which refer to localities outsideof present- day Hungary accordingto the referencesmentioned by him SiphLonurusaltema- lr.rs(Say, 1824),R hithrogena hybrida Eaton, 1885,Ecdy onurus macaniThomas er Sowa,I910, ElectrogenaquatlriLineata (Landa, 1969), Behningia ulrueriLestage, 1929 andPalingenia fuliginosa (Georgi, 1802). The occurrenceof some speciespreviously quoted for Hungary remains doubtful in the absenceof referencespecimens: Ecdyonurusforcipukt (Pictet, 1843) has been reportedby Pongrdcz(1914) from Nagymaros.At presentEcdyotxurus forcipula (Pictet, 1843)is considereda nomendubtum (seeBauernfeind & Humpesch2002 for further references)and it cannotbe ascertainedto which speciesthe specimenfrom Nagymarosactually re- ferred. EpeorusalpicolaEaton, 1871 has been recorded by Pongrdcz(1936) from the M6tra Mountains. Sdtori (1939a) again cited this record, but.the specieswas no longer mentionedin comprehensiveworks publishedlater (Ujhelyi 1959, 1966, T6th 1992b)and could not be found during recentextensive collectings in the Miitra Mountains (Kovdcs 2001). Larvae of Epeorus aLpicoladevelop in rather cold streams,usually above 1000m a.s.l.or at the foot of high mountains(Bauernfeind 1994). Rhithrctgenaferruginea(Navas, 1905) was mentioned by Andrikovics(1991a, b) following Sowa (1971). The taxon is presentlyconsidered to be restrictedto Spain (Sowa & Degrange1987 et auct. sequ.)and the Hungarianrecord rnay be- long to one of severaltaxa of the "RhitlLrogetnsemicolorata-grorp" .

FoLiaent. hnrs. 64, 2003 Checklist of tie Hungarianmayfly fauna (Ephemeroptera) 71

Leptophlel:iavespertina (Linnaeus,1758) was reported by Ujhelyi (1959)for the first time from the Bakony Mountains(without preciselocality data).Then, in 1966, he mentioned it from "Cuha-valley, Mts Bakony", with a note that the voucherspecimens had beenlost (destroyed).Later, during the faunistic explora- tion of the Bakony Mountains anotherspecies, Leptophlebia marginata was col- lectedin the Cuha Valley (Ujhelyi 1979).

RESULTSAND DISCUSSION

Basedon previouslypublished records the checklistfor Hungarypresented in this paperincludes 91 taxa of Ephemeroptera. Additional recordsof anothertwo taxa (Ecdyonurushelveticus, E. fluminum) are based on doubtful identification only. Six taxa listed previously by T6th (1992b)have been recorded from localitiesoutside the presentHungarian territory (Siphlnnurus altematus, hybrida, Ecdyonurus macani, Electrogena quadrilineata,B ehningia ulmeri andP alingenia fuli gino s a). Recordsof anotherfour species(Ecdyonurus forcipula, Epeorus alpicola, Rhithrogenaferrugineaand LeptophLebia vespertina) are based on insufficientev- idenceor dubiousnames. Although no confirmedrecords exist, at leastthree of the 12 taxalisted above may still occur in Hungary (Siphlonurusalternatus, Ecdyonurus macani, Lepto- phlebta vespertina).Furthermore some additional taxa (known presentlyto occur nearthe Hungarianborders) are expectedto be found in the future.

LIST OF SPECIES

SIPHLONURIDAEUlmer. 1920

Siphlonurusaestivalis (Earon, 1903) - Ujhelyi1959, Ujhetyi 1966, T6rh t9'72, T6th t9'13, T6th 1992r, Kov6cse t al.20Ol b,Kovics et al.2002a, Kov ics et a|.2002b,B^De:nfelt\det al.2004. Siphlonurus armatus (Eaton, I870) - Mocsdry 1899,Pillich 1914,PongrCcz 1914. Ujbelyi 1966,Ujhelyi I979,T6th 1992a,Szirrlki 1998a,Bauernfeind et al. 2004. Siphlonurus lacustris (Eaton, 1870) - Mocslry 1899, Pongrdcz 1914. Ujhelyi 1966, Andrikovics | 99la, T6th 1992b,K ovics et aI. 19990,Kovics & Amb ts2001,KoviLcset al.2OO2b, Kovdcs e/ al. 2003, B a:ue:rll.felndet al.2004.

Foltuent. hung.64.2U3 72 T. Kovdcs& E. Bauemfeind

AMELETIDAEMCCAffErtV. 1991

Metreletus balcanicus (Ulmer, 1920r - Ultretyl IOOO(Merrelerus /lrlrg4l"icri see Puthz 197?),Ujhelyi 1966 (Metreletushungaricus), T6th 1973 (M. hwryaricus),T6th 1992a (Ameleatr inopinatussee Kovacs 2001b)KovS.cs et al. 1999r, Kovdcs 2001a,KoviLcset al. 2003, Bauernfeind et al.2OO4.

AMETROPODIDAEBenstsson. 1913

Am€tropusfragilis Albarda,1878 Mocsary1899, Pongri.cz 1914, Sziri.kl 1995, Juhisz e/ al. 1998,Szirdki 1998a, Kovecs et al. 1999r,Andrikovics & Turcsinyi2001, Kovdcs & Ambrus 200I,Kovics et a|.2001b,Kovdcs ?, al. 2002b,KoviLcs et a\.2003,Wendling & Haybach2003.

BAETIDAELeach. 1815

Baetisalpinus (Pictet, 1843) - Sziiiki 1998a,Kovdcs 2001a. Baetisbuceratus Eaton, 1870 - Szir6ki1998b, Szirdki 2002, Bauernfeind er dl. 2004. Baetisfuscatus (Linnaeus, l76l) - Mocsdry1899 (Baetis binocliatus),Pillich 1914 (8. bino- culatus),Pon$icz 1914(8. binoculaus),Ujhelyi 1966(8. bioculatus),Szirdki 1998a, Gildean 1999,Gt1y6set al. 1999,Kovicset al. 1999b,Andlikovics & Kiss2000, Kiss ?, al. 2001,Kovrics & Ambrus2001, Kov{cs et al. 2001b,Kovdcs & Ambrus2002, Kovics et al. 2002a,Kov6,cs et al. 2002b,Szirik:,2002, Kov1cs et al. 2003.Bauemfeind et dl 2004. Baetisgracilis Bogoescu et Tabacaru, 1957 - Kovdcset al. 1998,Kov1cset al.2O01r, Kovacs et al. 2002b,Ko'/acs et al. 2003. Baetislutheri Miiller-Liebenau,196? - T6th 19920. Baetismuticus (Linnaeus, 1758) - Uj helyi 1959\Bact is puDtilut). Uj helyi 1961(8. pumilus), t1neryi ]9de1n. pu^ilu.r), Glldean 1999, Kovrics 2001a, KovScs et al.2O02b,Kovi&set al.2003. Bauemfeindet al. 2004. Baetisniger (Linnaeus, l76l ) - Kovdcser dl. 1998,Kovics & Ambrus2001, Kovdcs & Amb- .us2002, Kovdcs e/ al. 2003. Baetispentaphlebodes Ujhelyi, 1966- Ujhelyi, 1966(see Alba-Tercedor 2002), Snith & Adrikovics2O0O (8. penraphlebodes),Bauernfeind et al. 2004 (8. pentalthlebodet).lB. penta- phlebodeshas been rccently synonymized by Alba-Tercedor(2002) with Bdeti.rrei&.r Navas 1918. Pendingfurther decisions the latter name is herestill consideredto representanomen dubian.l Baetisrhodani (Pictet,1843) Pillich 1914,Pongrdcz 1914, Sdtori 1939a, Sitori 1939r, tiitr"tyi t9ee, Andrikovics1988r, Anddkovics 1991a, Andrikovics & K6ri 1991,Andrikovics & Hadnagy1994, Kriska st dl. 1998&,Andrikovics & Kiss1999, Gdldean 1999, Gulyds etal. 1999,Ko- vics erdl. 1999r,Kiss & Andrikovics2000, Kov6cs 2001a, Kovdcs & Ambrus2001, Kolacs et al. 2001r,Kov6cs & Ambrus2002, Koviics et a1.2002b, Szit6 2002, Kovdcs er al. 2003,Bauernfeind er al.2004. Baetisscambus Eaton, 1870 - KovdcseI dl. 1998,Kovdcs e/ a/. 19990,Kovrics & Ambrus 2001,Szir5ki 2002. Kov4cs et al. 2003.Barernfeind et a\.2004. Baetistracheatus Keffermiiller et Machel.1967 - Bauernfeindet al. 2004.

