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I I Rivista Italiana di Paleon roìor,iaeSrrltrqrriia I rolur ;;- -, | f;-' "oo, l UPPER OLENEKIAN (SPATHIAN) AMMONOIDS FROM CHIOS (LO\T/ER TRIASSIC, GREECE): TAXONOMY AND STRATIGRAPHIC POSITION. DOROTHEE MERTMANN' & VOLKER TACOBSHAGEN' Receh:ed July 1st,2002; accepted June 25,20A3 Key-raords: Ammonoidea, Triassic, Olenekian, Chios, Greece. There, the fossils are included in red limestones of rhe Hallstatt facies (Marmarorrapeza. Formation). Renz & Abstact. On the Greek island of Chios, Lower Triassic sections Renz (1948) have delivered rich ammonoid faunas from red limestones of the Hall- published a palaeontological description of statt facies. Here, we present a palaeontological description of Spatl-rian its ammonoids in a monograph. Bender (1970) was the ammonoids, which have been collected during two field campaigns by first to collect fossils from measured sections (CM i, kalian-German teams (Assereto et al. 1980; Gaetani et al. 1992). The II, III) in order to establish coordinared ammonoid and ammonoid assocìations are composed of Ceratitida and Phyllocerati- conodont stratigraphies. This author recognized, more- da, each order containing species of the Noritaceae, Hedenstroemia- ceae, Xenodiscaceae, Dinaritaceae, Pinacocerataceae, Phyllocerataceae, over, that the Spathian beds are overlain by lowermost Ptychitaceae, lJssuritaceae and Megaphyllitaceae. Long ranging gen- Anisian ones s/ith ammonoids comparable with faunas era, e.g. Procarnites and Leioplrylliter, are present as well as endemic of the so-called Hydaspian substage from the Himalayas ones, e.g. Cbiotites, which occurs exclusively in the Lower Trìassic of and Timor. Furthermore, Jacobshagen & Tietze (1974) Chios. The ammonoìd association is indicative of the Probungarites- described an associarion of both Spathian and Lower Subcolwmbites zone sensu Kumn.rel (1973a,). -
Paleontological Contributions
THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS May 15, 1970 Paper 47 SIGNIFICANCE OF SUTURES IN PHYLOGENY OF AMMONOIDEA JURGEN KULLMANN AND JOST WIEDMANN Universinit Tubingen, Germany ABSTRACT Because of their complex structure ammonoid sutures offer best possibilities for the recognition of homologies. Sutures comprise a set of individual elements, which may be changed during the course of ontogeny and phylogeny as a result of heterotopy, hetero- morphy, and heterochrony. By means of a morphogenetic symbol terminology, sutural formulas may be established which show the composition of adult sutures as well as their ontogenetic development. WEDEKIND ' S terminology system is preferred because it is the oldest and morphogenetically the most consequent, whereas RUZHENTSEV ' S system seems to be inadequate because of its usage of different symbols for homologous elements. WEDEKIND ' S system includes only five symbols: E (for external lobe), L (for lateral lobe), I (for internal lobe), A (for adventitious lobe), U (for umbilical lobe). Investigations on ontogenetic development show that all taxonomic groups of the entire superorder Ammonoidea can be compared one with another by means of their sutural development, expressed by their sutural formulas. Most of the higher and many of the lower taxa can be solely characterized and arranged in phylogenetic relationship by use of their sutural formulas. INTRODUCTION Today very few ammonoid workers doubt the (e.g., conch shape, sculpture, growth lines) rep- importance of sutures as indication of ammonoid resent less complicated structures; therefore, phylogeny. The considerable advances in our numerous homeomorphs restrict the usefulness of knowledge of ammonoid evolution during recent these features for phylogenetic investigations. -
Late Triassic and Early Jurassic Phylloceratids from the Salzkammergut (Northern Calcareous Alps)
