Journ. Hattori Bot. Lab. No . 69: 1- 19 (Jan. 1991)

NEW HEPATIC RECORDS FROM NEW GUINEA

HEINAR STREIMANN 1

Reported here are new provincial records together with 12 species previously not known from New Guinea and 3 species which are new to , but were previously recorded from West Irian. These records were checked against the species published by Grolle and Piippo (l984a) and later publications as listed in the references. The records of these species (Table I) resulted from extensive field work during 1981 - 1983 by the author in various Papua New Guinea porvinces, especially the highland areas. Many of the new records for the provinces visited by the author are not surprising, as previous collectors tended to visit provinces with better access. Even now, most provinces are still under-collected, as can be seen from Table 1 in Grolle and Piippo (1984a) and with further collecting the range of many species will be extended. Names of common geographical features are abbreviated as is the name of the author (HS) in collection citations. Where there are more than three different localities from a province then only the number of localities is given. Habit and habitat data have been standardized (only a slight change from the label data) for better comparison. Distribution is based on the administration areas of the Indonesian province of West Irian and the provinces of Papua New Guinea as outlined by Grolle and Piippo (1984a: 2, 3). However, the Papuan Islands are treated as Milne Bay, the province to which they belong. Where the records are new to New Guinea the closest occurrence of that species to New Guinea is given, not its total range. Or R. Grolle kindly identified the collections to species level where possible (otherwise to generic level) and made available the Jungermannia determinations of

TABLE 1. New records for New Guinea and Papua New Guinea (* ).

* Anastrophyllum piligerum Lophoc()lea kurzii Chei/o/ejeunea paroica * Metzgeriopsis pusilla C. serpentina Radu/a decurrens Drepanolejeunea /evicornua R. subsimilis D. tenera * Rhaphidolejeunea bischlerae Lejeunea eifrigii Steno/ejeunea thallophora L. fleischeri Stictolejeunea bal[ourii Lepto/ejeunea e/liptica

1 Cryptogamic Herbarium, Australian National Botanic Gardens, Box 1777, Canberra City, ACT 2601, . 2 Journ. Hattori Bot. Lab. No. 69 199 I

Dr J. Vana. Collections identified only to generic level were sent to appropriate specialists. Drs. M . Mizutani and K. Yamada (NICH) determined most of the Lejeuneaceae and Radula respectively.

Acrolejeunea pycnoclada (Tay!.) Schiffn. Western Highlands: Kagamuga, 10 km E of , 1600 m, on exotic Eucalyptus at Forestry Station, HS 21749,30 Jun 1982 . Eastern Highla nds: Lapegu, 6km SW of , 1520m, on Taxodium trunk in Taxodillnl distichllnl Plantation, HS 18441, II Apr 1982; Kassam Pass, 19 km NE of Kainantu, 1600m, on exposed log in Lauraceae and Fagaceae dominated forest, HS 11524, 15 Jan 198 1. Southern Highlands: Nogoti, Tagari R., Tari-Komo Rd, 20km SW ofTari. 1250m, on tree trunk in lower montane forest, HS 32608, 16 Dec 1982. Distribution: Jayawijaya, Merauke, Morobc, Simbu, E. Highlands, Central, Milne Ba y. Acromastigum bifidum (Steph.) Evans West Sepik: Headwaters of Kong Kong Ck, 5 km S of . 250 m, lowland fo rest, A. Mundlla 286,24 Feb 1986; Amanab, 400m, R. Brown I, Jan 1985. Distribution: Sorong, Cenderawasih, Marokwari, E. Sepik, S. Highlands, Milne Bay. Anastrophyllum piligerum (Nees) Steph. Morobe: Mt Kaindi Rd, 5 km WNW ofWau, 1450 m. earth embankment in montane forest, HS 33409, 9 Jan 1983 . Distribution: Manokwari; new to Papua New Guinea; Indonesia. Anastrophyllum prionophyllum Hatt. Southern Highlands: Tari Gap, 26 km SE of Tari, 2260 m, on tree stump in remnant Phyllocladlls dominated forest , HS 32466, 15 Dec 1982. Distribution: Paniai, Jayawijaya, Morobe, Simbu, E. H ighlands, Milne Bay. Anomacaulis jiaccidus (Steph.) Grolle Southern Highlands: from 4 localities, 1800 2300 m, on tree trunks and rotting logs in poor and montane forests, HS 23290, 8 Sep 1982, HS 23398,9 Sep 1982, HS 23812,1 1 Sep 1982, HS 32753, 17 Dec 1982. Distribution: Jayawijaya, Morobe. Balantiopsis ciliaris Hatt. SSp. novoguineensis Hatt. Southern Highlands: Andawe R. , 6 km SE of lalibu, 1840 m, on root plate of fallen tree and bare ground in NotllOj'aXlIs and Podocarpaceae dominated fo rest, HS 26481 ,26484 , II Dec 1982. Distribution: Jayapura, W. Sepik, E. Sepik, Morobe. Caudalejeunea recurvislipula (Gott.) Schiffn. Morobe: , 10 m, on Spathodea campanlllata trunk in garden, HS 10788, 1I Jan 1981; Pouyu ViiI., 2km SE of Aseki, 1500m, on Sloanea streimannii trunk in advanced secondary forest. HS 12711,24 Jan 1981; Slate Ck and Gumi Ck Divide, 2100 m, on Pandanus leaf in No/ho/aXlIs dominated forest, HS 13992, 30 Jan 198 1. Milne Bay: Kaurai logging area, 9km N of Kulumadau, Woodlark I. , lOOm, on leaves in lowland forest , R. KlImei 45, 46, 47, 48, 8 Oct 1984; between Kropan ViiI. and Gidaloga, 3.5 km SW of Kulumadau, 5 m, on Pillosporunl stem in regrowth, R. KlImei 94 ( + Lejeunea sordida), 12 Oct 1984. Distribution: , Central. Caudalejeunea reniloba (Gott.) Steph. East Sepik: But ViiI. , on Aitape Rd, on fallen branch in rainforest, K . Naoni 38,6 Aug 1981. New Ireland: Sali ViII. , IOm, on tree trunk in secondary growth, W. Apelis 14,30 Dec 1982. Distribution: Jayapura. Jayawijaya, Madang, Morobe, S. Highlands, Central. Milne Ba y, New Britain, H. STREIMANN: New hepatic records from New Guinea 3

Manus. Ceratolejeunea belangeriana (Gott.) Steph. Gulf: Hepataewa, 5 km S of Kaintiba, 1000 m, on treelet (Trichospermum) stem in regrowth, HS 33900, 26 Jan 1983. Milne Bay: Kulumadau, Woodlark I., 60m, on teak (Tectona grandis) stem in six year old plantation, R. Kumei 7, Oct 1984. Distribution: Jayapura, Merauke, Morobe. Chandonanthus hirtellus (Web.) Mitt. Enga: Mape Ck, Mount Hagen - Wapenamanda Rd, 17 km SE of Wapenamanda, 2500 m, tree crown in poor montane forest, HS 21618,29 Jun 1982. Western Highlands: Jimi-Waghi Divide, 9 km N of Banz, 2400 m, on felled Nothofagus in regrowth, HS 22324, 7 Jul 1982; Mur Mur Pass, 7 km NE of Tambul, 2800m, disturbed montane forest. on fa llen tree branch, HS 21216,27 Jun 1982; Kagamuga, 10 km E of Mount Hagen, 1580m, trunk of exoIic Eucalyptus at Agricultural College, HS 20416,20417,20419, 20457, 20458, 22 Jun 1982. Southern Highlands: from 8 localities, 1820-4 110m, on ground, rocks, tree trunks and crowns in lower and montane forests, regrowth and subalpine grasslands, D. McVean 267366, 1967, HS 23292,8 Sep 1982, HS 23571 , 10 Sep 1982, HS 23725,23814,23892, 23894,23957, 23997, 11 Sep 1982, HS 24073.10 Sep 1982, HS 24081 . 24109, 24270, 24325,12 Sep 1982, HS 24396, 13 Sep 1982. HS 24576, 24637. 14 Sep 1982, HS 26447, 1I Dec 1982, HS 26816,26826. 12 Dec 1982. HS 32287, 14 Dec 1982, HS 32951.19 Dec 1982. Comments: Appears to be quite common at higher altitudes. Distribution: Manokwari, Jajawijaya, Morobe. E. Highlands, Northern. Chandonanthus pilifer Steph. Southern Highlands: Tari Gap. 23 km SE of Tari. 2740 m. on fallen NotllOfagus hranch in montane forest. HS 24458, 13 Sep 1982; Munia logging area. 2300m. on tree branches in Notho/agus and Podocarpaceae dominated forest, HS 23205,23318.8 Sep 1982, HS 23660,23662, 10 Sep 1982. Distrihution: Jayawijaya. Morobe. Central. Cheilolejeunea ceylanica (Gott.) Schust. & Kachroo Western Highlands: Baiyer R.- Ruti Rd, 42 km N of Mount Hagen. 1600 m, on leaf in lower montane forest, HS 22247, 5 Jul 1982. Distrihution: Manokwari, Morobe. Central, Milne Bay. Cheilolejeunea gigantea (Steph.) Schust. & Kachroo Morobe: Mt Kaindi. 5km W of Wau. 2360m, on small Elaeocarpus branch in Notho{agus and Cunoniaceae dominated regrowth. HS 17593 ( + C. imbricata, Lejeunea flava & Frullania), 13 Mar 1982. Distribution: Jayawijaya, Central. Cheilolejeunea imbricata (Nees) Hatt. Western Highlands: Tabibuga Rd, Jimi Valley, 14 km N of Banz, 1900 rn, on Casuarina trunk in regrowth, HS 20797,25 Jun 1982; Karpena Plantation, 28 km NE of Mount Hagen, 1650 m, on Eucalyptus stem in Eucalyptus plantation, HS 21917, I Jul 1982; Kagamuga, 10km E of Mount Hagen, 1580m, on branch of planted mango in Agricultural College, HS 20434,22 Jun 1982. Simbu: Yagle ViII., 4 km N of Mingende, 1800m, on Cordyline in advanced regrowth, A. Kawagle 15, 17 Apr 1982; Dirima ViII., 3km NW of Gumine, 1680m, on coffee stem in cultivated area, M . Toia 120,14 Dec 1982. Eastern Highlands: Daulo Pass, 18km WNW of Goroka. 2250m, on shrub in remnant forest, HS 18141 , 7 Apr 1982; Lapegu, 6 km SW ofGoroka, 1520 m, on Wendlandia stem in grasslands, HS 18402, I1 Apr 1982. Southern Highlands: Tambul-Mendi Rd, 18 km SW of Tambul, 2840 m, on shrub leaf in disturbed montane fo rest, HS 26758, 12 Dec 1982; Andawe R., 6 km SW of Ialibu, 1840 m, crown of dead tree in Nothofagus and Podocarpaceae dominated forest, HS 26500, 11 Dec 1982. Distribution: Manokwari, Morobe, Central. 4 Journ. Hattori Bot. Lab. No. 69 I 9 9 I