Fotia etlt. hunP.64.2U3 Checklist of the Hungarianmayfly fauna (Ephemeroptera) 73

Baetis tricolor Tshemova, 1928 - Gildean 1999. Kovaicset d/. 2001a. Kovtics & Antbrus 2001, Kovdcs et aL. 2o0lb, Koviics & Ambrus 2002, Kovdcs et al. 2002a, Koyics et al. 2002b, Kovlcs et al. 2003. Baetis vardarensis Ikonomov, 1962 - Kovics et al. 1998,Koviics el dl. 1999b,Kovdcs & Ambrus 2001, Kovdcs e/ dl. 2001,, Kovdcs & Ambrus 2002, Kovdcs er dl- 2002a, Kovics et al. 2OO2b,Koyics et al.2003. Baetisvernus currjs, 1834-fjhelyi 1959,Ujhelyi 1961,fjhelyi 1966,Andrikovics l99la, T6th 1991,T6th 1992a,Szir,kl 1995,ADdrikovics & Kiss 1999,Kiss & Andrikovics 2000. Sziriki 2002, Ba]J€tnfeind et al. 2004 . Baetopus tenellus (Albarda, l8?8) Szirdki 1995, Kovics ?r 41. 19991r,Kovics & Ambrus 2001, Kov6cs er al. 2001r, Kovdcs & Ambrus 2002, Kovrcs el d|.2002b, Ko't6cs et a|.2003. Centroptilum luteolurn (Mtiller, 1776) Mocsrlry 1899, Pongr6cz 1914, Ujhelyi 1966, Erdelics 1968,tijnetyi t919, .Lndrltovics 19880,Andrikovics l99la, cildean 1999,Gulyris er a/. 1999, Kovdcs et al. 1999b, Andrikovics & Kiss 2000, Kovdcs 2001a, Kovdcs & Ambrus 2001, Kovics & Ambrus 2002, Kovlcs et al. 20O2b,Kov'cs et al. 2003. Bauernfeinde/ al. 2004. Centroptilum nanum Bogoescu,1951 - Kovlcs et a|.2002b (Pieudocentroptiloidesshadili). Centroptilum pennulatum Earon, 1870 Sitori 1939a,Ujhetyi 1959,Ujhelyi 1966,Andri- kovics l99la, Ko,t{cs et al. 1999r, Kovdcs & Ambrus 2002, Kovdcs er al. 2002a, Kovics et al. 2002b, Kov'cs et at. 2003. Centroptilum pulchrum Eaton, 1885- Kovdcs et al. 1998,Kovics & Ambrus 2001, Kovdcs et al. 20olb, Kovlcs & Ambrus 2002, Kovdcs et ol.2O02b, KoviLcset al.2003. Cloeon dipterum (Linnaeus, 176I) (sensulato) - Mocsdry 1899,Mocsdry 1899(Cloeon ru- fulum),Sztli'dy 1912,Pillich 1913,Pjllich 1914,Pongrdcz 1914 (C rufulLun),S1tori 1939a,Sdtori & Nagy 1940,djhelyi 1959(C. inscnpttolt),Ilhelyi I961, Ujhelyi 1966,Ulhelyr 1966(C. inscriptunl, tinay;, DSe g. nt|rtarr), Erdelics 1968,Erdelics 1968(C. nfllrr, T6th 1972,Andrikovics i9?3, Toth 1973.Andrikovics lq?9. Andrikovics& Kefldsz197s. Ujhelyi 1979.Ujhelli 1083.Ujhelyi 1986,Andrikovics 1981a, Andrikovics 1981r, Andrikovics 1981c, Andrikovics 1988a, Andrikovics 1988r,Andrikovics 19914, T6th 1991,T6th 1992d,Szirdki 1995, Sziriki 1998lr,Szjt6 1998. Gulyds et al. 1999, Kovi'cs et al. 1999r, Andrikovics & Kiss 2000, Csctrgits2000, Csdrgits2000 (C /rt- ll,,rr),Smith & Adrikovics 2000, Kovdcs 2001a, Koviics & Ambrus 2001, Kovecs ?r al 2001r, Ko- vdcs& Ambrus 2002,Kovics et al.2Do2a,Koyics eI aL.2O02b,Szitiki 2002,Varga 2002,Kovdcs e, al 2003,Bauernfeind et al.2004. Clo€on simile Eaton, 1870(sensu lato.l - Mocscry 1899,Pongrccz 19 14. U|helyi 1959(Cloeon praetextultl),tijtretyi tget (C. praetextwnl,Ujt'"tyi tlO6. AnJrikovics1973. Andrjkovics I9?9, Ujhelyi 1979,Andrikovics & Kiss 1999,Kovrics er al. 1999b,Kiss & Andrikovics 2000. Procloeonbifidurn (Bengtsson,1912) - Ujbelyi I959.Ujhetyi t 96t. Ujhelyi 1966,Andriko- vics 1988b,Andrikovics 1991a, Szirdki 1998a, Andrikovics & Kiss 1999,Gildean 1999, Kovrics et al. 1999b,Andrikovics & Kiss 2000, Cstirgits 2000, Kiss & Andrikovics 2000, Kiss et a\.2001, Kovdcs & Ambrus 2001,Ko\6cs et al. Z00lb, Kovdcs & Ambrus 2002,Kov^cs et al.20O2a,Kovics et al. 2oo2b,Koyacs er al 2003, Bauernfeindet d\.2004. Procloeon macronyx Kluge et Novikova, 1992 - Kovrics et al. 1998 (Pseudocentroptilunt nanum), Kovics et al. 2001b (Procloeon nana), Kovtcs & Ambrus 2002 (Centroptilum nanum), Kovics et al. 2002a (C. nanutn),Kovacs et al.20O2b (C. nanwn), KovlLcset al.2003.

