Dragons of the Deep
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Cranial Anatomy, Taxonomic Implications
[Palaeontology, Vol. 55, Part 4, 2012, pp. 743–773] CRANIAL ANATOMY, TAXONOMIC IMPLICATIONS AND PALAEOPATHOLOGY OF AN UPPER JURASSIC PLIOSAUR (REPTILIA: SAUROPTERYGIA) FROM WESTBURY, WILTSHIRE, UK by JUDYTH SASSOON1, LESLIE F. NOE` 2 and MICHAEL J. BENTON1* 1School of Earth Sciences, University of Bristol, Wills Memorial Building, Queen’s Road, Bristol BS8 1RJ, UK; e-mails: [email protected], [email protected] 2Geociencias, departamento de Fisica, Universidad de los Andes, Bogota´ DC, Colombia; e-mail: [email protected] *Corresponding author. Typescript received 5 December 2010; accepted in revised form 6 April 2011 Abstract: Complete skulls of giant marine reptiles of the genera. The two Westbury Pliosaurus specimens share many Late Jurassic are rare, and so the discovery of the 1.8-m- features, including the form of the teeth, but marked differ- long skull of a pliosaur from the Kimmeridge Clay Forma- ences in the snout and parietal crest suggest sexual dimor- tion (Kimmeridgian) of Westbury, Wiltshire, UK, is an phism; the present specimen is probably female. The large important find. The specimen shows most of the cranial size of the animal, the extent of sutural fusion and the and mandibular anatomy, as well as a series of pathological pathologies suggest this is an ageing individual. An erosive conditions. It was previously referred to Pliosaurus brachy- arthrotic condition of the articular glenoids led to pro- spondylus, but it can be referred reliably only to the genus longed jaw misalignment, generating a suite of associated Pliosaurus, because species within the genus are currently in bone and dental pathologies. -
First Occurrence of a Gigantic Pliosaurid Plesiosaur in The
Bull. Soc. géol. Fr., 2003, t. 174, no 3, pp. 271-278 First occurrence of a gigantic pliosaurid plesiosaur in the late Jurassic (Kimmeridgian) of Mexico MARIE-CÉLINE BUCHY1,EBERHARD FREY2,WOLFGANG STINNESBECK1 &JOSÉ GUADALUPE LÓPEZ-OLIVA3 Key words. – Kimmeridgian, Pliosauridae, Mexico, Palaeobiogeography. Abstract. – Reinvestigation of a partial vertebral column from the Kimmeridgian La Caja Formation of Mexico, housed in the University of Linares (Mexico), and previously attributed to a dinosaur, proves to be from a very large pliosaurid plesiosaur. This specimen represents the first plesiosaur described from the Jurassic of Mexico. Its length has been esti- mated at 15 metres and, as a juvenile, is considered to be one of the largest Jurassic marine reptiles. The remains of this animal are here described. The morphology of the vertebral column is not diagnostic beyond family level. Large pliosaur vertebrae of a similar size are known from the Upper Jurassic of Europe, and are often referred to the genera Liopleurodon or Simolestes but these identifications are based only upon the size of the centra and have no taxonomic justification. A portion of rostrum with teeth was discovered together with the vertebral column but is unfortunately now lost. The Mexican pliosaur fills geographical and chronological gaps between western Tethys and South American pliosaurs, and is an additional support to the hypothesis of a Hispanic corridor linking at least temporarily the NW Euro- pean marine province with the western South American marine (Pacific) realm during the late Jurassic. Première occurrence d’un plésiosaure pliosauride géant dans le Jurassique supérieur (Kimméridgien) du Mexique Mots clés. -
A Revision of the Classification of the Plesiosauria with a Synopsis of the Stratigraphical and Geographical Distribution Of
