Holism and Evolution
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Rhythms of the Brain
Rhythms of the Brain György Buzsáki OXFORD UNIVERSITY PRESS Rhythms of the Brain This page intentionally left blank Rhythms of the Brain György Buzsáki 1 2006 3 Oxford University Press, Inc., publishes works that further Oxford University’s objective of excellence in research, scholarship, and education. Oxford New York Auckland Cape Town Dar es Salaam Hong Kong Karachi Kuala Lumpur Madrid Melbourne Mexico City Nairobi New Delhi Shanghai Taipei Toronto With offices in Argentina Austria Brazil Chile Czech Republic France Greece Guatemala Hungary Italy Japan Poland Portugal Singapore South Korea Switzerland Thailand Turkey Ukraine Vietnam Copyright © 2006 by Oxford University Press, Inc. Published by Oxford University Press, Inc. 198 Madison Avenue, New York, New York 10016 www.oup.com Oxford is a registered trademark of Oxford University Press All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted, in any form or by any means, electronic, mechanical, photocopying, recording, or otherwise, without the prior permission of Oxford University Press. Library of Congress Cataloging-in-Publication Data Buzsáki, G. Rhythms of the brain / György Buzsáki. p. cm. Includes bibliographical references and index. ISBN-13 978-0-19-530106-9 ISBN 0-19-530106-4 1. Brain—Physiology. 2. Oscillations. 3. Biological rhythms. [DNLM: 1. Brain—physiology. 2. Cortical Synchronization. 3. Periodicity. WL 300 B992r 2006] I. Title. QP376.B88 2006 612.8'2—dc22 2006003082 987654321 Printed in the United States of America on acid-free paper To my loved ones. This page intentionally left blank Prelude If the brain were simple enough for us to understand it, we would be too sim- ple to understand it. -
A Limiting Factor
Published on Plants in Action (http://plantsinaction.science.uq.edu.au/edition1) Home > Printer-friendly > Printer-friendly Chapter 15 - Water: a limiting factor [1] A superb stand of flooded gums, (Eucalyptus grandis) near Coffs Habour, northern New South Wales, 'each tall because of each' (Les Murray (1991), Collected Poems) (Photograph by Ken Eldridge, supplied by Peter Burgess, CSIRO Forestry and Forest Products) With perspective phrasing, Les Murray (1991) summarises structural aspects of a gum forest as: 'Flooded gums on creek ground, each tall because of each' and on conceptualising water relations, 'Foliage builds like a layering splash: ground water drily upheld in edge-on, wax rolled, gall-puckered leaves upon leaves. The shoal life of parrots up there.' (Les Murray, Collected Poems, 1991) Introduction Life-giving water molecules, fundamental to our biosphere, are as remarkable as they are abundant. Hydrogen bonds, enhanced by dipole forces, confer extraordinary physical properties on liquid water that would not be expected from atomic structure alone. Water has the strongest surface tension, biggest specific heat, largest latent heat of vaporisation and, with the exception of mercury, the best thermal conductivity of any known natural liquid. A high specific grav-ity is linked to a high specific heat, and very few natural substances require 1 calorie to increase the temperature of 1 gram by 1ºC. Similarly, a high heat of vaporisation means that 500 calories are required to convert 1 gram of water from liquid to vapour at 100ºC. This huge energy requirement (latent heat of vaporisation, Section 14.5) ties up much heat so that massive bodies of water contribute to climatic stability, while tiny bodies of water are significant for heat budgets of organisms. -
Interpreting the History of Evolutionary Biology Through a Kuhnian Prism: Sense Or Nonsense?
