Psocodea: 'Psocoptera': Troctomorp
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André Nel Sixtieth Anniversary Festschrift
Palaeoentomology 002 (6): 534–555 ISSN 2624-2826 (print edition) https://www.mapress.com/j/pe/ PALAEOENTOMOLOGY PE Copyright © 2019 Magnolia Press Editorial ISSN 2624-2834 (online edition) https://doi.org/10.11646/palaeoentomology.2.6.1 http://zoobank.org/urn:lsid:zoobank.org:pub:25D35BD3-0C86-4BD6-B350-C98CA499A9B4 André Nel sixtieth anniversary Festschrift DANY AZAR1, 2, ROMAIN GARROUSTE3 & ANTONIO ARILLO4 1Lebanese University, Faculty of Sciences II, Department of Natural Sciences, P.O. Box: 26110217, Fanar, Matn, Lebanon. Email: [email protected] 2State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China. 3Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France. 4Departamento de Biodiversidad, Ecología y Evolución, Facultad de Biología, Universidad Complutense, Madrid, Spain. FIGURE 1. Portrait of André Nel. During the last “International Congress on Fossil Insects, mainly by our esteemed Russian colleagues, and where Arthropods and Amber” held this year in the Dominican several of our members in the IPS contributed in edited volumes honoring some of our great scientists. Republic, we unanimously agreed—in the International This issue is a Festschrift to celebrate the 60th Palaeoentomological Society (IPS)—to honor our great birthday of Professor André Nel (from the ‘Muséum colleagues who have given us and the science (and still) national d’Histoire naturelle’, Paris) and constitutes significant knowledge on the evolution of fossil insects a tribute to him for his great ongoing, prolific and his and terrestrial arthropods over the years. -
Insecta: Psocodea: 'Psocoptera'
Molecular systematics of the suborder Trogiomorpha (Insecta: Title Psocodea: 'Psocoptera') Author(s) Yoshizawa, Kazunori; Lienhard, Charles; Johnson, Kevin P. Citation Zoological Journal of the Linnean Society, 146(2): 287-299 Issue Date 2006-02 DOI Doc URL http://hdl.handle.net/2115/43134 The definitive version is available at www.blackwell- Right synergy.com Type article (author version) Additional Information File Information 2006zjls-1.pdf Instructions for use Hokkaido University Collection of Scholarly and Academic Papers : HUSCAP Blackwell Science, LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082The Lin- nean Society of London, 2006? 2006 146? •••• zoj_207.fm Original Article MOLECULAR SYSTEMATICS OF THE SUBORDER TROGIOMORPHA K. YOSHIZAWA ET AL. Zoological Journal of the Linnean Society, 2006, 146, ••–••. With 3 figures Molecular systematics of the suborder Trogiomorpha (Insecta: Psocodea: ‘Psocoptera’) KAZUNORI YOSHIZAWA1*, CHARLES LIENHARD2 and KEVIN P. JOHNSON3 1Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo 060-8589, Japan 2Natural History Museum, c.p. 6434, CH-1211, Geneva 6, Switzerland 3Illinois Natural History Survey, 607 East Peabody Drive, Champaign, IL 61820, USA Received March 2005; accepted for publication July 2005 Phylogenetic relationships among extant families in the suborder Trogiomorpha (Insecta: Psocodea: ‘Psocoptera’) 1 were inferred from partial sequences of the nuclear 18S rRNA and Histone 3 and mitochondrial 16S rRNA genes. Analyses of these data produced trees that largely supported the traditional classification; however, monophyly of the infraorder Psocathropetae (= Psyllipsocidae + Prionoglarididae) was not recovered. Instead, the family Psyllipso- cidae was recovered as the sister taxon to the infraorder Atropetae (= Lepidopsocidae + Trogiidae + Psoquillidae), and the Prionoglarididae was recovered as sister to all other families in the suborder. -