©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at Gedenkband zum 100. Todestag von Dionys Stur Redaktion: Harald Lobitzer & Albert Daurer Jb. Geol. B.-A. ISSN 0016-7800 Band 136 Heft4 S.933-963 Wien, Dezember 1993 Late Triassic and Early Jurassic Phylloceratids from the Salzkammergut (Northern Calcareous Alps) By MILO~ RAKlJS*) With 19 Text-Figures and 5 Plates Oberösterreich Steiermark Satzkammergut Nördtiche Ka/ka/pen Ober/rias Osterreichische Kartei: 50.000 Lias Blätter 96,97 Ammoniten Contents Zusammenfassung 933 Abstract. .. 933 1. Introduction 933 2. Systematic Part 934 3. The Phyllogeny of the Late Triassic and Early Jurassic Phylloceratids and their Possible Relationship with Jurassic Ammonites 950 Acknowledgements. .. 952 References 962 Phylloceratiden aus der Obertrias und dem Unterlias des Salzkammergutes (Nördliche Kalkalpen) Zusammenfassung Gegenstand dieser Arbeit ist eine Revision der obertriadischen und liassischen phylloceratiden Ammoniten, die anhand des Originalmaterials von HAUER(1846), E. MOJSISOVICS(1873) and 1902), M. NEUMAYR(1879) und F. WÄHNER(1882-1898) durchgeführt wurde. Die beschriebenen Taxa werden in den Sammlungen der Geologischen Bundesanstalt und des Naturhistorischen Museums in Wien sowie der Bayerischen Staatssammlung in Mün- chen aufbewahrt. Außerdem wird das neue Ammonitengenus Togaticeras und Fergusonites neumayri sp.nov. beschrieben. Weiters werden phyllogenetische Probleme der obertriadischen und liassischen Phylloceratiden und deren mögliche Verwandtschaftsbeziehungen zu jurassischen Taxa diskutiert. Abstract This article deals with the reassessment of the Late Triassic and Early Jurassic Phylloceratid cephalopods which was carried out on the original specimens of F. HAUER(1846). E. MOJSISOVICS(1873 and 1902), M. NEUMAYR(1879) and F. WÄHNER(1882-1898). The subject specimens are deposited in the collections of GBA and NHM in Vienna (Austria) as well as in BSM in Munich (Germany). -
Stratigraphy and Ammonoids from the Middle Triassic Botneheia Formation (Daonella Shales) of Spitsbergen
Mitt. Geol.-Paläont. Inst. S. 27-54 Univ. Hamburg Heft 54 Hamburg, Dezember 1983 Stratigraphy and ammonoids from the Middle Triassic Botneheia Formation (Daonella Shales) of Spitsbergen WOLFGANG WEITSCHAT & ULRICH LEHMANN*) With plates 1-6, 2 tables and 9 text-figures Contents Abstract 27 Zusammenfassung 28 I. Introduction 28 II. Stratigraphy 28 a) Previous work 28 b) Descriptionof thesections 30 c) Age and correlation 33 III. Systematic Paleontology 36 IV. References 53 Abstract Sections of the Middle Triassic Daonella Shales (Botneheia Formation) of the Is- fjorden region of Spitsbergen are measured and their fossil contents of ammonoids are described and stratigraphically analyzed. A revised biostratigraphical Interpretation of this sequence is given. The Daonella Shales yield f ossilif erous nodule layers with dif f er- ent ammonoid f aunas which permit a threef old subdivision into the f ollowing ammonoid zonation: T T ^;„;^„ zone of Indigirites tozeri Lower Ladinian zpne pf Tsvetkovites vaHus Upper Anisian zone of Frechites laqueatus A correlation of this zonal sequence with the Middle Triassic beds of Taymyr Penin- sula, NE USSR, and NE British Columbia is discussed. Seven genera with eight species are described in detail. Tsvetkovites varius n. sp. and Indigirites tozeri n. sp. are new to science. A special attention is given to the ex- tremely high infraspecific Variation of the ammonoid faunas investigated. *) Address of authors: Dr. W. WEITSCHAT and Prof. Dr. U. LEHMANN, Geologisch-Paläon tologisches Institut und Museum der Universität Hamburg, Bundesstr. 55, D-2000 Hamburg 13, Germany (F.R.) 27 Zusammenfassung Profile der Mitteltriassischen Daonellen-Schiefer der Eisfjord-Region Spitzber gens wurden vermessen und die in ihnen enthaltenen Ammoniten beschrieben und stra- tigraphisch ausgewertet. -
