Jonathan M. Brown Department of Biology, Noyce Science Center Grinnell College Grinnell, IA Voice: (641) 269-3096 e-mail: [email protected] http://web.grinnell.edu/individuals/brownj

Education

Ph.D., September 1989 Michigan State University, East Lansing, MI. Dept. of Zoology and W.K. Kellogg Biological Station Interdepartmental Graduate Specialization in Ecology and Evolutionary Biology

B.A. magna cum laude, 1983 Carleton College, Northfield, MN. Major: Biology Honors: -Phi Beta Kappa -Distinction in Biology -Dean's List (3 years) -Certificate of Superior Achievement in German Language and Area Studies

Positions held

8/08- Professor of Biology, Grinnell College, IA. 6/02-6/04 Biology Department Chair, Grinnell College, IA. 9/00-08. Associate Professor, Department of Biology, Grinnell College, IA. 8/95-00 Assistant Professor, Department of Biology, Grinnell College, IA. 6/94-6-95 Visiting Assistant Professor, Department of Biology, Bucknell University. 6/91-5/94 David Burpee Post-doctoral Associate, Department of Biology, Bucknell University. 12/89-6/91 Post-doctoral Research Associate, Section of Ecology and Systematics, Cornell University.

Recent Course Titles

Bio 150 - Introduction to Biological Inquiry: Prairie Restoration Bio 150 - Introduction to Biological Inquiry: Sexy Beast Bio 252 - Organisms, Evolution, and Ecology Bio 240 - Behavior Bio 301 - History of Biological Thought Bio 373 - Mechanisms of Evolution Bio 297 – The Ecology of Place in Great Britain (Grinnell-in-London) Tutorial – Envisioning Nature

Scholarships, Fellowships and Grants

2015 Collaborative Research: RUI: Evolution of color variation in Hawaiian damselflies: causal links for an ecological selection hypothesis. [PIs: J.M. Brown and I.A. Cooper] National Science Foundation Division of Environmental Biology, Evolutionary Processes Cluster. $189,057 (Grinnell portion).

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2005 RUI: Generating the entangled bank: Biogeography and host use in a Hawaiian herbivore radiation. [PI: J. M. Brown] National Science Foundation Division of Environmental Biology, Systematic Biology and Biodiversity Investigations Cluster. $185,350. (through 4/2010)

2003 MRI/RUI: Acquisition of instrumentation for nutrient analyses in ecological research. National Science Foundation [PI: Peter Jacobson; CO-PI’s: Kathy Jacobson, Vince Eckhart and Jonathan Brown]. $135,887.

2000 ROA: Evolution of Wing Patterns in Hawaiian Picturewing Flies. National Science Foundation Population Biology Program. [PI: Donald Price; supported J. Brown’s collaboration] $16, 332

00-01 RUI: Acquisition of a capillary electrophoresis system for DNA analysis in investigations of adaptation, speciation, species interactions and DNA repair." National Science Foundation Multi-user Equipment and Instrumentation Resources for Biological Sciences Program. $67,351 (Co-Author and Principal Investigator)

99-02 "Reform of Undergraduate Biology Education: Biological Inquiry and Integrative Biology" Co- author and co-PI. National Science Foundation Curriculum, Classroom and Laboratory Improvement (CCLI) grant. $150,000.

95-7 "The role of ecological interactions in diversification: rates of habitat shifts and within-habitat diversification in the Enallagma damselflies of two communities." Co-P.I. with Dr. Mark A. McPeek, Dartmouth College. National Science Foundation. $70,000.

95-96 “Supplement to host expansion and speciation in phytophagous (Host association patterns of cynipid gallmakers and oaks).” Co-PI’s Warren Abrahamson (Bucknell University) and George Melika (Uzhgorod State University, Ukraine). National Science Foundation, Division of International Programs/Research in Undergraduate Institutions. $10,000.

85-88 National Science Foundation Graduate Fellowship, Michigan State University.

