Delivered by Publishing Technology to: Bibliothek der Karl-Franzens-Universitaet Graz IP: 143.50.232.80 on: Mon, 17 Jun 2013 06:17:31 Copyright (c) American Society for Plant Taxonomists. All rights reserved. nhams rea h ae(cf., peri- base the the e.g. at definition, free within this almost with anth groups agree not several do Urgineoideae However, the in Urgineoideae. taxa most by capsule subfamily terms general congruent developing in nearly shared the characters is the on which with 2007), cup Goldblatt a at and Manning as abscising (cf., withering bracts, perianth, and floral circumscissile base spurred the and the base, with caducous the day), at the a united to and less up or to more hours tepals several from (living flowers circumscriptions. generic similar consistent more A render alter- to possible rank. evaluate natives to generic Urgineoideae in the needed these still at and is study accepted clades, satisfactorily with be congruent can fully are syndromes morpho- consistent logic analyses, phylogenetic the regions in DNA included nuclear are and plastid sufficient where (2011), al. et Azorı prahs(pt 98 ann ta.20,20;Martı 2009; 2004, al. et different Manning on 1998; (Speta based approaches proposed sce- of recently taxo- This were contrasting treatments where definition unsatisfactory.” nomic Ornithogaloideae, often in “the similar is is that subfamily nario 2012). this commented al. in et himself Pfosser genera (Manning 2004; polyphyletic 2001, (1998) Speta or and Speta para- Pfosser 2004; be al. to et proven these have of several genera However, accepted. were Ornithogaloideae) O 10.1600/036364413X666831 DOI © Botany Systematic ae nmrhlg Sea19) nwihu o1 dif- 10 to up (excluding which in genera approaches 1998), analytical (Speta ferent morphology more on favoured based authors other hand, Thuranthos as Salisb., tradition- such genera, several accepted includes ally and morphology, vegetative and of concept and expanded resulting 1990), al. whole et been the for accepted have are genera subfamily: two treatments (Manning only studies where broad 2004), last phylogenetic al. et the hand, on based in one proposed controversy recently the other of in in On matter a As circumscription decades. been east. generic has the Hyacinthaceae, Urgineoideae to Europe, India of Africa, reaching subfamilies in range. Asia, distributed geographic western are wide and subfamily its this throughout of group knowledgeSpecies poor plant the to this due higher of considerably be taxa of to number likely the but is 2004), al. et (Manning species 100 about oyih 03b h mrcnSceyo ln Taxonomists Plant of Society American the by 2013 Copyright Drimia h aiyHaitaeesbaiyUgnoda includes Urgineoideae subfamily Hyacinthaceae family The rmaciliata Drimia pce.Hr epoieadtie opooia ecito o hsseis nldn clgcladcoooia aa fiiisand Affinities data. chorological and ecological including species, this for discussed. description also morphological are allies detailed close a with divergences provide we Here species. ´ 1 ta.21) oee,a hw yMartı by shown as However, 2011). al. et n II IsiuoUiestrod aBoiesdd,Uiesddd lcne .O o 9 -38 lcne Spain. Alicante, E-03080 99, Box O. P. Alicante, de Universidad Biodiversidad), la de Universitario (Instituto CIBIO Abstract— Keywords— Rhodocodon 2 scaatrzdi eea em yteshort-lived the by terms general in characterized is emrShnadHraim eateto oay hdsUiest,Gaason64,SuhAfrica. South 6140, Grahamstown University, Rhodes Botany, of Department Herbarium, Schonland Selmar Bowiea .H rgt and Wright, H. C. 21) 82:p.332–338 pp. 38(2): (2013), e pce of species new A and rmacochlearis Drimia Distribution, Drimia av xHo.f ihtoseis(Reid species two with f. Hook. ex Harv. .sclerophylla D. Baker, ai Martı Mario aq ihaot10seis The species. 