Palaeontographica Beitrage Zur Naturgeschichte Der Vorzeit
SONDER-ABDRUCK aus PALAEONTOGRAPHICA BEITRAGE ZUR NATURGESCHICHTE DER VORZEIT Pal. A. Bd. 207 ANISIAN-LADINIAN BOUNDARY IN BOREAL REGION BASED ON AMMONOEDEA BY V. V. ARKADIEV and M. N. VAVILOV With 2 plates, 19 figures and 2 tables in the text STUTTGART E. SCHWEIZERBART'SCHE VERLAGSBUCHHANDLUNG (NAGELE u. OBERMILLER) 1989 Palaeontographica Abt. A 207 Lfg. 1-3 49-78 Stuttgart, April 1989 ANISIAN-LADINIAN BOUNDARY IN BOREAL REGION BASED ON AMMONOIDEA BY V. V. ARKADIEV and M. N. VAVILOV^ With 2 plates, 19 figures and 2 tables in the text Zusammenfassung Beschrieben werden die Ammonoideen der Anis-Ladin-Randablagerungen vom NE der UdSSR, die den Arten Longobardites, Indigirites, Eutomoceras, Frechites, Monophyllites, Arctoptychites angehoren. Untersucht wird die Morphologie der Schalen von Longobardites,, Indigirites in Queranschliffen sowie die stratigraphische Verteilung dieser Arten. Angegeben wird auch die ontogenetische Entwicklung der Lobenlinie. Eingehend wird die Lage der Grenze zwischen Anis und Ladin in den Borealgebieten betrachtet. Eine neue Moglichkeit der Grenzziehung zwischen Anis und Ladin wird angegeben. Schlusselworter: Ammonoidea — Trias — UdSSR — Biostratigraphie — Ontogenese. Summary The paper describes Ammonoidea from the Anisian-Ladinian boundary deposits in the North-East of the USSR, belonging to Longo- bardites, Indigirites, Eutomoceras, Frechites, Monophyllites, Arctoptychites. The morphology of shells belonging to Longobardites and Indigirites is studied in polished sections, stratigraphic distribution of these genera is analyzed and data on the ontogeny of their lobe line development are given. The location of the boundary between the Anisian and Ladinian stages in Boreal region is discussed in detail and a new version of its establishment is suggested. Key words: Ammonoidea — Triassic — USSR — ontogeny — biostratigraphy. -
64. the Akiyoshi Orogenic Cycle in the Mongolian Geosyncline*. by Teiichi KOBAYASHI
306 [Vol. 18, 64. The Akiyoshi Orogenic Cycle in the Mongolian Geosyncline*. By Teiichi KOBAYASHI. (Comm.by T. KATO,M.I.A., June 12, 1942.) Since Schonmann had expressed the opinion in 1929 that the Mongolio-Amur Faltengurtel had been a geosyncline which was disturbed by late Mesozoic orogenses, it was a moot question among geologists, as discussed by Stille, Obrutschew and othersl'. It is the object of this paper to demonstrate that the Akiyoshi, instead of the Sakawa, cycle of orgenesis2' is responsible for the oronization of the Mongolian geosyncline. A thick Palaeozoic complex for which I proposed the name , Mammo ( i) group3', is the sediment in the geosyncline existing between the Altenseheitel and the Yenshan orogenic zone. Because it is one of the least known but nevertheless one of the most important f ormational unit in the geology of Eastern Asia, I have paid special attention to it since I studied a small collecl,ion of fossils from Soron in the middle part of the Great Khingan range in 19314'. On the basis of fresh evidences which we obtained in Manchoukuo, its stratigraphic succession can now be roughly sketched as follows : 1. The lowest of the group so far known is the Ordovician Orthis sandstone of the Tyktaminda on the Siberian side of the Upper Amur valley where it is overlain by the Gotlandian Galymene manly shale. In the same area there is a good display of the fossiliferousDevonian formation composedmostly of shale and limestone in the lower part, of marl, limestone and reef limestone in the middle and of limestone in the upper parts). -
Middle Triassic)
