Insects of Samoa and Other Samoan Terrestrial Arthropoda
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IK KM].ML.. SllSBftSSf BRITISH MUSEUM (NATURAL HISTORY) INSECTS OF SAMOA AND OTHER SAMOAN TERRESTRIAL ARTHROPODA PART IV. COLEOPTERA FASC. 2. Pp. 67-174 HETEROMERA, BOSTRYCHO IDEA, MALACODERMATA and BUPRESTIDAE. By K. G. Blair, B.Sc. ELATERIDAE. By R. H. van Zwaluwenberg. MELASIDAE (EUCNEMIDAE). By E. Fleutiaux. CERAMBYCIDAE. By Chr. Aurivillius. BRENTHIDAE. By R. Kleine. ANTHRIBIDAE. By Karl Jordan, PhD. PROTERHINIDAE. By R. C. L. Perkins, D.Sc, F.R.S. WITH THIRTY-NINE TEXT-FIGURES AND ONE PLATE LONDON PRINTED BY ORDER OF THE TRUSTEES OF THE BRITISH MUSEUM SOLD AT The Bbitish Museum (Natubal Histoby), Obomwell Road, S.W. 7 AND BY B. Quabitch, Ltd. ; Dulau & Co., Ltd. ; The Oxfobd Univebsity Pbess ; and Whbldon & Wesley, Ltd., London; also by Oliveb & Boyd, Edinbubgh 1928 Issued 26th February, 1928.] [Price Five Shillings. INSECTS OF SAMOA AND OTHER SAMOAN TERRESTRIAL ARTHROPODA Although a monograph, or series of papers, dealing comprehensively with the land arthropod fauna of any group of islands in the South Pacific may be expected to yield valuable results, in connection with distribution, modification due to isolation, and other problems, no such work is at present in existence. In order in some measure to remedy this deficiency, and in view of benefits directly accruing to the National Collections, the Trustees of the British Museum have undertaken the publication of an account of the Insects and other Terrestrial Arthropoda collected in the Samoan Islands, in 1924-1925, by Messrs. P. A. Buxton and G. H. E. Hopkins, during the Expedition of the London School of Hygiene and Tropical Medicine to the South Pacific. Advantage has been taken of the opportunity thus afforded, to make the studies as complete as possible by including in them all Samoan material of the groups concerned in both the British Museum (Natural History) and (by courtesy of the authorities of that institution) the Bishop Museum, Honolulu. It is not intended that contributors to the text shall be confined to the Museum Staff or to any one nation, but, so far as possible, the assistance of the leading authorities on all groups to be dealt with has been obtained. " The work will be divided into eight " Parts (see p. 3 of wrapper), which will be subdivided into " Fascicles." Each of the latter, which will appear as ready in any order, will consist of one or more contributions. On the completion of the work it is intended to issue a general survey, summarising the whole and drawing from it such conclusions as may be warranted. E. E. AUSTEN, Keeper of Entomology, British Museum (Natural History), Cromwell Road, S.W.7. 1 * ^1 lyan f not nlea, IV. 2 G7 1 INSECTS OF SAMOA Part IV. Fasc. 2. HETEKOMEKA, BOSTRYCHOIDEA, MALACO- DEEMATA AND BUPRESTIDAE By K. G. Blair, B.Sc. (With 14 Text-figures.) From the distributional standpoint, the Coleoptera of the groups here dealt fall : with seem to into three categories (a) peculiar to the Samoan Group ; (6) of wider distribution in the Pacific, in some cases apparently limited to neighbouring groups of islands, in others extending throughout the tropics of the old world (c) cosmopolitan, in most cases store-pests ; and undoubtedly introduced by commerce ; this latter category is of course of no value in considering zoogeographical distribution. Out of about sixty species discussed in the present paper, twenty-two belong to the first category, while five genera, Apteromerus, n., Callistroma Fairm., Scolytocis, n., Melaneros Fairm. and Samoaneros, n., are also peculiar to Samoa. A further ten species, together with the genus Menandris Haag, are known in addition only from the adjacent groups of Wallis Is., Fiji, Tonga, and the Ellice Is. Of these autochthonous genera, Apteromerus is no doubt an offshoot from Bradymerus Perr., a genus mainly Indo-Malayan and Melanesian in distribution, and Callistroma from Paracupta Deyr., largely developed in Melanesia. For Scolytocis it is difficult to assign any particular origin ; Melaneros and probably Samoaneros are offshoots from the large and somewhat indeterminate genus Plateros, and probably also of Indo-Malayan origin. A consideration of the distribution of some of the more extensive but not too generally distributed genera, Bradymerus, Menimus, Uloma, Thesilea, IV. 2 67 1 68 INSECTS OF SAMOA. Chariotheca, Amarygmus among the Tenebrionidae, Pelecotomoides in the Rhipi- phoridae, Callirhipis, etc., appears to support the inference that the fauna of our region is a development of a branch originating in the Indo-Malay region and running eastward through Melanesia to Polynesia. So far as it is possible to judge from the limited fauna here considered, there seems little evidence of any important contribution having been received from Australia, no genera peculiar to that fauna, or their