DOCSLIB.ORG

A New Species of the South American Genus Metachaetodus (Coleoptera: Scarabaeoidea: Hybosoridae: Hybosorinae)

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
A New Species of the South American Genus Metachaetodus (Coleoptera: Scarabaeoidea: Hybosoridae: Hybosorinae) ISSN 0373-5680 (impresa), ISSN 1851-7471 (en línea) Rev. Soc. Entomol. Argent. 69 (3-4): 201-207, 2010 201 A new species of the South American genus Metachaetodus (Coleoptera: Scarabaeoidea: Hybosoridae: Hybosorinae) OCAMPO, Federico C. Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Áridas, CCT-CONICET-Mendoza, C.C. 507, 5500 Mendoza, Argentina; e-mail: [email protected] Una nueva especie del género sudamericano Metachaetodus (Coleoptera: Scarabaeoidea: Hybosoridae: Hybosorinae) RESUMEN. Se describe una nueva especie del género sudamericano de hybosórido Metachaetodus, M. carolineae Ocampo sp. nov. Se provee una clave para las especies del género y caracteres diagnósticos de cada una. Se suministra un mapa de la distribución de las especies, y se cita por primera vez en Brasil la especie M. discus De Borre. PALABRAS CLAVE. Hybosoridae. Neotrópico. ABSTRACT. A new species of the South American hybosorid genus Metachaetodus is described, M. carolineae Ocampo sp. nov. A key and diagnostic characters for all known species of the genus are presented. A map with the known distribution for the species is provided. The species M. discus De Borre is cited for Brazil for the first time. KEY WORDS. Hybosoridae. Neotropic. INTRODUCTION was supported by Ocampo & Hawks (2006) in a phylogenetic analysis of the family The genus Metachaetodus was described Hybosoridae. In his contribution, Martínez by De Borre (1886) for two species: D. discus (1994) also indicated that M. discus and M. De Borre (type species) from Argentina and brunneicollis are probably color variations Uruguay and M. brunneicollis De Borre of the same species, but he preferred to keep from Argentina (De Borre, 1886; Allsopp, them as different species. 1984; Ocampo & Ballerio, 2006). Schmidt As a result of a survey and inventory (1913) provided a short reference to the of the scarab fauna of the Chacoan region genus without providing further details in Northern Argentina, a new species of on these species. Many years after that, Metachaetodus was found and it is described Martínez (1994) refers to Metachaetodus and here as part of my continuous work on the briefly discussed its systematic relationships, New World hybosorid fauna. The purpose suggesting that this genus is closer to Coilodes of this contribution is to describe the new Westwood, Apalonychus Westwood, and species of Metachaetodus, provide a key and Hapalonychoides Martínez (Hybosorinae) diagnostic characters for the three species of than it is to Chaetodus Westwood the genus, and summarize what it is known (Anaidinae) as it was originally proposed about their distribution including a new by De Borre (1886). The latter hypothesis country record for Brazil. Recibido: 10-VI-2010; aceptado: 20-VIII-2010 202 Rev. Soc. Entomol. Argent. 69 (3-4): 201-207, 2010 MATERIAL AND METHODS Type species: Metachaetodus discus De Borre, 1886 (original designation). Body measurements, puncture density and size, and density of setae are based on Diagnosis. Hybosorid genus that can be the following standards. Body length was recognized from all other New World genera measured from the anterior margin of the of the family by the following combination pronotum (at the middle) to the apex of the of characters: Mandibles dorsoventrally elytra. Body width was measured across the flat, broadly rounded, lacking dorsal teeth; elytra at the widest point. Puncture density labrum transverse, with anterior margin was considered “dense” if punctures were slightly emarginated; antenna with nine nearly confluent to less than two puncture antennomeres, antennal club with three diameters