GSSP of base of Ladinian The Global Boundary Stratotype Section and Point (GSSP) for the Base of the LADINIAN STAGE (Middle Triassic) A proposal for the GSSP at the base of the curionii zone in the Bagolino section (Southern Alps, Northern Italy) Peter Brack1, Hans Rieber2, Alda Nicora3 1Departement Erdwissenschaften, ETH-Zentrum, CH-8092 Zürich 2Paläontologisches Institut, Universität Zürich, CH-8006 Zürich 3Dipartimento di Scienze della Terra, Università di Milano, Via Mangiagalli 34, I-20133 Milano ABSTRACT the South Alpine Buchenstein Beds and siliceous The Global boundary Stratotype Section and Point limestones of Bakony (Hungary). Because (GSSP) for the Base of the LADINIAN STAGE Mojsisovics erroneously equated this interval with (Middle Triassic) is defined at the top of a distinctive parts of the ammonoid-rich Hallstatt-limestones, 20-cm-thick groove (= “Chiesense groove”) of he used the term “Norian” as the stage name. limestone nodules in a shaley matrix, located about “Norian” refers to the Norian Alps around Hallstatt 5 m above the base of the Buchenstein Beds in the near Salzburg (Austria). Later, the Hallstatt- Caffaro river bed (45°49ʼ09.5ʼʼN, 10°28ʼ15.5ʼʼE), ammonoids turned out to be much younger. Bittner south of the village of Bagolino (Province of (1892) therefore proposed the term “ladinisch” Brescia, northern Italy). The lower surface of (Ladinian) as a new label of the stratigraphic the overlying thick limestone bed has the lowest interval of South Alpine Buchenstein and Wengen occurrence of the ammonoid Eoprotrachyceras Beds. Although not accepted by Mojsisovics et al. curionii (base of the E. curionii zone; onset of the (1895), “Ladinian” was subsequently a generally Trachyceratidae ammonoid family). -
THE EVOLUTION of the CEPHALOPODA Bv L
THE EVOLUTIONOF THE CEPHALOPODA BY L. F. SPATH,D.Sc. [Rennrr.rruoeno*r BIOLOGICAL REVIEWS, Vot. VIII, No.4, Octonr.n19331 - CAMBRIDGE AT THE UNIVERSITYPRESS. THE EVOLUTION OF THE CEPHALOPODA Bv L. F. SPATH, D.Sc. (ReceiaedFebruary zz, 1933.) (With Thirteen Text-figures.) CONTENTS. PAGE I. Introduction +r8 II. Distinctions between Ammonoidea and Nautiloidea +2r (r) The initial chambers +2r (z) The coiling 42+ (3) The siphuncle 430 (4) Other characters 439 III. The ammonoid ancestor 4+2 IV. " Bactrites " and recapitulation 445 V. The primitive Cephalopoda 450 vI. The supposed cephalopod Volborthella 4s3 VII. The phylogeny of the Cephalopoda 455 VIII. Thb Dibranchiata 457 IX. bummary 459 References 46o I. INTRODUCTION. Tun development of the ammonites cannot be discussed independently of the evolution of their Palaeozoic forerunners, the goniatites, and the earlier types of these, again, are very intimately allied to the ancestral nautiloids. Although some- what diffident of casting so wide a net, I feel that the problem of the evolution of the Cephalopoda must be restated. In the past, what in my opinion are erroneous views of the development of the Ammonoidea have been popularisedlargelythrough the enthusiasm of workers who were attracted to the subject but had little practical experience of ammonites, and who too often were inclined to treat these fossils not as the remains of natural organisms but as material for speculation. This article, however, not only criticises the prevalent view that the ammonites, like the ancestral goniatites and Clymenia, arose from a straight nautiloid (Bactrites) and then hurried through the early stages of coiling before uncoiling again towards the end of their career; it also discusses various views put forward by workers on the primitive nautiloids. -
Lower to Middle Triassic) K@Ra Section, Albania
