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Zootaxa 4787 (1): 001–377 ISSN 1175-5326 (print edition) https://www.mapress.com/j/zt/ Monograph ZOOTAXA Copyright © 2020 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4787.1.1 http://zoobank.org/urn:lsid:zoobank.org:pub:FE69E0FD-5A02-45A4-9CC4-3B76FA7D1D9A ZOOTAXA 4787 Catalog of the Biting Midges of the World (Diptera: Ceratopogonidae) ART BORKENT1 & PATRYCJA DOMINIAK2 1 Research Associate of the American Museum of Natural History, 691-8th Ave. SE, Salmon Arm, British Columbia, V1E 2C2, Canada. email: [email protected] 2 Norges arktiske universitetsmuseum og akademi for kunstfag, UiT Norges Arktiske Universitet, NO-9037 Tromsø, Norway. email: [email protected] Magnolia Press Auckland, New Zealand Accepted by J. Moulton: 5 Nov. 2019; published: 5 Jun. 2020 ART BORKENT & PATRYCJA DOMINIAK CataLOG OF THE BITING MIDGES OF THE WORLD (DIPTERA: CERatoPOGONIDAE) (Zootaxa 4787) 377 pp.; 30 cm. 5 Jun. 2020 ISBN 978-1-77670-947-2 (paperback) ISBN 978-1-77670-948-9 (Online edition) FIRST PUBLISHED IN 2020 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: [email protected] https://www.mapress.com/j/zt © 2020 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use. ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition) 2 · Zootaxa 4787 (1) © 2020 Magnolia Press BORKENT & DOMINIAK Table of Contents Abstract. 9 Introduction. 9 Format and Coverage. 18 Nomenclatural Changes, Status of Names and Commentary on Placement of some Taxa. 20 Current Numbers of Species of Ceratopogonidae and their General Distribution. 23 Commentary on Culicoides. 25 Location of Type Material. 27 Acknowledgements. 34 Subfamily Lebanoculicoidinae Borkent. 34 Genus Lebanoculicoides Szadziewski. 34 Subfamily Leptoconopinae Noè. 35 Genus Archiaustroconops Szadziewski. 35 Genus Austroconops Wirth and Lee. 35 Genus Fossileptoconops Szadziewski. 35 Genus Jordanoconops Szadziewski. 35 Genus Leptoconops Skuse. 36 Subgenus Brachyconops Wirth and Atchley. 36 Subgenus Holoconops Kieffer . 36 Subgenus Leptoconops Skuse. 38 Subgenus Megaconops Wirth and Atchley. 39 Subgenus Palaeoconops Borkent. 39 Subgenus Proleptoconops Clastrier . 40 Subgenus Styloconops Kieffer . 40 Genus Minyohelea Borkent . 40 Subfamily Atriculicoidinae Szadziewski . 40 Genus Atriculicoides Remm. 40 Subfamily Forcipomyiinae Lenz. 41 Tribe Dasyheleini Lenz. 41 Genus Dasyhelea Kieffer. 41 Nomina dubia. 55 Tribe Forcipomyiini Lenz. 56 Genus Atrichopogon Kieffer . 56 Nomina dubia. 68 Genus Forcipomyia Meigen. 68 Subgenus Atopomyia Yu and Liu . 69 Subgenus Baliohelea Yu and Liu . 69 Subgenus Bassoforcipomyia Debenham. 69 Subgenus Blantonia Wirth and Dow. 69 Subgenus Caloforcipomyia Saunders. 69 Subgenus Collessohelea Debenham. 70 Subgenus Dycea Debenham. 70 Subgenus Euprojoannisia Brèthes . 71 Subgenus Forcipohelea Clastrier and Delécolle. 73 Subgenus Forcipomyia Meigen . 74 Subgenus Gampsohelea Yu and Liu. 82 Subgenus Herakleohelea Debenham . 82 Subgenus Ixodehelea Yu and Liu. 82 Subgenus Japygahelea Yu and Liu. 82 Subgenus Kattangomyia Debenham. 82 Subgenus Lasiohelea Kieffer . 82 Subgenus Lepidohelea Kieffer. 87 Subgenus Metaforcipomyia Saunders. 89 Subgenus Microhelea Kieffer. 89 Subgenus Nicothohelea Liu, Tang and Hao. 92 Subgenus Oreinohelea Yu and Liu. 92 Subgenus Panhelea Remm. 92 Subgenus Pedilohelea de Meillon and Wirth. 93 Subgenus Phytohelea Remm. 93 Subgenus Pterobosca Macfie. 94 Subgenus Rhinohelea de Meillon and Wirth. 95 Subgenus Rhynchoforcipomyia Wirth and Dow. 95 Subgenus Saliohelea Wirth and Ratanaworabhan. 95 CataloG OF THE BITING MIDGES OF THE WORLD Zootaxa 4787 (1) © 2020 Magnolia Press · 3 Subgenus Schineromyia Debenham. 95 Subgenus Schizoforcipomyia Chan and LeRoux . 96 Subgenus Synthyridomyia Saunders. 96 Subgenus Thyridomyia Saunders . 97 Subgenus Trichohelea Goetghebuer. 98 Subgenus Trithicomyia Yu and Song . 100 Subgenus Typhonomyia Debenham . 100 Subgenus Warmkea Saunders. 100 Species of Forcipomyia unplaced to subgenus. 101 Nomina dubia. 101 Subfamily Ceratopogoninae Newman . 102 Tribe Culicoidini Kieffer. 102 Genus Culicoides Latreille. 102 Subgenus Amossovia Glukhova . 102 Subgenus Anilomyia Vargas. 102 Subgenus Avaritia Fox . 103 Subgenus Beltranmyia Vargas. 106 Subgenus Cotocripus Brèthes. 108 Subgenus Culicoides Latreille. 108 Subgenus Diphaomyia Vargas. 110 Subgenus Drymodesmyia Vargas . 110 Subgenus Fastus Liu. ..