Publications in preparation/revision

Messing, R., A. Vorsino, J.M. Brown, and D. Rubinoff. Distribution and Conservation Status of Endemic Tephritidae of the Hawaiian Islands. Conservation and Diversity. In preparation.

Rasmussen, R., P. Cook, J.M. Brown, and I.A. Cooper. Male harassment and female color dimorphism in Megalagrion calliphya damselflies.

Mamantov, M., R. Rasmussen, E. Hsieh, L. Lyons and J.M. Brown. Love thy neighbor: Cooperation without kinship in a native, unicolonial ant species (Formica exsectoides).

Publications (undergrad student co-authors)

Cooper, I. A., Brown, J. M. and Getty, T. (2016), A role for ecology in the evolution of colour variation and sexual dimorphism in Hawaiian damselflies. J. Evol. Biol., 29: 418–427. doi:10.1111/jeb.12796

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Viale, E., I. Martinez-Sañudo, J.M. Brown, M. Simonato, A. Squartini, A. Bressan, M. Faccoli , L. Mazzon. 2015. Pattern of association between endemic Hawaiian fruit flies (Diptera, Tephritidae) and their symbiotic bacteria: evidence of cospeciation events and proposal of “Candidatus Stammerula trupaneae.” Molecular Phylogenetics and Evolution. 90: 67–79.

Mittelbach, G.G., D. Schemske, H.V. Cornell, A.P. Allen,. J. M. Brown, M. Bush, S.P. Harrison, A. Hurlbert, N. Knowlton, H. A. Lessios, C.M. McCain, A.R. McCune, L.A. McDade, M.A. McPeek, T.J. Near, T.D. Price, R.E. Ricklefs, K. Roy, D.F. Sax, D. Schluter, J.M. Sobel, M. Turelli. 2007. Evolution and the latitudinal diversity gradient: Speciation, extinction, and biogeography. Ecology Letters 10: 315-331.

McPeek, M.A. and J.M. Brown. 2007. Clade age and not diversification rate explains species richness among animal taxa. American Naturalist 169:E97-106.

Heard, S.B, J.O. Stireman, J.D. Nason, G.H. Cox, C. Kolacz, and J.M. Brown. 2006. On the elusiveness of enemy-free space: Spatial, temporal, and host-plant-related variation in parasitoid attack rates on three gallmakers of goldenrods. Oecologia 150:421-434.

Brown, J.M., M. Todd-Thompson, A. McCord, A. O’Brien, and B. O’Fallon 2006. Phylogeny, host association, and wing pattern variation in the endemic Hawaiian tephritids (Tephritidae: Tephritini). Instrumenta Biodiversitatis VII:259-274.

Brown, J.M., and I. Cooper. 2006. Evolution of wing pigmentation patterns in a tephritid gallmaker: divergence and hybridization. Pp. 253-261 in Galling and Their Associates – Ecology and Evolution, K. Ozaki, J. Yukawa, T. Ohgushi, and P.W. Price, eds. Springer-Verlag, Tokyo.

Turgeon, J. R. Stoks, R.A. Thum, J.M. Brown, and M.A. McPeek. 2005. Simultaneous quaternary radiations of three damselfly clades across the Holarctic. American Naturalist. 165:E78-107.

McGranahan, D.A., S. Kuiper, and J.M. Brown. 2005. Temporal patterns in use of an Iowa woodlot during the autumn bird migration. American Midland Naturalist 153:61-70.

Cooper, I., E. Roeder, and J.M. Brown. 2003. response to burning and mowing in a reconstructed prairie. Ecological Restoration 21:204-5.

Brown, J.M, M.A. McPeek, and M. May. 2000. A phylogenetic perspective on habitat shifts and diversity in the North American Enallagma damselflies. Systematic Biology 49:697-712.