Geophys. J. Int. (1996) 127,503-514 Magnetobiostratigraphy of the Spathian to Anisian (Lower to Middle Triassic) K@ra section, Albania Giovanni Muttoni,'92Dennis V. Kent,' Selam Me~0,~Alda Nicora,2 Maurizio Gaetani,2 Marco Balini,2 Daniela Germani2 and Roberto Rettori4 ' Lamont-Doherty Earth Observatory, Palisades 10964 NY, USA 'Dipartimento di Scienze della Terra, tiia Mangiagalli 34,20133 Milan, Italy Fakultety Geolojia i Miniera, Polythekniki Unitiersiteti, Tirana, Albania Dipartimento di Scienze della Terra, Piazza Universita, 06100 Perugia, Italy Accepted 1996 July 18. Received in original form 1996 March 26 SUMMARY Magnetobiostratigraphic data are presented from three Early/Middle Triassic Han- Bulog Limestone successions from Kqira, northern Albania. A total of 206 standard palaeomagnetic samples were obtained for thermal demagnetization and statistical analysis from the 42, 10 and 5m thick sections. The reversal-bearing characteristic component, carried by haematite and magnetite, defines a composite sequence of six main polarity intervals (Kqln to Kq3r) in which are embedded four short polarity intervals, one at the base of Kqln and three towards the top of Kqlr. The early acquisition of the characteristic remanence is supported by the lateral correlation of magnetozones between sections. The Early/Middle Triassic boundary, approximated by the first occurrence of the conodont Chiosella timorensis, falls close to the Kqlr/Kq2n polarity transition. This is in good agreement with recently published magneto- biostratigraphic data from the coeval Chios (Greece) sections. The palaeomagnetic pole calculated from the Kqira characteristic directions lies close to the Triassic portion of the apparent polar wander path for Laurussia (in European coordinates). However, a 40-45" clockwise rotation of the external zone of the Albano-Hellenic Belt to the south of the Scutari-Pec Line is thought to have occurred since the Early-Middle Miocene. -
Department of the Interior U.S. Geological Survey To
DEPARTMENT OF THE INTERIOR TO ACCOMPANY MAP MF-ZOO7 U.S. GEOLOGICAL SURVEY FOSSIL LOCALITY MAP AND FOSSIL DATA FOR THE SOUTHEASTERN PART OF THE CHARLEY RIVER QUADRANGLE. EAST-CENTRAL ALASKA Compiled by Ronny T. Miyaoka INTRODUCTION This table and location map are compiled from fossil information collected within the Charley River A-1. A-2, A-3. 6-1, 8-2, 8-3, C-I, and C-2 1:63.360-scale quadrangles. Sources of information for the campilation are mainly published reports of the U.S. Geological Survey, as well as unpublished fieldnotes. maps. and fossil reports of USGS workers conducting geologic studies in the area. The fossil data in the table are listed chronologfcally, by collector, according to when the fossils were collected. This information includes, where available, map number. geologic map unit, collector's field number, USGS collection number, fossil list, and fossil age designation. Sources of information and names of paleontologists Hho identified fossils are keyed with superscripts and cited at the end of the table. Reports for most of the collections that have been assigned a USGS collection number are on file at the U.S. National Museum of Natural History in Washington. D.C. The fossil lists are reproduced as originally prepared by the paleontologists indicated. However, many of the older collections have been restudied by later workers, and because of improved taxonomic and stratigraphic frameworks, restudy has resulted in revised identification, correlation, and(or) age designation for many collections. Consequently, care should be taken in using the appraisals of early workers and most attention should be given to the more current reports and publications. -
The Triassic Marine Biota of Eastern Indonesia and Its Interregional and Global Correlation: a Review