100 about with Jacq. rmaciliata Drimia Schizobasis Igidia Drimia Litanthus Drimia u tdfesi ubro lrladvgttv hrcesta lal upr t eonto sadistinct a as recognition its support clearly that characters vegetative and floral of number a in differs it but , Urginea Urginea pt htblnsto belongs that Speta shr ecie rmSuhAfrica. South from described here is svr aibei flower in variable very is ´ nez-Azorı , ,ntpritn ntop on persistent not ), 3 rmasclerophylla Drimia Baker, tih nteother the On Steinh. Harv., uhrfrcrepnec ([email protected]) correspondence for Author Haitaee,aNwSeisfo ot Africa South from Species New a (Hyacinthaceae), omnctn dtr et agKlitgaard Bang Bente Editor: Communicating Rhadamanthus Tenicroa ´ ´ n, nez-Azorı 1,3 aulB Crespo, B. Manuel clg,txnm,Urgineoideae. taxonomy, ecology, , ´ Raf., nez- ´ n 332 hlg ftpl,admrhlg flae n ub) the mor- bulbs), and and color leaves species (e.g. of new morphology genus and the been tepals, in of have phology taxa that differentiate characters to morphological used to according and Nordenstam 1977; Jessop of Mu (cf., any group 1970; in the found of not taxa are described the that characters of syndrome peculiar non Baker, sclerophylla D. fia ehv eetyfudpat tfrtsgtsimilar sight first at plants found both recently to have we Africa, treatments. and multigeneric alternative broader of a allowing undertaken re-evaluation This until is subfamily accepted 2009). the provisionally in Oliver study here comprehensive is and Manning concept (cf., Manning genus unclear brownish 2002; or al. grey et cream, dull undefined colored inconspic- more rather uously mostly even flowers, caducous short-lived, an the by in Hyacinthaceae, Moreover, within concept with. genus work to and 2003). make Wyk van facts and these Jaarsveld All (van day one than more (cf., lasting withering after capsule the of 4 lnsi n ouain swl as well as population) one in plants (47 and ns uhas related such other and ones, taxa both between existed differences that many revealed characters relevant of resembled examination careful which a ever, Africa, South Cape Western of Karoo, Province Little the in Calitzdorp of surroundings odi eedescribed. here is Dold R,K B,adPE(coysacrigt hes21) Authorities (2012). 2012). IPNI Thiers the ABH, to follow according taxa studied: cited (acronyms of PRE other were and herbaria for NBG, following K, detail GRA, the in from described specimens been Herbarium Martı have by Ornithogaloideae and in groups seeds, related and fruits, leaves, bulbs, flowers, of features quantitative and qualitative both included studies Fg.13 al )i ecie eeiht accommodate to herewith described is plants. distinct 1) these Table 1–3; (Figs. ntefaeo aooi oko rieieei southern in Urgineoideae on work taxonomic of frame the In rmacochlearis Drimia uigtecus ffedok lnswr on nthe in found were plants fieldwork, of course the During ealdmrhlgclsuyo idlvn lnsof plants living wild of study morphological detailed A .sclerophylla D. ¨ rmaciliata Drimia lrDbise l 01.Bsdo u observations, our on Based 2001). al. et ller-Doblies 1 n nhn .Dold P. Anthony and rmarigidifolia Drimia .cochlearis D. .C ann odlt (= Goldblatt & Manning C. J. 3 nfu ouain)wsudrae.Morphological undertaken. was populations) four in (36 .sclerophylla D. aeil n Methods and Materials eut n Discussion and Results p o.i tfrtsgtcoeyrltdt both to related closely sight first at is nov. sp. Drimia L . .C ann odlt and Goldblatt & Manning C. J. f.) (L. Mart.