derivatives, being represented ; such features as they have in common, e.g. Carphurus, would appear to be due to a common origin in the Papuan region, rather than to an independent origin in Australia. Collections of greater or less extent from the Samoan Group have been received at the British Museum from the Rev. S. J. Whitmee, 1875, 1876, 1890, and 1897 ; Dr. H. Swale, 1917, 1918, 1920 ; Dr. F. W. O'Connor, 1920 ; while for the purposes of this present work further collections have been kindly lent by the authorities of the Bishop Museum, Honolulu (Coll. Swezey and Wilder, E. H. Bryan, Jr., A. F. Judd), and by Mr. F. Muir of Honolulu (Coll. H. C. Kellers). For bibliography, the reader is referred to the respective parts of Junk's Coleopterorum Catalogus where published, only the original citation, with a reference to a figure, and to papers subsequent to Junk's work being given in the present paper. The types of new species, except when otherwise stated, are in the British Museum. The following table shows the distribution, so far as it is known, of the Samoan fauna among the Samoan Is., and in other parts of the Pacific. The names of species and genera described below as new are shown in heavier type. U hi TuTUILA. Further Distribution. o Savaii. Manua. P Tenebrionidae Mesomorphus villiger Blanch. X Melanesia, Hawaii, Indo-Malaya, etc. Bradymerus granaticollis Fairm. X Melanesia. „ amicorum Fairm. X Tonga and Fiji. ? lobicollis Geb. X New Guinea. Apteromerus convexus Fairm. X X Wallis Is. Menimus samoensis, n. X Tagalus swalei, n. 7 — HETEROMERA, MALACODERMATA, ETC. 69 < fc> rUILA. hi <3 Further Distribution. O '< Ph HP m Tenebrionidae . continued. Gnathocerus cornutus F. ? Cosmopolitan. Tribolium castaneum Hbst. x X Palorus austrinus Champ. x Australia, Damma Is. ,, upoluensis, n. x TJlonid cct'Vicollis Fairm. • Av Wallis Is., Ellice Is. Alphitobius diaperinus Pz. x Cosmopolitan. Ictevic/atus F. X Sciophagus pandanicola Bdv x Melanesia, Polynesia. Thesileci puncticeps Fairm. V x Fiii (?) Chnriothecci plcinicollis Fairm. v X x Wallis Is. samoensis, n. x Ql 1 If^l T^f^n Y\ 1 C Tl 1 1_ 1 1 1 1 1 • v v o 1 b, 11. • , , U | A A Menandris aenea Haag. X X Ovalau (Fiji), N. Australia (?). A.Wj<irygmu$ sctTnoensis Haag. x X x Vavau (Tonga). fl/nPTPitfiflPv Fairm Kin xnvio'^i t/VVUZ'l Is f.K/Myo/ X'Ctli-lll. • A X Av Melandryidae Bryancllci samoGHsis, n. Anthicidae Anthicus oceanicus Laf. x Marquesas Is., Society Is., Hawaii China, Ceylon, Seychelles, etc. Mordellidae Mordella novemguttatct Montr. x Woodlark Is., Philippines, Borneo etc. XIX '( L ('! [_'(< /{If' [.(vlt'l//t'H.t'C' . TVpw dalprloma T^m Australia W J.TJ_VJlJUi. Av Av AV -ll W \JC\l*^\.l.\J lllCv . J. Ill, XI LID Ul tXlld). » • l^ty IVO VI fVVV VO J ) 1 J. * Av X Fiii j samoensis, n. x X Av X swezeyi, n. x X x TCfllorsi n Av X nigrescens, n. x X x Rhipiphoridae iplppofo}iifi)(Jp^ j ulvft ^pr H'fi n <\ T^aiTm J- \JV\s\j\J V\J IIV\J Ht'C-O / lyoCI t'vlwto j. (til III, Av X Fiji, Tonga, Ellice. Oedemeridae Ananca kanack Fairm. x X Melanesia, Polynesia. bicolor Fairm. ? X Tahiti, Tonga, Ellice, N. Hebrides. ,, decolor Fairm. X X Tahiti, Ellice, Marquesas. Pselaphanca lateritia Fairm. X Society Is. Sessinia livida F. X X Tahiti, Ellice, Tonga, Fiji. ClIDAE Cis savanensis, n. X ,, tutuilensis, n. X Scolytocis samoensis, n. X . 70 INSECTS OF SAMOA. u 3 a Further Distribution. o H > Mantja. P EH CO Lyctidae Lyctus brunneus Steph. X X Cosmopolitan. Minthea rugicollis Wlk. X BoSTRYCHIDAE Rhizopertha dominica P. X Xylothrips religiosus Bdv. X Melanesia, Polynesia. Xylopsocus castanoptera Fairm. ? Tropics of Old World. Anobiidae Lasioderma serricome F. X Cosmopolitan. Lycidae Melaneros atroviolaceus Fairm. X X quadraticollis Fairm. X X X Tonga (?). Samoaneros aculicollis Fairm. X X ,, muin, n. X Melyridae Carphurus flavipes, n. X Carphuroides pectinatus Shp. ? Hawaii, Borneo, N. India. Oleridae ? Cylidrus cyaneus F. X Tropics of Old World. Tarsostenus univittatus Rossi X Cosmopolitan. Necrobia ntfipes De G. X X X X Rhipiceridae Callirhipis femorala Waterh. X X Nassau Is. Buprestidae Callistroma samoensis Sndrs. X X Cyphogastra abdominalis Waterh. X D. of York Is., New Britain. Chrysobothris chrysomela Deyr. X Agrilus indignus Fairm. X X Vavau (Tonga), Polynesia. ,, samoensis, n. X j The above are represented in the material before me as follows : — Family : TENEBRIONIDAE. Gebien, Coleopt. Catal. Tenebr. (Junk, parts 15, 22, 28, 37), 1910-11. The beetles of this family occurring in Melanesia have been well mono- graphed by H. Gebien in " Resultats de l'Expedition Scientifique Neerlandaise a la Nouvelle-Gruinee," Vol. xiii., Zoologie. Livr. 3, 1920 (subsequently referred to as Tenebr. N. Guin.). In this work a full synonymy is given, in addition to HETEROMERA, MALACODERMATA, ETC. 71 a complete account of the distribution of each species as known to the author ; many records from Samoa are included.