Psocoptera Em Cavernas Do Brasil: Riqueza, Composição E Distribuição
PSOCOPTERA EM CAVERNAS DO BRASIL: RIQUEZA, COMPOSIÇÃO E DISTRIBUIÇÃO THAÍS OLIVEIRA DO CARMO 2009 THAÍS OLIVEIRA DO CARMO PSOCOPTERA EM CAVERNAS DO BRASIL: RIQUEZA, COMPOSIÇÃO E DISTRIBUIÇÃO Dissertação apresentada à Universidade Federal de Lavras, como parte das exigências do programa de Pós-Graduação em Ecologia Aplicada, área de concentração em Ecologia e Conservação de Paisagens Fragmentadas e Agroecossistemas, para obtenção do título de “Mestre”. Orientador Prof. Dr. Rodrigo Lopes Ferreira LAVRAS MINAS GERAIS – BRASIL 2009 Ficha Catalográfica Preparada pela Divisão de Processos Técnicos da Biblioteca Central da UFLA Carmo, Thaís Oliveira do. Psocoptera em cavernas do Brasil: riqueza, composição e distribuição / Thaís Oliveira do Carmo. – Lavras : UFLA, 2009. 98 p. : il. Dissertação (mestrado) – Universidade Federal de Lavras, 2009. Orientador: Rodrigo Lopes Ferreira. Bibliografia. 1. Insetos cavernícolas. 2. Ecologia. 3. Diversidade. 4. Fauna cavernícola. I. Universidade Federal de Lavras. II. Título. CDD – 574.5264 THAÍS OLIVEIRA DO CARMO PSOCOPTERA EM CAVERNAS DO BRASIL: RIQUEZA, COMPOSIÇÃO E DISTRIBUIÇÃO Dissertação apresentada à Universidade Federal de Lavras, como parte das exigências do programa de Pós-Graduação em Ecologia Aplicada, área de concentração em Ecologia e Conservação de Paisagens Fragmentadas e Agroecossistemas, para obtenção do título de “Mestre”. APROVADA em 04 de dezembro de 2009 Prof. Dr. Marconi Souza Silva UNILAVRAS Prof. Dr. Luís Cláudio Paterno Silveira UFLA Prof. Dr. Rodrigo Lopes Ferreira UFLA (Orientador) LAVRAS MINAS GERAIS – BRASIL ...Então não vá embora Agora que eu posso dizer Eu já era o que sou agora Mas agora gosto de ser (Poema Quebrado - Oswaldo Montenegro) AGRADECIMENTOS A Deus, pois com Ele nada nessa vida é impossível! Agradeço aos meus pais, Joaquim e Madalena, pela oportunidade e apoio. -
Insecta: Psocodea: Psocomorpha
1 for submission to Insect Systematics & Diversity (ISD-2020-0035) 2 3 Systematic Position of the Enigmatic Psocid Family Lesneiidae (Insecta: 4 Psocodea: Psocomorpha), with Description of Two New Species 5 6 Kazunori Yoshizawa,1,6 Yuri M. Marusik,2,3,4 Izumi Yao,1 and Charles Lienhard 5 7 8 1 SystematiC Entomology, SChool of AgriCulture, Hokkaido University, Sapporo 060-8589, Japan 9 2 Institute for BiologiCal Problems of the North RAS, Portovaya Str.18, Magadan, Russia 10 3 Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South 11 AfriCa 12 4 ZoologiCal Museum, Biodiversity Unit, University of Turku, FI-20014, Finland 13 5 Geneva Natural History Museum, CP 6434, CH-1211 Geneva 6, Switzerland 14 15 6 Corresponding author, e-mail: [email protected] 16 17 18 Abstract 19 The systematiC plaCement of an enigmatiC psocid family restriCted to AfriCa, Lesneiidae, 20 was estimated by using a multiple gene data set. The candidates for its close relatives are now 21 Classified under two different infraorders, the family Archipsocidae of the infraorder Archipsocetae 22 or the families Elipsocidae/Mesopsocidae of the infraorder Homilopsocidea. The maximum 23 likelihood and Bayesian analyses of the moleCular data set strongly suggested that the Lesneiidae 24 belongs to Homilopsocidea and forms a clade with Elipsocidae/Mesopsocidae/EolaChesillinae 25 (LaChesillidae). However, the relationships among these (sub)families and Lesneiidae, including the 26 monophyly of Elipsocidae and MesopsoCidae, were ambiguous or questionable, showing the 27 neCessity of further investigations for elucidating their relationships and validating the status of 28 these families. -
Morphology of Psocomorpha (Psocodea: 'Psocoptera')