Jurnal Geologi Indonesia, Vol. 5 No. 1 Maret 2010: 31-47 The Triassic Marine Biota of Eastern Indonesia and its Interregional and Global Correlation: A Review F. HASIBUAN Geological Survey Institute Jln. Diponegoro 57, Bandung 40122, Indonesia ABSTRACT The interregional and global correlation of the Triassic biota of Indonesia was based on the review of previous workers and the author himself. Scythian Epoch (Early Triassic) in Timor is subdivided into Early Scythian with Ophiceras demisso, Meekoceras sp., Pseudomonotis subaurita, Gervillia subpannonica, and Myophoria sp., whilst Late Scythian is indicated by the presence of Owenites egrediens and Sibirites sp. The presence of Anisian Stage (Middle Triassic) in Misool is indicated by ammonite Beyrichites and bivalve Daonella lilintana. In Timor, this stage is pointed out by the presence of Joannites cymbiformis, Monophyl- lites wengensis, Protrachyceras archaelus, Daonella indica, Tracyceras cf. aon, Brochidium timorense, and Lima subpunctatoides. Terebellina mackayi found above Beyrichites-bearing bed in Misool has an age range from Anisian to Ladinian. It is concluded that the boundary between Anisian and Ladinian lies between beds with Beyrichites and Terebellina mackayi. Early Carnian Stage (Late Triassic) in Timor is indicated by the presence of Joanites cymbiformis, Waldhausenites sp., Miltites sp., and Halogyra cipitiensis; whereas Late Carnian is indicated by the presence of Cladicites crassestriatus and Tropites subbulatus. The presence of Halobia verbeeki, Pinacoceras parma, Neobetites sp., Parabetites sp., Malayites sp., Amarassites sp., and Halorites sp., indicates the Early Norian Stage, whilst the presence of Cladiscites tornatus, Cyrtopleurits malayicus, and Trachypleuraspidites sp. implies the Late Norian. The Rhaetian Stage in Timor contains Choristoceras indoaustralicum, whereas in Misool it contains Choristoceras sp. -
The Ammonoid Recovery After the End−Permian Mass Extinction: Evidence from the Iran−Transcaucasia Area, Siberia, Primorye, and Kazakhstan
The ammonoid recovery after the end−Permian mass extinction: Evidence from the Iran−Transcaucasia area, Siberia, Primorye, and Kazakhstan YURI D. ZAKHAROV and NASRIN MOUSSAVI ABNAVI Zakharov, Y.D. and Moussavi Abnavi, N. 2013. The ammonoid recovery after the end−Permian mass extinction: Evidence from the Iran−Transcaucasia area, Siberia, Primorye, and Kazakhstan. Acta Palaeontologica Polonica 58 (1): 127–147. Investigations of the Upper Permian strata in the Iran−Transcaucasia resulted in identification of 32 ammonoid genera. The majority of ammonoids in this collection belong to the order Ceratitida (75%). Among Dzhulfian ceratitid ammonoids representatives of the family Araxoceratidae (Otoceratoidea) are most abundant. The assemblage structure changed radically during latest Permian (Dorashamian) time, bringing a domination of the family Dzhulfitidae. The Induan (Lower Triassic) succession in the Verkhoyansk area provided a few groups of ammonoids which are Palaeozoic in type: families Episageceratidae (Episageceras), Xenodiscidae (Aldanoceras and Metophiceras), and Dzhulfitidae (Tompophiceras) and superfamily Otoceratoidea (Otoceras and Vavilovites). It demonstrates the survival of ammonoids belonging to these groups the Permian–Triassic (P–T) boundary extinction event and their quick migration to the vast ar− eas of higher latitudes (together with some representatives of the Mesozoic−type families). Induan–Olenekian ammonoid successions in South Primorye, Mangyshlak, and Arctic Siberia illustrate the high rate of Early Triassic ammonoid recov− ery in both the Tethys and the Boreal realm. New ammonoid taxa are described: Proptychitina subordo nov., Ussuritina subordo nov., Subbalhaeceras shigetai gen. and sp. nov. (Flemingitidae), Mesohedenstroemia olgae sp. nov. (Heden− strormiidae), and Inyoites sedini sp. nov. (Inyoitidae). Key words: Ammonoidea, recovery, Permian, Triassic, Russia, Azerbaijan, Kazakhstan, Iran.