Cheilolejeunea intertexla (Lindenb.) Steph. Western HighJands: Karpena Plantation, 28 km NE of Mount Hagen, 1650m on Dodonaea stem in regrowth, HS 21831 , I Jul 1982 . Eastern Highlands: Koblugi, 12 km SE of Okapa, 1400 m, on treelet stem in lower montane forest, HS 26283,9 Dec 1982. New Britain: Wangowango, Willaumez Peninsula, 20km NNE of Talasea, on Cycas at seashore, S. Kolema 37, 8 Dec 1984. Distribution: Madang, Morobe, Central, Manus, N. Solomons. Cheilolejeunea acclusa (Herz.) Kodama & Kitag. Western Highlands: Baiyer R.- Ruti Rd, Jimi Valley, 42km N of Mount Hagen, 1600m, on a vine in lower montane forest, HS 22024, 22025, 2 Jul 1982. Eastern Highlands: near Hagobi ViiI. , 6 km SW of Lufa, 1850m, on leaves in montane forest , HS 18600,14 Apr 1982. Southern Highlands: Andawe R., 6km SE of Ialibu, 1840m, on Prunus trunk in NOlhofagus and Podocarpaceae dominated forest, HS 26564, I1 Dec 1982. Distribution: Morobe, Central. Cheilolejeunea paroica Mizut. Morobe: Oomsis, 21 km W of Lae, 50m, on planted Acacia auriculiformis in nursery area, HS & T. Umba 11575, 16 Jan 1981 ; Slate Ck, 15km W of Bulolo, 1200m, on Ficus branches in regrowth and grasslands, HS 14050, 30 Jan 1981; Korepa ViiI. , 7 km SE of Garaina, 750 m, on tree trunk in Anisoptera and Eucalyptus dominated forest, Kairo 309, I1 ov 1980. Distribution: new to New Guinea; Indonesia (Borneo, Celebes). Cheilolejeunea serpentina (Mitt.) Mizut. New Britain: Wangowango, Willaumez Peninsula, 20 km NN E of Talasea, sea level, on Calophyllum stem, S. Kolema 43, 8 Dec 1984. Distribution: new to New Guinea; Australia (Queensland). Cheilolejeunea tri/aria (Reinw. et al.) Mizut. Simbu: Makaua Mountain, 3 km NW of Gumine, 1800m, on ScheJJera stem in regrowth, M . Toia 16, II Dec 1982. Eastern Highlands: Kassam Pass, 19km NE of Kainantu, 1600m, on log in Lauraceae and Fagaceae dominated forest, HS & T. Umba 11 517, 15 Jan 1981. Gulf: Hepataewa, 6 km S of Kaintiba, 1200m, on rotting log in old garden site, HS 33891, 26 Jan 1983 . Distribution: Morobe, Central. Colura ari (Steph.) Steph. Western Highlands: Gernt R., Jimi Valley near Ruti, 54 km N of Mount Hagen, 600m, on shrub leaf in lowland forest, HS 21983, 2 Jul 1982. Distribution: Manokwari, Morobe, Central. Colura imperfecta Steph. Western Highlands: Gernt R. , Jimi Valley near Ruti, 54km N of Mount Hagen, 600m, on Syzygium leaves in lowland forest, HS 21892, 2 Jul 1982. Distribution: Manokwari , Morobe, Northern. Conoscyphus trapezioides (Sde Lac.) Schiffn. Southern Highlands: Mt Giluwe, 3450-3500 m, on ground and base of Papuacedrus [ = Libocedrus] in subalpine grasslands, HS 24104,24148, 12 Sep 1982. Distribution: Manokwari, W. Sepik, Morobe, W. Highlands. Denotarisia lingui/olia (De Not.) GroJle Jayawijaya: Mt Antares, Star Mountains, 300- 3300 m, on ground amongst heathy vegetation, B. van Zantefl 636, 678, Jul 1959. Eastern Highlands: Famin, Kainantu-Okapa Rd, 28 km SW of Kainantu, 2080 m, H. STREIMANN: New hepatic records from New Guinea 5 on Sloallea trunk in montane forest, HS 26839, 9 Dec 1982. Southern Highlands: from 4 localities, 1650-2400 m, on stumps, tree crowns and trunks in swampy grasslands and montane forest, HS 23995, 11 Sep 1982, HS 24707, 14 Sep 1982, HS 26028,5 Dec 1982, HS 26458, 26464,26570,26633, 11 Dec 1982, HS 26839,9 Dec 1982, HS 26998, 14 Dec 1982, HS 32517, 15 Dec 1982. Dis tribution: Sorong, W. Sepik, Morobe, W. Highlands, Central, Milne Bay.

Dicrano/ejeunea inoueana Mizut. & Piippo Southern Highlands: laro R., Onim, 14 km N W of lalibu, 2230 m, on Pal/danus prop root in old garden site, HS 23944, 11 Sep 1982; Andawe R., 6 km SE of lalibu, 1840 m, on Pandanus stem in Nothofagus and Podocarpaceae dominated forest, HS 26512, 11 Dec 1982. Distribution: Morobe, Central. Dicranolejeunea javanica Steph. Southern Highlands: Anggura R., Mendi-Mount Hagen Rd, 23 km SE of Mendi, 1800 m, on Casuarina stem in old garden si te, HS 32343, 14 Dec 1982. Di stribution: Jayawijaya, Morobe. Dip/asio/ejeunea cavifolia Steph. Western Highlands: Kagamuga, 10 km E of Mount Hagen, 1580 m, on planted mango in grounds of Agricultural College, HS 20448 (+ Lejeunea), 22 lun 1982. Distribution: Morobe. Drepanolejeunea intermedia Zwick. Western Highlands: Tabibuga Rd , limi Valley, 11 km N of Banz, 1900m, on Xylosma? leaf in dense regrowth, HS 20984,25 Jun 1982. Eastern Highlands: Koblugi, 12 km SE of Okapa, 1400m, on shrub leaf in lower montane forest, HS 26292, 9 Dec 1982; near Hagobi ViII ., 6 km SW of Lufa, 1850 m, on orchid leaves in montane forest. HS 18599 , 14 Apr 1982. Southern Highlands: Piribu Sawmill, Tari-Komo Rd, 3 km SW of Tari, 1650m, on trcelet leaf in Castanopsis dominated forest, HS 32530, 15 Dec 1982. Distribution: Manokwari, Jayapura, Morobe, Central. Drepanolejeunea levicornua Steph. Gulf: Hepataewa, 5 km S of Kaintiba, 1000 m, on Araceae leaf in poor lowland forest, HS 33806, 26 Jan 1983. Comments: Bonner (1965) reports New Guinea, but Piippo (pers. comm.) does not know on what it is based. Distribution: new to New Guinea; Indonesia (Ceram). Drepano/ejeunea spinoso-cornuta Steph. Morobe: Wagau-Malolo track, 26 km NE of Mumeng, 1600- 1700 m, on Kibara, Psychotria, Saurauia and fern leaves in montane forest, HS 19588, 19594, 19617, 19621 , 15 May 1982; Angabena Ridge, Aseki-Bulolo Rd, 1750m, on shrub leaves in montane forest, HS 25926,4 Dec 1982. Distribution: previous report (Grolle & Piippo 1984a) as 'Nova Guinea'. However, Bonner ( 1965) cites as type 'New Guinea, Moroka, s. d., L. Loria S.II., hb Levier 115, hb G-8325'. This is in Central Province. Drepanolejeunea tenera Goeb. Morobe: Logging area, 15 km E of Bulolo, 1500 m, on Pandanus leaf in Castanopsis and Engelhardtia dominated forest, HS 13327, 27 Jan 1981. Eastern Highlands: Waiopa, Aiyura-Omaura Rd, 13km SE of Kainantu, 1450m, on shrub leaf in disturbed Castanopsis forest, HS 26233, 8 Dec 1982. Dis tribution: new to New Guinea; Indonesia (Sumatra). Drepanolejeunea vesiculosa (Mitt.) Steph. Eastern Highlands: track to Mt Michael, 1800m, on Castanopsis trunk in advanced regrowth, HS 6 Journ. Hattori Bot. Lab. No. 69 I 9 9 I