McPeek, M.A. and J.M. Brown. 2000. Building a regional species pool: Diversification of the Enallagma damselflies in eastern North American waters. Ecology 81: 904–920.

Drown, D.M., and J.M. Brown. 1998. Molecular phylogeny of North American oak-galling Cynipini (Hymenoptera: Cynipidae) supports need for generic revision. Pp. 241-246. The Biology of Gall-Inducing Arthropods (Csoka, G., W.J. Mattson, G.N. Stone and P.W. Price eds.). Gen. Tech. Rep. NC-199. St. Paul, MN: U.S. Department of Agriculture, Forest Service, North Central Research Station.

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Faust, L., and J.M. Brown. 1998. Sexual selection via female choice in the gall-making fly Eurosta solidaginis Fitch (Diptera: Tephritidae). The Biology of Gall-Inducing Arthropods (Csoka, G., W.J. Mattson, G.N. Stone and P.W. Price eds.). Gen. Tech. Rep. NC-199. St. Paul, MN: U.S. Department of Agriculture, Forest Service, North Central Research Station.

Brown, J.M., J. Leebens-Mack, O. Pellmyr, J.N. Thompson, and R.G. Harrison. 1997. Phylogeography and host association in a pollinating seed parasite, politella (: ). Molecular Ecology 6:215-224.

Pellmyr, O., J.N. Thompson, J. M. Brown and R.G. Harrison . 1996. Evolution of pollination and mutualism in the yucca lineage. American Naturalist 148:827-847.

Hess, M.D. W.G. Abrahamson and J.M. Brown. 1996. Intraspecific larval competition in the herbivorous ball-gallmaker, Eurosta solidaginis (Diptera: Tephritidae). American Midland Naturalist 136:121-133.

McPeek, M.A., A. K. Schrot, and J. M. Brown. 1996. Adaptation to predators in a new community: swimming performance and predator avoidance in damselflies. Ecology 77:617-629.

Brown, J.M, W.G. Abrahamson and P.A. Way. 1996. mtDNA phylogeography of host races of the goldenrod ball gallmaker (Diptera: Tephritidae: Eurosta solidaginis). Evolution 50:777-786.

Brown, J.M., W.G. Abrahamson, R. Packer, and P.A. Way. 1995. The role of enemy escape in a gallmaker host-plant shift. Oecologia 104:52-60.

Brown, J.M., O. Pellmyr, J.N. Thompson, and R.G. Harrison. 1994. mtDNA phylogeny of the Prodoxidae (Lepidoptera: Incurvarioidea) indicates a rapid ecological diversification of the yucca . Annals of the Entomological Society of America 87:795-802.

Brown, J.M., O. Pellmyr, J.N. Thompson, and R.G. Harrison. 1994. Phylogeny of Greya (Lepidoptera:Prodoxidae) based on nucleotide sequence variation in cytochrome oxidase I and II: congruence with morphological data. Molecular Biology and Evolution 11:128-141.

Abrahamson, W.G. J.M. Brown, S.K. Roth, D.V. Sumerford, J.D. Horner, M.D. Hess, S.T. How, T.P. Craig, R.A. Packer, and J.K. Itami. 1994. Gallmaker speciation: an assessment of the roles of host-plant characters and phenology, gallmaker competition, and natural enemies. In P. Price, W. Mattson, and Y. Baranchikov, eds. Gall-forming Insects. USDA Forest Service, North Central Experiment Station. General Technical Report. pp. 208-222.

Brown, J.M. and D.S. Wilson. 1994. Poecilochirus carabi: Behavioral and life-history adaptations to different hosts and the consequences of geographical shifts in host communities. pp. 1-22 in M. Houck, ed., Mites -- Ecological and Evolutionary Analyses of Life History Patterns. Chapman and Hall, New York.

Brown, J.M. and D.S. Wilson. 1992. Local specialization on sympatric hosts: phoretic mites on carrion beetles. Ecology 73:463-478.

