Curriculum Vitae

M. Songül Alpaslan-Roodenberg Department of Genetics (Reich Laboratory) Harvard Medical School New Research Building, Room 260 77 Avenue Louis Pasteur Boston, MA 02115, USA e-mail: [email protected]

Academic Position

Scientific Associate and Consulting Anthropologist at Harvard University (Ancient DNA Laboratory of Professor David Reich). My role is to lead the laboratory’s work on joint anthropological and genetic studies in Turkey.

Education

1995 Ph.D. (summa cum laude) University of Ankara. Thesis title: “Morphometric Study on the Occlusal Surfaces of Hominoid Molar Teeth from Paşalar, Turkey”, 738 p. Supervisor Prof. Peter Andrews, Natural History Museum, London. 1988 MA degree (summa cum laude), University of Ankara. Thesis: “Comparative Study of the Neogene Fauna (Mammalia) in Turkey”, 288p. (in Turkish). 1980 – 1984 B.A. degree, Dept. of Physical and Palaeoanthropology, Univ. of Ankara.

Professional Experience

Active Genetic Projects

2008 – present, working with Ron Pinhasi (beginning 2008) and later David Reich as well (beginning of 2013) on ancient DNA studies of population transformations in Anatolia and Southeastern Europe. My collaboration with Prof. Reich’s lab has become a central component of my work, and as a result I became a Scientific Associate of the laboratory in 2017 and travel extensively as part of this collaborative set of projects. [https://reich.hms.harvard.edu/people]. This work has already produced 5 co authored papers, including our last paper at Nature in which I am second listed-author.

1998 – present, coordination of anthropological and genetic analyses of ancient skeletons (I am an expert not only in morphological analysis of ancient skeletons, but also in identifying skeletal material for DNA analysis). I was involved in an important way in a key innovation in ancient DNA that was published in 2015, which was the discovery of the petrous bone of the skull as a highly useful type of bone for DNA analysis.

1993 – present, working as a physical anthropologist focusing on palaeodemography, palaeopathology, behavior, diet, and mortuary practices, especially in early sedentary communities of Anatolia and Southeast Europe. In addition participating in ancient DNA and isotope studies. In this capacity I take part in various prehistoric settlement excavations (see ‘present projects’) directed by Zafer Derin, Aegean University, Izmir; Necmi Karul, Univ. of Istanbul; Krassimir Leshtakov, Univ. of Sofia; by Nedko Elenski, Museum of Veliko Tarnovo, Georgi Ganetovski, Museum of Vratsa; Stanislav Iliev, Museum of Haskovo. I furthermore cooperate with Vanica Veljanovska, anthropologist at the Nat. Hist. Museum of Skopje, Macedonia.

Since 2016, I have participated in excavations headed by Nihat Erdoğan, director of the Mardin Museum (Southeast Turkey), Ayşe Acar from the Anthropology Dept. of Artuklu Univ. in Mardin and in a (Late) Antique research project in collaboration with Mustafa Şahin, head of the Archaeological Dept., Uludağ Univ. Bursa, including a Roman theatre and an Early Christian basilica. Since 2018, I have also collaborated on ancient DNA studies with colleagues from the Anthropology Department of Ankara University.

2000 Guest lecturer giving a workshop for advanced undergraduate classes on the “Introduction to anthropology” and “Anthropological method and theory" at the Depart. of Archaeology and History of Art, Bilkent Univ., Ankara.

1997 – 1999 Guest researcher at the Anatomy Dept., Medical Faculty, Leiden Univ., The Netherlands.

1990 – 1993 Scientific staff member at the Department of Research, Planning and Coordination in the Ministry of Energy and Natural Resources, Ankara.

1984 –‘90, ’93 – ‘97 Palaeontologist, curator of the Palaeontology section at the Natural History Museum of the Institute of Mineral Research and Explorations (MTA General Directorate) Ankara. Between 1994-1997 I was abroad working on my PhD thesis and scientific research.

Past field work and projects Osteo-archaeological research

2009-2015 Collaboration with professor Malcolm Lillie and his team from Hull University (UK) on stable isotope studies of prehistoric bone material from Turkey and Bulgaria. We are continuing to work together on publishing the results.

