HISTORICAL PERSPECTIVES

Copeia, 2005(3), pp. 693–700

DANIEL MORRIS COHEN

J. RICHARD DUNN AND THEODORE W. PIETSCH

ANIEL MORRIS COHEN, best known to marine research laboratory. The course was in- D ichthyologists for his taxonomic studies of tensive, meeting all day three times a week, and deep-sea salmoniform, gadiform, and ophidi- one that Dan himself would teach in 1962. He iform fishes over the past 45 years, was born on considered Bolin ‘‘a pretty good guy.’’ In 1951– 6 July 1930 in Chicago, Illinois (Anon., 2003a). 52, Dan took an advanced ichthyology course Dan’s father, Leonard U. Cohen, graduated from George Sprague Myers (1905–1985). from the University of Illinois. He spent one Dan then decided to study fishes. He spent year in law school at the University of Michigan the summer of 1952 with Professor Donald Eu- but, acceding to his father’s desire, followed the gene Wohlschlag (1918–) out on the tundra family business and became a furrier. Dan’s south of Point Barrow, Alaska, conducting field mother, Myrtle Gertz, also worked in the fur work on the age and growth of ciscoes and business as well as being a housewife who cared whitefishes, genus Coregonus, as part of a pro- for Dan and his sister Judith (1933–), an accom- gram on the population biology of whitefishes plished artist who now resides in Santa Rosa, funded by the Office of Naval Research. He California. The family relocated to California in took his M. A. degree (Cohen, 1954) and re- 1953 where Leonard went into the chicken-egg turned to Point Barrow the following year to business in the Carmel Valley. Dan’s father later assist Norman Joseph Wilimovsky (1925–1997) became a mortgage broker in Monterey. in his dissertation research on the ecology of Dan’s early interests in biology were encour- the Arctic Ocean ice pack. aged by grade school and high school teachers At this stage of his graduate education, Dan as well as by frequent trips to the Field Museum. was in a quandary searching for an appropriate He particularly enjoyed sport fishing and in his topic for his own doctoral dissertation. With an youth he accompanied his father on angling interest in trout, he thought of pursuing re- trips to lakes in northern Wisconsin, Michigan, search on salmonids, possibly studying under and Minnesota, and on weekends in Illinois. His Professor Paul Robert Needham (1919–1964) at curiosity about biology led him to Stanford Uni- the University of California, Berkeley. He was versity in Palo Alto, California, where relatives dissuaded from that venture by his fellow grad- had studied chemistry and English. Stanford uate students and his professors, so he next con- University (SU) was also a long way from Illi- sidered a study of the fishes of San Francisco nois, a fact that helped lure Dan to the West in Bay, but he learned from Myers that Earl Stan- 1948. In contrast to his father, Dan was encour- nard Herald (1914–1973) was conducting that aged to follow a career path of his own choos- work. He then consulted with Bolin, who sug- ing. As an undergraduate (Fig. 1) he considered gested tropical blennies, only to discover that a number of potential majors, including crea- Wilbert McLeod Chapman (1910–1970) was tive writing, pre-medical instruction, and geol- working on that group. At the time, Dan was a ogy, but he decided to specialize in biology, re- curatorial assistant in the Stanford fish collec- ceiving his B. A. in biological sciences in 1952. tion and he was ultimately led to the deep-sea In the spring of that year he spent three ‘‘fun’’ fishes from the Stanford-Crocker Expedition months on board the ORCA out of San Diego (Brittan, 1997) and Charles William Beebe’s collecting fishes and herps (amphibians and (1877–1962) Pacific fishes housed in the collec- reptiles) in the Gulf of California with fellow tion. As Dan put it, he then ‘‘got the call’’ to students, including James Erwin Bo¨hlke (1930– study deep-sea fishes and began his doctoral 1982) and Jay Mathers Savage (1928–), among work under Professor Myers, receiving his Ph.D. others. In 1953, he took ichthyology from Rolf degree in 1958. His dissertation on the system- Ling Bolin (1901–1973) at the Hopkins Marine atics of the deep-sea argentinoids (Cohen, Station in Pacific Grove, Stanford University’s 1957) ultimately led to the publication of 17 pa-