LUNDS UNIVERSITETS ARSSKRIFT. N. F. Avd. 2. Bd 59. Nr l. KUNGL. FYSIOGRAFISKA SÅLLSKAPETS HANDLINGAR, N. F. Bd 74. Nr 1. A REVISION OF THE CLASSIFICATION OF THE PLESIOSAURIA WITH A SYNOPSIS OF THE STRATIGRAPHICAL AND GEOGRAPHICAL DISTRIBUTION OF THE GROUP BY PER OVE PERSSON LUND C. W. K. GLEER UP Read before the Royal Physiographic Society, February 13, 1963. LUND HÅKAN OHLSSONS BOKTRYCKERI l 9 6 3 l. Introduction The sub-order Plesiosauria is one of the best known of the Mesozoic Reptile groups, but, as emphasized by KuHN (1961, p. 75) and other authors, its classification is still not satisfactory, and needs a thorough revision. The present paper is an attempt at such a revision, and includes also a tabular synopsis of the stratigraphical and geo graphical distribution of the group. Some of the species are discussed in the text (pp. 17-22). The synopsis is completed with seven maps (figs. 2-8, pp. 10-16), a selective synonym list (pp. 41-42), and a list of rejected species (pp. 42-43). Some forms which have been erroneously referred to the Plesiosauria are also briefly mentioned ("Non-Plesiosaurians", p. 43). - The numerals in braekets after the generic and specific names in the text refer to the tabular synopsis, in which the different forms are numbered in successional order. The author has exaroined all material available from Sweden, Australia and Spitzbergen (PERSSON 1954, 1959, 1960, 1962, 1962a); the major part of the material from the British Isles, France, Belgium and Luxembourg; some of the German spec imens; certain specimens from New Zealand, now in the British Museum (see LYDEK KER 1889, pp. -
A Large Rhomaleosaurid Pliosaur from the Upper Lias of Rutland Richard Forrest
A large Rhomaleosaurid Pliosaur from the Upper Lias of Rutland Richard Forrest Abstract: The fragmentary remains of a very large rhomaleosaurid pliosaur were retrieved during building works at Barnsdale Hall, Rutland. The limited material prevents clear identification at specific level, though on the basis of similarities of ratios of dimensions it shows closer affinity to Rhomaleosaurus arcuatus and R.victor than to R.cramptoni. Although scaling up from such fragmentary material is unreliable, the estimated length of this animal at 7.5 to 8 metres makes it possibly the largest Rhomaleosaurid pliosaur described to date. The fossil material broken end the shaft is oval in section, 148 mm wide and 96 mm deep. The head is 153 mm broad and The bones were excavated in 1988 by Mr. Roy 160 mm deep. Orientation can be determined by Draycott during construction of a retaining wall at rugosities from ridges for muscle attachment on the Barnsdale Hall, east of Rutland Water, in the county posterior side and the ventral surface. A deep hole in of the same name. An outer whorl of the ammonite the posterior muscle attachment presumably marks Hildoceras bifrons was found in association with the where a ligament was connected to the bone. There bones. It can therefore be placed with confidence in is slight taphonomic crushing around the trochanter. the bifrons Zone of the Upper Lias (Lower Jurassic, The surface is encrusted in places with a pyritised Toarcian, Whitbian). It is probable that much more deposit, which shows traces of tracks left by extensive remains of the animal were present at the scavengers post-mortem. -
How Plesiosaurs Swam: New Insights Into Their Underwater Flight Using “Ava”, a Virtual Pliosaur
Preprints (www.preprints.org) | NOT PEER-REVIEWED | Posted: 9 October 2019 doi:10.20944/preprints201910.0094.v1 How Plesiosaurs Swam: New Insights into Their Underwater Flight Using “Ava”, a Virtual Pliosaur Max Hawthorne1,*, Mark A. S. McMenamin 2, Paul de la Salle3 1Far From The Tree Press, LLC, 4657 York Rd., #952, Buckingham, PA, 18912, United States 2Department of Geology and Geography, Mount Holyoke College, South Hadley, Massachusetts, United States 3Swindon, England *Correspondence: [email protected]; Tel.: 267-337-7545 Abstract Analysis of plesiosaur swim dynamics by means Further study attempted to justify the use of all four flippers of a digital 3D armature (wireframe “skeleton”) of a simultaneously via the use of paddle-generated vortices, pliosauromorph (“Ava”) demonstrates that: 1, plesiosaurs which require specific timing to achieve optimal additional used all four flippers for primary propulsion; 2, plesiosaurs thrust. These attempts have largely relied on anatomical utilized all four flippers simultaneously; 3, respective pairs studies of strata-compressed plesiosaur skeletons, and/or of flippers of Plesiosauridae, front and rear, traveled through preconceived notions as pertains to the paddles’ inherent distinctive, separate planes of motion, and; 4, the ability to ranges of motion [8, 10-12]. What has not been considered utilize all four paddles simultaneously allowed these largely are the opposing angles of the pectoral and pelvic girdles, predatory marine reptiles to achieve a significant increase in which strongly indicate varied-yet-complementing relations acceleration and speed, which, in turn, contributed to their between the front and rear sets of paddles, both in repose and sustained dominance during the Mesozoic. -