-Azorı .l eydfiutt understand to difficult very l. s. ae) oee,te hwa show they However, Baker). e species, new A . .ciliata D. Rhodocodon ´ 2 ´ nez-Azorı ,M .Cep .P. A. & Crespo B. M. n, Drimia 6 nfv populations) five in (64 rie rigidifolia Urginea ´ ta.(07 2010). (2007, al. et n .ciliata D. ,o h flowers the or ), scharacterized is rmacochlearis Drimia .cochlearis D. How- . Delivered by Publishing Technology to: Bibliothek der Karl-Franzens-Universitaet Graz IP: 143.50.232.80 on: Mon, 17 Jun 2013 06:17:31 Copyright (c) American Society for Plant Taxonomists. All rights reserved. .Bl ihlae,aia iw clsa m :1c;c m :1m;ef cm. 1 e–f: mm; 1 d: mm; 2 c: cm; 1 b: mm; 5 a: Scales view. apical leaves, with Bulb F. 03 MARTI 2013] Fig. 1. rmacochlearis Drimia ´ NEZ-AZORI Mart.-Azorı ´ ,M .Cep .P od .Foe.B nlrsec.C asl.D ed.E ub ihlae,ltrlview. lateral leaves, with Bulbs E. Seeds. D. Capsule. C. Inflorescence. B. Flower. A. Dold. P. A. & Crespo B. M. n, ´ TA. RMACCLAI,ANWSEISFO OT FIA333 AFRICA SOUTH FROM SPECIES NEW A COCHLEARIS, DRIMIA AL.: ET N Delivered by Publishing Technology to: Bibliothek der Karl-Franzens-Universitaet Graz IP: 143.50.232.80 on: Mon, 17 Jun 2013 06:17:31 Copyright (c) American Society for Plant Taxonomists. All rights reserved. oshrcl ltee ttetp 1.6–2.5 top, the at flattened spherical, to pce ninsfoiu rm dpcuab adspectu primu floribus insignis Species evs49dsoe nabslrste otywithered mostly rosette, basal a in disposed 4–9 Leaves h px ot lsy hc,bace,u o50 to up branched, reaching thick, not slightly fleshy, tunics, Roots or outer apex. membranous hypogeal the pale clumps, with dense epigeal, forming proliferous, rmacochlearis Drimia 3 YTMTCBTN Vlm 38 [Volume BOTANY SYSTEMATIC 334 ot Africa. South eiuu ubu lns pt –3c al ubovoid Bulb tall. cm 8–23 to up plants, bulbous Deciduous Fig. uqa iiombssbeeiu rigidioribusque. subteretibus filiformibus brevioribusque, nunquam latioribus multo albidis discrepat obovatis tamen coriaceis foliis posteriore tepalis A instructis, reflexis. purpurascenti valde necnon nunquam interioribus viridi et erectis media viridibus carina pallide floribus foliis pedicellis non longioribus; et differt scarioso-eroso inflorescentia priore margine ciliato; A cucullatis distinguitur. apice cochleatis facile et distincta sclerophylla D. MMA941 2/121 nGaason,33 Grahamstown), in hort. ex flowers (28/01/2012 plants), 29/09/ wild from m, leaves with 408 (bulbs 2011 Reserve), Reserve Nature Nature Gamkaberg Groenefontein Calitzdorp, (former of SW km 11 Cape,ca. Western AFRICA. SOUTH —HOLOTYPE: nov. sp. E, .Gnrlve of view General 2. .Martı M. hltp:GA;ioye:AH K). ABH, isotypes: ; GRA (holotype: ´ nez-Azorı aooi Treatment Taxonomic Mart.-Azorı euas e utscaatrbssat characteribus multos sed aemulans, ´ rmacochlearis Drimia n , .P Dold P. A. ´ ,M .Cep .P Dold, P. A. & Crespo B. M. n, , .Vlok J. Mart.-Azorı
+ 37 .–. m usually cm, 1.5–2.3 & 0 16 .Martı A. rmaciliata Drimia 00 ´ ,M .Cep .P oda h yelclt,GmaegNtr eev,WsenCape, Western Reserve, Nature Gamkaberg locality, type the at Dold P. A. & Crespo B. M. n, ,21 S,