Folia cat. hmg.64.2003 1,1 T. Kovdcs& E. Bauernfeind

OLIGoNEURIIDAE Ulmer. 1914

Oligoneuriellakeffermuellerae Sowa, 19'73 -Kov6cs et al. 1999d,Kovdcs & Ambrus2001, Kovdcs& Ambrus2002, Kovics et a|.2003. Oligoneuriellapallida (Hagen, 1855) Hagen1855, Mocsdry 1899, Pongrdcz 1914. Ujhelyi 1966(OLigoneuriella rhenana see Mol 1984),Mol 1984,Szidki 1995,Szirdki 1998a, Kovics et a[. 1999a.Kotdcs & Antbrus2001. Kovdcs et al. 2003. Oligoneuriellapolonica Mol, 1984- Kovicsetal.1999a, Ka ics et al.2l0lb, Kovlcs et al. 2002D,Kor ics rt a/.2003. Oligoneuriellarhenala (lmhoff,1852 )- Ujhelyit959. Ulhelyr I966 (unreliable data see Ko- v6.cset al. l999ar,Szirdki 1995, Jrh'sz et al. 1998,Szirdki 1998a, Gildean 1999, Kovdcs et al 1999a,Kordcs &Ambrus 2001, Kovdcs et aL2001b,Kovdcs & Amb s 2002,Kov4cs et al.200Zb, Kovicset a|.2003,Bauemfeind el dl. 2004.

ISONYCHIIDAEBurKs, 1953

Isonychiaignota (Walker, 1853) - Mocsdry1899, Pongrdcz 1914, Kovrics & Ambrus2001, Kov6cs& Ambrus2002, Kov{cs et al.2003.

HEPTAGENIDAENeedham. I9O1

Ecdyonurusaurantiacus (Burmeister, 1839) - Szirdki1995, Cildean 1999,Ko.tics et al. 2002a,Wendling & Haybach2003, Bauernfeind et al.2004. Ecdyonurusdispar (Curtis, 1834) - Cildean1999, Wendling & Haybach2003, Bauernfeind eta|.2004. Ecdyonurusinsignis (Eaton, 1870) Erdelics1968, cildean 1999,Kovdcs et al. 1999b, Kovdcs& Ambrus200I,KolJ(cs et al.20Olb, Kovdcs & Ambrus2002, Koyics et al.20O2a,Kov6,cs et a\.2002b,Kov{,cs et al 2003,Bauernfeind et al.2004. Ecdyonurussiveci Jacob et Braasch,1984 Bauernfeindet al. 2004. Ecdyonurusstarmachi Sowa, 1971 - Andrikovics19880, Andrikovics 1991a, Andrikovics & K6ri 1991,Kiss et al. 2001,Bauernfeind ct al. 2004. Ecdyonurussubalpinus Klapdlek, 1905 Andrikovics& Kiss2000. Ecdyonurussubmontanus Landa, 1969 Andrikovics1991a,B^\re:irrfeind et al. 2004. Ecdyonurustorrentis Kimmins, 1942 Bauernfeindel41. 2004. Ecdyonurusvenosus,Fabricius. 1775) PongrdcT lgl4.SJlori lgjqd.Ujhelyi lg5q.Ujhelyi 1966,Bdba & Ferencz197 | , T6th 1992b,Ktiska et ol. 1998b,Andrikovics & Kiss2000, Csijrgits 2000.Kiss ei al 2001. Electrogenaaffinis (Eaton, 1883) Ujhelyi1979. Kovics c! al. 1998lElectrosend fasciocu- lcra),Koviics & Ambrus2001 (E. fascioculatn),KovScs et al. 2001b(E. fctscioculata),Kovrcs & Anrbrus2002 (E /dsciocLiata),Koylcs et al. 2002a(E. fasciocular.r), Koviics ?r al 2002b(E. fas- ciocliata),Kovi.cs et a|.2003. Electrogenalateralis (Curris, 1834) - fjhelyi 1959,I]jhelyi 1961,Ujhelyi 1966,B6ba & Ferencz1971, Ujhelyi 1979, Andrikovics l99la, Sziriki1995, Kriska er al. 1998a,Szirdki 1998a, Andrikovics& Kiss 1999,Gildean 1999, Csdrgits 2000, Kiss & Andrikovics2000.

Folia ent. hung. 64, 2403 '75 Checklist of the Hungarianmayfly fauna (Ephemeroptera)

Electrogenaujhelyii (Sowa,1981) tinetyi Oe6 {Ecdyonurutsubalpinus see Sowa l98l ), Sowa1981, Sziriki 1998d, Gulyas e/al. 1999,Kiss erdl.200l, Bauernfeind ?/al 2004. Heptageniacoerulans Rostock, 187'7 - Pongrlcz1914, Erdelics 1968, Juhesz et al. 1998, Kovicset al. 1999b,Andrikovics & Kiss2000, Koviics & Amb rs 2001,Kov6cs et al.2OOlb, Kovics & Ambrus2002, Ko\ics et a|.2002a,KoviLcs et al. 2002b.Ko\ecs et al. 2003. Heptageniallava Rostock, l8?7 Mocs6ry1899, Pillich l914,Pongricz19l4, Ujhelyi 1959, Ujhelyi1966, Erdelics 1968, Bdba & Ferencz1971, Andrikovics 1988a, T6th 1992a, Szir6ki 1995, Sziriki 1998a,Gdldean 1999, Kov6cs er al. 1999r,Andrikovics & Kiss2000, Andrikovics & Tur- csdnyi2001, Kovdcs & Ambrus2001, Kovdcs et al. 20llb, Koy{cs& Ambrus2002, Kovdcs er ol 2002a,Kovi.cs et al. 2002b,Szir6ki 2002, Kovdcs el 41.2003, Wendling & Haybach2003, Bauernfeinde/ al. 2004. Heptageniafuscogrisea (Retzius, 1783) - Erdelics1968, Kovr{cs er a/. 2002b,Kovics et al. 2003. Heptagenialongicauda (Stephens,1836) - Mocsiliy 1899 (Heptageniaflayipennis), Pon€ricz1914 (H. flafipennis), S1tori 1938 (H. flatipennis), Inhisz et al. t998,Szirdki 1998a, Ko- vecse[ al. 1999b,Andrikovics & Turcsdnyi2001, Kovdcs & Ambrus2001, Kovics et al.2O01b, Kovircs& Ambrus2002, Kov{cs et al.20O2b,Kov1cset a|.2003,Wendling & Haybach2003. Ileptageniasulphurea (Mulier, 1776) - Mocsdry1899, Pongrdcz l9l4,Sitori&Nagy 1940, U.lhetyit 9S9,tl.ltretyi 1966, Bdba & Ferenczl9? I, CsdnyiI994, Szirdki 1995, Cs6nyi, 1996, Juhdsz et al. 1998,Szirikt 1998a,Gildean 1999, Kovics erdl 1999b,Andrikovics & Kiss2000, Andriko vics& Turcsdnyi2001, Kov6cs & Ambrus2001, Kovics er dl. 2001&,KovScs & Ambrus2002, Ko- !ics etal.2O02a,Kovacs etal.2002b,Szlrtrkrz0lz,Koydcs etal.2003, Wendi;ng & Haybach2003, Bauernfeindet dl. 2004. Epeorusassimilis (Eaton, 18?1) - Ujhelyi1959. Ujhelyj 1961. Ujhelyi 1966, Andrikovics 1988b(Epearus sllyicola), And kovics1991a (E. sylvicolo),Andrikovics & K6ri l99l (E syi,i- cola),Andrikovics & Hadnagy1994 ('. sllyicola),Kri,ska et al. 1998b(E. sylvicola),Kovacset al. 1999b(E. sytvicola),Andrikovics & Kiss2000 (E sylvicola),CsiSrgits 2000, Kiss et dl. 2001(A rll- vicola),Kovdcs 2001a (E. sylvicola),Kovdcs & Ambrus2OO1 (E. sylvicola),Kovdcs ?t al. 2003, Bauemfeindeldl. 2004. Rhithrogenabeskidensis Alba-Tercedor et Sowa,1987 - Kovics e, al 1998,Kov6cs & Ambrus2001, Kovics et al. 2001r,Kovrcs & Ambrus2002, Kovdcs e/ al 20OZb,Kov'cs et aL. 2003. Rhifhrog€nagennanica Eaton, 1885 - Mocsdry1899, Pongrdcz 1914, Juhrisz ?r a/ 1998.Ac- cordingto tjhelyi (1966)the record from Pilismar6t (Mocsdry 1899, Pongricz 1914) is uncertaiD. Rhithrogenairidina (Kolenati,1859) - Andrikovics1991a, Andrikovics 1991b, T6th 1992r, Kisset al. 2001,Bauetnfetnd et al. 2004. Rhithrogenapicteti Sowa, 1971 - Andrikovics1991d, Andrikovics 1991r, Bauernfeind et al 2004. Rhithrogenapuytoraci Sowa et Degrange,1987 Kisseral.2001. Rhithrogenasemicolorata (Curtis, 1834) - Mocsdry1899, Pongrdcz 1914, Sdtori 1938, Sdtori19394, tjltretyi teSl, flletyi tlee, Ujhelyi1979, Andrikovics 1991d, Andrikovics 1991r, Kriskaer al 19980,Gildean 1999, Csitrgits 2000, Kiss er al. 2001,Bauernleind et al. 2004.