18528, 12 Apr 1982. Distribution: Morobe, Simbu, Central. Drepanolejeunea yulensis Steph. Southern Highlands: Mt Gilu we, 3450 m, on shrub stem in subalpine grasslands, ifS 24100, 12 Sep 1982. Distribution: previous report (Grolle & Piippo 1984a) as 'Nova Guinea' as given by Stephani 1913. However, Bonner (1965) cites the type as 'New Guinea, Mt. Yule, 1891, Roy. Geogr. Soc. Australia Exped., in hb G'. This is in Central Province. Dumortiera hirsuta (Sw.) Nees sensu amp\o East Sepik: Kairiru I. , on ground, stones and base of tree, W. Borrell16, J. Klei I, 2, 3, 4, Jul & Sep 1979. Distribution: Sorong, Manokwari, Paniai, Madang, Morobe, Enga, W. Highlands, Simbu, S. Highlands, Central. Northern. Gottschelia schizopleura (Spruce) Grolle Western Highlands: Rd from Banz to Jimi-Waghi Divide, 2200 m, beside road in regrowth, ifS 20933, 20934,20934, 25 Jun 1982. Eastern Highlands: Daulo Pass, 2200m, exposed road cutting in regrowth and grasslands. HS 20412,21 Jun 1982. Southern Highlands: Tari Gap, 27- 29 km SE-ESE of Ta ri , 2560- 2660 m, on ground in grasslands, HS 32354.32356. 15 Dec 1982. HS 32774,32775, 17 Dec 1982; Tambul-Mendi Rd, 25 km SW of Tambul, 2400 m, road cutting in regrowth. ifS 26832. 12 Dec 1982; Mt Giluwe, 3450-4100m, on rock and ground in subalpine grasslands, P. Mumna 2, HS 241l7. 24128, 24207, 24208, 24329, 12 Scp 1982. Distribution: Manokwari, Jayawijaya, Morobe, Enga, Simbu, Central. Harpalejeunea constricta Grolle Southern Highlands: Mt. Giluwe, 3450 m. on shrub stem in subalpine grasslands, ifS 24 J08, 12 Sep 1982. Distribution: Manokwari, Morobe, Simbu, Central, Milne Bay. Heteroscyphus argutus (Reinw. et at. ) Schiffn. Western Highlands: Tumbang ViiI., 7km SE of Minj. 1800m, tree roots in remnant Araucaria and NOlhofagus dominated forest, HS 21346, 28 Jun 1982. Central: from 5 localities, 400--600 m, on rocks, tree trunks and buttresses in advanced regrowth, Dipterocarpaceae dominated and lowland forests, HS & A. Vinas 14442,4 Fcb 1981, HS & A. Vinas 14539, 5 Feb 1981, HS & K. Naoni 14721, 9 Feb 1981, HS & K. Naoni 15245, 15247,12 Feb 1981. HS & K. Naoni 16173,16180,14 Feb 1982. New Britain: Mopili-Galai No. I, 30 km SE of . 60 m, on log in lowland forest, S. Kolema 75, 25 Jan 1985. Distribution: Sorong. Manokwari, Jayawijaya, W. Sepik, Madang, Morobe, Western, New Ireland. Heteroscyphus aselliformis (Reinw. et al.) Schiffn. Southern Highlands: Tari Gap, 23- 27km SE of Tari, 2660--2740 m, on rotting wood, humus rich ground and shrub stem on edge of grasslands and subalpine forest and in montane forest , HS 24494, 13 Scp 1982, HS 32398, 32403, 32407, 15 Dec 1982. Distribution: Jayawijaya. Madang, Morobe, W. Highlands. Simbu, Milne Bay. Heteroscyphus coalitus (Hook.) Schiffn. Western Highlands: M ur M ur Pass, 7 km NE ofTambul, 2800 m, on rotting log in disturbed Cryplocarya and Elaeocarpus dominated forest, HS 22402,6 Jul 1982. Simbu: Kotdame, 8 km SE of Mt Wilhelm, 2650 m, on tree trunk in secondary forest, A. Mundua 230, 15 Dec 1982. Junction Keglsugl & Bundi Rds, 9 km SE of Mt Wilhelm, 2350 m, on shaded rock in grasslands, HS 18562, 13 Apr 1982; Makaua Mountain, 3 km NW of Gumine, 1860m, on ground in regrowth, M. Toia 92, 13 Dec 1982. Southern Highlands: Iaro R., Onim, 14km NNW of Ialibu, 2230m, on boulder in poor montane forest, HS 23771, 11 Sep 1982; Lai R., Mendi-Tari Rd, 1500m, on rotting log in advanced secondary forest, HS 22175,22191, 3 Jul 1982; Lama Sawmill,S km SE of lalibu, 2000 m, on rotting log in Notho(agus and Podocarpaceae dominated forest, HS H. STREIMANN: New hepatic records from New Guinea 7

24701, 14 Sep 1982. Distribution: Manokwari, layapura, layawijaya, Morobe, Enga, E. Highlands, Central, Northern, Milne Bay. lackiella javanica Schiffn, Enga: Mape Ck, Mount Hagen - Rd, 17 km SE of Wapenamanda, 2500 m, on shaded ground in disturbed area beside road, HS 22120, 29 lun 1982. Western Highlands: Tabibuga Rd, limi Valley, 14 km N of Banz, 1900 m, on road cutting in regrowth, HS 2076 7, 20800, 25 lun 1982; Mondomil Mission, 5 km SE of Minj, 1700 m, on roadside rock in poor primary forest, HS 21469, 28 lun 1982. Simbu: Yagle ViiI. , 1800m, shaded ground in regrowth forest, A. Kawagle 7,17 Apr 1982. Southern Highlands: laro R. , Onim, 14 km NNW of Ialibu, 2230 m, on rotting log in poor montane forest, HS 23722, II Sep 1982; Lama Sawmill, 6 km SE of lalibu, 1850 m, on track cutting in Cas/anopsis forest, HS 26600, I I Dec 1982. Central: Road to K B Sawmill, 9km SW of Sogeri, 580m, on ground in regrowth, HS 16678, 16 Feb 1981. Distribution: Manokwari, layawijaya, W. Sepik, E. Sepik, Morobe, E. Highlands, Northern. lamesoniellaflexicaulis (Nees) Schiffn. Southern Highlands: Piribu Sawmill, Tari-Komo Rd, 3 km SW of Tari, 1650 m, on rotting log in Cas/anopsis dominated forest, HS 32510, 15 Dcc 1982. Distribution: Morobe, W. Highlands, Simbu. lubula hutchinsiae (Hook,) Dum, ssp, javanica (Steph,) Verd. Eastern Highlands: Daulo Pass, 18 km WNW of Goroka, 2700 m. on Pandanus stem in advanced secondary forest, HS 17980, 7 Apr 1982. Southern Highlands: Munia logging area, 2300 m, on Pandanus prop root in No/hoj'agus and Podocarpaceae dominated forest , HS 23221 , 8 Sep 1982. Distribution: layapura, Morobe, Milne Ba y, New Ireland. lungermannia ariadne Tay!. Morobe: Aseki-Mdamna track, I km SW of Ase ki . 1350 m, on ground in advanced secondary vegetation, HS 12498, 23 l an 1982. Di stribution: W. Sepik, Central, Milne Bay. lungermannia hasskarliana (Nees) Mitt. Southern Highlands: Piambil Vi iI. , 20km ESE of Mt Giluwe, 2100m, on riverbank in regrowth, R. Yalo 2, 22 Dec 1982. Distribution: Morobe, W. Highlands, E. Highlands, Central. Lejeunea albescens (Steph,) Mizut. Eastern Highlands: track to Mt Michael, 5 km SW of Lufa, 2000 m, on ground in disturbed montanc forest, HS 18468, 12 Apr 1982. Southern Highlands: Tari Gap, Mendi-Tari Rd, 2660m, on ground in Phyllocladus dominated forest. HS 32457, 15 Dec 1982. Distribution: Madang, Morobe, Central. Lejeunea catanduana (Steph,) Miller et al. East Sepik: Victoria Bay. Kairiru I., on rain tree, J. Klei 33. 25 Jul 1979. Distribution: Morobe, E. Highlands, Central. Lejeunea cucullata (Reinw. et al.) Nees Eastern Highlands: Daulo Pass, 18 km WNW of Goroka, 2700 m, on small tree trunk in advanced secondary forest, HS 17956, 7 Apr 1982. Distribution: Morobe, Central. 8 Journ. Hattori Bot. Lab. No. 69 1 99 I