Mosasaurs Continuing from Last Time…
Pliosaurs and Mosasaurs Continuing From Last Time… • Pliosauridae: the big marine predators of the Jurassic Pliosauridae • Some of the largest marine predators of all time, these middle Jurassic sauropterygians include such giants as Kronosaurus, Liopleurodon, Macroplata, Peloneustes, Pliosaurus, and Brachauchenius Pliosaur Mophology • While the number of cervical vertebrae is less than in plesiosaurs, there is still variation: Macroplata (29) vs. Kronosaurus (13) Pliosaur Morphology • Larger pliosaurs adopted a more streamlined body shape, like modern whales, with a large skull and compact neck, and generally the hind limbs were larger than the front, while plesiosaurs had larger forelimbs Pliosaur Morphology • Powerful limb girdles and large (banana sized) conical teeth helped pliosaurs eat larger, quicker prey than the piscivorous plesiosaurs Liopleurodon • NOT 25 m long in general (average of 40 feet), though perhaps certain individuals could reach that size, making Liopleurodon ferox the largest carnivore to ever live • Recent skull studies indicate that Liopleurodon could sample water in stereo through nostrils, locating scents much as we locate sound Cretaceous Seas • Breakup of Gondwana causes large undersea mountain chains to form, raising sea levels everywhere • Shallow seas encourage growth of corals, which increases calcium abundance and chalk formation • Warm seas and a gentle thermal gradient yield a hospitable environment to rays, sharks, teleosts, and the first radiation of siliceous diatoms Kronosaurus • Early Cretaceous -
By Natalie Lunis
by Natalie Lunis DINO_COVERS.indd 5 4/25/08 3:33:24 PM [Intentionally Left Blank] by Natalie Lunis Consultant: Luis M. Chiappe, Ph.D. Director of the Dinosaur Institute Natural History Museum of Los Angeles County 1630_OceanMonsters_FNL.indd 1 5/16/11 4:41:40 PM Credits Title Page, © Luis Rey; TOC, © Phil Wilson; 4-5, © John Sibbick; 6, © Phil Wilson; 7, © John Bindon; 8, © Spencer Brinker; 9, © Thomas Miller; 10, © Chris Butler/Photo Researchers, Inc.; 11, © John Bindon; 12, © Terry Mckee; 13, © by Karen Carr and Karen Carr Studio; 14T, © Pete Von Sholly; 14B, © Kristian Sekulic/Shutterstock; 15, © John Bindon; 16, © SeaPics.com; 17, © Tick Tock Media; 18, © Ken Lucas/Visuals Unlimited; 19, © Phil Wilson; 20, © Anness Publishing/The Natural History Museum, London; 21, © Tick Tock Media; 23TL, © Kat Ayer/Bearport Publishing; 23TR, © Falk Kienas/ Shutterstock; 23BL, © John Sibbick; 23BR, © Chris Butler/Photo Researchers, Inc. Publisher: Kenn Goin Editorial Director: Adam Siegel Creative Director: Spencer Brinker Design: Dawn Beard Creative Cover Illustration: Luis Rey Photo Researcher: Omni-Photo Communications, Inc. Library of Congress Cataloging-in-Publication Data Lunis, Natalie. Ocean monsters / by Natalie Lunis. p. cm. — (Dino times trivia) Includes bibliographical references and index. ISBN-13: 978-1-59716-713-0 (library binding) ISBN-10: 1-59716-713-4 (library binding) 1. Marine reptiles, Fossil—Juvenile literature. I. Title. QE861.5.L855 2009 567.9’37—dc22 2008010638 Copyright © 2009 Bearport Publishing Company, Inc. All rights reserved. No part of this publication may be reproduced in whole or in part, stored in any retrieval system, or transmitted in any form or by any means, electronic, mechanical, photocopying, recording, or otherwise, without written permission from the publisher. -
The First Record of Freshwater Plesiosaurian from the Middle