apart, “moderately dense” if antennomeres, basal antennomere of club punctures were two to six diameters apart, cup-shaped; pronotum with lateral margins and “sparse” if punctures were more than six convergent toward apex; pronotal anterior diameters apart. Puncture size was defined angles acute and posterior angles right- as “small” if punctures were 0.02 mm or angled; pronotal and elytral surface glabrous, smaller, “moderate” if 0.02-0.07 mm, and punctate, punctures moderately dense; “large” if 0.07 mm or larger. Setae were meso- and metatibiae with one transversal defined as “sparse” if there were few setae, carinae, tarsal claws simple, curved; and “moderately dense” if the surface was visible male parameres strongly asymmetrical. but with many setae, and “dense” if the surface was not visible through the setae. Metachaetodus brunneicollis De Borre, Elytral carinae were counted from the elytral 1886 suture, with the elytral suture included. (Figs. 1, 2, 9) Holotype specimen label data was copied verbatim using “/” between the lines of the Metachaetodus brunneicollis De Borre, labels. 1886: 119. This study was based on specimens from the following institutions (with curators given Diagnosis. Metachaetodus brunneicollis in brackets): can be recognized from other species in CMNC – Canadian Museum of Nature, the genus by the following combination of Ottawa, Canada (R. S. Anderson, F. characters: Body color uniform light brown; Génier). pronotum with lateral margins smooth; CNCI – Canadian National Collection of elytral surface punctate lacking carinae; Insects, Ottawa, Canada (Y. Bousquet) male genitalia as in figures 1, 2. FMNH – Field Museum of Natural History, Chicago, USA (A. Newton, M. Thayer) Distribution (Fig. 9). ARGENTINA. IADIZA – Colección del Laboratorio de Córdoba: Anisacate (1), Cruz del Eje (2). Entomología (CEI), Instituto Argentino La Rioja: Chepes (6 km E) (2), Quebrada de Investigaciones de las Zonas Áridas, del Rosillo (3). Mendoza: Ñacuñán (1). San Mendoza, Argentina (F. C. Ocampo). Luis: El Milagro (1), La Toma (1), Quines (3). UNSM – University of Nebraska State Museum, Lincoln, Nebraska, USA (B. C. Remarks. Aside from the differences in Ratcliffe). color that are consistent between M. discus and M. brunneicollis, I found differences in the male genitalia that justify to keep these RESULTS two as separate species (Figs. 1-4), contrary to what was suggested by Martínez (1994) Metachaetodus De Borre, 1886 who probably, at that time, did not study the genitalia of Metachaetodus. Metachaetodus De Borre, 1886: 118. OCAMPO, F. C. A new species of Metachaetodus 203 Figs. 1-4. Aedeagus of Metachaetodus species. M. brunneicollis, 1, left view, 2, right view; M. discus, 3, left view, 4, right view. Metachaetodus discus De Borre 6-8). Length 6.12 mm; width 3.85 mm. (Figs. 3-5, 9) Color: Head, pronotum, elytra, legs, and venter brown. Head (Figs. 7, 8): Frons Metachaetodus discus De Borre, 1886: 118. slightly convex; surface rugo-strigulate. Frontoclypeal suture obsolete. Clypeal Diagnosis. Metachaetodus discus can be shape subtrapezoidal, anterior margin recognized from other species of the genus straight, surface rugo-strigulate. Clypeal by the following combination of characters: anterior declivital surface developed (Fig. Body bicolored; head, pronotum, and elytral 8), surface smooth. Labrum transverse, disc brown, elytral margins light brown; anterior margin indented; dorsal surface pronotum with lateral margins smooth; with sparse setae. Mandibles protruding elytral surface punctate, lacking carinae; and beyond labral apex, dorsoventrally flat. male genitalia as in figures 3-4. Labial surface sparsely setose, setae long, labial palps with 4 palpomeres. Maxillary Distribution (Fig. 9). ARGENTINA. palps with 4 palpomeres. Eyes visible in Buenos Aires: José Carlos Paz (2), Felipe Solá dorsal view. Antennae with 9 antennomeres; (1), La Plata (1), Sierra de la Ventana (1). Entre antennomere 2 cylindrical, 3 slightly Ríos: Pronunciamiento (16). BRAZIL. Santa conical, 4-6 moniliform; antennal club with Catarina: Nova Teutonia (2). URUGUAY. 