Lejeunea cuculliflora (Steph.) Mizut. Morobe: Wagau-Malolo track, 26 km NE of Mumeng, 1600m, on Bubhia [ = Zygogynum] leaves in montane forest, HS 19618, 15 May 1982. Gulf: Hepataewa, 5 km S of Kaintiba, 1000 m, on fallen tree branch in poor lowland forest, HS 33764, 26 Jan 1983. Distribution: Central. Lejeunea discreta Lindenb. Western Highlands: Kagamuga Forestry Station, 9 km E of Mount Hagen, 1560-1600 m, on tree stump and trunks in plantation of local and exotic trees, HS 21507, 28 Jun 1982, HS 21702, 21704, 21 722, 21 730, 21 746, 21 753, 21756, 21 757, 30 Jun 1982; Kagamuga, 10km E of Mount Hagen, 1600 m, side of ditch in Agricultural College, HS 21964,2 Jul 1982; Tumbang ViiI. , 7km SE of Minj, 1800m, on tree trunk in regrowth, HS 21410,28 Jun 1982 . Southern Highlands: Margarima-Tari Rd , 3 km N of Margarima, 1820 m, on Caslanopsis trunk in disturbed Caslanopsis forest, HS 24356, 13 Sep 1982; Onim Forestry Station, 14 km NNW of lalibu, 2250 m, on tree stump, base of Eucalyptus and Pandanus trunk in grasslands planted with exotic trees and in disturbed montane forest, HS 23549, 24663, 24670, Sep 1982. Distribution: Morobe, Central, Northern. Lejeunea eifrigii Mizut. Morobe: Mt Kaindi Rd, 5 km WNW of Wau, 1450 m, on roadside drain in disturbed montane forest, HS 33336, 9 Jan 1983. Western Highlands: Kagamuga Forestry Station, 9km E of Mount Hagen, 1600m, on Grevillea robusta trunk in plantation, HS 21 737; 30 Jun 1982. Southern Highlands: Anggura R., Mendi-Mount Hagen Rd , 23 km SE of Mendi, 1800 m, on CaslIarina stem in regrowth, HS 32344, 14 Dcc 1982. Distribution: new to New Guinea; Indonesia (Sumatra, Java). Lejeunea exilis (Reinw. et al.) Grolle Eastern Highlands: Wopeia, Aiyura-Omara Rd, 10 km SE of Aiyura, 1550 m, on small stump in Castanopsis forest, HS 18331, 10 Apr 1982. Distribution: Morobe, Simbu, Central. Lejeunea jiava (Sw.) Nees Southern Highlands: Kagamuga Forestry Station, 9 km E of Mount Hagen, 1600 m, on Gre villea robusla trunk and base of Melaleuca in plantation, HS 21 761 , 21685, 30 Jun 1982. Eastern Highlands: near Hagobi ViiI., 6km SW of Lufa, 1850m, on shrub in montane forest, HS 18717, 14 Apr 1982. Southern Highlands: Anggura R., Mendi-Mount Hagen Rd, 23 km SE of Mendi, 1800 m, on CaslIarina stem in regrowth, HS 32345,14 Dcc 1982. Gulf: Bema-Kaintiba Rd, 3 & 4km N of Kaintiba, 1170- 1180m, on Cordy line stem in regrowth, HS 33674, 33733,25 Jan 1983. Distribution: Jayawijaya, Morobe, Central. Lejeunea jieischeri (Steph.) Mizut. Morobe: Ekuti Divide, Bulolo-Aseki Rd, 30 km WSW of Bulolo, 2100 m, on shrub (Fagraea) leaves in montane forest, HS 20240, 16 Jun 1982 . Distribution: new to New Guinea; Indonesia (Java). Lejeunea herzogii Mizut. Western Highlands: Gernt R., Jimi Valley near Ruti, 54 km N of Mount Hagen, 600 m, on palm leaf in lowland forest , HS 21977 ( + Cololejellnea), 2 Jul 1982. Distribution: Morobe, Central. Lejeunea kinabalensis Mizut. Western Highlands: Tabibuga Rd, Jimi Valley, II km N of Banz, 1900 m, on treelet branch in dense H . STREIMANN: New hepatic records from New Guinea 9 regrowth, HS 20983,25 Jun 1982. Eastern Highlands: Daulo Pass, 18km WNW of Goroka, 2700m, on humus rich ground in advanced secondary forest, HS 17972, 7 Apr 1982. Distribution: Morobe. Lejeunea leratii (Steph.) Mizut. Western Highlands: Kagamuga, 9 km E of Mount Hagen, 1600 m, base of Melaleuca in Forestry plantation, HS 21674,21699,30 Jun 1982; Kagamuga, 10 km E of Mount Hagen, 1580 m, on mango branch in Agricultural college, HS 20436, 22 Jun 1982; Tumbang ViII., 7 km SE of Minj, 1800 m, on Casuarina trunk in regrowth, HS 21394,28 June 1982. Eastern Highlands: Lapegu, 6km SW of Goroka, 1520m, on Taxodium dislichum trunk in Taxodium plantation, HS 18415, II Apr 1982; track to Mt Michael, 4km SW of Lufa, 1900 m, on shrub stem in disturbed montane forest, HS 18773, 12 Apr 1982; Entabindi-Pusarasa area, 2 km NNW of Okapa, 1720 m, on shrub in Castanopsis forest, HS 26309, 9 Dec 1982. Distribution: Jayawijaya, Morobe, Simbu, Central.

Lejeunea lumbricoides (Nees) Nees Enga: Mape Ck, Mount Hagen-Wapenamanda Rd, 17 km SE of Wapenamanda, 2500 m, on ground in regrowth, HS 21509, 29 Jun 1982. Western Highlands: Mondomil Mission, 5 km SE of Minj, 1700 m, on roadside rockface in poor primary forest, HS 21493,28 Jun 1982. Simbu: Makaua Mountain, 3 km NW ofGumine, 1760-1860m, on rock and tree branch in grasslands and secondary growth, M . Toia 57A. 84, 13 Dec 1982. Southern Highlands: Tari Gap, Tari-Mendi Rd, 27 km SE of Tari, 2660 m, on Vaccinium at edge of grasslands and subalpine forest, HS 32385, 15 Dec 1982; Mt Giluwe, 3500m, on creek bank in sUbalpine grasslands, HS 24138, 12 Sep 1982. Distribution: Jayawijaya, Morobe, Central. Lejeunea microloba Tay!. Western Highlands: Baiyer R.-Ruti Rd, Jimi Valley, 42 km N of Mount Hagen, 1500 m, on tree buttress and trunk in lower montane forest, HS 21998,22006,2 Jul 1982. Southern Highlands: Lama Sawmi ll , 6 km SE of lalibu, 1850 m, on old tree stump in disturbed Caslanopsis forest, HS 26430, II Dec 1982. Distribution: Morobe, Centra l, Milne Bay. Lejeunea microstipula Steph. Gulf: Hepataewa, 5 km S of Kaintiba, 1000 m, on fallen tree branch in poor lowland forest, HS 33768, 26 Jan 1983. Distribution: Morobe. Lejeunea sordida (Nees) Nees East Sepik: Victoria Bay, Kairiru I., on coconut palm in village, J. Clay 8,27 Jul 1979. Simbu: Dirima Vill., 3 km NW of Gumine, 1680 m, on coffee stem in cultivated area, M . Toia J22, 14 Dec 1982. Southern Highlands: Det Mission, 15km SSW of Mendi, 1600m, on a vine in montane forest, HS 32919,18 Dec 1982; Lai R., Mendi-Tari Rd, 15km SSE of Mendi, 1500m, on treelet stem in advanced secondary forest, HS 22210,3 Jul 1982. Gulf: Bema-Kaintiba Rd, 3 & 4km N of Kaintiba, 1170-1200m, on Pandanus and Cordyline stems in regrowth, HS 33661,33685,25 Jan 1982. Milne Bay: Kulumadau, Woodlark I., 60m, on teak stem in plantation, R. Kumei 8, 15 (+ Lopholejeunea eulophal, 16,22,4 Oct. 1984; between Kropan ViII. and Gidaloga, 3.5 km SW of Kulumadau, Woodlark, I., 5 m, on Pittosporum stem in disturbed forest, R. Kumei 94 ( + Caudalejeunea recurvislipulal, 12 Oct 1984. New Ireland: Sali ViII. , 46 km SW of , 10m, on tree trunk in secondary growth, W. Apelis 3, 5, 6, 29 Dec 1982. Distribution: Sorong, Morobe, Central, Manus. Lejeunea umbilicata (Nees) Gott. et at. East Sepik: Kairiru I., on tree trunk, W. Borrell 33, Sep 1979. Distribution: Morobe. 10 Journ , Hattori Bot. Lab, No, 69 I 9 9 I