Folia an hun8.6a.2003 76 T. Kovdcs& E. Beuernfeiod

LEPTOPHLEBIIDAEBanKs. 1 9OO

Choroterpes picteti (Eaton, l87l) Kovdcs e.4/. 1999b,Kovlcs et al.2OOlb,Kovlcs et at. 2002b, Kovics et a|.2003. Habroleptoides coDfusa Sartori et Jicob. I986 Ulhelyr 1959 (HabrolePtoidesmodesta), Ujhelyi 1961(H. modesra),IJ1hdyi 1966(H. no.lesta), Andrikovics I988, (H. nodetta), Aidtiko vtcs 1991a(H. nrodesta),Andrikovics & Hadn^Ey 1994(H. nlodesta),Kriska et./. 1998r, Kovdcs et dl. 1999r, Csitrgits2000 (H. nodesra), Kiss er al. 2001, Kovdcs2001a, Kovlcs er al. 2001r, Kovdcs & Arnbrus 2002, Kovics e/ al. 2003, Bauernfeinder al. 2004. Habrophlebia fusca (Curtis, 183,1)- S6tori 1939D,Ujhelyi 1966,Ujhelyi I979, Andrikovics i9914, Kovics et al. 1999b,Csdrgits 2000, Kovdcs 2001a, Koviics & Ambrus 2001,Kovics & Ambrus 2002, Koviics et al. 2002b, Kovi,cs er al. 2003, Bauernfeind et t . 2004. Habrophlebialauta Eaton,188,1 - Sirori l9lso. Ujhelyr t959.Ujhelyi I966, Andrikovics 1991c,Kovrics et al. 1999b,Csijrgits 2000, Kovrcs 2001a,Kovics & Ambrus 2001,Kovdcs & Ambrus 2002, Kovdcs ?t 41 2003, Bauemfeind et al.2004. Leptophlebia marginata (Linn6, 176'7\ Ponl"lcz 191,1,Ujhelyi 19'/9, l:uhisz er dl. 1998, Kovics erdl. 19990,Kovics & Ambrus2002. Kovdcs erdl. 2003. Paraleptophlebiacincta (Retzius,1783) - Mocsdry 1899,Pongriicz 1914. Ujhelyi I95q. fjhelyi 1966,Kovdcs & Ambrus2001. Paraleptophlebia submarginata (Stephens.I 835) - Mocsiry 1899,Pongricz I I | 4. Uj hely i 1966,Ujhelyi 1979,Andrikovics 1988b, Andrikovics 199 I d, Andrikovics& Keri 199l,Inh szet al. 1998,Kovics el al. i 999r, Kovics 2001.?,Kovdcs & Ambrus200I , Kovics zr dl. 2001r,Kovics & Ambrus 2002,Kovdcs et al. 2002a,Ko\6,cs et al.2002b,Koylcs et a1.2003, Bauernteind etal. 2004. Paraleptophlebiawerneri Ullner, lSl0 - Sitori & Nrgy 1910.Ujhelyi tq66,T6th 1973,T6th 1992b,Sziriki 1998a,Koviics er dl. 1999r. Smith & Adrikovics 2000, Kovrcs 2001a, Kovdcs ?l al. 2003. Bauernfeinder al. 2004.

EPHEMERIDAELatreiIIe. I8I O

BphemeradaDica Miiller, 1764- Mocsrify1899, Pongficz 1914, Sdtori 1938, Sdtori 1939d, Sitori 1939r,Ujhelyi 1966,Uihelyi 19?9,Andrikovics 1988b, Andrikovics 1991a, Kriska 4 al. 1998l',Andrikovics & Kiss 1999,Cildean 1999,Kovdcs zral. 1999r,Csijrgits 2000, Kiss & Andri- kovics2000, Kovdcs 2001.?, Koviics & Ambrus2001, Kovics & Antbrus2002, Kovdcs zt al.2002a, Kovilcs er d/. 2002b. Kovics et al. 2003. Bauernleinder d1.2004. Ephernera glaucops Pictet, 1843 - Sziriki 1995, Kovics eI al. 1999b,Koviics & Arnbrus 2402. Ephemera lineata Eaton, 1870 - Mocsdry 1899,Pongrrcz i 91,1,Jnhesz et al. 1998,Kovics et al 1999b, Kovics et at. 2o0ll:, Koviics ct d1. 2002d, Koviics et al. 2002b, Kovdcs el al. 2003, Wendling & Haybach2003, Barernfeind er.?1.2004. Ephemeravulgata Linnaeus, lT5U Mocsdry1899, Pillich 1914,Pongricz 1914, Sdtori & Nagy 1940,Ujhelyi 1966,Lljhelyi 19?9,T6rh 1991,T6rh l9?2, fjhetyi 1983,Juhiiszerd1.1998,S2i- rfki 1998a,Kovdcs €r al 19990,Andrikovics & Kiss2000, Kovrics & Ambrus2001, Kovics er dl. 20010,Kovics & Arnbrus2002, Kovics eI dl.2002b, Szit'ki20O2, Koviics er.?1.2003, Bauernfeind et al.2004.