Title MORPHOLOGY OF PSOCOMORPHA (PSOCODEA: 'PSOCOPTERA') Author(s) Yoshizawa, Kazunori Insecta matsumurana. New series : journal of the Faculty of Agriculture Hokkaido University, series entomology, 62, 1- Citation 44 Issue Date 2005-12 Doc URL http://hdl.handle.net/2115/10524 Type bulletin (article) File Information Yoshizawa-62.pdf Instructions for use Hokkaido University Collection of Scholarly and Academic Papers : HUSCAP INSECTA MATSUMURANA NEW SERIES 62: 1–44 DECEMBER 2005 MORPHOLOGY OF PSOCOMORPHA (PSOCODEA: 'PSOCOPTERA') By KAZUNORI YOSHIZAWA Abstract YOSHIZAWA, K. 2005. Morphology of Psocomorpha (Psocodea: 'Psocoptera'). Ins. matsum. n. s. 62: 1–44, 24 figs. Adult integumental morphology of the suborder Psocomorpha (Psocodea: 'Psocoptera') was examined, and homologies and transformation series of characters throughout the suborder and Psocoptera were discussed. These examinations formed the basis of the recent morphology-based cladistic analysis of the Psocomorpha (Yoshizawa, 2002, Zool. J. Linn. Soc. 136: 371–400). Author's address. Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060-8589 Japan. E-mail. [email protected]. 1 INTRODUCTION Psocoptera (psocids, booklice or barklice) are a paraphyletic assemblage of non-parasitic members of the order Psocodea (Lyal, 1985; Yoshizawa & Johnson, 2003, 2005; Johnson et al., 2004), containing about 5500 described species (Lienhard, 2003). They are about 1 to 10 mm in length and characterized by well-developed postclypeus, long antennae, pick-like lacinia, reduced prothorax, well-developed pterothorax, etc. Phylogenetically, Psocoptera compose a monophyletic group (the order Psocodea) with parasitic lice ('Phtiraptera': biting lice and sucking lice) (Lyal, 1985; Yoshizawa & Johnson, 2003, in press; Johnson et al., 2004). The order is related to Thysanoptera (thrips) and Hemiptera (bugs, cicadas, etc.) (Yoshizawa & Saigusa, 2001, 2003, but see also Yoshizawa & Johnson, 2005). -
In Search of the Sister Group of the True Lice: a Systematic Review of Booklice and Their Relatives, with an Updated Checklist of Liposcelididae (Insecta: Psocodea)
Arthropod Systematics & Phylogeny 181 68 (2) 181 – 195 © Museum für Tierkunde Dresden, eISSN 1864-8312, 22.06.2010 In search of the sister group of the true lice: A systematic review of booklice and their relatives, with an updated checklist of Liposcelididae (Insecta: Psocodea) KAZUNORI YOSHIZAWA 1, * & CHARLES LIENHARD 2 1 Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo 060-8589, Japan [[email protected]] 2 Natural History Museum, c. p. 6434, CH-1211 Geneva 6, Switzerland * Corresponding author Received 23.ii.2010, accepted 26.iv.2010. Published online at www.arthropod-systematics.de on 22.06.2010. > Abstract The taxonomy, fossil record, phylogeny, and systematic placement of the booklouse family Liposcelididae (Insecta: Psoco- dea: ‘Psocoptera’) were reviewed. An apterous specimen from lower Eocene, erroneously identifi ed as Embidopsocus eoce- nicus Nel et al., 2004 in the literature, is recognized here as an unidentifi ed species of Liposcelis Motschulsky, 1852. It represents the oldest fossil of the genus. Phylogenetic relationships within the family presented in the recent literature were re-analyzed, based on a revised data matrix. The resulting tree was generally in agreement with that originally published, but the most basal dichotomy between the fossil taxon Cretoscelis Grimaldi & Engel, 2006 and the rest of the Liposcelididae was not supported. Monophyly of Liposcelis with respect to Troglotroctes Lienhard, 1996 is highly questionable, but the latter genus is retained because of lack of conclusive evidence. Paraphyly of Psocoptera (i.e., closer relationship between Liposcelididae and parasitic lice) is now well established, based on both morphological and molecular data. -