Invited lectures (since 1996) Jonathan M. Brown -- 5

1/16 Reconstructing adaptive change in two radiations of Hawaiian insects. Biology Department. James Madison University.

2/12 Evolution of Host Relationships in Endemic Hawaiian Tephritids. (Invited symposium speaker). 14th Pacific Entomology Conference. Honolulu, HI. 6/11 Does being a host generalist promote speciation on islands? Evolutionary Ecology Across Trophic Levels -- An International Symposium in Honor of Warren Abrahamson. Bucknell University, Lewisburg, PA. 3/11 Evolution in your backyard. Day of Insects, Reimann Gardens, Ames, IA. 7/10 Phylogeography of the goldenrod ball gallmaker (Eurosta solidaginis) revisited. Genetics and the Origin of Species: The Continuing Synthesis. A symposium in honor of Richard G. Harrison. Cornell University, Ithaca, NY. 4/10 Food, Sex, and the Origin of Species. Department of Biology. Macalester College. St. Paul, MN 4/10 Generating the Entangled Bank. Department of Biology. Grinnell College. Grinnell, IA 4/10 Food, Sex, and the Origin of Species Department of Biology. Bucknell University, Lewisburg, PA. 11/09 Food, Sex, and the Origin of Species. “Spend a Day with Darwin” Grinnell College Alumni Event. San Diego Natural History Museum, San Diego, CA. 3/07 Convocation address. Grinnell College, Grinnell, IA. 9/05 Zoology Department and Center for Ecological Research, Kyoto University, Kyoto, Japan 5/04 Biology Department, St. Olaf College, Northfield, MN. 3/04 Biology Department, Luther College, Decorah, IA 4/03 Biology Department, Grinnell College, Grinnell, IA. 2/03 Biology Department, Dartmouth College, Hanover, NH. 8/02 Symposium: “Restoration Education” Annual Meeting for the Ecological Society of America/Society for Ecological Restoration. 2/02 Department of Biology, University of Hawaii-Hilo. 11/01 Department of Biology, University of Washington-Bellingham. 10/01 Department of Animal Ecology, Iowa State University, Ames, IA. 6/00 Workshops on teaching evolution. Society for the Study of Evolution Annual Meeting. University of Indiana, Bloomington, IN. 11/98 Dept. of Biology, University of North Dakota. 4/98 Dept. of Biology, University of Iowa, Iowa City, IA. 9/97 Dept. of Biology, University of Missouri. Columbia, MO. 5/97 Dept. of Biology, Carleton College. Northfield, MN. 3/97 Dept. of Animal Ecology, Iowa State University, Ames, IA. 7/96 ACM Wilderness Field Station. Ely, MN. 4/96 Dept. of Entomology, University of Illinois.

Contributed Papers and Posters (since 1999; undergraduate research students in bold)

6/16 Cook, P., R. Rasmussen, E. Hsieh, J. Brown, I. Cooper. Female dimorphism and male harassment in the Hawaiian damselfly Megalagrion calliphya. Evolution 2016 (Annual Meeting of the Society for the Study of Evolution) Austin, TX.

6/16 Brown, JM, R. Rasmussen, E. Hsieh, M. Mamantov, L. Lyons. Native unicolonial ants show Jonathan M. Brown -- 6