᭧ 2005 by the American Society of Ichthyologists and Herpetologists 694 COPEIA, 2005, NO. 3

Fig. 1. Daniel Morris Cohen as an undergraduate at Stanford University, 1950. Fig. 2. Dan and Anne in the Rose Garden of the White House, celebrating the 200th anniversary of the birth of James Smithson, 1965. pers and to a long and productive career in penhagen, and Charles Richard Robins (1928–), which Dan continued to pursue deep-sea fishes. University of Miami, Florida, now at the Univer- Dan considered Myers a bit difficult at times. sity of Kansas. He tended to leave his students very much Dan met Anne Carolyn Constant (1935–) at alone, but he was very loyal to them. He credits Stanford University when she was an undergrad- Myers for his encyclopedic knowledge of fish lit- uate biology major (Fig. 2). They became better erature as well as his portrayals of fish and herp acquainted when both were participants in a diversity and evolution. Dan acknowledges his field trip in 1954 to the San Pedro Martir Moun- fellow graduate students in the Stanford Natural tains of Baja California, Mexico, an expedition History Museum for much that he learned Dan termed ‘‘a lot of fun.’’ They were married about fishes, citing in particular Jim Bo¨hlke, in 1955. Anne received her B. A. from Stanford who Dan said ‘‘had phenomenal vibes for fish- in 1956, her M. S. (1972) at the University of es,’’ Norm Wilimovsky, who knew a ‘‘lot about Maryland, and her Ph.D. (1987) degree from cold-water fishes,’’ and Stanley Howard Weitz- George Washington University. She is an expert man (1927–), for his ‘‘knowledge of fish anat- on the taxonomy, systematics, and biology of omy.’’ Dan credits Elbert Halvor Ahlstrom crustaceans, particularly ostracodes (Anon., (1910–1979) and H. Geoffrey Moser (1938–) 2003b), on which she continues to publish. for passing along their insights into the world They have two daughters, Carolyn Annette of early life history stages of fishes. Others from Leech (1956–) of Naperville, Illinois, who is an whom he learned much were Rolf Bolin, Nor- engineering manager for Lucent Technologies man Bertram (Freddy) Marshall (1915–1996), and the mother of three children, and Cynthia of the British Museum (Natural History), Lon- Sarah Cohen (1959–), an assistant professor of don, as well as Dan’s two colleagues in the study biology at San Francisco State University and of ophidiiform fishes, Jørgen G. Nielsen the mother of one. (1934–), Zoological Museum, University of Co- Dan took a position in 1957 as assistant pro- HISTORICAL PERSPECTIVES 695

Fig. 3. Dan at his desk at the Systematics Laboratory, National Marine Fisheries Service, Smithsonian In- stitution, shortly before 1982. Courtesy of Bruce B. Collette. fessor of biology and curator of fishes at the to four full-time positions. Williams studied University of Florida, Gainesville, a position decapods and Pe´rez Farfante specialized in John Carmon Briggs (1920–) had recently va- shrimp. Dan’s own research focused on the tax- cated to teach anatomy in the medical school onomy of deep-sea salmoniform and ophidi- of that university. Dan found the southern racial iform fishes and on the general biology of deep- attitude of the era a distinct culture shock and sea fishes. He also spent considerable time dur- he did not consider his year at the University a ing those years with gadiform fishes, as urged pleasant one. In 1959, Dan accepted a position by Professor Henry Bryant Bigelow (1879–1967) as a systematic zoologist with the U.S. Bureau of of Harvard University, who said that studying Commercial Fisheries Ichthyological Laborato- deep-sea fishes would end with Dan ‘‘measuring ry housed in the U.S. National Museum mackerel in Marblehead’’ or even worse, (USNM), Washington, D. C. Giles Willis Mead, ‘‘counting smelt on the Miramichi.’’ Dan en- Jr. (1928–2003) was then the director of the lab- joyed the relative freedom of research that he oratory. Dan stayed with the Bureau in Wash- had with the Bureau and highly valued his con- ington for 23 years (Fig. 3), becoming director tact with the ichthyologists at the USNM, but his of the newly formed National Systematics Lab- laboratory was perpetually starved for funds. In oratory in 1960 after Mead moved to Harvard the early days it lacked secretarial help and, ac- University in Cambridge, Massachusetts. cording to Dan, the administrative arrangement After Mead left the laboratory, Dan worked with Bureau headquarters (unaffectionately alone with the aid of an illustrator, but during named by Mead the ‘‘goat barn’’) across town his tenure as director he oversaw the expansion was ‘‘a lousy way to run a ball team.’’ Later, in of the laboratory and hired first a malacologist logic evident only to the bureaucracy, Dan was Kenneth Jay Boss (1935–) and, then, in 1963, required to report to the Northeast Fisheries another ichthyologist, Bruce Baden Collette Center at Woods Hole, Massachusetts. (1934–). Under Dan’s leadership the laboratory Dan conducted field work aboard a number added two carcinologists, Austin Beatty Williams of research vessels, including the Bureau’s JOHN (1919–) and Isabel Pe´rez Farfante (1916–), to N. COBB, bottom trawling on the continental the staff bringing the professional complement slope off Washington State; the OREGON, work- 696 COPEIA, 2005, NO. 3 ing in the Gulf of Mexico and in the Caribbean; was nearly time for afternoon tea. Upon com- and the DELAWARE in the North Atlantic. He pletion of tea, there was little time to work be- participated also in a number of oceanic expe- fore quitting time and heading home. Such a ditions, including the International Indian routine schedule frustrated Dan, as well as oth- Ocean Expedition in 1964 on Cruise 6 of the er visiting scientists of the period. Dan, along United States Research Vessel ANTON BRUUN with others, wondered with envy at how Freddy (Cohen, 1972, 1986). This cruise worked from Marshall could be so productive while following Bombay, India, through the Arabian Sea and the ‘‘normal’’ Museum routine. south to Mauritius, and then south to the ‘‘roar- In addition to field work and his stint in Lon- ing 40s.’’ Other ichthyologists on that particular don, Dan traveled widely to visit fish collections segment of the expedition included, among in Denmark, Norway, Sweden, Germany, The others, Nielsen and Eric Bertelsen (1912–1993) Netherlands, Belgium, France, Austria, Russia, of the University of Copenhagen, Giles Mead Portugal, Italy, Monaco, South Africa, India, and Richard Lee Haedrich (1938–) of Harvard China, New Zealand, Australia, and Japan. In University, and Basil George Nafpaktitis (1929–), 1969–1970, Dan visited Panama as a member of then a student at Harvard University. In the fall the Committee on Ecological Research for an of 1973, he took part in a cruise of the German Interoceanic Canal of the National Academy of fisheries research vessel WALTHER HERWIG II Sciences. In 1978, he was part of an oceanog- across the Denmark Strait from Iceland to raphy delegation of the National Academy of Greenland and then to Portugal. This expedi- Sciences to the People’s Republic of China (Co- tion was led by Gerhard Krefft (1912–1993), hen, 1979), among other consulting assign- along with Alfred Post (1935–) and Matthias F. ments. Dan was also Editor-in-Chief of Part 6 of W. Stehmann (1943–) of the Sea Fisheries In- Fishes of the Western North Atlantic, published by stitute, Hamburg, in which Haedrich, Nielsen, the Sears Foundation for Marine Research of and Bertelsen also took part. The vessel fished Yale University (Cohen, 1973). a large commercial herring trawl (the so-called The year 1981 brought a severe funding cri- Rectangular Mid-water Trawl, with an 8 m2 sis–one of seemingly ‘‘endless crises,’’ according mouth opening) in mid-water at 2,000 meters to Dan–to the National Marine Fisheries Service and Dan termed this cruise a ‘‘most wonderful (name changed in 1980 from the Bureau of experience.’’ Dan also taught and collected fish- Commercial Fisheries) and the Systematics Lab- es aboard the Stanford research vessel TE VEGA oratory was slated for closure. Positions were of- in 1965 for three months, sailing from Singa- fered at other Service laboratories for the four pore to Guadalcanal (Cohen and Davis, 1969; scientific staff members. Dan was made a Senior Davis and Cohen, 1969). Along with Tomio Iwa- Fisheries Scientist and detailed to the Northwest moto (1939–), California Academy of Sciences, and Alaska Fisheries Center (now the Alaska San Francisco, and Mamoru Yabe (1952–), Hok- Fisheries Science Center) in Seattle. The fund- kaido University, Hakodate, Japan, he also col- ing crisis passed quickly, however, and the lab- lected fishes aboard the Japanese YAKUSHI– oratory remained open but, because of the dif- MARU NO. 21 in the Bering Sea in the summer ficulty entailed in transferring Dan to Seattle, it of 1979 (Yabe et al., 1981), among other expe- was not feasible to transfer him back to Wash- ditions. ington, D. C., where Bruce Collette then be- Dan was a visiting researcher at the British came laboratory director. Dan remained in Museum (Natural History) in London during Seattle for one year, but soon recognized that 1966–67, where he was introduced to the rather he was misplaced in a fisheries laboratory as he unusual customs practiced by a few prominent ‘‘was very much a museum person.’’ members of the zoology and paleontology de- A position became available at the Los An- partments at the Museum of that era. According geles County Museum of Natural History to Dan, and well remembered by other visitors (LACM) where, in 1982, Dan became Chief Cu- during that time, most of the ichthyology staff rator of Life Sciences and then Deputy Director routinely arrived at the Museum at about 10:00 for Research and Collections, holding the latter in the morning, just in time for morning tea. position until his retirement in 1995. During his After an extended tea break, and a brief at- tenure at the Museum, some hard times not- tempt at work, the pub opening time of 11:30 withstanding, he was able to recruit scientists for approached and off they went for lunch at a ten curatorial positions, six of which were new. nearby pub. Several pints would be consumed, Dan also continued his research on gadoid and along with such items as ‘‘bangers and mash,’’ ophidioid fishes during those years, authoring pork pie, and Scotch eggs. Upon their return or co-authoring over 20 papers on these fishes, to the Museum, after 2:30 pub closing time, it including two major contributions on which he HISTORICAL PERSPECTIVES 697 served as a co-author (Cohen et al., 1991; Niel- authored two or more papers include Jim sen et al., 1999). Bo¨hlke, William Pierce Davis (1939–), Richard Dan organized a very successful invitational Haedrich, Tomio Iwamoto, Robert Karl John- Workshop on Gadiform Systematics (WOGADS) son (1944–2000), Giles Mead, H. Geoffrey Mos- at the LACM in January 1986, which drew to- er, C. Richard Robins, Richard Heinrich Rosen- gether those ichthyologists whose major inter- blatt (1930–), and John P. Wourms (1937–). ests included the codfishes and their allies (Co- Dan is most proud of his work on the ophi- hen, 1989). A reception and dinner (at which dioids, in which he examined the historical re- the wine flowed freely) for the participants, sults of previous workers on these fishes in light held by Giles Mead around the swimming pool of recent collections. This work by Dan and col- at his home in Beverly Hills will never be for- leagues resulted in the recognition of two sub- gotten by those in attendance. After dinner, cer- orders in the order Ophidiiformes, based on tain renowned ichthyologists stripped to their their modes of reproduction, the oviparous ‘‘BVDs’’ and tumbled into the pool, marking Ophidioidei and the ovoviviparous Bythitoidei one of the true highlights of the symposium. (Cohen and Nielsen, 1978; Nielsen et al., 1999). While