Falia dt. htunt.64.2NJ Checklist of the Hungarianmayfly fauna (Ephemeroptera) 77

PALINGENIIDAE Albarda. 1888

Palingenialongicauda (Olivier, 1?91) - Mocstuy1899, Vurskirs 1902, Szitddy 1904, Pillich 1913,Pillich l9l4,Ponglicz 1914,Pongrdcz 1926, Unger 1927, Sdtori 1939a, Csoknya & Ferencz 1972,Csoknya l9?3, Csoknya & Halasy1974, Ferencz 1974 a, Fercnczl9'7 4b, Csoknya& Ferencz 1975,Csoknya & Halasy1975, Andikovics 1988a,Andrikovics et aL.1992, Fink & Andrikovics 199'7,JlJhesz et dl. 1998,SchmidlKloiber 1998, Gildean 7999, Kovlcs et al. 1999b,Hanar 2oo0, And kovics& Turcs6nyi2001, Kovdcs & Ambms2001, Kovdcs et (tl.20olb,Kov6cs etal.20O2b, Szit62002, Kovdcs er .'l 2003,Wendling & Haybach2003, Bauernfelnd et al.2OO4.

POLYMITARCYIDAE BanKs. 19OO

Ephoronyirgo (Olivier,1791) - Mocsery1899, Pongrdcz 1914, Pongrdcz 1933, S6tori & Nagy1940, Ujhelyi 1966, Andrikovics l988a.hhisz etal. 1998,Cildean lggg,Kovics et al. lg99b, Kovics& Ambrus200l, Kovi.cset al.200lb, Kovdcs& Ambrus2002, Kovlcs et al.2002a, Kov6cs et al.2002b,Szit6 2002, Kovlcs et al.2003,Wendling & Haybach2003, Bauernfelnd et al.2OO4.

POTAMANTHIDAE Albarda, 1888

Potamanthusluteus (Linn6,116'7) - Szil dy 1912,Mocsdry 1899, Pongr6cz 1914, Sdtori 19394,Ujhelyi 1966,Erdelics 1968, Juhdsz et al. 1998,Kovics er d/ 1999r,Andrikovics & Kiss 2000,Kovdcs & Ambrus2001, Kovacs et al. 2001r,Kovdcs & Ambrus2002, Kovdcs et al.2002a, Kovicset al.2O02b,Szirdki 2002, Kovics e/ al. 2003,Bauernleind et al.2OO4.

EPHEMERELLIDAEKlaorilek. 1909

Ephemerellaignita (Poda,176l) Mocsrry 1899(Eplrcnerella gibba), Pongricz 1974 (E. gibba),Ujhelyi1966, Ujhelyi 1979, Sziriki 1995,Andrikovics 1988r, Andrikovics l99la,Inh{sz et al. 1998,Szidki 1998a,Andrikovics & Kiss 1999,Kov6cs et dl. 1999r,Andrikovics & Kiss2000, Csdrgits2000, Kiss & Andrikovics2000, Kovrics 2001a, Kovdcs & Ambrus2001, Kovacs et al. 2001b,Kovrics & Ambrus2002, Kovecs et al. 2002a,Kovdcs er al. 2002b,Kov6cs et al. 2003, Bauenfeindet a/. 2004. Ephemerellamesoleuca (Brauer, 1857) - Kovics et al. 1998,Kovrics & Ambrus2001, KovAcs& Ambrus2002,Kov{cs et al.2OO2a,Koyacset al.20O2b.KovAcset a|.2003. Ephemerellamucronata (Bengtsson, 1909) - Kovdcs?t dl. 1998,Kovdcs & Ambrus2001, Kovrics& Ambrui2002, KoyiLcs et al.2003. EpheDerellanotata Eaton,1887 Sziraki1995, Kovics et al. 1999b,Kovdcs & Ambrus 2001,Kov6cs & Ambrus2002, Szirdki 2002, Kovics et a|.2003. Torleyamajor (Klapdlek,1905) - Ujhelyi I959.Iljhely' 1966,Jlhlsz et al. 1998,Kovdcs et al. l999b,Kov1cset al.2O0Ib,Ko'/6cset al.2002b,Kovacs et al.2003. Eurylophellakarelica Tiensuu, 1935 - Kovics & Ambrus1999, Kovics & Ambrus2002, KovAcset a|.2003,Bauernleind et dl. 2004.

Folia ent. hu^s. 64, 2403 78 T. Kovics & E. Beuernfeind

NEOEPHEMERIDAETraver, 1935

Neoephemerarnaxima (Joly, 1870) - Kovrcser dl. 1998,Kov:ics & Ambrus2001, Kovics & Ambrus2002, Kovrics et a|.2003.

CAENIDAENewman. 1853

Brachycercus europaeusKluge, 1991-Kovics & Arnbrus2001, Ko,r'6cs et al.2001,1,KorAcs & Ambtus 2002, Kovdcs et al.2002a.Kovics et al.2003. Brachycercus harrisellus Curtis, 1834- Andrikovics 1988a.Anddkovics & TurcsSnyi2001, Kovdcs & Ambrus 2001, Koviics et dl. 2002a, Kovlcs et al. 2003. Brachycercus minutus Tshernova, 1952 Juhlsz et al. 1998 (BrcLchyetcusharirelld see Kovecs et dl. 2001r), Kov6.s et al. 1998,Kovdcs & Ambrus 2001, Koviics el dl. 2001r, Kovdcs & Ambrus 2002, Kov6cs et al.20O2b.Kov'cs et a|.2003. Caenis beskidensisSowa, 1973- Szirdki 2000. Caenishoraric (Linnaeus.1758i - Srilddy lql2 rCaenit la,rtllot. Ujhelli Iq5q. UJhelyi 1961,Ujhelyi 1966,Andrikovics 1973, Anddkovics 1979, Andrikovics & Kertdsz1979. Ulhelyi 1983,Ujhelyi 1986,Anddkovics 1988a,Andrikovics 1988r, T6rh 1992b,Szirriki 1998a,Glldean 1999,Kovdcs et al. 1999r, Andrikovics & Kiss 2000, Kovrcs 20jla,Kov{cs et al.20Ol r, Kovi4cset al. 2002b, Szn1kr 2002, KoviLcset .tl. 2003, Bauernfeind al. 2004. "r Caenislactea (Burmeister, 1839) - Andrikovics1988d, T6th 1992b. Caenisluctuosa (Burmeister,1839) Andrikovics 19880,Andrikovics 1991d,Glldean 1999, Andrikovics & Kiss 2000. Bauernfeind€t ai. 2004. Caenis macrura Stephens,1835 - Mocsiry 1899 (Caeni.shaltenta), Por\gt^cz l9l4 (C. lulteruta),Ujhelyi 1966,Bdba & Ferencz 1971,Ferencz 1971b. Ujhelyi, 1979,A ddkovics 1988r, Andrikovics1991d, Szit6 1998, Aodrikovics & Kiss 2000,Cs6rgits 2000. CaeDismartae Beltiore, 1984 Szirlki 1998a. CaeDispseudorivulorum Keffermtiller, 1960 Szirriki 1998d,Kovics ?/ al. l999r,Kovrcs& Amblrjs 2001, Kovics et a\.2001r, Kovdcs & Ambrus 2002, Kovdqs er a1. 2002a, Kovecs et al. 2OO2b.Szir 6ki 2002. Kov 6cse t a[. 2003. Bauernfeind?/ a1. 2004. 9ri Caenisrobusta Eaton, 1884 - Mocsdry1899, Pongrdcz 1914, And.ikovics 1979, Andrikovics & Kert6sz1979, Andrikovics I98lb, Andrikovicsl98lc, Ujhelyi 1983,Ujhelyi 1986,Andrikovics 1988d,Andrikovics 1988r, T6th 1992b, Szirriki 1998d, Gulyis et al. 1999, Kov6cs et al. 1999b, Andrikovics & Kiss 2000, Smith & Ad kovics 2000, Andrikovics 200i , Andrikovics & Turcsinyi 2001,Kovdcs 2001a, Kovics & Ambrus 2001, Kovdcs& Ambrus 20O2,Kov1cset al.2002b,Szir'ki 2002. Varea 2002. Kovics et a[.2003.