A Review of Recent Phylogenetic Contributions
ACTA ENTOMOLOGICA MUSEI NATIONALIS PRAGAE Published 8.xii.2008 Volume 48(2), pp. 217-232 ISSN 0374-1036 Four chapters about the monophyly of insect ‘orders’: A review of recent phylogenetic contributions Jan ZRZAVÝ Department of Zoology, Faculty of Science, University of South Bohemia, and Biology Centre, Czech Academy of Science, Branišovská 31, CZ-370 05 České Budějovice, Czech Republic; e-mail: [email protected] Abstract. Recent phylogenetic analyses, both morphological and molecular, strongly support the monophyly of most insect ‘orders’. On the contrary, the Blattaria, Psocoptera, and Mecoptera are defi nitely paraphyletic (with respect of the Isoptera, Phthiraptera, and Siphonaptera, respectively), and the Phthiraptera are possibly diphyletic. Small relictual subclades that are closely related to the Isoptera, Phthiraptera, and Siphonaptera were identifi ed (Cryptocercidae, Lipo- scelididae, and Boreidae, respectively), which provides an enormous amount of evidence about the origin and early evolution of the highly apomorphic eusocial or parasitic ex-groups. Position of the enigmatic ‘zygentoman’ Tricholepidion Wygodzinsky, 1961, remains uncertain. Possible non-monophyly of the Megalo- ptera (with respect of the Raphidioptera) and the Phasmatodea (with respect of the Embioptera) are shortly discussed. Key words. Insecta, Zygentoma, Tricholepidion, Blattaria, Isoptera, Cryptocercus, Psocoptera, Phthiraptera, Liposcelididae, Mecoptera, Siphonaptera, Boreidae, Nannochoristidae, Timema, phylogeny, monophyly Introduction The goal of modern systematics is twofold: to provide a biological ‘lingua franca’ that facilitates an exchange of information among researchers, and to provide a hierarchical system that is meaningful in the context of our understanding of phylogenetic history. However, both goals are often in confl ict. Phylogenetics is about a nested hierarchy of clades, without any privileged ‘rank’ (like ‘order’ or ‘family’). -
Redalyc.A List of Psocoptera (Insecta: Psocodea) from Brazil
Revista Mexicana de Biodiversidad ISSN: 1870-3453 [email protected] Universidad Nacional Autónoma de México México García Aldrete, Alfonso N.; Mockford, Edward L. A list of Psocoptera (Insecta: Psocodea) from Brazil Revista Mexicana de Biodiversidad, vol. 80, núm. 3, 2009, pp. 665-673 Universidad Nacional Autónoma de México Distrito Federal, México Available in: http://www.redalyc.org/articulo.oa?id=42515996009 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative Revista Mexicana de Biodiversidad 80: 665- 673, 2009 A list of Psocoptera (Insecta: Psocodea) from Brazil Listado de Psocoptera (Insecta: Psocodea) de Brasil Alfonso N. García Aldrete1* and Edward L. Mockford 2 1Instituto de Biología, Universidad Nacional Autónoma de México, Apartado postal 70-153, 04510 México, D. F., Mexico. 2Department of Biological Sciences, Illinois State University, Campus Box 4120, Normal, Illinois 61790-4120, USA. *Correspondent: [email protected] Abstract. The species of Psocoptera currently known for Brazil are listed, with state distribution and biogeographic status. The categories of geographic distribution are discussed, as well as some of the evidence indicating that the present size of the Brazilian fauna is underestimated. Key words: Psocodea, Psocoptera, Brazil, geographic distribution. Resumen. Se listan las especies de Psocoptera actualmente registradas en Brasil, incluyendo la distribución por estado y su categoría biogeográfi ca. Se presenta alguna de la evidencia que hace suponer que el tamaño actual de la fauna de Psocoptera de Brasil está subestimada. -