Interpreting the History of Evolutionary Biology through a Kuhnian Prism: Sense or Nonsense? Koen B. Tanghe Department of Philosophy and Moral Sciences, Universiteit Gent, Belgium Lieven Pauwels Department of Criminology, Criminal Law and Social Law, Universiteit Gent, Belgium Alexis De Tiège Department of Philosophy and Moral Sciences, Universiteit Gent, Belgium Johan Braeckman Department of Philosophy and Moral Sciences, Universiteit Gent, Belgium Traditionally, Thomas S. Kuhn’s The Structure of Scientific Revolutions (1962) is largely identified with his analysis of the structure of scientific revo- lutions. Here, we contribute to a minority tradition in the Kuhn literature by interpreting the history of evolutionary biology through the prism of the entire historical developmental model of sciences that he elaborates in The Structure. This research not only reveals a certain match between this model and the history of evolutionary biology but, more importantly, also sheds new light on several episodes in that history, and particularly on the publication of Charles Darwin’s On the Origin of Species (1859), the construction of the modern evolutionary synthesis, the chronic discontent with it, and the latest expression of that discon- tent, called the extended evolutionary synthesis. Lastly, we also explain why this kind of analysis hasn’t been done before. We would like to thank two anonymous reviewers for their constructive review, as well as the editor Alex Levine. Perspectives on Science 2021, vol. 29, no. 1 © 2021 by The Massachusetts Institute of Technology https://doi.org/10.1162/posc_a_00359 1 Downloaded from http://www.mitpressjournals.org/doi/pdf/10.1162/posc_a_00359 by guest on 30 September 2021 2 Evolutionary Biology through a Kuhnian Prism 1. -
3 Limiting Factors and Threats
LOWER COLUMBIA SALMON RECOVERY & SUBBASIN PLAN December 2004 3 Limiting Factors and Threats 3 LIMITING FACTORS AND THREATS........................................................................3-1 3.1 HABITAT –STREAMS .....................................................................................................3-2 3.1.1 Background..........................................................................................................3-2 3.1.2 Limiting Factors...................................................................................................3-3 3.1.3 Threats................................................................................................................3-22 3.2 ESTUARY AND LOWER MAINSTEM HABITAT ..............................................................3-26 3.2.1 Background........................................................................................................3-26 3.2.2 Limiting Factors.................................................................................................3-27 3.2.3 Threats................................................................................................................3-36 3.3 HABITAT – OCEAN ......................................................................................................3-38 3.3.1 Background........................................................................................................3-38 3.3.2 Limiting Factors.................................................................................................3-38 -
Population Growth Is a Critical Factor in Specie’S Ability to Maintain Homeostasis Within Its Environment
Chapter 4-B1 Population Ecology Population growth is a critical factor in specie’s ability to maintain homeostasis within its environment. Main themes ~Scientists study population characteristics to better understand growth and distribution of organisms. ~Populations have different distributions and densities depending on the species represented in the population ~Organisms in a population compete for energy sources, such as food and sunlight ~Homeostasis within a population is controlled by density-dependent and density-independent limiting factors ~Human population varies little overall, but can change greatly within small populations. I. Population dynamics A. 1. Population density a. defined- b. eg. deer in Newark deer in 5 mi. sq. area 2. a. defined-organism pattern in area (eg. more deer on Londondale than on Green Wave Drive) b. c. Types p. 93 -UNIFORM dispersion-relatively even distribution of a species. Eg. pine forest -CLUMPED dispersion-organism are clumped near food. Eg. schools of fish, eg. mushrooms on a fallen log -_________________-unpredictable & changing. Eg. deer, large fish, etc. 3. a. area where organisms CAN/ARE living b. Meaning of Distribution -in Science~it is area where something lives and reproduces -in everyday language ~it means distribution (passing out) eg. of report cards. B. 1. Limiting factors-biotic or abiotic factors that keep a population from increasing forever (indefinitely). a. b. amount of space=abiotic c. 2. Density-independent factors a. b. independent=doesn’t rely on the number of organisms, would happen regardless of the number of organisms c. it is usually abiotic (limits population growth) -flood (eg. no matter how many deer in an area) - -blizzard - -fire (Ponderosa pine needs fire to kill undergrowth which takes all the soil nutrients from them. -
Macroevolutionary Diversification Rates Show Time Dependency