3 antennomeres; basal antennomere of club Montevideo (De Borre 1886). cup-shaped, capable of receiving penultimate and ultimate antennomeres. Pronotum (Figs. Metachaetodus carolineae sp. nov. 6, 8): Shape convex, pronotum 0.57 times (Figs. 6-9) as long as wide; surface mostly moderately punctate, slightly rugo-punctate on apical Diagnosis. Metachaetodus carolineae half in middle; punctures moderate in can be recognized from other species of size. Anterior margin with weak bead and the genus by the following combination of membrane well developed; lateral margins characters: Body color uniformly dark brown; nearly straight, beaded, denticulate with pronotum with lateral margins denticulate; long setae; posterior margin beaded. Anterior elytra with 10 well developed, punctate angles acute; posterior angles right-angled. carinae. Scutellum: Shape sub-triangular. Elytron (Fig. 6): Globose, apex rounded; surface Description. Holotype Female (Figs. with 10, punctate striae, lateral striae only 204 Rev. Soc. Entomol. Argent. 69 (3-4): 201-207, 2010 Fig. 5. Metachaetodus discus, frontolateral view. Fig. 6. Metachaetodus carolineae, holotype, dorsal view. OCAMPO, F. C. A new species of Metachaetodus 205 Fig. 7. Metachaetodus carolineae, holotype, frontolateral view. Fig. 8. Metachaetodus carolineae, holotype, head and pronotum. 206 Rev. Soc. Entomol. Argent. 69 (3-4): 201-207, 2010 Fig. 9. Distribution map of Metachaetodus species. evident in middle; interval surface punctate, not erected. Six abdominal sternites exposed, punctures moderately dense. Lateral margin basal sternite only visible on lateral view. glabrous. Epipleuron only developed on Legs (Figs. 7, 8): Protibia with 3 teeth and 3 elytral basal half, surface smooth, wider denticles on outer margin; basal tooth small,
Load more

Recommended publications
  • The Beetle Fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and Distribution
    INSECTA MUNDI, Vol. 20, No. 3-4, September-December, 2006 165 The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and distribution Stewart B. Peck Department of Biology, Carleton University, 1125 Colonel By Drive, Ottawa, Ontario K1S 5B6, Canada stewart_peck@carleton. ca Abstract. The beetle fauna of the island of Dominica is summarized. It is presently known to contain 269 genera, and 361 species (in 42 families), of which 347 are named at a species level. Of these, 62 species are endemic to the island. The other naturally occurring species number 262, and another 23 species are of such wide distribution that they have probably been accidentally introduced and distributed, at least in part, by human activities. Undoubtedly, the actual numbers of species on Dominica are many times higher than now reported. This highlights the poor level of knowledge of the beetles of Dominica and the Lesser Antilles in general. Of the species known to occur elsewhere, the largest numbers are shared with neighboring Guadeloupe (201), and then with South America (126), Puerto Rico (113), Cuba (107), and Mexico-Central America (108). The Antillean island chain probably represents the main avenue of natural overwater dispersal via intermediate stepping-stone islands. The distributional patterns of the species shared with Dominica and elsewhere in the Caribbean suggest stages in a dynamic taxon cycle of species origin, range expansion, distribution contraction, and re-speciation. Introduction windward (eastern) side (with an average of 250 mm of rain annually). Rainfall is heavy and varies season- The islands of the West Indies are increasingly ally, with the dry season from mid-January to mid- recognized as a hotspot for species biodiversity June and the rainy season from mid-June to mid- (Myers et al.