no between-colony aggression across 1000 km, despite genetic and CHC differentiation. Evolution 2016 (Annual Meeting of the Society for the Study of Evolution) Austin, TX. 3/15 Riedl, R. and J.M. Brown. Population Structure of Two Nearctic Calopteryx Damselfly Species. Midwest Ecology and Evolution Conference, Bloomington, IN. 3/15 Rasmussen, R, E. Hsieh, L. Lyons, and J.M. Brown. Lack of aggression across large distances in in a North American mound-building ant. Indiana University Animal Behavior Conference. Bloomington, IN. 3/15 Mamantov, M. and J.M. Brown. Population structure of a N. American unicolonial ant, Formica exsectoides. Midwest Ecology and Evolution Conference. Indiana University, Bloomington, IN. 7/14 Brown, J.M., I.A. Cooper, S. Sherwood, M. Cloud, and M.A. McPeek. Adaptation and the tempo of phenotypic change during radiation in the Hawaiian damselflies (Coenagrionidae: Megalagrion). Island Biology Conference. Honolulu, HI. 6/14 Brown, J.M., I.A. Cooper, S. Sherwood, M. Cloud, and M.A. McPeek. Adaptation and the tempo of phenotypic change during radiation in the Hawaiian damselflies (Coenagrionidae: Megalagrion). Annual meeting of the Society for the Study of Evolution. Raleigh, NC. 3/14 Gunshanen, C. and J.M. Brown. Behavioral tests for supercolony boundaries in a native ant, Formica exsectoides Forel. Midwest Ecology and Evolution Conference. Dayton, OH. 6/13 Cooper, I.A, Brown, J.M., Getty, T. Ecological selection drives sexual dimorphism and species variation in Hawaiian damselflies. Annual meeting of the Society for the Study of Evolution. Snowbird, Utah. 8/12 Cooper, I.A, Brown, J.M., Getty, T. Rapid evolution of color and sexual dimorphism in a radiation of Hawaiian damselflies. Annual meeting of the Society for the Study of Evolution. Ottawa, Canada. 2/12 Cooper, I.A. and J.M. Brown. Sexual dimorphism, color variation, and habitat association in Megalagrion damselflies. (Contributed paper). 14th Pacific Entomology Conference. Honolulu, HI. 6/11 J.M. Brown. Generating the entangled bank – herbivore evolution in a Pacific archipelago – Smithsonian conference Evolution of Life on Pacific Islands and Reefs: Past, Present, and Future. Honolulu, Hawaii. 6/10 J.M. Brown. Phylogeography of the goldenrod ball gallmaker (Eurosta solidaginis) revisited. Annual Meeting of the Society for the Study of Evolution. Portland, OR. (Poster) 8/09 Brown, J.M. Evolution of gallmaking during the radiation of the endemic Hawaiian tephritid flies. 5th Symposium on the biology of gall inducing arthropods. Serra do Cipo – Brazil. 8/09 Brown, J.M. Revisiting the phylogeography of Eurosta solidaginis: host races and subspecies. 5th Symposium on the biology of gall inducing arthropods. Serra do Cipo – Brazil. 6/08 Funk, M. and J.M Brown. A nuclear phylogeny of Hawaiian tephritids. Annual meeting of the Society for the Study of Evolution, Minneapolis, MN. (Poster) 6/08 Fenster, M. and J.M Brown. Between-island speciation of an endemic Hawaiian herbivore. Annual meeting of the Society for the Study of Evolution, Minneapolis, MN. (Poster) 6/08 Brown, J.M. Evolution of host association in endemic Hawaiian tephritids. Annual meeting of the Society for the Study of Evolution, Minneapolis, MN. 6/08 Brown, J.M. Evolution of host association in endemic Hawaiian tephritids. 4th International meeting of Tephritoid Systematics and Ecology. Gatlinburg, TN. 8/07 Brown, J.M. Evolution of melanin pigmentation patterns during speciation. Meeting of the European Society of Evolutionary Biology. Uppsala, Sweden (Poster) 6/07 Brown, J.M. Evolution of host association in endemic Hawaiian tephritid flies. Society for the Study of Evolution Annual Meeting, Christchurch, New Zealand. (Poster) Jonathan M. Brown -- 7