Macroevolutionary diversification rates show time dependency L. Francisco Henao Diaza,b, Luke J. Harmonc, Mauro T. C. Sugawaraa,b, Eliot T. Millerd, and Matthew W. Pennella,b,1 aDepartment of Zoology, University of British Columbia, Vancouver, BC, Canada V6T 1Z4; bBiodiversity Research Centre, University of British Columbia, Vancouver, BC, Canada V6T 1Z4; cDepartment of Biological Sciences, University of Idaho, Moscow, ID 83844; and dCornell Lab of Ornithology, Cornell University, Ithaca, NY 14850 Edited by David M. Hillis, The University of Texas at Austin, Austin, TX, and approved February 28, 2019 (received for review October 22, 2018) For centuries, biologists have been captivated by the vast disparity time dependence, with the fastest rates estimated over the shortest in species richness between different groups of organisms. time scales. This time scale dependence is apparent in rates of Variation in diversity is widely attributed to differences between molecular (12) and trait evolution (13, 14) and has even been groups in how fast they speciate or go extinct. Such macroevolu- observed when estimating long-term rates of sedimentation (15). tionary rates have been estimated for thousands of groups and The prevalence of time scaling of rate estimates in other types of have been correlated with an incredible variety of organismal data, along with previous hints of a possible similar pattern in traits. Here we analyze a large collection of phylogenetic trees and speciation rates (16–20), suggest that a broader comparison is fossil time series and describe a hidden generality among these needed. Here we explore the time scaling of diversification rates seemingly idiosyncratic results: speciation and extinction rates using both phylogenetic data from the tree of life and paleobio- follow a scaling law in which both depend on the age of the group logical data from the fossil record and find evidence that there are in which they are measured, with the fastest rates in the youngest indeed general scaling laws that govern macroevolutionary dy- clades. -
Memory, History, and the Preservation of Archival Records*
The Past that Archives Keep: Memory, History, and the Preservation of Archival Records* BRIEN BROTHMAN RÉSUMÉ Est-ce que « le passé » prend des sens différents dans les contextes de l’histoire et de la mémoire? Est-ce que la réponse à cette question a une relation avec les archives? Cet article répond par l’affirmative à ces deux questions. Son argument principal est que d’attribuer un sens distinct au « passé » dans le cadre de la mémoire permet le développement d’une perspective sur le travail archivistique qui rehausse la valeur des documents anciens pour les organisations et la société contemporaines. Le raisonnement de l’auteur se dévoile en trois sections. La première veut démontrer que certains éléments du concept australien de continuum des documents sont plus compat- ibles avec l’idée de la mémoire sociale et organisationnelle que ne l’est celui du cycle de vie des documents et que de plus, sur un plan archivistique, le continuum des docu- ments est un concept plus cohérent au point de vue temporel que le cycle de vie des documents. La deuxième section se base sur des recherches de différentes disciplines pour démontrer que, dans le cadre de la mémoire, faire référence au passé représente simplement une autre façon de parler du présent. La troisième section se fonde sur les idées des deux premières pour proposer dix questions d’ordre conceptuel, organisation- nel et technologique qui requièrent l’attention en ce qui concerne les programmes de conservation à long terme des documents d’archives. ABSTRACT Does “the past” take on different meanings in the contexts of history and memory? Does the answer to this question have any bearing on archives? This arti- cle answers affirmatively to both these questions. -
Holism Agriculture and Natural Resources
Holism Agriculture and Natural Resources An Inaugural Lecture Given in the University of Rhodesia by Professor J. M. de Yilliers UNIVERSITY OF RHODESIA l.'nirersity of Rhodesia Printed in Rhodesia by Sebri Printers, Martin Drive, Msasa, Salisbury HOLISM, AGRICULTURE, AND NATURAL RESOURCES I am unable to resist the temptation to recall for you tonight what Mark Prestwich, on acceding to the Chair of History and Political Science in the University of Natal, had to say about the Inaugural Lecture — this ordeal to which new Professors are subjected. The occasion, said Professor Prestwich, affords his learned colleagues the opportunity to enjoy one of the pleasures of malice — the pleasure of noting his superficiality, his deficiencies of oratory and power of reasoning, the abysmal deeps of his ignorance. His students are able to taste revenge for what he has done or will do to them, and the lay public is treated to a display of academic nakedness which bears remarkable similarities to the initiation rites practised by primitive tribes. The purpose of all this, no doubt, is to ensure that he will not become too big for his boots. And there is the prospect of direr perils still. Such as the fate which befell a certain ninth-century philosopher — one Johannes Scotus Erigena — who so exasperated his audience that they rose up and slew him with their styluses. One is told also of Nicolas Cop who was compelled to flee for his life after his Inaugural Lec ture as Rector of the University of Paris in 1533. And of Ernest Renan whose Inaugural Lecture as Professor of Hebrew in 1862 led to his instant suspension from the Chair. -
Successive Range Expansion Promotes Diversity and Accelerates Evolution in Spatially Structured Microbial Populations
The ISME Journal (2017) 11, 2112–2123 © 2017 International Society for Microbial Ecology All rights reserved 1751-7362/17 www.nature.com/ismej ORIGINAL ARTICLE Successive range expansion promotes diversity and accelerates evolution in spatially structured microbial populations Felix Goldschmidt1,2, Roland R Regoes1 and David R Johnson2 1Department of Environmental Systems Science, ETH Zürich, Zürich, Switzerland and 2Department of Environmental Microbiology, Eawag, Dübendorf, Switzerland Successive range expansions occur within all domains of life, where one population expands first (primary expansion) and one or more secondary populations then follow (secondary expansion). In general, genetic drift reduces diversity during range expansion. However, it is not clear whether the same effect applies during successive range expansion, mainly because the secondary population must expand into space occupied by the primary population. Here we used an experimental microbial model system to show that, in contrast to primary range expansion, successive range expansion promotes local population diversity. Because of mechanical constraints imposed by the presence of the primary population, the secondary population forms fractal-like dendritic structures. This divides the advancing secondary population into many small sub-populations and promotes intermixing between the primary and secondary populations. We further developed a mathematical model to simulate the formation of dendritic structures in the secondary population during succession. By introducing mutations in the primary or dendritic secondary populations, we found that mutations are more likely to accumulate in the dendritic secondary populations. Our results thus show that successive range expansion can promote intermixing over the short term and increase genetic diversity over the long term. Our results therefore have potentially important implications for predicting the ecological processes and evolutionary trajectories of microbial communities. -