    [Show full text]
  • The Evolution and Genomic Basis of Beetle Diversity
    The evolution and genomic basis of beetle diversity Duane D. McKennaa,b,1,2, Seunggwan Shina,b,2, Dirk Ahrensc, Michael Balked, Cristian Beza-Bezaa,b, Dave J. Clarkea,b, Alexander Donathe, Hermes E. Escalonae,f,g, Frank Friedrichh, Harald Letschi, Shanlin Liuj, David Maddisonk, Christoph Mayere, Bernhard Misofe, Peyton J. Murina, Oliver Niehuisg, Ralph S. Petersc, Lars Podsiadlowskie, l m l,n o f l Hans Pohl , Erin D. Scully , Evgeny V. Yan , Xin Zhou , Adam Slipinski , and Rolf G. Beutel aDepartment of Biological Sciences, University of Memphis, Memphis, TN 38152; bCenter for Biodiversity Research, University of Memphis, Memphis, TN 38152; cCenter for Taxonomy and Evolutionary Research, Arthropoda Department, Zoologisches Forschungsmuseum Alexander Koenig, 53113 Bonn, Germany; dBavarian State Collection of Zoology, Bavarian Natural History Collections, 81247 Munich, Germany; eCenter for Molecular Biodiversity Research, Zoological Research Museum Alexander Koenig, 53113 Bonn, Germany; fAustralian National Insect Collection, Commonwealth Scientific and Industrial Research Organisation, Canberra, ACT 2601, Australia; gDepartment of Evolutionary Biology and Ecology, Institute for Biology I (Zoology), University of Freiburg, 79104 Freiburg, Germany; hInstitute of Zoology, University of Hamburg, D-20146 Hamburg, Germany; iDepartment of Botany and Biodiversity Research, University of Wien, Wien 1030, Austria; jChina National GeneBank, BGI-Shenzhen, 518083 Guangdong, People’s Republic of China; kDepartment of Integrative Biology, Oregon State
    [Show full text]
  • Current Classification of the Families of Coleoptera
    The Great Lakes Entomologist Volume 8 Number 3 - Fall 1975 Number 3 - Fall 1975 Article 4 October 1975 Current Classification of the amiliesF of Coleoptera M G. de Viedma University of Madrid M L. Nelson Wayne State University Follow this and additional works at: https://scholar.valpo.edu/tgle Part of the Entomology Commons Recommended Citation de Viedma, M G. and Nelson, M L. 1975. "Current Classification of the amiliesF of Coleoptera," The Great Lakes Entomologist, vol 8 (3) Available at: https://scholar.valpo.edu/tgle/vol8/iss3/4 This Peer-Review Article is brought to you for free and open access by the Department of Biology at ValpoScholar. It has been accepted for inclusion in The Great Lakes Entomologist by an authorized administrator of ValpoScholar. For more information, please contact a ValpoScholar staff member at [email protected]. de Viedma and Nelson: Current Classification of the Families of Coleoptera THE GREAT LAKES ENTOMOLOGIST CURRENT CLASSIFICATION OF THE FAMILIES OF COLEOPTERA M. G. de viedmal and M. L. els son' Several works on the order Coleoptera have appeared in recent years, some of them creating new superfamilies, others modifying the constitution of these or creating new families, finally others are genera1 revisions of the order. The authors believe that the current classification of this order, incorporating these changes would prove useful. The following outline is based mainly on Crowson (1960, 1964, 1966, 1967, 1971, 1972, 1973) and Crowson and Viedma (1964). For characters used on classification see Viedma (1972) and for family synonyms Abdullah (1969). Major features of this conspectus are the rejection of the two sections of Adephaga (Geadephaga and Hydradephaga), based on Bell (1966) and the new sequence of Heteromera, based mainly on Crowson (1966), with adaptations.