Lepicolea rara (Steph.) Grolle Simbu: Lake Aunde, 5 & 6 km SE of Mt Wilhelm, 3520- 3700 m, on ground and branches in subalpine woodland, D, McVean 26695, 266136A, 266277 (+ Lepicolea ocl!roleuca), May 1966; Makaua Mountain, 3 km NW of Gumine, 1860 m, on humus rich ground in NOlhofagus fo rest, M . Toia 67B, 13 Dec 1982, Southern Highlands: from 4 localities, 1740- 3600 m, on ground, tree stump, shrub and tree crown in suba lpine grasslands, regrowth and montane forest, HS 24218, 12 Sep 1982, HS 26904, 13 Dec 1982, HS 26965, 14 Dec 1982. HS 32448, 15 Dec 1982. HS 32765, 17 Dec 1982. Distribution: Madang, Mo robe, W , Highl a nds, E. Highlands, Central, Northern, Milne Bay, Lepidolejeunea graeffei (Jack & Steph.) Schust. Milne Ba y: Kaurai logging area, 9 km N of Kulumada u, Woodlark I., 100 m, on Dysoxylum bra nches in lowland forest, R, Kumei 65,66, 9 Oct 1984. Distribution: Morobe. New Britain, New Ireland, Lepidozia ferdinandi-muelleri Steph. Eastern Highlands: track to M t Michael. 4 km SW of Lufa, 1900 m, on rotting log in disturbed montane forest. HS 18777, 12 Apr 1982, Comments: a very common hepatic. Distribution: Manokwa ri , Jayapura, Jayawijaya, W, Scpik , Morobe, W, Highlands, S, Highlands, Central. Northern, New Ireland , Lepidozia laceri/olia Steph. Morobe: Ekuti Divide, 33 km WSW of Bu lolo, 2260 m, on rotting log in Lauraceae-Myrtaceae dominated forest, HS 20065, 16 Jun 1982. Distribution: Mano kwari. W. Sepik, E, Sepik , Lepidozia quadridens Nees Morobe: Kaisinik ViiI., 7 km SE of Wa u, 1100 m, on rocks in st ream in remnant Caslallopsis forest, HS 13752, 29 Jan 198 1. Distribution: E. Highla nds. Lepidozia wallichiana Gott. Western Highlands: Mur Mur Pass, 7 km NE of Tambul. 2800m. on ground in disturbed Cryplocarya and Elaeocarpus dominated fo rest, HS 21205 (+ Zoopsis liukiuensis) , Southern Hi ghlands: Mt Bosavi, 760 m, on log in rainforest, G, 0 'Hara 37, May 1976. Distribution: Manokwari , Jayapura, Jayawijaya, Merauke, W, Sepik, E, Sepik, Morobe. Central, M ilne Bay, Leplolejeunea elliptica (Lehm. & Lindenb.) Schiffn. Central: Imila R" Kwiki la-Kupia no Rd, 26 km from K upiano, 40 m, on vine in monsoon forest, HS & K. Naoni 16089, 13 Feb 1981. Distribution: new to New Guinea; Solomon Is, Leplolejeunea maculata (Mitt,) Schiffn, Milne Ba y: Kaurai logging a rea. 9km N of Kulumadau, Woodlark I., lOOm . on climber (Araceae) leaf in lowland forest, R . Kumei 82.85.9 Qct 1984, Distri bution: Manokwari. Madang. Morobe, Central. Lophocolea kurzii Sde Lac. Morobe: Middle Ck·Watut R, Divide, 8 km NW of Bulolo, 900 m. on rotting log in dense regrowth, HS 14277, 2 Feb 198 1. Central: Hombrom Bluff Rd, 26 km NE of , 650 m, on tree buttress, lower montane forest , HS & K. Naoni 15014, 10 Feb 1981. H. STREIMANN: New hepatic records from New Guinea 11

Distribution: new to New Guinea; Indonesia (Java), Australia. Lopho/ejeunea dent(folia Mizut. & Piippo Southern Highlands: Lama Sawmill, 6 km SE of lalibu, 1850 m, on Pandanlls and Pillosporllm stems in Castallopsis forest, HS 26673, 26684, I I Dec 1982. Distribution: Morobc. Lopholejeunea eulopha (Tay!.) Schiffn. Western Highlands: Nebilyer R. , 28 km WNW of Mount Hagen, 2760 m, on Palldanlls stem in logged montane forest, HS 22261, 6 Jul 1982; Karpena Plantation, 28km NE of Mount Hagen, 1650 m, on Welldlalldia stem in regrowth, HS 21872, I Jul 1982; Kagamuga Forestry Station, 9 km E of Mount Hagcn, 1600m, on Gre villea robusta trunk in plantation, HS 21735, 30 Jun 1982. Southern Highlands: Ta ri Ga p, 23km SE of Tari, 2740m. on shrub branch in montane forest, HS 24444, 13 Scp 1982; Andawe R ., 6km SE of lalibu, 1840 m, on tree trunk in Nothoj'af!,us and Podoca rpaceae dominated forest, HS 26508, II Dec 1982. Milne Bay: Kulumadau, Woodlark I., 60 m, on teak stem in plantation, R. Kumei 15 (+ Ll!jeunea sordida), 4 Oct 1984. Di stribution: Manokwari, Cenderawasih, Jayawijaya. Morobe, Simbu, Central, New Britain. Lopholejeunea herzogiana Verd. Western Hi ghlands: Tabibuga Rd, Jimi Valley, 14 km N of Ba nz, 1900 m, on Cordyline stem in regrowth, HS 20798.25 Jun 1982. Distribution: Morobe. Lopholejeunea subfusca ( ees) Schiffn. East Sepik: Victoria Ba y, Kairiru I., on coconut palm, J. Klei 35,25 Jul 1979. Western Highlands: Kagamuga, 9km E of Mount Hagen, 1600m, base of Melall!uca in plantation, HS 21689,21693, 21703, 21705, 30 Jun 1982; Kagamuga, 10 km E of Mount Hagen. 1580 m, base of mango in Agricultural College, HS 20425, 22 Jun 1982. Eastern Highlands: Lapegu, 6km SW of Goroka, 1520m, on Taxodillm trunk in Taxodilllll distichum plantation, HS 18411, II Apr 1982. Southern Highlands: laro R., Onim, 14 km NNW of lalibu, 2230 m. on dead tree trunk in regrowth, HS 23982. II Sep In2. Gulf: Hepataewa. 5 km S of Kaintiba, 1000m, on trcelet (TricllOspermllm) stem in regrowth, HS 33902. 26 Jan 1983. Milne Bay; Kulumadau, Woodlark I. , 60 m, on teak stem in plantation, R. Kumei 17 (+ Caudalejeullea reniloha), 4 Oct 1984. Comments: very common in Morobe a nd Central Provinces. Distribution: Sorong, Manokwari. Jayawijaya, Moro be, Central, ew Britain, New Ireland. Marsupella subinlegra S. Am. Southern Highlands: MI. Giluwe, 4000 m, on rock in subalpine grasslands HS 24307, 12 Sep 1982. Distribution: Simbu. Masligo/ejeunea humilis (Gott.) Schiffn. Gulf: Bema-Kaintiba Rd, 3 km N of Kaintiba, I 170 m, on vine (Araliaceae) in regrowth, HS 33715, 25 Ja n 1983. Central: from 9 locali ties, 10- 600 m, on rotting log, rock and tree trunks in plantation, monsoon, lowland and Castanopsis forests, HS 14674, 5 Feb 1981, HS & K. Naoni 14920, 10 Feb 1981 , HS & K. Naoni 15109, II Feb 1981, HS & K. Naoni 15192, 12 Feb 1981, HS & K. Naoni 16058, 16061 , 16085, 13 Feb 1981, HS & K. Naofli 16250, 14 Feb 1981, HS & K. Naolli 16413, 15 Feb 1981 , HS & K. Naoni 16686, 16 Feb 1982. Comments: common in Morobe a nd Central Provinces. Distribution: Manokwari, Madang, Morobe. W. Highlands, E. Highlands, Milne Bay, New Britain. Mastigolejeunea ligulata (Lehm. & Lindenb.) Schiffn. Central: Angabanga R. , 39km ENE of Bereina, 600m, on tree trunk and bullress in lowla nd forest, 12 Journ. Hattori Bot. Lab. No. 69 I 9 9 I

HS & K. Naoni, 16222, 16224, 14 Feb 1981. Distribution: E. Sepik, Madang, Morobe, New Ireland.