PROSOPISTOMATIDAELameere, l9 l7

Prosopistorna pennigerurn (Muller, 1785) Kesselydk 1943 (Prosoltistonn foliaceunt), tllnety; loss {r. ptiaceuml, (Jlhelyi 1966(P. fotiaceunl.

Foltucnt. huns.64,2403 Checklist of the Hungarianmayfly fauna (Ephemeroptera) 79

Acknowledgements-W e aremost grateful to Dr Arne Haybach(Mainz) andDr Gyijrgy Szir:i- ki (HungarianNatural History Museum,Budapest) for their valuableinformation. Thanks a.e due to Dr Ott6 Merkl (HungarianNatural History Museum, Budapest)for his linguistic help in a prelimi nary draft of the manuscript.

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Palingeniafuliginosa (Greorgi) Insecta,Ephemeroptera. - In: Landolt p. & Sartori M. (eds): Ephemeroptem& Plecoptero:BiologJ Ecology-Systematics.Maron+Tinguely & LachatSA, CH-Fribout€,pp. 326-331. Gdldean,N. (1999)Some considerations about the reophilic elements ofthe beDticfauna (ord. Ephe- meroptera,Plecoplera and Trichoptera.)of rhe Upper Tisza Region. In: Hamar, J. & Sdr- kiny-Kiss, A. (eds):??e Upper Tisa Valley,Tiscir nlotrcgraphseries. Szeged, pp.413 425. Gulyds,P., N6merh,J., Csinyi, B. & Juhdsz,P. (1999)A Balaronttipldl6 kjsvizfolydsokvizmi- n6s6ge6s 6l6vildga. [Water qrality and living worid of the sma]l running watersfeeding the Lake Balaton.l - Viziigyi Kiizletndtryef81(3): 405-452. Hagen,H. A. (1855) Die Ephemeren-CattlrngOligoneuria. - Srerilrer Entunok)gischeZeitung l6i 262-2'70. Hamar,J. (2000) Tiszavir6gzds 2000. lMass swarmingofPalingenia longicaudain 2000.] fard (rr, Kiadrdtl!, pp. 14. Haybach,A. (1998) Eintagsfliegen(lnsectat Ephemeroptera) von Rheiniand-Pfalz.Dissertation Jo- hannes-Gutenberg-Universit:it-Mainz:1 4t7, XIII + 129pp. Anhang. JuhdszP., Turcslnyi I., Kovdcs T., Olajos P., Turcsinyi B. & Kiss B. ( 1998)Vizi makroszk6pikus gerinctelen61616nyegyiittesek vizsgilata a Fels6 Tiszdn. (Study on the lnacroscopicwater in- vertebratecommunities of the Upper Tisza.) - Hidrol'giai Ktjzltjtq,TS(5 6):346 34'7. Kesselydk,A. (1943) Uj magyar k6r6sz-csaltrd.[A new family of mayflies in Hungffy.] , Allrrrdlri Kiizlemdnyek 40l.213-214. Kiss, O. & Andrikovics, S. (2000) Functionalfeeding groupsalong a lowland stream(Eger Stream, - Hongary) Ve Mndhutgen der ltttentationalen Verehigung fiir Theoretische urul Angewandte Li11t1ologie 27i 1489-l 493. Kiss, O., Andrikovics,S., Murinyi, D. & Lippdczy,A. (200t) A M6rrahegysegi Csdrg6-parak vizirovar (Trichoptera,Plecoptera, Ephemeroptera) faundja. (The aquaticinsect fauna (Tricho- ptem,Plecoptera, Epheneroptera) in the Csdrg6Stream of the Miitra Mountains(Hungary).) - Hidro16giai Ki;zLijry 81(5 6): 392-393. Kluge, N. Ju., & Novikova, E. A. ( 1992)Revision of the Palearcticgenera and subgeneraof mayflies in the subfamily Cloeoninae(Ephemeroptera, Baetidie) with descriptionof new speciesfrom the USSR.- EnrorrologiceskoeObcozranie 71(l):60 83. (in Russian) Kovtrcs, T. (2001a) K6r€sz lirvdk a Mdtribdl (Ephemeroprera).( lafvae from rhe Mirra Mountains (Ephemetopten).)- Folia llistorico-naturaLiaMusei nlatruensis2St 163-169. Kovdcs,T. (2001r) Somogy l11egyek616szeinek karal6gusa (Ephemeroprera). (Checklisr olrhe may- fly fauna of Somogy county (Ephemeroptera).)- NLture Sanog)iensis 1;8'l-92. Kovics, T. & Ambrus, A. (1999)Eurylophella karelica Tiensuu, 1935 in rheCaryathian Basin (Ephe- mercptera:Ephemerellidae). - Folia historico-tnturalia Musei natraenris 23l. 153 156. Koviics, T. & Ambrus, A. (2001) Ephemeroptera,Odonata and Plecopteralarvae f.om the River Rribaand River Lapincs. - Folia llistorico-neturalia Musei matrctensis25,t 115-162. Kovics, T. & Ambrus,A. (2002)Ldrva adarokuz drseg dsa KeLkr vjdik (Herds)k6rdsz, szirxkijr6 €s dlk6r6szfaundjihoz (Ephemeroptera,Odonata, Plecoptera). (Data of larvae to the mayfly, dragonfly and stonefly faunaof the 6rqeg and Kerka-videk tHeres)(Ephemeroprera, Odonata, Plecopteq).) - Praenofica, Folia llistorico-naturalia Musei matraensis6t 2340. Koviics, T., Ambrus, A. & Blnkuti, K. (1999a) Dara on the distriburion of Oligoneuriella larvae in (Epheneroptera: - Hungary Oligoneuriidae). Folia entonologica hungarita 6O:149-154. l:- Kovdcs,T., Ambrus,A. & Bdnkuti,K. (1999b)Dara to theHungariar mayfly (Ephemeroprerir I t;ullx€ arisingfrom collectingsol laryae.- Folia listotico-naturalia M useimatraensis 23: | 5i 1'10. Koviics,T., Ambrus, A. & Juhisz, P. (2001a)New Hungarianmayfly (Ephemeroprera)species aris- jng f.om collectings of larvae II.. - Miscellaneapologica hungarica 13(2000):8l-83.