    [Show full text]
  • Quick Guide for the Identification Of
    Quick Guide for the Identification of Maryland Scarabaeoidea Mallory Hagadorn Dr. Dana L. Price Department of Biological Sciences Salisbury University This document is a pictorial reference of Maryland Scarabaeoidea genera (and sometimes species) that was created to expedite the identification of Maryland Scarabs. Our current understanding of Maryland Scarabs comes from “An Annotated Checklist of the Scarabaeoidea (Coleoptera) of Maryland” (Staines 1984). Staines reported 266 species and subspecies using literature and review of several Maryland Museums. Dr. Price and her research students are currently conducting a bioinventory of Maryland Scarabs that will be used to create a “Taxonomic Guide to the Scarabaeoidea of Maryland”. This will include dichotomous keys to family and species based on historical reports and collections from all 23 counties in Maryland. This document should be cited as: Hagadorn, M.A. and D.L. Price. 2012. Quick Guide for the Identification of Maryland Scarabaeoidea. Salisbury University. Pp. 54. Questions regarding this document should be sent to: Dr. Dana L. Price - [email protected] **All pictures within are linked to their copyright holder. Table of Contents Families of Scarabaeoidea of Maryland……………………………………... 6 Geotrupidae……………………………………………………………………. 7 Subfamily Bolboceratinae……………………………………………… 7 Genus Bolbocerosoma………………………………………… 7 Genus Eucanthus………………………………………………. 7 Subfamily Geotrupinae………………………………………………… 8 Genus Geotrupes………………………………………………. 8 Genus Odonteus...……………………………………………… 9 Glaphyridae..............................................................................................
    [Show full text]
  • Insect Classification Standards 2020
    RECOMMENDED INSECT CLASSIFICATION FOR UGA ENTOMOLOGY CLASSES (2020) In an effort to standardize the hexapod classification systems being taught to our students by our faculty in multiple courses across three UGA campuses, I recommend that the Entomology Department adopts the basic system presented in the following textbook: Triplehorn, C.A. and N.F. Johnson. 2005. Borror and DeLong’s Introduction to the Study of Insects. 7th ed. Thomson Brooks/Cole, Belmont CA, 864 pp. This book was chosen for a variety of reasons. It is widely used in the U.S. as the textbook for Insect Taxonomy classes, including our class at UGA. It focuses on North American taxa. The authors were cautious, presenting changes only after they have been widely accepted by the taxonomic community. Below is an annotated summary of the T&J (2005) classification. Some of the more familiar taxa above the ordinal level are given in caps. Some of the more important and familiar suborders and families are indented and listed beneath each order. Note that this is neither an exhaustive nor representative list of suborders and families. It was provided simply to clarify which taxa are impacted by some of more important classification changes. Please consult T&J (2005) for information about taxa that are not listed below. Unfortunately, T&J (2005) is now badly outdated with respect to some significant classification changes. Therefore, in the classification standard provided below, some well corroborated and broadly accepted updates have been made to their classification scheme. Feel free to contact me if you have any questions about this classification.