2011-2016 Collaborator of the European Research Council project No. 263441 entitled From the earliest modern humans to the onset of farming (45,000-4,500 BP)" Project director professor Ron Pinhasi, Dept. of Anthropology, University of Vienna: "the role of climate, life-style, health, migration and selection in shaping European population history”. In this project I am involved in DNA and stable isotope analysis of human skeletons from Neolithic populations in West Anatolia and Southeast Europe. Collaboration on publishing the results continues.

2004-2015 Excavations at the multi-period site Çadır höyük, Yozgat / Turkey (in collaboration with various American Universities).

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2011-2015 Excavations at the Late Neolithic site Barcın höyük (The Netherlands Institute for the Near East, Leiden / Istanbul).

2007 – 2013 Excavations at the Early Neolithic site of Yabalkovo / Bulgaria (Univ. of Sofia)

2005 – 2006 Excavations at Barcın höyük, Bursa (The Netherlands Institute for the Near East, Leiden / Istanbul).

2000 – 2004 Excavations at the Bronze Age cemetery of Harmanören, Göndürle höyük, Isparta (Univ. of Istanbul).

1989 – 2002 Excavations at the Late Neolithic site Ilıpınar höyük, Bursa (The Netherlands Institute for the Near East, Leiden / Istanbul)

1996 - 2000 Excavations at the Late Neolithic site Menteşe höyük, Bursa (The Netherlands Institute for the Near East, Leiden / Istanbul.

Palaeontology

1993 Excavations at a Pliocene site near Muğla (Univ. of Ankara). 1989 – 1990 Excavations at the Late Miocene Locality Sinap, Ankara (Univ. of Ankara). 1984 – 1992 Excavations at the Middle Miocene site Paşalar, Bursa (Univ. of Ankara).

Active Excavation Projects I performed osteo-archaeological research as well as DNA and Isotope studies for ongoing excavations: • 2004- present, Late Neolithic site Aktopraklık höyük, Bursa (Univ. of Istanbul). • 2007- present, Late Neolithic site Yeşilova höyük, İzmir (Aegean Univ., Izmir - Bornova). • 2008- present, Early Neolithic site Dzhulyunitsa / Bulgaria (Museum of Veliko Tarnovo). ! 2010- present, Early Neolithic site Ohoden / Bulgaria (Museum of Vratsa). ! 2016- present, Pre-Pottery Neolithic excavations, Mardin Museum (Ilisu Dam Project SE Turkey). ! 2016- present, (Late) Antique excavation project, Archaeological Dept., Uludağ Univ. (Bursa), comprising a Roman theatre and an Early Christian basilica .

Languages Turkish (native speaker) Dutch (advanced) English (advanced) German (elementary)

Conferences, seminars (2011-2017)

! Harvard medical School, Dept. of Genetics, Boston. ! Yeşilova excavations, Aegean University, İzmir-Bornova (bi-annual seminars for students). ! Symposium at the Netherlands Institute for the Near East, Leiden. ! Symposium organized by the Uludag University and Bursa Municipality . ! New Bulgarian University, Sofia.

3 ! National Museum of History, Bucharest. ! Institut für Anthropologie, Johannes Gutenberg-Unversität Mainz.

Publications

Mathieson, I., S. Alpaslan Roodenberg, C. Posth, A. Szécsényi-Nagy, N. Rohland, S. Mallick, et al., 2018 - The Genomic History of Southeastern Europe. Nature, 555. 197-203.

Lazaridis, I., A. Mittnik, N. Patterson, S. Mallick, N. Rohland, S. Pfrengle, A. Furtwängler, A. Peltzer, C. Posth, P.J.P. McGeorge, E. Konsolaki-Yannopoulou, G. Korres, H. Martlew, M. Michalodimitrakis, M. Özsait, N. Özsait, A. Papathanasiou, M. Richards, S. Alpaslan Roodenberg, Y. Tzedakis, A. Vasilakis, R. Arnott, D. M. Fernandes, J.R. Hughey, D.M. Lotakis, P. A. Navas, Y. Maniatis, J. A. Stamatoyannopoulos, K. Stewardson, P. Stockhammer, R. Pinhasi, D. Reich, J. Krause, G. Stamatoyannopoulos, 2017 - Genetic origins of the Minoans and Mycenaeans. Nature 548. 214-8.