at LACM, Dan collected shore fishes in Dan joined the ASIH as a graduate student in the Galapagos Islands in 1984, along with Rob- the early 1950s, has attended many of the So- ert James Lavenberg (1937–), Curator of Fishes ciety’s annual meetings, and has presented at the Museum, and William Albert Bussing about a dozen papers at these gatherings. He (1933–) from the University of Costa Rica, on a was vice-president of the Society in 1969–70, trip that among others included John William president in 1985–86, and retains life member- Wright (1936–), who collected herps. Dan also ship on the Board of Governors. Dan has pub- participated in a cruise to the Arabian Sea and lished about 19 papers in Copeia and helped edit around Madagascar in 1988 aboard the Soviet a book published by the Society, in which he research vessel VITIAZ, which he remembers as contributed three articles in whole or in part memorable because of the ‘‘wonderful’’ deep- (Moser et al., 1984). sea collections that were made as well as be- Asked about what he remembers from the cause of the collegiality of the other ichthyolo- many ASIH meetings he has attended, Dan par- gists aboard (Fig. 4). The chief scientist on this ticularly recalls a paper presented by Bruce Col- expedition was Nikolas Vasilyevich Parin lette, while he was still a student, which was crit- (1932–) of Moscow and the ‘‘foreign’’ (i.e., non- icized by Carl Leavitt Hubbs (1894–1979). Col- Russian) scientists, besides Cohen and Bruce lette calmly and firmly defended his conclu- Collette, included Nielsen and Bertelsen of sions, a confident and mature response from a Denmark, Nigel Robert Merrett (1940–) of Eng- student to the then most highly respected ich- land, Michael Eric Anderson (1946–) of South thyologist. Dan values attending ASIH meetings Africa, John Richard Paxton (1938–) of Austra- where he can talk with like-minded people. He lia, and Kenneth J. Sulak (1946–), then of Can- is most honored by his election to the presiden- ada. cy of the ASIH and by his selection in 1997 for Having spent more than a decade looking at the Robert H. Gibbs, Jr. Memorial Award for the corpses of deep-sea fishes, Dan was deeply Excellence in Systematic Ichthyology, the Soci- interested in using submersibles to look at living ety’s highest honor for ichthyologists. ones. His first experience in such a vessel was Questioned about the major changes in ich- aboard the DEEPSTAR off New England in the thyology that he has observed during his career, summer of 1972. Subsequent dives in the ALVIN Dan responded that there has been a ‘‘sea- took place in Hudson Canyon in 1975, the Ba- change’’ over the years in the basic approach to hamas in January 1977, and the Galapagos rift systematics, from basing phylogenetic classifica- zone in December 1979. He was chief scientist tion largely on subjective criteria, or to use and co-chief scientist on the latter two expedi- Dan’s descriptor, ‘‘vibes,’’ to the present use of tions. cladistics, which purports to be a more objective Dan is the author or co-author of some 124 method. Further, Dan feels that hypotheses of scientific publications, generally divided into pa- relationships will have to be re-examined in pers on the taxonomy and biology of salmoni- light of molecular techniques. A surprise is that form, gadiform, and ophidiiform fishes as well apparently many fish species remain to be de- as on the general biology of deep-sea fishes. He scribed. About 200 species are currently being has forged a number of productive research al- named every year (Eschmeyer, 1998) and that liances with other ichthyologists; for example, figure seems to be holding steady. Forty-five co-authoring more than nine papers with years ago the figure was about 100 a year. Jørgen Nielsen. Others with whom he has co- Dan has long been known for his genuine 698 COPEIA, 2005, NO. 3