    [Show full text]
  • A New, Well-Preserved Genus and Species of Fossil Glaphyridae (Coleoptera, Scarabaeoidea) from the Mesozoic Yixian Formation of Inner Mongolia, China
    A peer-reviewed open-access journal ZooKeysA 241: new, 67–75 well-preserved (2012) genus and species of fossil Glaphyridae (Coleoptera, Scarabaeoidea)... 67 doi: 10.3897/zookeys.241.3262 RESEARCH ARTICLE www.zookeys.org Launched to accelerate biodiversity research A new, well-preserved genus and species of fossil Glaphyridae (Coleoptera, Scarabaeoidea) from the Mesozoic Yixian Formation of Inner Mongolia, China Zhuo Yan1,†, Georgiy V. Nikolajev2,‡, Dong Ren1,§ 1 Key Laboratory of Insect Evolution and Environmental Changes, College of Life Sciences, Capital Normal University, Beijing 100048, P R. China 2 Al-Farabi Kazakh National University (Dept. of Biology and Bio- technology), al-Farabi Prospekt, 71, Almaty 050038, Kazakhstan † urn:lsid:zoobank.org:author:EAD752A4-96E0-4B7E-9F18-C4A5B6F0E863 ‡ urn:lsid:zoobank.org:author:DE41D0D0-A693-47D5-8A06-AD076CB74124 § urn:lsid:zoobank.org:author:D507ABBD-6BA6-43C8-A1D5-377409BD3049 Corresponding author: Dong Ren ([email protected]) Academic editor: A. Frolov | Received 20 April 2012 | Accepted 6 November 2012 | Published 14 November 2012 urn:lsid:zoobank.org:pub:794F0502-41FA-420F-B374-3FD16E68A021 Citation: Yan Z, Nikolajev VG, Ren D (2012) A new, well-preserved genus and species of fossil Glaphyridae (Coleoptera, Scarabaeoidea) from the Mesozoic Yixian Formation of Inner Mongolia, China. ZooKeys 241: 67–75. doi: 10.3897/ zookeys.241.3262 Abstract A new genus and species of fossil Glaphyridae, Cretohypna cristata gen. et sp. n., is described and illus- trated from the Mesozoic Yixian Formation. This new genus is characterized by the large body; large and strong mandibles; short labrum; elytra without longitudinal carina; and male meso- and possible metati- bia apically modified.
    [Show full text]
  • From Tethyan-Influenced Cretaceous Ambers
    Geoscience Frontiers 7 (2016) 695e706 HOSTED BY Contents lists available at ScienceDirect China University of Geosciences (Beijing) Geoscience Frontiers journal homepage: www.elsevier.com/locate/gsf Research paper Evolutionary and paleobiological implications of Coleoptera (Insecta) from Tethyan-influenced Cretaceous ambers David Peris a,*, Enrico Ruzzier b, Vincent Perrichot c, Xavier Delclòs a a Departament de Dinàmica de la Terra i de l’Oceà and Institut de Recerca de la Biodiversitat (IRBio), Facultat de Geologia, Universitat de Barcelona, Martí i Franques s/n, 08071 Barcelona, Spain b Department of Life Science, Natural History Museum, Cromwell Rd, SW7 5BD London, UK c UMR CNRS 6118 Géosciences & OSUR, Université de Rennes 1, 35042 Rennes cedex, France article info abstract Article history: The intense study of coleopteran inclusions from Spanish (Albian in age) and French (AlbianeSantonian Received 23 September 2015 in age) Cretaceous ambers, both of Laurasian origin, has revealed that the majority of samples belong to Received in revised form the Polyphaga suborder and, in contrast to the case of the compression fossils, only one family of 25 December 2015 Archostemata, one of Adephaga, and no Myxophaga suborders are represented. A total of 30 families Accepted 30 December 2015 from Spain and 16 families from France have been identified (with almost twice bioinclusions identified Available online 16 January 2016 in Spain than in France); 13 of these families have their most ancient representatives within these ambers. A similar study had previously only been performed on Lebanese ambers (Barremian in age and Keywords: Beetle Gondwanan in origin), recording 36 coleopteran families. Few lists of taxa were available for Myanmar Fossil (Burmese) amber (early Cenomanian in age and Laurasian in origin).