Lazaridis, I., D. Nadel, G. Rollefson, D.C. Merrett, N. Rohland, S. Mallick, D. Fernandes, M. Novak, B. Gamarra, K. Sirak, S. Connell, K. Stewardson, E. Harney, Q. Fu, G. Gonzalez-Fortes, E.R. Jones, S. Alpaslan Roodenberg, G.Lengyel, F.Bocquentin, ….R. Pinhasi & D. Reich. 2016 - Genomic insights into the origin of farming in the ancient Near East. Nature. P.536 (7617), 419-424.

Mathieson, I., I. Lazaridis, N. Rohland, S. Mallick, N. Patterson, S. Alpaslan Roodenberg e.a., 2015 – Genome-wide patterns of selection in 230 ancient Eurasians. Nature 528, 499-503.

Gokhman, D., L. Agranat-Tamir, G. Housman, R. Garcia-Perez, M. Nissim Rafinia, S. Mallick, M. Nieves- Colón, H. Li, S. Alpaslan-Roodenberg, M. Novak, H. Gu, M. F. Bernal, P. Gelabert, I. Lipende, I. Kondova, R. Bontrop, E. Quillen, A. Meissner, A. C. Stone, A.E. Pusey, D. Mjungu, L. Kandel, M. Liebergall, M. E. Prada, J. M. Vidal, K. Prüfer, J. Krause, B. Yakir, S. Pääbo, R. Pinhasi, C. Lalueza-Fox, D. Reich, T. Marquez, Bonet, E. Meshorer, L. Carmel, 2017 - Extensive Regulatory Changes in Genes Affecting Vocal and Facial Anatomy Separate Modern from Archaic Humans, bioRxiv October (preprint).

Pinhasi, R., D. Fernandes, K. Sirak, M. Novak, S. Connell, S. Alpaslan-Roodenberg, F. Gerritsen, V. Moiseyev, A. Gromov, P. Raczky, A. Anders, M. Pietrusewsky, G. Rollefson, M. Jovanovic, H. Trinhhoang, G. Bar-Oz, M. Oxenham, H. Matsumura, M. Hofreiter, 2015 - Optimal Ancient DNA Yields from the Inner Ear Part of the Human Petrous Bone. PLoS ONE 10(6), p. 1-13.

Budd, C., N. Karul, S. Alpaslan-Roodenberg, R. Schulting, M. Lillie, 2017- Diet uniformity in Late Neolithic and Early Chalcolithic Northwest Anatolia (Turkey): Carbon and nitrogen isotope studies of bone collagen at Aktopraklık. Archaeological and Anthropological Sciences. Volume 9, Issue 40: p. 1-13.

Steadman, S., G. McMahon, J.C. Ross, M. Cassis, T. E. Şerifoğlu, B. S. Arbuckle, S.E. Adcock, S. Alpaslan Roodenberg, M. von Baeyer, and A. J. Lauricella, 2015 - The 2013 and 2014 excavation seasons at Çadır Höyük on the Anatolian North Central Plateau. Anatolica XLI.

N. Todorova, V. Petrova, M.S. Alpaslan Roodenberg, 2014 - The Early Neolithic human skeletons: an overview (chapter VIII). In: Maritsa Project Volume II, Yabalkovo Volume I. Editors: J. Roodenberg, K. Leshtakov, V. Petrova. Sofia University St. Kliment Ohridski, Sofia, p. 297-304.

Roodenberg, J., S. Alpaslan Roodenberg, 2014 - Neolithische bewoning van het oostelijke Marmara-

4 gebied in Noordwest-Anatolië. In: Waar de geschiedenis begon. Nederlandse onderzoekers in de ban van spijkerschrift, hiërogliefen en aardewerk. Uitgave naar aanleiding van het 75-jarig bestaan van het Nederlands Instituut voor het Nabije Oosten, 1939-2014. Editors: O.E. Kapers, J.G. Dercksen. Leiden. p. 248-259.

Roodenberg, J., and S. Alpaslan-Roodenberg, 2013 – Ilıpınar and Menteşe: Two Neolithic Sites in the Eastern Marmara Region. In: The Neolithic in Turkey. New excavations & new developments. The Marmara region. M. Özdoğan - N. Başgelen - P. Kuniholm. Archaeology and Art Publications, Istanbul, 69-91.

Alpaslan Roodenberg, S., N. Todorova, V. Petrova, 2013 – The Human Burials of Yabalkovo. Prähistorische Zeitschrift, Berlin (88 / 2), p. 23-37.

Alpaslan Roodenberg, M.S., F.A. Gerritsen, R. Özbal, 2013 - Neolithic burials from Barcın Höyük: The 2007-2012 excavation seasons. Anatolica 39, p. 93-111.