Shelford's Law & Concept of Limiting Factor
Shelford’S Law & Concept of Limiting Factor Dr. Md. Osaid Alam Guest Faculty (Environmental Science & Management) Department of Zoology Patna University 1 Shelford’s Law of Intolerance • Shelford's law of tolerance is a principle developed by American zoologist Victor Ernest Shelford in 1911. • Shelford's law of tolerance states that an organism’s success is based on a complex set of conditions and that each organism has a certain minimum, maximum, and optimum environmental factor or combination of factors that determine success. • According to the law of tolerance, populations have optimal survival conditions within critical minimal and maximal thresholds. • As population is exposed to the extremes of a particular limiting factor, the rates of survival begin to drop. 2 Shelford’s Law of Intolerance • The distribution of a species in response to a limiting factor can be represented as a bell-shaped curve with three distinct regions: 1. Optimal zone: Central portion of curve which has conditions that favour maximal reproductive success and survivability. 2. Zones of stress: Regions flanking the optimal zone, where organisms can survive but with reduced reproductive success. 3. Zones of intolerance: Outermost regions in which organisms cannot survive (represents extremes of the limiting factor). 3 Shelford’s Law of Intolerance • According to the Shelford’s law of tolerance, there are upper and lower threshold value on the gradient beyond which the species cannot survive. Tolerance range differ for one species to another. 4 Definition of Limiting Factor • A limiting factor is a resource or environmental condition which limits the growth, distribution or abundance of an organism or population within an ecosystem. -
The Structure of Evolutionary Theory: Beyond Neo-Darwinism, Neo
1 2 3 The structure of evolutionary theory: Beyond Neo-Darwinism, 4 Neo-Lamarckism and biased historical narratives about the 5 Modern Synthesis 6 7 8 9 10 Erik I. Svensson 11 12 Evolutionary Ecology Unit, Department of Biology, Lund University, SE-223 62 Lund, 13 SWEDEN 14 15 Correspondence: [email protected] 16 17 18 19 1 20 Abstract 21 The last decades have seen frequent calls for a more extended evolutionary synthesis (EES) that 22 will supposedly overcome the limitations in the current evolutionary framework with its 23 intellectual roots in the Modern Synthesis (MS). Some radical critics even want to entirely 24 abandon the current evolutionary framework, claiming that the MS (often erroneously labelled 25 “Neo-Darwinism”) is outdated, and will soon be replaced by an entirely new framework, such 26 as the Third Way of Evolution (TWE). Such criticisms are not new, but have repeatedly re- 27 surfaced every decade since the formation of the MS, and were particularly articulated by 28 developmental biologist Conrad Waddington and paleontologist Stephen Jay Gould. 29 Waddington, Gould and later critics argued that the MS was too narrowly focused on genes and 30 natural selection, and that it ignored developmental processes, epigenetics, paleontology and 31 macroevolutionary phenomena. More recent critics partly recycle these old arguments and 32 argue that non-genetic inheritance, niche construction, phenotypic plasticity and developmental 33 bias necessitate major revision of evolutionary theory. Here I discuss these supposed 34 challenges, taking a historical perspective and tracing these arguments back to Waddington and 35 Gould. I dissect the old arguments by Waddington, Gould and more recent critics that the MS 36 was excessively gene centric and became increasingly “hardened” over time and narrowly 37 focused on natural selection. -
AVID for Higher Education: High Engagement Practices for Teaching and Learning
AVID for Higher Education: High Engagement Practices for Teaching and Learning DEVELOPED BY: Debra Shapiro, J.D. Joseph Cuseo, Ph.D. I DEVELOPED WITH SUPPORT FROM: Instructional Strategists: Rachel Juarez-Torres, Ed.D. Ben Solomon Shannon McAndrews, Ed.D. Project Lead: Jaime Rae Sandoval Contributors: Julia Howe Alma Montemayor Sandigo, Ed.D. Gloria Pindi, Ph.D. Aaron Thompson, Ph.D. Editor: Moon Wilkman Proofreaders: Mariyama Scott Megan Zid Graphic Designer: Tracy Castle Copyright © 2018 AVID Center • San Diego, CA All rights reserved. HEPTL-20171221 II AVID for Higher Education: High Engagement Practices for Teaching and Learning Table of Contents AVID History and Philosophy ........................................................................V Foreword by Dr. Joseph Cuseo ............................................................... VIII Chapter 1: Relational Capacity: The Importance of Building Positive Student–Instructor and Student–Student Relationships ...................................... 1 Chapter 2: Writing to Learn ...................................................................... 29 Chapter 3: Inquiry ............................................................................................57 Chapter 4: Collaboration .......................................................................... 103 Chapter 5: Organization of Thought and Materials ................. 123 Chapter 6: Reading to Learn .................................................................. 153 Resources ..........................................................................................................205