    [Show full text]
  • Insect Egg Size and Shape Evolve with Ecology but Not Developmental Rate Samuel H
    ARTICLE https://doi.org/10.1038/s41586-019-1302-4 Insect egg size and shape evolve with ecology but not developmental rate Samuel H. Church1,4*, Seth Donoughe1,3,4, Bruno A. S. de Medeiros1 & Cassandra G. Extavour1,2* Over the course of evolution, organism size has diversified markedly. Changes in size are thought to have occurred because of developmental, morphological and/or ecological pressures. To perform phylogenetic tests of the potential effects of these pressures, here we generated a dataset of more than ten thousand descriptions of insect eggs, and combined these with genetic and life-history datasets. We show that, across eight orders of magnitude of variation in egg volume, the relationship between size and shape itself evolves, such that previously predicted global patterns of scaling do not adequately explain the diversity in egg shapes. We show that egg size is not correlated with developmental rate and that, for many insects, egg size is not correlated with adult body size. Instead, we find that the evolution of parasitoidism and aquatic oviposition help to explain the diversification in the size and shape of insect eggs. Our study suggests that where eggs are laid, rather than universal allometric constants, underlies the evolution of insect egg size and shape. Size is a fundamental factor in many biological processes. The size of an 526 families and every currently described extant hexapod order24 organism may affect interactions both with other organisms and with (Fig. 1a and Supplementary Fig. 1). We combined this dataset with the environment1,2, it scales with features of morphology and physi- backbone hexapod phylogenies25,26 that we enriched to include taxa ology3, and larger animals often have higher fitness4.
    [Show full text]
  • Burmese Amber Taxa
    Burmese (Myanmar) amber taxa, on-line checklist v.2018.1 Andrew J. Ross 15/05/2018 Principal Curator of Palaeobiology Department of Natural Sciences National Museums Scotland Chambers St. Edinburgh EH1 1JF E-mail: [email protected] http://www.nms.ac.uk/collections-research/collections-departments/natural-sciences/palaeobiology/dr- andrew-ross/ This taxonomic list is based on Ross et al (2010) plus non-arthropod taxa and published papers up to the end of April 2018. It does not contain unpublished records or records from papers in press (including on- line proofs) or unsubstantiated on-line records. Often the final versions of papers were published on-line the year before they appeared in print, so the on-line published year is accepted and referred to accordingly. Note, the authorship of species does not necessarily correspond to the full authorship of papers where they were described. The latest high level classification is used where possible though in some cases conflicts were encountered, usually due to cladistic studies, so in these cases an older classification was adopted for convenience. The classification for Hexapoda follows Nicholson et al. (2015), plus subsequent papers. † denotes extinct orders and families. New additions or taxonomic changes to the previous list (v.2017.4) are marked in blue, corrections are marked in red. The list comprises 37 classes (or similar rank), 99 orders (or similar rank), 510 families, 713 genera and 916 species. This includes 8 classes, 64 orders, 467 families, 656 genera and 849 species of arthropods. 1 Some previously recorded families have since been synonymised or relegated to subfamily level- these are included in parentheses in the main list below.
    [Show full text]
  • New Scarabaeoidea (Coleoptera) from the Lower Cretaceous Yixian Formation, Western Liaoning Province, China: Elucidating the Systematics of Mesozoic Hybosoridae
    Cretaceous Research 86 (2018) 53e59 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes Short communication New Scarabaeoidea (Coleoptera) from the Lower Cretaceous Yixian Formation, western Liaoning Province, China: Elucidating the systematics of Mesozoic Hybosoridae ** Yuanyuan Lu a, b, 1, Ruie Nie a, 1, Chungkun Shih c, d, Dong Ren c, , Xingke Yang a, * Ming Bai a, a Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 92 Box, No. 1 Beichen West Road, Chaoyang District, Beijing, 100101, China b University of Chinese Academy of Sciences, Yuquan Road, Shijingshan District, Beijing, 100049, China c College of Life Sciences, Capital Normal University, Beijing, 100048, China d Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013-7012, USA article info abstract Article history: Two new genera are created to accommodate two new Hybosoridae (Coleoptera: Scarabaeoidea) species: Received 18 September 2017 Sinohybosorus cheni gen. et sp. n. and Sinochaetodus tridentatus gen. et sp. n.. The new species are Received in revised form described and illustrated based on two nearly complete fossil specimens from the Lower Cretaceous 2 January 2018 (Barremiane lower Aptian) Yixian Formation of western Liaoning Province, China. A key of described Accepted in revised form 6 February 2018 species of Hybosoridae from the Mesozoic and a catalog of extinct Hybosoridae are provided. Based on Available online 7 February 2018 the morphological characters, Fortishybosorus ericeusicus Yan, Bai et Ren, 2013 is transferred back to Fortishybosorus instead of Lithohypna in Glaphyridae. The discovery and description of these two new Keywords: fi Fossil taxa provide further evidence for the Jurassic and Cretaceous diversi cation of lineages of Hybosoridae.