Fig. 4. Dan standing next to chief scientist Nik Parin aboard the Soviet research vessel VITIAZ in the Western Indian Ocean, 1988, with (left to right) Matthias Stehmann, Bruce Collette, Jørgen Nielsen, Erik Bertelsen, Eric Anderson, Nigel Merrett, and John Paxton. Courtesy of Jury A. Rudyakov. concern for students, always interested to know some initial hesitation and discussion, a meet- what they are doing and always doing his best ing was arranged and they shared a fine evening to help them along the way. Over the years, together. The two European ichthyologists sub- many a recent Ph.D. in ichthyology can remem- sequently became fast friends and colleagues for ber that it was Dan who first got them interested many years. in the ASIH. This deep concern has not waned Dan and Anne retired to Bodega Bay, Cali- even in retirement. During a visit to Seattle just fornia, where they tend to their garden. Dan is last year, Dan was invited for pizza and beer pleased to be a Fellow of the California Acade- along with a group of a dozen or so University my of Sciences as well as a Research Associate of Washington graduate students. All were sur- of the Academy’s Ichthyology Department, prised and deeply impressed that Dan made the which provides logistical support as he contin- effort to question each one about his or her ues to write papers. He serves NOAA as a mem- research in a friendly, non-threatening way that ber of the Sanctuary Advisory Committee for made them all feel that they were doing some- research for the Cordell Bank National Marine thing truly important. For them it was a brief Sanctuary. connection, but one that they will no doubt long remember. Dan is a fine raconteur, able to regale people Postscript.—Dan was interviewed by Ted Pietsch with stories of ichthyology and ichthyologists, and Jean Dunn on 27 March 2004 at the Cohen past and present. Many of these stories cannot residence in Bodega Bay, California. We thank reasonably be put to paper or are too long and Dan and Anne for their warm hospitality on involved to be repeated here, but one brief ex- that occasion. We were particularly entertained ample need suffice. In a small European coun- by numerous but, alas, unprintable stories told try there were only two active ichthyologists in by Dan (with tape recorder turned off) about the nation, but bureaucratic and political insti- ichthyology and ichthyologists. Dan’s profes- tutions kept the two men from meeting each sional papers will eventually be deposited in the other. While visiting in that country, Dan sug- Smithsonian Institution Archives, Washington, gested that the three meet for dinner. After D. C. HISTORICAL PERSPECTIVES 699

LITERATURE CITED ROBINS. 1999. Ophidiiform Fishes of the World (Order Ophidiiformes). FAO species catalogue, ANON. 2003a. Cohen, Daniel Morris, p. 312. In: Amer- Vol. 18. FAO, Rome. ican Men and Women of Science, 21st ed., vol. 2, YABE, M., D. M. COHEN,K.WAKABAYASHI, AND T. IWA- C–F. Gale, Farmington Hills, Michigan. MOTO. 1981. Fishes new to the eastern Bering Sea. ——— 2003b. Cohen, Anne Constant, p. 311. In: U.S. Fish. Bull. 79:353–356. American Men and Women of Science, 21st ed., vol. 2, C–F. Gale, Farmington Hills, Michigan. BRITTAN, M. R. 1997. The Stanford school of ichthy- SELECTED PUBLICATIONS OF ology: eighty years (1891–1970) from Jordan DANIEL MORRIS COHEN (CHRONOLOGICAL) (1851–1931) to Myers (1905–1985), p. 233–268. In: Collection Building in Ichthyology and Herpetolo- COHEN, D. M. 1960. New records of the opisthoproc- gy. T. W. Pietsch and W. D. Anderson, Jr. (eds.). tid genus Bathylychnops, with a notice of neoteny in Amer. Soc. Ichthyol. Herpetol., Spec. Publ. 3. the related genus Dolichopteryx. Copeia 1960:147– COHEN, D. M. 1954. Age and growth studies on two 149. species of whitefishes from Point Barrow, Alaska. ———. 1964. Bioluminescence in the Gulf of Mexico Stanf. Ichthyol. Bull. 4:168–187. anacanthine fish Steindachneria argentea. Ibid. 1964: ———. 1957. Contributions to a classification of the 406–409. fishes of the suborder Opisthoproctoidei. Unpubl. ———. 1964. A review of the ophidioid fish genus Ph.D. diss., Stanford University, Palo Alto, Califor- Oligopus with the description of a new species from nia. West Africa. Proc. U.S. Nat. Mus: 116:1–22. ———. 