Gao et al. Journal of Palaeogeography (2019) 8:27 https://doi.org/10.1186/s42501-019-0043-5 Journal of Palaeogeography ORIGINALARTICLE Open Access The first record of freshwater plesiosaurian from the Middle Jurassic of Gansu, NW China, with its implications to the local palaeobiogeography Ting Gao, Da-Qing Li* , Long-Feng Li and Jing-Tao Yang Abstract Plesiosaurs are one of the common groups of aquatic reptiles in the Mesozoic, which mainly lived in marine environments. Freshwater plesiosaurs are rare in the world, especially from the Jurassic. The present paper reports the first freshwater plesiosaur, represented by four isolated teeth from the Middle Jurassic fluviolacustrine strata of Qingtujing area, Jinchang City, Gansu Province, Northwest China. These teeth are considered to come from one individual. The comparative analysis of the corresponding relationship between the body and tooth sizes of the known freshwater plesiosaur shows that Jinchang teeth represent a small-sized plesiosaurian. Based on the adaptive radiation of plesiosaurs and the palaeobiogeographical context, we propose a scenario of a river leading to the Meso-Tethys in the Late Middle Jurassic in Jinchang area, which may have provided a channel for the seasonal migration of plesiosaurs. Keywords: Freshwater plesiosaur, Middle Jurassic, Jinchang, Gansu Province, Palaeobiogeography 1 Introduction Warren 1980;Satoetal.2003; Kear 2012). Up to now, Plesiosaurs are one of the most familiar groups of Mesozoic the taxonomic affinities of most freshwater plesio- marine reptiles, which mainly lived in marine environ- saurs have remained unclear; some of them are re- ments. The records of plesiosaurs in non-marine deposits ferred to Plesiosauroidea (Cruickshank and Fordyce are sparse in comparison to those from marine sediments. -
A Cladistic Analysis and Taxonomic Revision of the Plesiosauria (Reptilia: Sauropterygia) F
Marshall University Marshall Digital Scholar Biological Sciences Faculty Research Biological Sciences 12-2001 A Cladistic Analysis and Taxonomic Revision of the Plesiosauria (Reptilia: Sauropterygia) F. Robin O’Keefe Marshall University, [email protected] Follow this and additional works at: http://mds.marshall.edu/bio_sciences_faculty Part of the Aquaculture and Fisheries Commons, and the Other Animal Sciences Commons Recommended Citation Frank Robin O’Keefe (2001). A cladistic analysis and taxonomic revision of the Plesiosauria (Reptilia: Sauropterygia). ). Acta Zoologica Fennica 213: 1-63. This Article is brought to you for free and open access by the Biological Sciences at Marshall Digital Scholar. It has been accepted for inclusion in Biological Sciences Faculty Research by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected], [email protected]. Acta Zool. Fennica 213: 1–63 ISBN 951-9481-58-3 ISSN 0001-7299 Helsinki 11 December 2001 © Finnish Zoological and Botanical Publishing Board 2001 A cladistic analysis and taxonomic revision of the Plesiosauria (Reptilia: Sauropterygia) Frank Robin O’Keefe Department of Anatomy, New York College of Osteopathic Medicine, Old Westbury, New York 11568, U.S.A Received 13 February 2001, accepted 17 September 2001 O’Keefe F. R. 2001: A cladistic analysis and taxonomic revision of the Plesio- sauria (Reptilia: Sauropterygia). — Acta Zool. Fennica 213: 1–63. The Plesiosauria (Reptilia: Sauropterygia) is a group of Mesozoic marine reptiles known from abundant material, with specimens described from all continents. The group originated very near the Triassic–Jurassic boundary and persisted to the end- Cretaceous mass extinction. This study describes the results of a specimen-based cladistic study of the Plesiosauria, based on examination of 34 taxa scored for 166 morphological characters. -
Novel Anatomy of Cryptoclidid Plesiosaurs with Comments on Axial Locomotion
NOVEL ANATOMY OF CRYPTOCLIDID PLESIOSAURS WITH COMMENTS ON AXIAL LOCOMOTION. A Thesis submitted to the Graduate College of Marshall University In partial fulfillment of The requirements for the degree of Master of Science In Biological Science by Benjamin C. Wilhelm Approved by Dr. F. Robin O’Keefe, Ph.D., Advisor, Committee Chairperson Dr. Brian L. Antonsen, Ph.D. Dr. Victor Fet, Ph.D. Dr. Suzanne G. Strait, Ph.D. Marshall University May 2010 Acknowledgements This thesis was greatly improved by the comments of my advisor Dr. F. R. O’Keefe and committee members Dr. B. Antonsen, Dr. V. Fet, and Dr. S. Strait. Description of the specimens presented in chapter 2 would not have been possible without the skilled preparation of J. P. Cavigelli. S. Chapman, H. Ketchum , and A. Milner arranged access to specimens at the BMNH. Chapter 2 was also improved by the comments of Dr. R. Schmeisser and an anonymous reviewer. This research was supported by a grant from the National Geographic Society (CRE 7627-04) and Marshall Foundation funds to Dr. F. R. O’Keefe and MURC student travel grants to B. C. Wilhelm. Many thanks are also due to A. Spriggs, my parents, George and Kris, and my brother Greg for all the love and support during the course of my Master’s studies. ii Table of Contents Acknowledgements…………………………………………………………………….