    [Show full text]
  • Mckenna2009chap34.Pdf
    Beetles (Coleoptera) Duane D. McKenna* and Brian D. Farrell and Polyphaga (~315,000 species; checkered beetles, Department of Organismic and Evolutionary Biology, 26 Oxford click beetles, A reP ies, ladybird beetles, leaf beetles, long- Street, Harvard University, Cambridge, MA 02138, USA horn beetles, metallic wood-boring beetles, rove beetles, *To whom correspondence should be addressed scarabs, soldier beetles, weevils, and others) (2, 3). 7 e ([email protected]) most recent higher-level classiA cation for living beetles recognizes 16 superfamilies and 168 families (4, 5). Abstract Members of the Suborder Adephaga are largely preda- tors, Archostemata feed on decaying wood (larvae) and Beetles are placed in the insect Order Coleoptera (~350,000 pollen (adults), and Myxophaga are aquatic or semi- described species). Recent molecular phylogenetic stud- aquatic and feed on green and/or blue-green algae ( 6). ies defi ne two major groups: (i) the Suborders Myxophaga Polyphaga exhibit a diversity of habits, but most spe- and Archostemata, and (ii) the Suborders Adephaga and cies feed on plants or dead and decaying plant parts Polyphaga. The time of divergence of these groups has (1–3). 7 e earliest known fossil Archostemata are from been estimated with molecular clocks as ~285–266 million the late Permian (7), and the earliest unequivocal fossil years ago (Ma), with the Adephaga–Polyphaga split at ~277– Adephaga and Polyphaga are from the early Triassic (1). 266 Ma. A majority of the more than 160 beetle families Myxophaga are not known from the fossil record, but are estimated to have originated in the Jurassic (200–146 extinct possible relatives are known from the Permian Ma).
    [Show full text]
  • Fossil Perspectives on the Evolution of Insect Diversity
    FOSSIL PERSPECTIVES ON THE EVOLUTION OF INSECT DIVERSITY Thesis submitted by David B Nicholson For examination for the degree of PhD University of York Department of Biology November 2012 1 Abstract A key contribution of palaeontology has been the elucidation of macroevolutionary patterns and processes through deep time, with fossils providing the only direct temporal evidence of how life has responded to a variety of forces. Thus, palaeontology may provide important information on the extinction crisis facing the biosphere today, and its likely consequences. Hexapods (insects and close relatives) comprise over 50% of described species. Explaining why this group dominates terrestrial biodiversity is a major challenge. In this thesis, I present a new dataset of hexapod fossil family ranges compiled from published literature up to the end of 2009. Between four and five hundred families have been added to the hexapod fossil record since previous compilations were published in the early 1990s. Despite this, the broad pattern of described richness through time depicted remains similar, with described richness increasing steadily through geological history and a shift in dominant taxa after the Palaeozoic. However, after detrending, described richness is not well correlated with the earlier datasets, indicating significant changes in shorter term patterns. Corrections for rock record and sampling effort change some of the patterns seen. The time series produced identify several features of the fossil record of insects as likely artefacts, such as high Carboniferous richness, a Cretaceous plateau, and a late Eocene jump in richness. Other features seem more robust, such as a Permian rise and peak, high turnover at the end of the Permian, and a late-Jurassic rise.
    [Show full text]