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Kovrcs, T., Ambrus, A. & Juhdsz,P. (2002a) Ephemeropteraand Odonatalarvae from the River lpoly (Hungary). - Fo lia historico-naturalio Musei natraensis 26|. 163-16'J. Kovdcs,T., Ambrus, A. & Juhdsz,P. (2002r) Ephemeroptera,Odonata and Plecopteralarvae from the River Tisza in the year of cyanid pollution (2000). - Folia historico-naruralia Musei matraensis26: 169 1'18. Kov{cs, T., Ambrus, A. & Juhdsz,P. (2003) Data to the Hungarianmayfly (Ephemeroptera)fauna arisingfrom collectingsof larvaeII. - Folia historico-naxtralia Musei nlatraensis27, 59 :72. KovAcs,T., Ambrus, A., Bdnkuti, K. & JuhAsz,P. (1998) New Hungarianmaytly (Ephemeroptera) speciesarising from collectings of larvae.- Miscellaneazoologica hungarica 12| 55-60. l Kovi{cs,T-, Juh6sz,P. & Turcsinyi I. (2001b) Ephemeroptera,Odonata and Plecopteralarvae liom the River Tisza ( 1997-1999\. - Folia listotico-naturclia Musei nntraensis 25i 135-143. Kriska, Gy, Andrikovics, S. & Szit6, A. (19984) Phenologicaldata on a parasiticrelationship be- . tweenElectrogena lateralis (Curtjs, 1834 ) (Ephenreroptera)and Symbiocladiusrhithrogenae '79-84. (Zavrel, 1924) (Chironomidae).- OpusculaZoologica 3l IGiska, Cy., Horydth, G. & Andrikovics, S. (1998r) Why do mayflies lay their eggsen masseon dry asphaltroads?Water-imilating polarized lightrcflected ftom asphaltattractsEphetneroptera. - The Joumol of ExperimentalBiology 201:22'73-2286. Malzacher,P. (1986)Diagnose, Verbreitung und Biologie der europaischenCaenis-Arten (Epheme- roptera:Caenidae).- Stuttgarter Beitrage zur Naurkunde, Serie A (Biologie),387. l4l. Mocsfuy, S. (1899) Ordo. Pseudo-neuroptera.-In A Magyar Birodalon Allatvildga (Farna Regni Hungaiae). K. M. TermdszettudominyiT6rsulat, Budapest, pp. 23-27. Mol, A. W. M. (1984) Oligoneuriella polonica n. sp., and a note on O. pallida (Hagen, 1855) (Ephemeroptera:Oligoneuriidae). - EntomologischeBeichtetl. 44r 126 128. Pillich, F. (1913) Bncsi 1912-t61.lFarewell to l9l2.l - Rovartani Inpok 20t 11-14. Pillich, F. (1914)AarderA hropodenwehSituotrtotnla's. Ein n@nogralhischerBeitraS. SirnontoF nya,112 pp. Pongrtrcz,S. ( 1914)Magyarorsziig Neuropteroidii. (EnuDeratioNeuropteroidurn Regni Hungariae.) - Rovartani LaDok 2| 109-155. Pongrtcz, S.( 1926)Adatok a Balaton ker6sz faon'j hoz. lData to the mayfly fauna of the Lake Ba- laton.f - Allattoni Kdztemanyek23t213 214. Pongrdcz,5.(1933) A Tisza "virdgai". l"Flowers" of the Tisza.] - Termis.eft Llotn.ib-i Kdzldny 65(ll I2\t 262110. Pongrdcz,S. (t 936) Helyesbit6seka magyarfauna.jegy26keben. (Berichtigungen in derEnumeration der ungaischen Fauna) - Allaxatli K(izlemdnyek33t 181-193. Puthz, V. (1977) Uber die europiiischenMetreletus Demoulin (Sjphlonuridae,Ephemeroptera). - Philippia 3: 199 205. Sitori, J. (1938) Adatok a Biikk-hegys6grovadauntrjinak ismeret6hez.[Contribution to the knowl- edgeof the insectsfauna of the Biikk Mountains.l AllunaniKoz.lenenyek 35: 5l-61- S{tori, J. (1939rt)Adatok a Biikk 6s a Mdtra iovadaunijihoz (Beilriigezur Insecten-Faunades Biikk- und M6tra-Gebirgesin Norgdungan.). Allattani.Kti.lenanlek 36: 156-168. Sdtori,J. (1939r) Insekten faunistischeNotizen aus dem Transdanubiumim Maj und Juli 1938.- Fragtnenn Faaistica Hungarica 2(2): 3l-32. Sdtori, J. & Nagy, H. (1940) lnsekten-faunistischeNotizen aus GrossenUngarischen tiefebene. - Fragmenta Faunistico Hungarica 3(4): 116-118. Schmidt-Kloiber,A. (1998) ?.4 Epheneroptera(Eintagsfliegen). - Inr Schmidt Kloiber, A., l*og, O. & Graf',W. (eds): Biozijnotische Charakteristikund natunziumlicheBewe ung der link- suftigen Donau-Auen des Tullner Beckens auf Basis makrozoobenthischerIndikatoren. SchriftenreiheForschung im Verbund 5O: 102-118.

Folia a;t. hunr. 64,2003 Checklist of the Hungarianmayfly fauna (Ephemeroptera) 83