1972. Zoogeography of the fishes of the In- MEAD, G. W., E. BERTELSEN, AND D. M. COHEN. 1964. dian Ocean, p. 451–463. In: The Biology of the In- Reproduction among deep-sea fishes. Deep-sea Res. dian Ocean. B. Zeitzschel (ed.). Ecol. Stud. 3. 11:569–596. ———. (ED.). 1973. Fishes of the Western North At- COHEN, D. M. 1968. The cyclopterid genus Paraliparis, lantic, Pt. 6, Order Heteromi (Notacanthiformes), a senior synonym of Gymnolycodes and Eutelichthys, Suborder Cyprinodontoidei, Order Berycomorphi with the description of a new species from the Gulf (Beryciformes), Order Xenoberces (Stephanober- of Mexico. Copeia 1968:384–388. yciformes), Order Anacanthini (Gadiformes). ———, AND S. ATSAIDES. 1969. Additions to a revision Mem. Sears Found. Mar. Res. 1. of argentinine fishes. Fish. Bull. 68:13–36. ———. 1979. Visit to the People’s Republic of China. ———. 1970. How many recent fishes are there? Copeia 1979:379–380. Proc. Cal. Acad. Sci. 4:341–346. ———. 1986. Latitudinal variation in diversity and ———, AND J. G. NIELSEN. 1972. A review of the vivip- biomass in IKMT catches from the western Indian arous ophidioid fishes of the genus Saccogaster. Ocean, p. 54–59. In: Pelagic Biogeography. A. C. Proc. Biol. Soc. Wash. 85:445–468. Pierrot-Bults, S. van der Spoel, B. J. Zahuranec, and MARSHALL, N. B., AND D. M. COHEN. 1973. Order An- R. K. Johnson (eds.). UNESCO Tech. Pap. Mar. Sci. acanthini (Gadiformes), characters and synopsis of 49. families in Fishes of the Western North Atlantic. ———. ED. 1989. Papers on the systematics of gadi- Mem. Sears Found. Mar. Res. 1,6:479–495. form fishes. Nat. Hist. Mus. Los Angeles Co., Sci. COHEN, D. M. 1974. A review of the pelagic ophidioid Ser., 32. fish genus Brotulataenia with descriptions of two ———, AND W. P. DAVIS. 1969. Vertical orientation in new species. Zool. J. Linn. Soc. 55:119–149. a new gobioid fish from New Britain. Pac. Sci. 23: JOHNSON,R.K.,AND D. M. COHEN. 1974. Revision of 317–324. the chiasmodontid fish genera Dysalotus and Kali, ———, T. INADA,T.IWAMOTO, AND N. SCIALABBA. 1991 with descriptions of two new species. Arch. Fis- (1990). Gadiform Fishes of the World. FAO species chwiss. 25:13–46. catalogue, Vol. 10. FAO, Rome. COHEN,D.M.,AND J. P. WOURMS. 1976. Microbrotula ———, AND J. G. NIELSEN. 1978. Guide to the iden- randalli, a new viviparous ophidioid fish from Sa- tification of genera of the fish order Ophidiiformes moa and New Hebrides, whose embryos bear tro- with a tentative classification of the order. NOAA photaeniae. Proc. Biol. Soc. Wash. 89:81–98. Tech. Rept. NMFS Circ. 417. ———, AND J. L. RUSSO. 1979. Variation in the four- DAVIS,W.P.,AND D. M. COHEN. 1969. A gobiid fish beard rockling (Enchelyopus cimbrius), a North At- and palaemonid shrimp living on an antipatharian lantic gadid fish, with comments on the genera of sea whip in the tropical Pacific. Bull. Mar. Sci. 18: rocklings. Fish. Bull. 77:91–104. 749–761. ———, AND R. L. HAEDRICH. 1983. The fish fauna of ESCHMEYER, W. N. 1998. Catalog of Fishes. Vol. 1, In- the of the Galapagos thermal vent region. Deep-sea troductory Materials, Species of Fishes A–L. Cali- Res. 30:371–379. fornia Academy of Sciences, San Francisco, Califor- ———, R. H. ROSENBLATT, AND H. G. MOSER. 1990. nia. Biology and description of a bythitid fish from MOSER, H. G., W. J. RICHARDS,D.M.COHEN,M.P. deep-sea thermal vents in the tropical eastern Pa- FAHAY,A.W.KENDALL, AND S. L. RICHARDSON cific. Ibid. 37:267–283. (EDS.). 1984. Ontogeny and Systematics of Fishes. ———, D. A. HENSLEY, AND J. KIMMEL. 1991. The Amer. Soc. Ichthyol. Herpetol., Spec. Publ. 1. deepsea ophidiid genus Lamprogrammus, a senior NIELSEN, J. G., D. M. COHEN,D.F.MARKLE, AND C. R. synonym of Bassobythites, with notes on the synony- 700 COPEIA, 2005, NO. 3

my and distribution of L. brunswigi. Japan. J. Ichth- VERSITY OF WASHINGTON,CAMPUS BOX 355020, ol. 38:125–132. SEATTLE,WASHINGTON 98195–5020. E-mail: ( JRD) [email protected]; and (TWP) SCHOOL OF AQUATIC AND FISHERY SCIENCES,COL- [email protected]. Send reprint requests LEGE OF OCEAN AND FISHERY SCIENCES,UNI- to TWP.