8/06 Homans, J. and J. M. Brown Diversification of seed-feeding tephritids in the Hawaiian archipelago. Society for the Study of Evolution Annual Meeting, SUNY-Stoneybrook. (Poster) 9/05 Brown, J.M., and I. Cooper. Evolution of wing pigmentation patterns in a tephritid gallmaker: divergence and hybridization. Biodiversity of galling arthropods and their associates. Kyoto, Japan. 9/05 Brown, J.M. Ecological diversity in a radiation of herbivorous Hawaiian flies. Biodiversity of galling arthropods and their associates. Kyoto, Japan. (Poster) 7/04 3rd International meeting of Tephritoid Systematists. Evolution of host association and wing pattern in the endemic Hawaiian tephritids. J.M. Brown and Megan Todd-Thompson. Geneva, Switzerland. 6/04 Annual Meeting of the Society for the Study of Evolution. Genetic evidence for speciation among island populations of endemic Hawaiian tephritids (J. Brown and M. Todd-Thompson). Colorado State University, Ft. Collins, CO. (Poster) 6/03 Annual Meeting of Society for the Study of Evolution. Evolution of host-plant use and wing pattern in endemic Hawaiian tephritids (J. Brown, A. McCord, and A. O’Brien) Cal. State at Chico. Chico, CA. (Poster) 8/02 Annual Meeting of Ecological Society of America. Effects of burning and mowing on arthropods in a reconstructed tallgrass prairie. (I. Cooper, E. Roeder and J. Brown). Tucson, AZ. (Poster) 6/02 Annual Meeting of the Council on Undergraduate Research. The Student as Scientist: A First Course in Biology at Grinnell College (C. Sullivan, C. Lindgren, J. Brown, and D. Lopatto) (Poster) 7/01 Annual Meeting of the Society for Conservation Biology. Prairie Restoration: An introduction to Biological Inquiry (co-author C. Caruso) University of Hawaii at Hilo. 6/01 Annual Meeting of the Society for the Study of Evolution. Evolution of wing pigmentation patterns in flies: divergence and hybridization (co-author Idelle Cooper). Univ. of Tennessee at Knoxville. 6/99 Annual Meeting of the Society for the Study of Evolution. Laboratory exercises to teach principles of heritability; Methods for quantifying wing variation in Hawaiian tephritids (Brendan O’Fallon and J. Brown).

Professional Societies

Associate Editor of Evolution – International Journal of Organic Evolution (Jan 2006-2009)

Society for the Study of Evolution (Member of Education Committee 1997-2001) Society of American Naturalists Organization for Tropical Studies Sigma Xi

Reviewer for

American Journal of Botany Evolution Annals Entomological Society of America Journal of Evolutionary Biology Biochemical Systematics and Ecology Molecular Phylogenetics and Evolution Ecology Proceedings of the Royal Society Molecular Ecology Entomologia Experimentalis et Applicata Jonathan M. Brown -- 8

American Midland Naturalist National Science Foundation Systematics and Population Biology Panels National Science Foundation Graduate Research Fellowship Program