+ ´ nez-Soler mm. 2 38 0 08 et 00 tmn ooopi,sbaetadsraig anthers spreading; reflexed. and of strongly 1 subpatent hairs outers subpatent, apical monomorphic, inners bearing Stamens long; apex, mm cucullate 0.5 slightly ca. or flat with ht,2 white, rdprat aal icmcsieadrmiigattached remaining and with- the circumscissile with basally apex, perianth the to ered valves base mature; the when from completely pale-brown splitting section, in subspherical to uia tie,1.1–1.5 stripes, tudinal h hl egh iaet uclnrcl ht,smooth, white, subcylindrical, filaments 1.9–2.5 length; whole the ueet easwiewt aegenmda tie ovate- stripe, median 4–6 green or oblong, pale erect a with Flowers white spur. mm tepals basal suberect; 1 evident to an up with pedicels, lowermost apex; than long, small the shorter very much to acuminate, inconspicuous, base and triangular, the to from ovate-lanceolate shortening bracts slightly patent green, suberect, pale long, or in mm papillate (13)15–20 green, pedicels pale part; long, basal cm the 5–10 peduncle flowers, long; 12–32(41) cm with 4–11 erose, raceme, mm erect 0.7–1 membranous, an ca. Inflorescence a of width. margin with undulate cucullate sometimes base, and with the translucent concave at surface, overlapping smooth apex, with green, glossy n nosiuu.Cpueoae a 5 ca. ovate, Capsule inconspicuous. and tfoeigtm,sbrc rsraig .–.(. nculti- in 1.9–3.3(4.5 spreading, or vation) suberect time, flowering at + . m etl n eicn ogtdnlyalong longitudinally dehiscing and fertile mm, 0.5 + +
+ longi- green pale with white ovate, Ovary mm. 0.5 .–. m bvt-bog im etey dark leathery, firm, obovate-oblong, cm, 0.7–0.9
. m sal et tgawie trigonous white, stigma bent; usually mm, 0.5 + .– m hrl ue tbs c.04mm), 0.4 (ca. base at fused shortly mm, 1.5–2
+ –. m esl;syefiliform, style sessile; mm, 1–1.3
+ m trigonous mm, 4 Delivered by Publishing Technology to: Bibliothek der Karl-Franzens-Universitaet Graz IP: 143.50.232.80 on: Mon, 17 Jun 2013 06:17:31 Copyright (c) American Society for Plant Taxonomists. All rights reserved. oae ihnanro aiuia tiernigfo ca. are from which ranging apart, stripe km latitudinal 33 narrow 400 a about within localities located several in Cape) (Eastern mountains Baviaanskloof to Province) Cape (Western ein fteScuetKroBoe(i.4 Mcn and (Mucina altitude. southern m 4) 900 the (Fig. and 150 in Biome between Karoo mostly 2006), Rutherford Succulent shrublands, the dry of rocky in on regions ledges, soils, sandy or deep in soils clumps dense forming usually April. to November from appear fruits and Cape), Eastern Grahamstown, in cultivation in (March January 1. Table 1–3, Figures black. to brown ie ogrta eas lwr rc rsbrc;tepals suberect; or erect flowers 4–6 4– tepals; about ovate-oblong, than mm, longer (13)15–20 at times of withered pedicels 5 long mostly 12–32(41) patent and with and raceme a–b), long flowers 3 a in (Fig. inflorescence width time; flowering mm 0.7–1 margin ca. undulate of sometimes and translucent green, membranous, with glossy apex, a dark the at are cucullate and that concave obovate, leaves leathery, firm, the by recognized easily is n reual opesd nld a 2–3 semidiscoidal ca. Seeds angled, cup. apical compressed, an irregularly as and capsule the of top the to MARTI 2013] ot fia n (c–d) and Africa) South Distribution— Habitat— Time— Flowering Fig. igotcCaatr n Relationships— and Characters Diagnostic o34.5 to .La agndtis (a–b) details: margin Leaf 3. ln h aeFl et(i.5). (Fig. Belt Fold Cape the along S rmacochlearis Drimia h pce skonfo Bredasdorp from known is species The rmaciliata Drimia tfoesfo aeSpebrt early to September late from flowers It