Budd, C., M. Lillie, S. Alpaslan-Roodenberg, N. Karul, R. Pinhasi , 2013 – Stable isotope analysis of Neolithic and Chalcolithic populations from the early farming settlement of Aktopraklik, Northern Anatolia. Journal of Archaeological Science. Vol. 40, issue 2, p. 860-867.

Lillie, M., C. Budd, S. Alpaslan-Roodenberg, N. Karul, R. Pinhasi, 2012 – Musings on Early Farming Communities in Northwest Anatolia and other flights of fancy. Paper dedicated to the memory of Professor Marek Zvelebil. IANSA (Interdisciplinaria Archaeologica, Natural Sciences in Archaeology) Volume III, Issue 1:11-22.

Leshtakov, K., V. Petrova, L. Leshtakov, G. Katsarov, I. Hadzipetkov, N. Tonkov, Ts. Pova, S. Alpaslan- Roodenberg, 2011 – Spasitelni arheologicheski pazkopki na neolitno selishte Yabalkovo. Natsionalniyat Arheologicheski Institut s Muzei pri BAN. Arheologicheski otkritiya i razkopki prez 2010g, 50-53.

Roodenberg, J., and S. Alpaslan-Roodenberg (editors), 2008 – Life and death in a prehistoric settlement in Northwest Anatolia. The Ilıpınar excavations with contributions on Hacılartepe and Menteşe, volume III. Pihans CX, Leiden, The Netherlands, 448 p.

Roodenberg, J., A. van As and S. Alpaslan-Roodenberg, 2008 – Barcın hüyük in the plain of Yenişehir (2005-2006). A preliminary note on the fieldwork, pottery and human remains of the prehistoric levels. Anatolica 34: 53-60.

Roodenberg, J., and S. Alpaslan Roodenberg, 2007 – Ilıpınar and Menteşe: Early settlement in the Eastern Marmara region. In: Living Well Together? Settlement and Materiality in the Neolithic of South-East and Central Europe. Douglas Bailey, A.W.R. Whittle and Dani Hofmann Eds. Oxbow Books, Oxford, 8-16.

Roodenberg, J., and S. Alpaslan-Roodenberg, 2007 – Vor 12.000 Jahren in Anatolien. In: Die ältesten Monumente der Menschheit. Badisches Landesmuseum, Karlsruhe, 154-155.

Alpaslan-Roodenberg, S., 2011 – A preliminary study of the burials from the Late Neolithic – Early Chalcolithic site of Aktopraklık. Anatolica 37: 17-43.

Alpaslan-Roodenberg, S., 2011 – Homicide in Aktopraklık, a prehistoric village in Turkey. Near Eastern

5 Archaeology , 74:1, 60-61.

Alpaslan-Roodenberg, S., 2009 – Demographic data from the Byzantine graveyard of Barcın. In: Archaeology of the countryside in Medieval Anatolia. T. Vorderstrasse and J. Roodenberg Eds. Pihans CXIII , Leiden,169-174.

Alpaslan-Roodenberg, S., 2008b – The Early Bronze Age human remains from Ilıpınar. In: Life and Death in a Prehistoric Settlement in Northwest Anatolia. With contributions on Hacılartepe and Menteşe. The Ilıpınar Excavations, Volume III. J. Roodenberg and S. Alpaslan Roodenberg Eds. Pihans CX, Leiden, 347-360.

Alpaslan-Roodenberg, S., 2008a – The Neolithic Cemetery – the anthropological view. In: Life and Death in a Prehistoric Settlement in Northwest Anatolia. With contributions on Hacılartepe and Menteşe. The Ilıpınar Excavations, Volume III. J. Roodenberg and S. Alpaslan Roodenberg Eds. Pihans CX, Leiden, 35-68.

Alpaslan-Roodenberg, S., 2006 – Death in Neolithic Ilıpınar, In: I. Gatsov, H. Schwarzberg Eds. Aegean – Marmara- Black Sea: Present state of the research of the Neolithic. Annual Meeting Proceedings Thessaloniki 2002. AK Monographs 5, Weissbach/Deutschland, 47-57.

Alpaslan-Roodenberg, S., 2002 – Preliminary report on the human remains from the Early Bronze Age cemetery at Ilıpınar – Hacılartepe, Anatolica, 28:91-107.