Mastigolejeunea undulata Gradst. & Grolle Southern Highlands: from 6 localities, 1450-2300 m, on rotting log, vine, tree trunk, Pandanus and Castanopsis trunks in regrowth, swampy grasslands, Nothofagus and Podocarpaceae dominated forest, Araucaria dominated depauperate forest and Castanopsis forest, HS 23208,8 Sep 1982, HS 23975,23977, 24031, II Sep 1982, HS 26417,26514,26656, II Dec 1982, HS 26911,26912, 13 Dec 1982, HS 32704, 16 Dec 1982. Distribution: Jayawijaya, Morobe, W. Highlands, Simbu, E. Highlands, Central. Mastigolejeunea virens (Aongstr.) Steph. New Ireland: Sali ViII. , 46 km SW of Kavieng, 10 m, on tree trunk in secondary growth, W. Apelis 12, 30 Dec 1982. Distribution: W. Highlands, Morobe, Central. Mastigophora diclados (Brid.) Nees West Sepik: Atkomfra, Eliptamin, 10km NNE of Telefomin, 1200m, on ground, B. Macrosimnok 42, 14 Jan 1983. Eastern Highlands: from 4 localities, 1720-2700m, on ground, on tree trunks and crowns in disturbed Castanopsis and montane forests, HS 17949, 7 Apr 1982, HS 18470,18517, 12 Apr 1982, HS 18661,14 Apr 1982, HS 18804,12 Apr 1982, HS 26346,26347,26348, 9 Dec 1982. Gulf: Hepataewa, 6km S of Kaintiba, 1200 m, on tree trunk in Castallopsis forest, HS 33852, 26 Jan 1983. Comments: common at middle and higher altitudes. Distribution: Manokwari, Madang, Morobe, W. Highlands, Simbu, S. Highlands. Metahygrobiella macgregorii (Steph.) Schust. Morobe: Wagau-Malolo track, 26km NE of Mumeng, 1600m, on tree roots in montane forest, HS 19652, 19655,15 May 1982. Distribution: Manokwari, Jayapura, E. Sepik, Northern, Milne Bay. Metzgeriopsis pusilla Goeb. Morobe: Slate and Gumi Cks Divide, 17km W of Bulo lo, 2JOOm, on Lauraceae leaves in montane forest, HS 13883, 30 Jan 1981. Distribution: Manokwari; new to Papua New Guinea. Phaeolejeunea latistipula (Schiffn.) Mizut. East Sepik: Kairiru I., on Artocarpus trunk, J. Klei 21,20 Jul 1979. Distribution: Morobe, New Britain, New Ireland, Manus. Plagiochilion giulianettii (Steph.) Inoue Southern Highlands: Mt Giluwe, 3500 m, on rotting stump in subalpine grasslands, HS 24154, 12 Sep 1982. Distribution: Jayawijaya. Simbu, Milne Bay. Plagiochilion oppositus (Reinw. et a\.) Hatt. Southern Highlands: Mt Giluwe, 3450 m, base of shrub in subalpine grasslands, HS & P. Garin 24113, 12 Sep 1982. Distribution: Manokwari, Jayawijaya, Morobe, Simbu, Central. Plagiochilion theriotanus (Steph.) Inoue Southern Highlands: Munia logging area, 14 km NW of Ialibu, 2300 m, on tree trunk in Nothofagus and Podocarpaceae dominated fores t, HS 23313,8 Sep 1982. H. STREIMANN : New hepatic records from New Guinea 13

Distribution: Morobe, Milne Bay. Pleurozia gigal1lea (Web.) Lindb. West Sepik: Misimin logging area, 4 km N of Tclefomin, 1400 m, on tree (Calophyllum) branch in montane forest , A. Bel/amy 1501. 13 Jul 1985. Gulf: Hepataewa, 6 km S of Kaintiba. 1200 rn , on tree trunk in CaSlanopsis dominated forest, HS 33851,26 Jan 1983 . Distribution: Manokwari, Paniai, Madang. Morobe. Enga, W. Highlands. Simbu, E. Highlands, S. Highlands. Northern, Milne Ba y. Ptychanthus striatus (Lehm. & Lindenb.) Nees Western Highlands: Banz to Jimi-Waghi Divide, 5km of Banz, 1900m, on Casuarina trunk in regrowth. HS 20963, 25 Jun 1982; Gernt R., Jimi Valley near Ruti. 54 km N of Mount Hagen, 600 m. on a vine in lowland forest. HS 21986. 2 Jul 1982; Baiyer R. Sanctuary, 39km N of Mount Hagen, 1160m. on shrub branches in lowland forest, HS 21004, 26 Jun 1982; Kagamuga, 10 km E of Mount Hagen, 1580 rn, on stem of exotic Hibiscus at Agricultural College, HS 20469, 22 Jun 1982. Simbu: Yagle Vill .. 4 km N of Mingende. 1800m, on rotting log in advanced regrowth, A. KOIragle 17.17 Apr 1982; Dirima Vii!. , 3km NW of Gumine. 1680 m, on rock in grasslands, M . Toia 133, 17 Dec 1982. Southern Highlands: from 4 locali ties, I 500-2840 m. on ground. on palm, Pandanus and treelet stems. HS 22144. 22170. 3 Jul 1982. HS 26647. II Dec 1982, HS 26744,12 Dec 1982, HS 32545. 16 Dec 1982. New Britain: Mopili-Galai No. I. 30 km SE of Kimhe, 60 m. on vine in lowland forest. S. Kolema 51 . 20 Dec 1984. New Ireland: Sali ViI!. . 46 km SE of Kavieng, 10 m. on tree trunk in sccondary forest. W. Ape/is 15. 30 Dec 1982. Distribution: Sorong, Jayawijaya. E. Sepik, Morobe. E. Highlands, Centra!' Pycnolejeunea grandiocellata Steph. Gulf: Bema-Kaintiba Rd. 4 km N of Kaintiba. 1200 m. on Canarium indicum trunk in regrowth. HS 33647. 25 Jan 1983. Distribution: Morobe. Centra!. Radula acutiloba Steph. Eastern Highlands: Lapegu. 6 km SW of Goroka. 1520 m. on Taxodium dis/iclrum trunk in Taxodium plantation. HS 18410. 11 Apr 1982. Distribution: Morobe. Sim bu. Radula anceps Sde Lac. West Sepik: Wutung Vill. area, epiphyte in rainforest. K. Naoni 23. 2 Aug. 1981. Southern Higlands: Tambul-Mendi Rd. 18 km SW of Tambul. 2840 m. on rotting log in disturbed montane forest. HS 26754 . 12 Dec 1982; Andawe R .• 6 km SE of lalibu, 1840 m, on ground in No/lrofagus and Podocarpaceae dominated forest. HS 26548. II Dec 1982. Distribution: Morobe, Central, Northern. Radula decurrens Mitt. Milne Bay: between Lakes Lelua and Boiak, 10km W of Kulumadau. Woodlark I.. 20m. on Cel/is branches in lowland forest, R. Kumei 97, 15 Oct 1984. Distribution: new to New Guinea; Samoa. Radula densifolia Castle West Sepik: near Kilifas, 550m, R. Brown 14,24, Jan 1985. Enga: Mount Hagcn-Wabag Rd, 17km SE of Wapenamanda, 2500m. on Cyr/andra roots in poor montane forest, HS 22093.29 Jan 1982. Gulf: Hepataewa. 6km S ofKaintiba. 1200m, on a vine in Caslanopsis dominated forest, HS 33873.26 Jan 1983. Central: Bereina-Angabanga R. Rd, 37 km ENE of Bcreina, 800 m, on vine and tree trunk. in lower montane forest J-/S & K. Naoni 16276. 16277. 16279. 14 Feb 1982; Ubcri Vill .• Kokoda Trail, on rotting branch in advanced regrowth. K. Naoni 15. Oct. 1981. 14 Journ. Hattori Bot. Lab. No. 69 I 9 9 I

Distribution: Manokwari, Jayapura, Jayawijaya, E. Sepik, Madang, Morobe, W. Highlands, Simbu, E. Highlands, S. Highlands, Milne Bay. Radula javanica Gott. Eastern Highlands: Yati R. , base of Kassam Pass, 24 km NE of Kainantu, 7S0 m, on Syzygium trunk in lower montane forest, HS & T. Umba JJ543, IS Jan 1981. Southern Highlands: Lai R., Mendi-Tari Rd , ISkm SSE of Mendi, ISOOm, on Pandanus stem in advanced secondary forest, HS 22212, 3 Jul 1982. Central: Varirata National Park, Sogeri, 22 km ENE of Port Moresby, 600 m, on tree trunk in Castanopsis and Schizomeria dominated forest, HS & A. Vinas 14596, S Feb 1981. Distribution: Manokwari, Madang, Morobe, N. Solomons. Radula madagascariensis Gott. Morobe: Pouyu Vi iI., I.S km SE of Aseki, 1300 m, on Wendlandia stem in regrowth, HS 26168, 6 Dec 1982. Southern Highlands: Piribu Sawmill, Tari-Komo Rd, 3 km SW of Tari, 16S0 m, on Caslanopsis trunk in Caslanopsis dominated forest , HS 32506, 32520, IS Dec 1982. Distribution: E. Highlands, Central. Radula multiflora Schiffn. Southern Highlands: Paunde logging area, IS km NNW of Ialibu, 2400 m, on tree trunk in montane forest, HS 32328, 14 Dec 1982. Distribution: Manokwari, Cenderawasih, Madang, Morobe, E. Highlands, Central, Northern, Milne Bay, New Britain, New Ireland, N. Solomons. Radula no vivrieseana Yamada Morobe: Logging area, IS km E of Bulolo, ISOO m, on tree trunk in Caslanopsis and Engelhardlia dominated forest , HS & A. Bellamy /3212, 27 Jan 198 1. Southern Highlands: Del Mission, ISkm SSW of Mendi, 1600m, on shrub in montane forest, HS 32805, 18 Dec 1982. Distribution: Manokwari; new to Papua New Guinea. Radula pinnulala Mitt. Morobe: Aseki-Mdamna track, I km SW of Aseki, 1370 m, on track cutting in advanced regrowth, HS 12555, 23 Jan 1981. Distribution: previous report (Grolle & Piippo 1984a) is 'Insulae Salomonae'. Radula protensa Lindenb. Western Highlands: Baiyer R. Sanctuary, 39 km N of Mount Hagen, 1160 m, on Citronella leaf and fallen tree branch in lowland fo rest, HS 21005,21078,26 Jun 1982; Kagamuga, 10km E of Mount Hagen, IS80 m, on cultivated mango at Agricultural College, HS 20432, 22 Jun 1982. Distribution: Sorong, Manokwari, Madang, Morobe, Central. Radula reflexa Nees & Mont. Central: Angabanga R ., 39km ENE of Bereina, 6S0m, on treelet stem in regrowth, HS & K. Naoni 16254, 14 Feb 198 1. Distribution: Madang, Morobe, Western. Radula retroflexa Tay\. Southern Highlands: Onim Forestry Station, 14 km NNW of lalibu, 22S0 m, on Pandanus roots in disturbed montane forest, HS 24651. 14 Sep 1982. Gulf: Hepataewa, S km S of Kaintiba, lOOOm , on treelet stem in lowland forest, HS 33793, 26 Jan 1983. Distribution: Morobe, W . Highlands, Simbu, Central, Milne Bay. H. STREIMANN: New hepatic records from New Guinea 15