'Psocoptera' (Psocodea) from Brazil
ARTICLE A checklist of ‘Psocoptera’ (Psocodea) from Brazil: an update to the list of 2009 of García Aldrete and Mockford, with an identification key to the families Alberto Moreira da Silva-Neto¹ & Alfonso Neri García Aldrete² ¹ Instituto Nacional de Pesquisas da Amazônia (INPA), Coordenação de Pesquisas em Entomologia (CPEN), Programa de Pós-Graduação em Entomologia. Manaus, AM, Brasil. ORCID: http://orcid.org/0000-0002-4522-3756. E-mail: [email protected] ² Universidad Nacional Autónoma de México (UNAM), Instituto de Biología, Departamento de Zoología, Laboratorio de Entomología. México, D.F., México. ORCID: http://orcid.org/0000-0001-7214-7966. E-mail: [email protected] Abstract. The described species of Psocoptera currently known for Brazil are listed, with state distribution and biogeographic status. An identification key to the families recorded in Brazil is presented. Key-Words. Geographic distribution; Psocids; Neotropics. INTRODUCTION (2009) is derived mostly from the ongoing study of the vast Psocoptera collection in the Psocoptera have no popular name in Brazil, Instituto Nacional de Pesquisas da Amazônia being known in other countries as book lice, bark (INPA), in Manaus, Amazonas; from the study of lice or psocids. These insects are small, measuring the Psocoptera collected through the program from 1 to 10 mm in length and feed on algae, li- PPBio-Semi-Árido, housed in the Entomological chens, fungi and organic fragments (Smithers, Collection Prof. Johann Becker of the Zoology 1991). Psocoptera is a paraphyletic group because Museum of the Universidade Estadual de Feira de the Phthiraptera are phylogenetically embed- Santana, in Feira de Santana, Bahia, Brazil (MZFS), ded in the Psocoptera infraorder Nanopsocetae from the study of the Psocoptera collected (Johnson et al., 2004; Yoshizawa & Johnson, through the program Cave invertebrates in Brazil: 2010; Yoshizawa & Lienhard, 2010). -
Aspects of the Biogeography of North American Psocoptera (Insecta)
15 Aspects of the Biogeography of North American Psocoptera (Insecta) Edward L. Mockford School of Biological Sciences, Illinois State University, Normal, Illinois, USA 1. Introduction The group under consideration here is the classic order Psocoptera as defined in the Torre- Bueno Glossary of Entomology (Nichols & Shuh, 1989). Although this group is unquestionably paraphyletic (see Lyal, 1985, Yoshizawa & Lienhard, 2010), these free-living, non-ectoparasitic forms are readily recognizable. In defining North America for this chapter, I adhere closely to Shelford (1963, Fig. 1-9), but I shall use the Tropic of Cancer as the southern cut-off line, and I exclude the Antillean islands. Although the ranges of many species of Psocoptera extend across the Tropic of Cancer, the inclusion of the tropical areas would involve the comparison of relatively well- studied regions and relatively less well-studied regions. The North America Psocoptera, as defined above, comprises a faunal list of 397 species in 90 genera and 27 families ( Table 1) . Comparisons are made here with several other relatively well-studied faunas. The psocid fauna of the Euro-Mediterranean region, summarized by Lienhard (1998) with additions by the same author (2002, 2005, 2006) and Lienhard & Baz (2004) has a fauna of 252 species in 67 genera and 25 families. As would be expected, nearly all of the families are shared between the two regions. The only two families not shared are Ptiloneuridae and Dasydemellidae, which reach North America but not the western Palearctic. Ptiloneuridae has a single species and Dasydemellidae two in North America. The rather large differences at the generic and specific levels are probably due to the much greater access that these insects have for invasion of North America from the tropics than invasion from the tropics in the Western Palearctic. -