……..ii Table of contents…………………………………………………………………………..iii-iv List of Figures, Tables, and Graphs………………………………………………………..v Abstract………………………………………………………………………………………...vi Chapter 1- Introduction to plesiosaurs………………………………………………...1-14 -
Anatomy and Systematics of the Rhomaleosauridae (Sauropterygia: Plesiosauria)
Anatomy and Systematics of the Rhomaleosauridae (Sauropterygia: Plesiosauria) Adam Stuart Smith BSc (hons) (Portsmouth), MSc (Bristol) A thesis submitted to the National University of Ireland, University College Dublin for the degree of Doctor of Philosophy November 2007 Supervisor: Dr Gareth J. Dyke Head of School: Professor Thomas Bolger School of Biology and Environmental Science University College Dublin Belfield Dublin 4 Ireland i Contents Frontispiece i Contents ii List of figures vi Acknowledgements xviii Declaration xx Abstract xxi Chapter 1. Introduction and objectives………………………………………….. 1 1.1 General introduction 1 1.2 Palaeobiology 4 1.3 Locomotion 7 1.4 Anatomy 10 1.5 Taxonomic diversity 12 1.6 Thesis objectives 14 1.7 Thesis structure 14 Chapter 2 - Historical background………………………………………………... 16 2.1 History of plesiosaurs 16 2.2 Plesiosaur systematics 16 2.3 Pliosauroidea 16 2.4 Rhomaleosauridae – taxon history 20 2.5 Rhomaleosauridae – previous research 24 2.6 Rhomaleosaurus or Thaumatosaurus? 25 Chapter 3 - Material and palaeontological approaches………………………. 27 3.1 Institutional abbreviations 27 3.2 Data collection –general 28 3.3 NMING F8785 Rhomaleosaurus cramptoni 30 3.3.1 History 30 3.3.2 Iconic specimen 33 3.4 NMING 10194 38 3.5 NMING F8749 38 3.6 NMNH R1336, NMING F8780, TCD.22931 Plesiosaurus 41 macrocephalus 3.7 NMING F8771 and TCD.22932 Thalassiodracon hawkinsi 41 3.8 BMNH 49202 45 3.9 BMNH R38525 Archaeonectrus rostratus 45 ii 3.10 BMNH R4853 Rhomaleosaurus thorntoni 45 3.11 BMNH R2028*, R2029*, R1317, R2061*, R2047*, R2027*, 49 R1318, R1319 and R2030* Eurycleidus arcuatus 3.12 BMNH R5488 Macroplata tenuiceps 50 3.13 BMNH R1310, TCD.47762a, TCD.47762b, 51 3.14 YORYM G503 Rhomaleosaurus zetlandicus 51 3.15 WM 851.S Rhomaleosaurus propinquus 54 3.16 SMNS12478 Rhomaleosaurus victor 54 3.17 LEICS G221.1851 54 3.18 WARMS G10875 56 3.19 TCD.57763 Attenborosaurus conybeari 56 3.20 Additional material 56 3.21 Possible rhomaleosaurids excluded from this study 58 Chapter 4 - Specimen descriptions………………………………………………. -
One Day in the Field.Pdf
ONE DAY IN THE FIELD This is a detailed account of our branch day-in-the-field, as seen through the eyes of our Branch Organiser, Mike Friday......Enjoy July 08, 2006 Introduction On July 8 2006, a good number of us met at the Bedford Museum for a talk and presentation by the Keeper of Natural History, Chris Andrew. This consisted of a potted history of the Oxford Clay Quest Brick-Pit at Stewartby, to the south west of Bedford, which was the location for our afternoon trip. Stories derived from a number of its workers, an insight into the kind of machinery used and the kinds of fossils that can be found at the site provided an entertaining morning. The essential details of the pit are explained by Chris, below, who very kindly supplied me with the text. The Jurassic Oxford Clay marine sediments extensively cover South East England from Dorset to Yorkshire. Although still appearing in the Late Jurassic as the Late Oxford Clay stage of the Oxfordian series, the majority occurs in the Middle Jurassic Callovian stage as the Early and Middle Oxford Clay, see Figure 1. Figure 1 The stratigraphy and age of the Oxford Clay Tom Miller, in Quest Pit Quest Pit (from Chris Andrew) Quest Pit is currently the only working Oxford Clay brick-pit in Bedfordshire. The pit exposes Lower Oxford Clay and the base of the Middle Oxford Clay; drainage trenches expose the very top of the more silty Kellaways Beds. The Oxford Clay is most famous for its vertebrate fauna.