+ ´ NEZ-AZORI .– m nessbaet outers subpatent, inners mm; 1.5–2 rw cainlysnl or singly occasionally grows rmacochlearis Drimia L . .C ann odlt Baesvle,Pr lzbt,EsenCape, Eastern Elizabeth, Port valley, (Baakens Goldblatt & Manning C. J. f.) (L. ´ TA. RMACCLAI,ANWSEISFO OT FIA335 AFRICA SOUTH FROM SPECIES NEW A COCHLEARIS, DRIMIA AL.: ET N Mart.-Azorı
rmacochlearis Drimia + – m dark mm, 1–2 ´ ,M .Cep .P od(yeseie:GmaegNtr eev,WsenCape, Western Reserve, Nature Gamkaberg specimen: (Type Dold P. A. & Crespo B. M. n, sal r arwyoln ooln-ieri outline, in observed varia- in been a also pseudopetiole, oblong-linear has a that leaves in tion to base cultivation, the oblong at In narrowing 1). sometimes narrowly Table are 1–3; usually (Fig. reflexed strongly rmtebs oteae in begins apex that the dehiscence to capsule base the the in Dold from between found 1897; is difference 1872, species remarkable Linnaeus both Another Baker 2000). 1782; 1843; Moberg fil. Kunth and Linnaeus 1823; (cf. Thunberg bear- keel are 1784; tepals brown-reddish pedicels white with a and nodding part ing are flowers fertile the its and stem, shorter, and pedicels maroon h omrpoue ayfoee nlrsecs with flowers larger inflorescences, usually 1). and (Table many-flowered pedicels reddish-brown produces shorter do Furthermore, species. which former new 1897), the with the 1878, all Baker at confusion (cf. allow not rigid and wiry, somewhat in cochlearis base D. the to sclerophylla apex the from and in flat, inflorescence The ground, c–d). long 3 the and However, (Fig. densely to margins with morphology. ciliate sprouting, appressed when are convex flower slightly latter even general the basal spreading in the a leaves in and disposed leaves rosette, smooth green, glossy rmacochlearis Drimia sas iia ntrso lwrmrhlg to morphology flower of terms in similar also is u t evsaevr ogadnro,terete, narrow, and long very are leaves its but , hrswith shares .cochlearis D. .ciliata D. Martı .ciliata D. .ciliata D. ´ nez-Azorı h hr n wide, and short the hra tbegins it whereas Tbe1). (Table ´ . 7 tal. et 979 n .ciliata D. GRA). Drimia has Delivered by Publishing Technology to: Bibliothek der Karl-Franzens-Universitaet Graz IP: 143.50.232.80 on: Mon, 17 Jun 2013 06:17:31 Copyright (c) American Society for Plant Taxonomists. All rights reserved. 3 YTMTCBTN Vlm 38 [Volume BOTANY SYSTEMATIC 1.6–2.5 Bulb 336 ot Africa. South ed a 2–3 ca. Seeds lwrn ielt etme oeryJnayJnayt erayNovember-January February to January January early to September late time Flowering evs4–9 Leaves nlrsec 23(1 flowers 12–32(41) Inflorescence asl 5 Capsule eas4–6 Tepals Fig. Table .Hbttof Habitat 4. .Mi igotccaatr between characters diagnostic Main 1. nessbaet uessrnl reflexed strongly outers subpatent, keel inners green pale a with white ihrda lwrn time flowering at withered margin translucent, erose, membranous, base the at overlapping apex cucullate with concave glossy dark leathery, firm, obovate-oblong, spreading or suberect avssltigcmltl rmbase from completely splitting valves .–.(. ncultivation) in 1.9–3.3(4.5 lwr rc tteanthesis the at erect flowers green pale stem and pedicels long mm (13)15–20 pedicels long cm 4–11 part fertile + rmacochlearis Drimia fc .– mwdh oeie undulate sometimes width, mm 0.7–1 c. of surface smooth with green, oapex to 4mm +
+ mm 1.5–2 + – m2–2.5 mm 1–2 .–. 2–2.5 1.5–2.3 rmacclai rmaclaaDii sclerophylla Drimia ciliata Drimia cochlearis Drimia Mart.-Azorı