Phylogenomics and the Evolution of Hemipteroid Insects
Phylogenomics and the evolution of hemipteroid insects Kevin P. Johnsona,1, Christopher H. Dietricha, Frank Friedrichb, Rolf G. Beutelc, Benjamin Wipflerc,d, Ralph S. Petersd, Julie M. Allena,e, Malte Petersenf, Alexander Donathf, Kimberly K. O. Waldeng, Alexey M. Kozlovh, Lars Podsiadlowskif,i, Christoph Mayerf, Karen Meusemannf,j,k, Alexandros Vasilikopoulosf, Robert M. Waterhousel, Stephen L. Cameronm, Christiane Weirauchn, Daniel R. Swansona, Diana M. Percyo,p, Nate B. Hardyq, Irene Terryr, Shanlin Lius, Xin Zhout, Bernhard Misoff, Hugh M. Robertsong, and Kazunori Yoshizawau aIllinois Natural History Survey, Prairie Research Institute, University of Illinois at Urbana–Champaign, Champaign, IL 61820; bInstitut für Zoologie, Universität Hamburg, 20146 Hamburg, Germany; cInstitut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität Jena, 07743 Jena, Germany; dCenter of Taxonomy and Evolutionary Research, Arthropoda Department, Zoological Research Museum Alexander Koenig, 53113 Bonn, Germany; eDepartment of Biology, University of Nevada, Reno, NV 89557; fCenter for Molecular Biodiversity Research, Zoological Research Museum Alexander Koenig, 53113 Bonn, Germany; gDepartment of Entomology, University of Illinois at Urbana–Champaign, Urbana, IL 61801; hScientific Computing Group, Heidelberg Institute for Theoretical Studies, 69118 Heidelberg, Germany; iInstitute of Evolutionary Biology and Ecology, University of Bonn, 53121 Bonn, Germany; jEvolutionary Biology and Ecology, Institute for Biology I (Zoology), University of -
Fossil Perspectives on the Evolution of Insect Diversity
FOSSIL PERSPECTIVES ON THE EVOLUTION OF INSECT DIVERSITY Thesis submitted by David B Nicholson For examination for the degree of PhD University of York Department of Biology November 2012 1 Abstract A key contribution of palaeontology has been the elucidation of macroevolutionary patterns and processes through deep time, with fossils providing the only direct temporal evidence of how life has responded to a variety of forces. Thus, palaeontology may provide important information on the extinction crisis facing the biosphere today, and its likely consequences. Hexapods (insects and close relatives) comprise over 50% of described species. Explaining why this group dominates terrestrial biodiversity is a major challenge. In this thesis, I present a new dataset of hexapod fossil family ranges compiled from published literature up to the end of 2009. Between four and five hundred families have been added to the hexapod fossil record since previous compilations were published in the early 1990s. Despite this, the broad pattern of described richness through time depicted remains similar, with described richness increasing steadily through geological history and a shift in dominant taxa after the Palaeozoic. However, after detrending, described richness is not well correlated with the earlier datasets, indicating significant changes in shorter term patterns. Corrections for rock record and sampling effort change some of the patterns seen. The time series produced identify several features of the fossil record of insects as likely artefacts, such as high Carboniferous richness, a Cretaceous plateau, and a late Eocene jump in richness. Other features seem more robust, such as a Permian rise and peak, high turnover at the end of the Permian, and a late-Jurassic rise.