Radula subsimilis Steph. Morobe: Manki Trig., Bulolo-Watut Divide, 10km SW of Bulolo, 1500m, on Syzygium trunk in disturbed Caslanopsis forest, /-IS & A. Bel/amy 13056, 26 Jan 1981. Central: Between head of Goldie R. and Owers Corner, 40 km ENE of Port Moresby, 500 m. on tree trunk in lower montane forest, /-IS & K. Naoni 14834, 9 Feb 1981. Distribution: new to New Guinea; Indonesia (Ceram). Raduta van-zantenii Yamada West Sepik: near Lumi, 600 m, R. Brown 27, Jan 1985. Di stribution: Manokwari, Jayawijaya, Morobe, Central. Raduta vrieseana Sde Lac. Wes tern Highlands: Gernt R. , Jim i Valley near Ruti, 600 m, on a vine in lowland forest, /-IS 21988, 2 Jul 1982. Di stribution: Morobe. Reboulia hemisphaerica (L.) Raddi Eastern Highlands: Rd to Frigano Sawmill, near Mopei ViII ., 34km SSW of Goroka, 1600m, on creek side in regrowth. /-IS 18859,15 Apr 1982. Distribution: Morobe. Enga, Simbu. Rhaphidolejeunea bischlerae Grolle Morobe: Nauti logging area, Upper Wa tut R., I I km SW of Bulolo, 1300 m, on Cyrlandra leaves a nd palm fronds, /-IS 17733,17735,6 Mar 1982. Distribution: Manokwari; new to Papua New Guinea. Riccardia argento-limbata Hewson & Grolle Eastern Highlands: Da ul o Pass. 18 km WNW of Goroka, 2700 m, o n Pandanus in advanced secondary fo rest, /-IS 17943, 7 Apr 1982. Distribution: Ma nokwari. PaniaL Morobe. Enga, W. Highlands. Simbu, S. Highlands. Saccogynidium muricellum (De Not.) Grolle Southern Highlands: Onim Forestry Station, 14km NNW of lalibu, 2250m, base of Notho/agus in distu rbed montane forest , /-IS 24573, 14 Sep 1982. Distribution: layawijaya, W. Sepik, Morobe. Scapania ja van ica Gott. Simbu: Komanegomambuno, 7 km SE of Mt Wilhelm, 3200 m, on dead stump in montane forest, A . Mundua 10, 13 Dec 1982; Yagle ViiI., 4km N of Mingende, 1800m, on ground in advanced regrowth, A. Kawagle. 13, 24, 17 Apr 1982; Dirima-Makaua Mountain Track, 3 km NW of Gumine, 1760 m, on road cutting in grasslands, M . Toia 56, 13 Dec 1982. Western Highlands: from 4 localities, 1700- 2800 m, on ground, on rock face , rotting tree branch and tree trunk in regrowth, disturbed and lower montane forests, /-IS 20813,25 lun 1982, /-IS 21168,27 Jun 1982, /-IS 21312, 21321 , 21322, 21491 ,21492, 28 lun 1982. Comments: in Grolle & Piippo (1984a) Wa bag should be in , not W. Highlands. Distribution: Jayawijaya, Madang, Morobe, Enga, E. Highlands, S. Highla nds, Milne Bay. Schiffneriolejeunea nymanii (Steph.) Gradst. & Terken Western Highlands: Rd from Banz to Jimi-Waghi Divide, 5 km N of Ba nz, 1900 m, on Casuarina trunk in regrowth, HS 20979, 25 1 un 1982; Karpena Plantation, 28 km NE of Mo unt Hagen, 1650 m, on Saurauia stem in regrowth, HS 21892, I lul 1982. Simbu: Dirima ViiI., 3 km NW of Gumine, 1680m, on tree trunk in grasslands, M. Toia 136, 17 Dec 1982. Distribution: Morobe. 16 Journ. Hattori Bot. Lab. No. 69 199 I

Schiffneriolejeunea tumida (Nees & Mont.) Gradst. var. haskarliana (Gott.) Gradst. Eastern HighJands: Kassam Pass, 19 km NE of Kainantu, 1600 m, on log in Lauraceae and Fagaceae dominated forest, HS & T. Umba 1l520, 15 Jan 1981. New Britain: Wangowango, Willaumez Peninsula, 20 km NNE of Talasea, sea leve l, on Ficus stem in strand vegetation, S. Kolema 24, 5 Dec 1984. Distribution: Merauke, Morobe, N . Solomons. Schistochila blumei (Nees) Trev. Eastern Highlands: Entabindi-Pusarasa area, 2 km NNW ofOkapa, 1720 m, on rotting log in disturbed Caslanopsis forest, HS 26326, 9 Dec 1982. Southern Highlands: Tari Gap, 23 km SE of Tari, 2740 m, on rotting wood in montane forest. HS 24460, 13 Sep 1982; Onim Forestry Station, 14km NNW of lalibu, 2250m, on tree root in disturbed montane forest, 24677, 14 Sep 1982; Fara Ra. , Kagua-Erave Rd, 11 km NW of Erave, 1800m, on log in advanced regrowth, A. Akivi la, 9 Sep 1982. Distribution: Manokwari, Jayapura, Jayawijaya, W . Sepik, Madang, Morobe, W. Hi ghla nds, Central, Northern, Milne Bay. Spruceanthus marianus (Gott.) Mizut. East Sepik: Kairiru I., 600m, on tree trunk in forest, W. BorreI/8I-20, 13 Nov 198 1. North Solomons: Buka I., Lonahan ViII., 14 km NNE of So ha no, 5 m, on Cel/is trunk in lowland forest, W. Korea 24,7 Jan 1983. Distribution: Sorong, Manokwari, Madang, Morobe, Central, New Britain. Spruceanthus pluriplicatus (Steph.) Gradst. Enga: Mape Ck, Mount Hagen-Wapenamanda Rd, 17km SE ofWapenamanda, 2500m, on vine and shrubs in advanced regrowth, HS 21563, 21638, 22099,29 Jun 1982. Western Highlands: Rd from Banz to Jimi-Waghi Divide, 5 km N of Banz, 1900 m, on Casuarina trunk, HS 20964, 25 Jun 1982; Tumbang ViII ., 7 km SE of Minj, 1800m, on Nothofagus trunk in regrowth, HS 21399, 28 Jun 1982. Southern Highlands: faro R. , Onim, 14 km NNW of lalibu, 2230 m, on tree trunk in poor montane forest, HS 23853, 11 Sep 1982. Distribution: Jayawijaya, Morobc, Central. Spruceanthus polymorphus (Sde Lac.) Verd. Western Hi ghlands: Baiyer R. Sanctuary, 39 km N of Mount Hagen, 1140 m, on Casuarina and Euodia stems in lowland forest, HS 21100, 21112, 26 Jun 1982 . Distribution: Jayawijaya. E. Sepik, M adang, Morobe, Central, N. Solomons. Stenolejeunea morobensis Grolle Western Highlands: Baiyer R.-Jimi Valley Rd at Sanctuary, 39 km N of Mount Hagen. 1180 m, on shrub in regrowth, HS 21138, 26 Jun 1982; Karpena Plantation, 28km NE of Mount Hagen, 1650m, on Trichospcrmum and Cordyline stems in regrowth, HS 21822,21883, I Jul 1982. Distribution: Morobe. Stenolejeunea thallop/lOra (Eifrig) Schust. Western Highlands: Karpena Plantation, 28km NE of Mount Hagen, 1650m, on Acalypha stem in regrowth, HS 21844, I Jul 1982. Distribution: new to New Guinea; Indonesia (Celebes). Stictolejeunea balfourii (M itt.) E. W. Jones Morobe: Busu R., 20 km NNW of Lac, 100 m, on small rocky mound in lowland rain forest on river flats, H. Streimann 19997, 10 Jun 1982. Distribution: new to New Guinea; Australia (Queensland). Syzygiella ovalifolia Inoue Morobe: Gumi Divide, 19 km W of Bulolo. 2000 m, on humus rich ground in Notholagus and H. STREIMANN: New hepatic records from New Guinea 17