Student research supervision at Grinnell

Jongbeom Park ’18 – Summer 2016 Julia Shangguan ’18 – Summer 2016 Amanda Bressoud ‘17 – Summer 2016 S. Jane Jordan ’18 – Summer 2016 Gabrielle Mercado ’16 – Spring 2016 Rebecca Rasmussen ’16 – Summer 2014, 2015 Edward Hsieh ’16 – Summer 2014, 2015 Margaret Mamantov ’16 -- Summer 2014 Ruth Riedl ’16 – Summer 2014 Clare Gunshanen ’14 – Fall 2013 Eric Streeder ’14 – Summer 2013 Madeline Cloud ’14 – Summer 2013 Sam Sherwood ’12 – Spring 2012 Karissa Fernandez ’12 - Spring 2012 Glenn Clark ’10 – Summer 2009 – Behavioral and genetic studies of a unicolonial ant Glenn Clark ’10 – Spring 2009 – Population structure of an endemic Hawaiian damselfly Allison Louthan ’08 – Fall 2007, Spring 2008 -- AFLP phylogeny of Hawaiian tephritid flies Mike Fenster ’08, Margarent Funk ’08 and Maria Namwanje ’09 – Summer 2007 – Molecular phylogenies of Hawaiian tephritids Mike Fenster ’08 and Emma Meade ’07 -- Summer 2006 – Speciation in Hawaiian tephritids Derrick Mitchell ’06 and Jonathan Homans ’06 -- Summer 2005 – Morphological and ecological variation among populations of endemic Hawaiian tephritids Anna Larimer ’05, Chris Mitros ’05, and Hanghang Wang ’07 -- Summer 2004 – Effects of burning and mowing on prairie soil and litter arthropods Megan Todd-Thompson ’05 – Summer 2003 -- Genetic differentiation among island populations of an endemic Hawaiian fly Anne O’Brien ’03 – Summer 2002 -- Molecular phylogeny of native Hawaiian herbivores Aleia McCord ’03 – Summer 2002 -- Genetic differentiation among island populations of a native Hawaiian herbivore Pam Marks ’02 – Summer 2001 -- Investigations for 4th grade science Idelle Cooper '01 – Spring 2001 -- Quantitative image analysis of a hybrid zone in gall flies Abby Laatsch '01 – Spring 2001 -- Genetic analysis of gall fly hybrid zone Monica Poelchau ’01 – Summer 2000 – Molecular systematics of S. African plants* Tor Janson '01 Summer 1999 -- Butterfly populations in Iowa prairie remnants and reconstructions Liz Roeder '01 Summer 1999 -- Effects of burning and mowing on above-ground prairie arthropods Idelle Cooper '01 Summer 1999 -- Effects of burning and mowing on below-ground prairie arthropods Brendan O'Fallon '99 Spring 1999 -- Methods for quantification of tephritid wing patterns Zoe McKiness '99 Spring 1999 -- Molecular phylogeny of oak gall wasps Shira Peltan ‘99, Summer 1998 -- Molecular markers for a gall fly hybrid zone Abby Laatsch ‘01, Summer 1998 -- Sexual selection in a gall fly Josh Rehmann ‘00, Summer 1998 -- Molecular phylogeny of oak gall wasps Jennifer Johnson ‘99, Summer 1997 -- Effects of ranching on Hawaiian cave arthropods* Jonathan M. Brown -- 9

Lisa Faust ‘97, Spring 1997 -- Sexual selection in a gall fly Devin Drown ‘97. Spring 1996-Spring 1997 -- Molecular phylogeny of oak gall wasps William Eichman ‘97, Spring 1996-Fall 1996 -- Comparison of growth in gall fly host races Marta Fisher ‘97, Summer 1996 -- Feeding ecology of macaws in Costa Rica* Aksel Casson ‘96, Summer 1996 -- Molecular phylogeny of N. American damselflies

College Service

14-16 Writing Advisory Committee 13-16 Faculty Director of the Conard Environmental Research Area (CERA) 12-15 Committee for the Support of Faculty Scholarship 12-13 Off-campus studies Advisory Board 12-14 Center for Prairie Studies Advisory Board 09-11 Personnel Committee 09-11 Center for International Studies Advisory Board 09-11 Ecocampus Committee 07-08 Tutorial Committee 07 EKI Earth Systems Science Search Committee 00-01, 02-05 Science Phase II Planning Committee 00-01, 4-5, 6-7 Center for Prairie Studies Advisory Committee 99- 01, 4-5, 6-7 Writing Advisory Committee 96- 01, 0 4-05 Faculty Director of the Conard Environmental Research Area (CERA) 97-Present Campus Advisor for the ACM and OTS Tropical Field Research program in Costa Rica 02-04 Chair, Department of Biology 96- 02 Campus Advisor for the ACM summer program at the Wilderness Field Station (Member of Executive Committee 1998) 96- 01 Health Professions Advisory Committee 98- 01 Environmental Studies Committee 00-01 Gender and Women's Studies Committee 97- 99 Academic Computing Committee