Alpaslan-Roodenberg, S., 2001 – Newly found human remains from Menteşe in the Yenişehir Plain: The season of 2000. Anatolica 27:1-14.

Alpaslan-Roodenberg, S, and G. Maat, 1999 – Human skeletons from Menteşe höyük near Yenişehir. Anatolica 25:37-51. ----- Alpaslan Roodenberg, M.S. (forthcoming) - Biyolojik antropoloji ve arkeolojide kullanılan yöntemleri. In: Arkeolojide Temel Yöntemler. Editörler: C. Çakırlar, S. Ünlüsoy, Ç. Çilingiroğlu. Ege Yayınları, İstanbul, (February 2018).

Alpaslan Roodenberg, S., 2017– Güneydoğu Marmara’nın İlk Çiftçilerine Genel Bir Bakış, Aktopraklık Halkı/A General Outlook at the Early Farmers of South-East Marmara: the People of Aktopraklık Odryses’ten Nilüfer’e Uluslararası Nilüfer Sempozyumu Nilüfer/ International Symposium 'From Odrysses to Nilüfer'. Editörler - Edited by M. Şahin • S. Sevim • D. Yavaş, Bursa Nilüfer Belediyesi • Bursa Nilüfer Municipality , Özyurt Matbaacılık, Ankara, p. 655-669.

Alpaslan-Roodenberg, S., 2016 - Tarih öncesi şiddet. Aktüel Arkeoloji, sayı 51: s. 40-49.

Alpaslan-Roodenberg, S., 2012 – Aktopraklık İnsanları: iskeletlerin anlattığı. Bursa’da Zaman, Sayı: 2: 46-50.

Alpaslan-Roodenberg, S., 2011 – Harmanören, Göndürle Höyük Mezarlığı İskeletlerinin Antropolojik Analizi. In: Özsait Armağanı. Studies Presented to Mehmet and Nesrin Özsait / Mehmet ve Nesrin Özsait Onuruna Sunulan Makaleler. Editörler H. Şahin – E. Konyar, G. Ergin. AKMED, Suna İnan Kıraç Akdeniz Medeniyetleri Araştırma Enstitüsü, yayınları. 1-8.

Roodenberg, J., ve S. Alpaslan-Roodenberg, 2011 – Ilıpınar ve Menteşe. In: Tarih Öncesinden Demir

6 Çağı’na, Anadolu’nun Arkeoloji Atlası. Necmi Karul ed. Arkeo-Atlas: 2011/01:116.

Alpaslan-Roodenberg, S., 2010 –Ok ucuyla gelen son. Atlas, Aralık , sayı 213.

Alpaslan-Roodenberg, S., 2009 – Kuzeybatı Anadolu’nun ilk çiftçileri: mezarlık ve gömütler. Arkeoloji ve Sanat, 131:1-12.

Roodenberg, J., ve S. Alpaslan-Roodenberg, 2007 – Ilıpınar ve Menteşe: Doğu Marmara’da Neolithik döneme ait iki yerleşme. In: Türkiye’de Neolitik dönem – yeni kazılar, yeni bulgular. Mehmet Özdoğan, Nezih Başgelen, editörler. Arkeoloji ve Sanat Yayınları, 392-400.

Alpaslan-Roodenberg, S., 2004 (2006) – Menselijke skeletresten afkomstig van de Kruisweg in Haarlem, Haarlems Bodemonderzoek, 38:33-36.

Roodenberg, J., and S. Alpaslan-Roodenberg, 2002 – Ilıpınar ve Menteşe. Arkeo-Atlas, sayı 1:125.

Roodenberg, J., and S. Alpaslan-Roodenberg, 1999 – Doğu Marmara Bölgesi, Ilıpınar Kazıları. Arkeoloji ve Sanat, 89:16-24 ------Alpaslan-Roodenberg, S., 1995 (Unpublished PhD thesis)– Morphometric Study on the Occlusal Surfaces of Hominoid Molar teeth from Paşalar, Turkey. PhD Thesis (in Turkish with English summary). Social Sciences Institute, University of Ankara Publications, 738p. (2 volumes).

Alpaslan, S., 1988 (Unpublished master thesis) – Türkiye ile Avrupa Karasal Neojen Faunalarının (Mammmalia) Korelasyonu. Yüksek Lisans Tezi, Antropoloji ABD. Sosyal BIlimler Enstitüsü (Ankara).

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