Smith M. E. & Adrikovics S. (2000) Benthic invertebratesin blackwaters:a comparisonof macro- andmesofaunal assemblages in south-eastemUnited StatesandMiddJe . Verlmndlun- gen der Intenationalen Vercinigungftir Lintnologie 27: 2556 2561. Sowa,R. (1971) Sur la taxonomiede Rhithrogenasemicolorata (Curtis) et quelqoesespbces voisines d'Europecontinental (Ephemeroptera: Heptageniidae). - 1?zvueSri.x e d" ZoologieTT| 895 920. Sowa, R. (1981) Taxonomy and ecology of Ecdyonurusujhelyii sp. n. (Ephemeroptera,Heptage- niidae) from the tributariesof Lake Balaton. - Acta Hydrobiologia ( Krakow) 23: 315-380. Sowa, R. & Degrange,Ch. ( 1987)Sur quelquesespbces europeennes de Rhithrogenadu groupese- micoforata(Ephemeroptera, Heptageniidae). Acta Hydrobiologia (Krakow) 29(4):523-534. Szll dy, Z. (1904) Tiszavirdg Debreczenben.lPalingenia longicauda in Debrecen.] - Rovartani htpok lL.: 195196. Szil6dy,Z. (1912) Magyarorszigi rovargyfijtdsemjegyz6ke.llnventory ofmy collection ofHungar- ian insects.]- Rovartani L.tpok l9r 53-58. Szirtki, Gy. (1995) Adatok a Dr6va-mell6kk6r6sz (Ephemeroprera) laundjdnak ismerer6hez. (Con- trjbution to the knowledge of maytlies (Ephemeroptera)of the Drdva Region, Hungary.) Dutviiltili Dolgozatok (A)Te 1lszexudomdnJiSorozat 8r 37-39. Szirdki, Gy. (1998a)A DrAvamente k6r6sz (Ephemeroptera) faun6ja. (The mayfly (Ephemeroprera) fauna olthe Dr6va region, Hungary.)- D/l'dntili Dolgozatok(A) Term4szettu(lomtinriSoro- zat 9| 131,134. Szirdki, Gy. ( 1998r) Baetisbuceratus Eaton, 1870(Ephemeroptera: Badtidae) 6s Chrysoperlarenoni (Lacroix, 1933) (Neuroptera:Chrysopidae) - Magyarorszig lhun6jdra 6j rovarfajok a Fer- t6-Hansiig Nemzeti Parkb6l. (Baetis buceratusEaton, 1870 (Ephemeroptera:Badtidae) and Chrysoperlarenoni (Lacroix, 1933) (Neuroptera:Chrysopidae) - insects,new to the fauna of Hungary from the Fert6-HansiigNational Park.) - Folia entonologicallungarica 59t 2'l Z-273. Szirdki, Gy. (2000) Magyarorszdgfauodjdra dj rovarfajok: Caenisbeskidensis, Sowa, 1973 (Ephe- meroptera:Caenidae) 6s Elipsocusnuptiatis Roesler, 1954 (Psocoptera: Elipsocidae). (Caenjs beskidensis,Sowa, 1973 (Ephemeroptera:Caenidae) and Elipsocusiuptialis Roesler, 1954 (Psocoptera:Elipsocidae), insects species, new to the faunaof Hungary.)- Foliae to'nolngica hungarica 611268 269. Szirtki, Gy. (2002) Ephemeropterafrom the Ferr6-HansegNational Park. - In: Mahunka, S. (ed.): The faunaofthe Fert6-Hans{g Natjonal Park.Hungarian Natural History Museum,Budapest, pp.2'71J15. Szi[6, A. (1998) A Cibakhtzi Holt-Tisza kiirnyezeti iillapotaaz iiled6kfaunaa]apjrn. (The environ- mental stagein the backwaterof the River Tisza by the macrozoobentosnear Cibalhiiza.). - H i drol6gi.ti Kdzldny 7a6--6): 323-324. Szit6, A. (2002)Az tiledekfaundtalkot6 dletkijzijssdgekdllapora aTisziiban a cianid szennyezdsutdn a hosszszelv6ny vizsgAlatok alapjrn 2000-ben.(The srageof the insect assemblagein River Tisza after the cyanide pollution by the longitudinal sectioninvestigations in 2000.)- flilro- I'giai Kdzldnr 82t l3O-133. Thomas.A., Mari, V. & Brulin, M. (2000)Corcctions i la Faunedes Eph6meres d'Europe occiden- tale: 2. Epeorusassimilis Eaton, 1885 est une espbcevalide, distincte d'E. sylvicolus (Pictet, 1865)(Ephemeroptera, Heptageniidae). - rp hemera l(2) 1999: 85-91. T6th, L. ( 1991)Contributions ro the Sronefly(Plecoptera) and Mayfly (Ephemeroptera)fauna ofrhe Biitorliget Nature Reserves.- Int Mahunka, S. (ed.\t The Bdtorliget Nature Reserves arter rtrrry years. Hungarian Natu.al History Museun, Budapest,Vol. 1. pp. 377-380. T6th, L. (1992a)Adatok a Boronka-mell6kiTejvddelmi Kiirzet kdr6sz(Ephemeroprera) 6s dlkdrdsz (Plecoptera) faunijdnak ismeretdhez.(Contribution to rhe mayfly (Ephemeroptera)and

F,'lia rn!. hunt. 61. 2003 84 T. Koviics & E. Bauernfeind

stonefly (Plecoptera)fauna of the Boronka-mell6kiNature Reserves .) Dundntili Dolgozatok (A) TennAszettudotlldnyiSororyt 7: 89 98. 'l6th, L. (1992b) Kev6sb6ismert rovarcsoportok(k6r6szek, dlk6rdszek) kutatisa a Bakony hegy s6gben(Ephemeroptera, Plecoptera). lResearch on lesserknown insectgroups 0nayf]ies, stone- flies) in the Bakony Mountains.]- aolra M&sei historico-totnnlis bako,yiersis 11: 145- 150. T6th, L. (1992c) Provisional checklist of Hungarian mayflies (Ephemercptela).- Anneles hi.ttori- co-tnturales Musei nationalis hungarici 84t 55-'14. T6th, S. (1972) Az oszldri Holt-Tisza 6l6villgir6l. [On the and plants of the Ho]t-Tisza at Oszllt.l - A Herman Ottrj Mizewn Evkdnlve ll:631 6'70. T6th, S. (1973) Adatok a Tardi-patakvolgye 6l6vilig6nak isrneretdhez.[Contribution to the knowl- edgeof the ljving wolld of lhe vxlley ol the Tard stream.] AHe natl Ott6 Mti:enm Evkijtryve lzt 549 582. tljtetyi, S. (tSSg) fer6szek Ephemeropte.n. h\t Mag)'arorszdBAllatvikiga (Fauna HLngariae). . Akad6miaiKiad6, Budapest,49: I 96. Ujhelyi, S. (1960) Metreletus hungaricussp. n., eine neueEintagsfliege (Epherneroptera) aus Un- . Eatn. Acta zoologicaAcademiae Scientiarlutr Hungaricae 6t 199-209. Ujhelyi, S. (1961)Adatok Magyarorszig k6rdszfaundjihoz.lData to Ihe mayfly faunaof Hungary.]- Allattani Kiizlendn\ek 48: 155-l 56. tijh"tyi, S. 1l9eOlt e mayflies ofHungary, with the descriptionof a new species,Baetis pentaphle- bodessp. n. (Epherneropleft).-ActaZoologica Acddeniae ScientiarLon Hungaricae 12:203210. ti.ltretyi,S. 1t OIO;laatok n6hiny rovanend bakonyi elterjedds6hez.(Data of someorde$ of Insecta - . occuring in Bakony Mountains.) A VeszprlmMegyei MLizewnokKdzlenftnyei 14| 85-93. Ujhelyi, S. ( 1983)The Ephemeropteraof the HortobdgyNational Park. - In: Mahunka, S. (ed.): The fauna of tlrc Hortobdgy National Pork. AkademiaiKiad6, Budapest,Vol. 2. p. 79. Ujhelyi, S. (1986) Ephemeropteraand Trichoptera from the KiskuDsiig National Park. - In: Mahunka, S. (ed.): Thefauna of the KiskunsdgNational Pdrt. Akaddmiai Kiad6, Budapest, Vol.2. pp.8l 83. Unger,E. (1927)Magyar tavak 6sfoly6k term6szeteshaltdpldldka. lNatural fbod lor fish in the Hun- garian lakes and rivers.l - KisArletiigyi Kiizlenlnyek 30t 555-569. Varga,l.(2002) Macroinve ebrateassemblages ofLake Fert6. - In:Mahunka,S. (ed.)rThe Faunaof theFert6-Hansiig National Park. Hungarian Natural History Museum, Budapest, pp. 171-182. Vutskits Gy. (1902) Tiszavirig (PalingenialongicaLrda Oliv.) a Zala rorkolatdn.[Palingenia longi caudiOliv. irt Ihe Zrh mouth.l- ALlattauiKiizlenteutct l: ll5 llo. Wendling, K. & Haybach,A. (2003)Notizen zu einigenEintagsfliegen (lnsecta: Ephemeroptera) aus der Theiss in Ungarn nach dem Cyanid-Unfall in Baja Mare (Rumanien) in Jahre 2000. - Leuteftornid 46t 1'1-B1.

(Received:28th Jub,, 2003)

Authors' addresses:Tibor KoVACS Mitra Museum H-3200 Cydngytjs, KossuthLajos u. 40, Hungary E-mail:[email protected]

B. BAUERNFEIND "N Naturhisto schesMuseum Wien, L ZoologischeAbteilung A-1014 Wien, Burgring 7, Austria E-nrail: [email protected]

Folia ekt. huhg. 61, 2003