+ .–. cm 0.7–0.9 .cochlearis D. ´ ,M .Cep .P oda h yelclt,GmaegNtr eev,WsenCape, Western Reserve, Nature Gamkaberg locality, type the at Dold P. A. & Crespo B. M. n, , .ciliata, D. 5–6 avssltigcmltl rmapex from completely splitting valves 0.8–3 10 3–7 ihrda lwrn time flowering at withered margins straight ciliate, long densely, base the at sprouting overlapping when convex slightly or flat firm, lorate, or ovate-elliptical ovate, prostrate nesptn,otr lgtyreflexed slightly outers patent, inners brown light a with white translucent lwr odn tteanthesis the at nodding flowers glossy reddish-brown, stem and pedicels long mm 4–14 pedicels long cm 2–6 part fertile flowers 10–15 ordihkeel reddish to obase to mohsurface smooth with green, glossy dark leathery, + + and + + .– mm 2.5–5 + .–. mm 1.9–2.6 mNtseen Not mm 2 2.5–3 cm 2–2.5 .–. cm 0.3–1.9 .sclerophylla D. . o seen Not 9–17 3–5 4–5 atal ihrdo re at green or withered partially margin glabrous base at overlapping not subterete nervose, wiry, rigid, linear, narrowly erect nessbaet uessubpatent outers subpatent, inners brown- a with white translucent lwr odn tteanthesis the at nodding flowers reddish-brown stem and pedicels long mm 5–13 pedicels long cm 10–15 part fertile flowers 30–56 lwrn time flowering glabrous green, osihl reflexed slightly to keel green to purple + + + .–. mm 1.9–2.8 . cm 0.1 .–. cm 2.8–3.5 Delivered by Publishing Technology to: Bibliothek der Karl-Franzens-Universitaet Graz IP: 143.50.232.80 on: Mon, 17 Jun 2013 06:17:31 Copyright (c) American Society for Plant Taxonomists. All rights reserved. 03 MARTI 2013] ndegrees. in s‘ as collection the ieo nsegEo rd,70m 3Sp1990, Sep 23 m, 720 Vrede, of E Anysberg of 1978, side May 09 Swellendam, of SW 720 km 6 (3420AB), Bredasdorp Cape: od(33,c.1 msuhtecosn ihR2 33 R62, with crossing Riversdale-Ladismith the the south on km 18 Ladismith, ca. of (R323), road south km 30 ca. Farm, Blouhuis es he ifrn pce,o hc hs aigbulbs having those at which of of material species, to vegetative correspond different only three includes least It 1990. December 3Dc18 n lwr B 60/92 evsNG07/07/1982, NBG leaves 06/01/1982, NBG hort. flowers ex flowers and 375 Snijman mountains, 1981 1980, Baviaanskloof Sep Dec of 28 13 side track, north Enkeldoorn on the of Wilgerivier side south 1431 the Perry from house, up managers gate third behind top 1980, hill 1998, Sep on Nov 30 Baviaanskloof, Farm, Geelhoutbos, Hoop Cape: Avontuur De Uniondale, towards off 1982, kilometres Jul road few 07 a NBG ex (3323CB), leaves Oudtshoorn, Willowmore 1981, (NBG); Dec of 13 hort. S ex flowers km Farm, 18 1985, (3322CB), Dec 03 Oudtshoorn 10–09–2012]; accessed WORLDS_BULBS/GRAPHICS/Ornithogalum/Ornithogalumlist.shtml); (http://www.bulbsociety.org/GALLERY_OF_THE_ website (http:/ collection website Grootscholten plant-collections/bulbs/britteni Ine as and labelled [pictures m, 250 ca. evs( leaves point. this clarify to of undertaken species being is other work in Molecular found not is regard- Drimia which aggregate distinct structure, a flower form vegeta- ing taxa in differ These they characters. but tive structure, flower similar very share te ersnaieSeiesExamined— Specimens Representative Other mn h tde aeil of materials studied the Among Fig. Etymology— species mentioned above three all that remarkable is It NG;LdsihDsrc 32B) nsegae,vle nnorth on valley area, Anysberg (3320BC), District Ladismith (NBG); rie pulchellomarginata Urginea .Kondsrbto of distribution Known 5. .l,ti vdnigcoeeouinr relationships. evolutionary close evidencing this l., s. cochlearis od43,Cak&Cocks & Clark 4432, Dold NG;Setevle(34B,aogtakfo nedonto Enkeldoorn from track along (3324CB), Steytlerville (NBG); (NBG). lk869 Vlok lvr9750 Oliver .cochlearis D. h aerfr otecnaesaeo the of shape concave the to refers name The spoon-like). = NG;Wlomr 32C) nodl,D Hoop De Uniondale, (3323CB), Willowmore (NBG); ´ NEZ-AZORI rihglmbritteniae Ornithogalum NG er nietfcto label identification an bears (NBG) . ehm)adPnsadSarsprivate Stavros and Panos and ae.html) rmacochlearis Drimia GA;Setevle(34B,na the near (3324CB), Steytlerville (GRA); .&D .D s’dtd17 dated msc’ M.-D. D. & U. rmacochlearis Drimia ´ /public.fotki.com/Grootscholten/ TA. RMACCLAI,ANWSEISFO OT FIA337 AFRICA SOUTH FROM SPECIES NEW A COCHLEARIS, DRIMIA AL.: ET N er 1448 Perry OT FIA Western AFRICA. SOUTH Mart.-Azorı 44 lvr9750 Oliver f.a ohteCok the both at aff. 0 06 NG.Eastern (NBG). 00 sebelow), (see S21 nja 372 Snijman ´ ,M .Cep .P odi otenSuhArc.Ae hwlniueadlatitude and longitude show Axes Africa. South southern in Dold P. A. & Crespo B. M. n, 09 (NBG); 0 59 Perry 00 E, od .P n .Mbr.20.Creto fahsoia ro nthe in error historical a of Correction 2000. Moberg. R. and P. A. Dold, Ramo esp .P 97 tde nteblosLlaeei ot fia 7. Africa: South in Liliaceae bulbous the in Studies 1977. P. http://www.ipni.org J. index. Jessop, names plant international The 2012. IPNI. 1897. G. J. Baker, inesfl,C 1782. C. fil., Linnaeus 1784. C. Linnaeus, 1843. S. C. Kunth, and Liliaceae. Scilleae known of little and species new of and Descriptions genera 1878. G. the J. of Baker, Revision 1872. G. J. Baker, of details locality providing for Grootscholten Ine cochlearis and D. Cok to Thanks nteWsenCp n atr aePoicsrsetvl.Susan of respectively. illustration the Provinces preparing Cape for Eastern thanked and is Development Cape specimens Abraham herbarium Economic Western collect of both the to permission Department also in for and Cacadu Affairs cited examined, Environment the the and specimens of and to curators Nature access the Cape thank provided We who for specimens. Reserve, herbaria, field Nature of with Gamkaberg of assisted collecting manager kindly Barry, facilitating and Tom and species 2011 new in the work discovered who Vlok Jan ann,J . .Glbat n .F a.20.Arvsdgeneric revised A 2004. Fay. F. M. and Goldblatt, P. C., J. 2002. Manning, Snijman. D. and Goldblatt, P. C., J. Manning, Acknowledgments. aooi ecito of description taxonomic Co. and Reeve London: Thiselton-Dyer. T. W. fia Botany African h aooyof taxonomy The 2012]. August [accessed Foreign and British Botany. of pce u hrceiu tdfeeti.Eii eiaquarta decima Dieterich. Editio C. J. differentiis. et characteribus cum species differentiis characteribus, quartus Tomus adjectis synonymis. disposita, et naturales familias secundum Chlorogaleae. yosso ycnhca nSbShrnArc,bsdon based Africa, Sub-Saharan in Hyacinthaceae of synopsis Bulbs Cape of ´ rcs(pi)adUiesddd lcne(pi) ethank We (Spain). Alicante de Universidad and (Spain) Areces n . otad ibrPress. Timber Portland: . ora fteLnenScey Botany Society. Linnean the of Journal 3 265–319. 43: Drimia ytm eeaiimscnu lse rie genera ordines classes secundum vegetabilium Systema nmrtopatrmonu uuqecognitarum, hucusque omnium plantarum Enumeratio upeetmplantarum Supplementum Drimia hswr a atyspotdb Fundacio by supported partly was work This p 3–4 in 436–443 Pp. . ieaueCited Literature rie ciliata Urginea n eti lidgenera. allied certain and ttada tTbna:J .Cottae. G. J. 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