Phyllocladus dominated forest, HS 22716, 13 Aug 1982; Ekuti Divide, Bulolo-Aseki Rd, 30 km WSW of Bulolo, 2100m, on base of shrub in montane forest, HS 20226,16 Jun 1982; Mt Kaindi, 4 & 5km W of Wau, 2230-2300 m, on humus rich ground, rotting log and tree stump in disturbed Nothofagus forest , HS 19831,19837,22 May 1982, HS 22445,26 Jul 1982. Distribution: Manokwari, Jayawijaya, W. Highlands, S. Highlands. Syzygiella subintegerrima (Reinw. et al.) Spruce Western Highlands: Tumbang ViII. , 18oom, on Araucaria trunk in remnant forest, HS 21319.28 Jun 1982. Distribution: Manokwari, Paniai, Jayawijaya, W. Sepik, Morobe, S. Highlands, Milne Bay. Telaranea kaindina Grolle Eastern Highland s: near Hagobi ViII. , 6 km SW of Lufa, 1850 m, on rotting log in montane forest, HS 18667, 14 Apr 1982. Distribution: Morobe. Telaranea la wesii (Steph.) Grolle Simbu: Komanegomambuno, 8 km SE of Mt Wilhelm, 3200 m, in moss forest, D. Mc Vean 267215, 1967 . Distribution: Manokwari, Jayapura, W. Sepik, Madang, Morobe, Central. Temnoma setigerum (Lindenb.) Schust. Eastern Highlands: Daulo Pass, 18 km WNW of Goroka, 2700 m, on humus rich ground in advanced secondary forest, HS 17967, 7 Apr 1982. Comments: common in . Dist ribution: Pania i, Jayawij aya, Morobe, W. Highlands, Simbu, S. Highlands. Thysananthus convolutus Lindenb. Southern Highlands: Lama Sawmill, 6 km SE of Iali bu, 1850 m, on Castanopsis trunk in Castallopsis dominated fo rest, HS 26660, II Dec 1982. Distribution: Sorong, Manokwari, Jayapura, W. Sepik, Morobe, E. Highlands, Central. Thysananthus fruticosus (Lindenb. & Gott.) Schiffn. West Sepik: Amisumbil, Eliptamin, 10km NNE of Telefomin, 12oom, on Garcillia trunk, 8. Macrosimnok 27, 29 Dec 1982. Eastern Highlands: Kassam Pass, 19 km NE of Kainantu, 1600m, on shrub branch in Lauraceae and Fagaceaedominated forest, HS & T. Umba 11451, 15 Jan 1981 . Southern Highlands: Bat!eri, Kagua-Erave Rd, 13km NW of Erave, 18oom, on rotting log in poor forest, HS 23433, 9 Scp 1982. New Britain: Willaumez Peninsula, 26 km NE of Talasea, 25 m, on young Planchonella branch in regrowth, S. Kolema 28, 5 Dec 1982. Comments: common in Morobe Province. Distribution: Sorong, Manokwari, Jayapura, Paniai, E. Sepik, Madang, Morobe, Western, Central, Milne Bay. Thysananthus mollis Steph. Western Highlands: Baiyer R.-Ruti Rd, Jimi Valley, 42km N of Mount Hagen, 1600m, on Dil/enia trunk in lower montane forest, HS 22075, 2 Jul 1982. Eastern Highlands: Daulo Pass, 18km WNW of Goroka, 2700 m, on tree trunk in advanced secondary forest, HS 18003, 7 Apr 1982. Southern Highlands: Lama Sawmill, 6 km SE of Ialibu, 1850 m, on Castanopsis trunk in Castanopsis dominated forest, HS 26631 , II Dec 1982. Distribution: Manokwari, Morobe, Simbu, Central, Milne Bay. Thysananthus spathulistipus (Reinw. et al.) Lindenb. West Sepik: Antingin, Eliptamin, 10 km NNE ofTelefomin, 1200 m, on stump in Caslanopsis dominated 18 Journ. Ha tt ori Bot. Lab. No. 69 I 9 9 I forest. B . Macrosimnok 4. 26 Dec 1982. Western Highlands: Baiye r R.-Ruti Rd, Jimi Valley. 42 km N of Mount Hagen. 1600m. on shrub in lower montane forest. l-/S 22061,2 Jul 1982, HS 22242,5 Jul 1982; Jimi-Waghi Divide. 9km N of Banz, 2400 m, on fallen tree branch in dense regrowth, l-/S 20916, 25 Jun 1982; Kum Magei Mo untain, 8km SE of Mount Hagen, 2550m, on Ardisia branches in Nothofagus dominated forest. l-/ S 20679, 24 Jun 1982. Simbu: Makaua Mountain , 3 km NW o f Gumine. 1860 m, on tree branch in NotllO{agus dominated fo rest. M . Toia 85. 13 Dec 1982. Eastern Highlands: Kassam Pass, 19 km NE of Kaina ntu, 1600 m. on ground, rotting log a nd fallen bra nch in Lauraceae and Fagaceae dominated forest. l-/S & T. Umha 11397, 11 398. 11469. 15 Ja n 198 1; track to Mt Michael, 4 km SW of Lufa , 1900m. on shrub in disturbed montane forest, lIS 18807, 12 Apr 1982; Daul o Pass, 18km WNW of Goroka, 2600m. o n treelet and tree stems in advanced re growth and remnant forest, l-/S 18093, 181 24, 7 Apr 1982. Southern Highlands: from 4 localities, 1450- 2250 m, on shrub, treelet and tree stems in di s­ turbed or poor forests. l-/S 24378. 24388,13 Sep 1982, l-/S 24740, 15 Sep 1982, HS 26902, 13 Dec 1982, HS 32712. 16 Dec 1982. Distribution: So ro ng. M a nokwari. Jayapura. E. Sepik. Morobe. Western, Central, Milne Bay. Trichocolea breviseta Steph. Southern Highlands: M unie logging a rea , 14 km NW of lalibu, 2300 m, on dead treelet in Notho{agus dominated forest, l-/S 23642. 10 Sep 1982. Distribution: Jayapura , Jayawijaya, W . Sepik, Morobe, W. Highlands. Simbu, E. Highlands. Central. Trichocolea plum a (Reinw. et at.) Mont. East Sepik: Kairiru I.. 500m. on ground and rotting log in lowland fo rest, W. 80rre1l4, 8 1-35, Sep 1979 and Nov 1981. Enga: Mape Ck, Mount H agen-Wapenamanda Rd. 17km SE of Wapenamanda. 2500 m , on shrub stem in advanced regrowth and poor montane forest, HS 21585, 2 1645,29 Jun 1982. Westcrn Highlands: Tabibuga Rd. Jimi Va lley. 14 km N of Banz, 2000 m. o n shaded rock in dense regrowth. l-/S 20835. 25 Jun 1982. Simbu: Kotdamc. 8 km SE of Mt Wilhclm, 2650 m, on ground in secondary forest, A. Mundlla 183.15 Dcc 1982. Distribution: Manokwari, Jayapura. Jayawijaya. Madang. Morobe, Central. Milne Bay. TuyarnaeIla serralislipa Hatt. Southern Highlands: Andawe R .. 6 km SE of la li bu, 1840 m. on SFygiw/1 leaf in Nothofaglls and Podoca rpaceae dominated forest, l-/S 26498. I I Dec 1982. Distribution: Manokwa ri . Jayapura, Morobe. Zoopsis liukiuensis Horik. Western Highl a nds: Mur Mur Pass, 7 km NE of Ta mbul. 2800 m. on ground in disturbed Cryptocarya and EllIl'ocarplIs dominated fo rest. HS 21205 ( + Lepidolejeunea wallichiana). 27 Jun 1982; Kagamuga, 10km E of Mount Hagen. 1580m. base of Eucalyptus in Agricultural College, HS 20460,22 Jun 1982. Eastern High lands: track to Mt Michael, 4km SW of Lufa , 1900m, on dead st ump in disturbed montane forest. l-/S 18735, 12 Apr 1982. Gulf: Hcpataewa. 5km S o f Kaintiba. 1000m, on dead Cyathea in poor lowland forest. HS 33820.26 Ja n 1983. Distribution: Manokwari . Jayapura. W. Scpik, Morobc, Simbu, S. Highlands, Western, Central.

ACKNOWLEDGEMENTS Specia l thanks are extended to Dr R. Grolle for assistance with determinations, queries and for checking this work. My gratitude is also extended to Drs M. Mizutani and K. Yamada for their willingness to determine collections and for allowing the publication of the results. Appreciation is also extended to R. Kumei, S. Kolema, A. Bellamy, and M. T oia for their keen and continuing interest in collecting cryptogams. A. Kairo, A. Bellamy, E. Tamba and T. Umba assisted the author with collecting in the Morobe Province in 198 1, in the Central Province the H. STREIMANN: New hepatic records from New Guinea 19 same assistance was willingly provided by K . Naoni and A. Vinas. During the author's term at the Papua New Guinea Forestry College (1982-1983) the principal, S. Kalogo, kindly consented to and assisted with the fie ld work.

R EFE RENCES

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