Atlas of Yorkshire Coleoptera (VCs 61-65)
Part 5a – Staphylinidae: Aleocharinae
Introduction
This is Part 5a…. This section of the atlas deals with the subfamily Aleocharinae being part of the family Staphylinidae.
Each species in the database is considered and in each case a distribution map representing records on the database (at 1/06/2018) is presented. The number of records on the database for each species is given in the account in the form (a, b, c, d, e) where 'a' to 'e' are the number of records from VC61 to VC65 respectively. These figures include undated records (see comment on undated records in the paragraph below on mapping). As recorders, we shall continue to use the vice- county recording system, as the county is thereby divided up into manageable, roughly equal, areas for recording purposes. For an explanation of the vice-county recording system, under a system devised in Watson (1883) and subsequently documented by Dandy (1969), Britain was divided into convenient recording areas ("vice-counties"). Thus Yorkshire was divided into vice-counties numbered 61 to 65 inclusive, and notwithstanding fairly recent county boundary reorganisations and changes, the vice-county system remains a constant and convenient one for recording purposes; in the text, reference to “Yorkshire” implies VC61 to VC65 ignoring modern boundary changes.
“Recent” records are those considered to be later than 1 January 2000 – this is also the break point used in the production of the distribution maps. (In previous parts of this Atlas, the break point for recent and otherwise was 1 January 1990).
Habitat A brief statement on habitat is included where known and relevant.
Taxon status Inclusion of a statement on the status of species presents problems in that those designations in Recorder 6 are often out of date. Recently Natural England has been publishing reports on various groups of Coleoptera which bring the national status of species in line with present knowledge and recording. Recorder 6 statuses are based largely on Hyman and Parsons (1992 and 1994), see below for status category explanations. RED DATA BOOK CATEGORY 1. RDB1 – ENDANGERED - Species which are known or believed to occur only as a single population within one 10km square of the OS National Grid. RED DATA BOOK CATEGORY 3. RDB3 – RARE - Species which are estimated to exist in only 15 or fewer 10km squares; this criterion may be relaxed where populations are likely to exist in more than 15 squares but occupy small areas of especially vulnerable habitat. RED DATA BOOK CATEGORY I. RDB I – INDETERMINATE - Taxa considered to be Endangered, Vulnerable or Rare, but where there is not enough information to say which of the three categories (RDB1 to 3) is appropriate. RED DATA BOOK CATEGORY K. RDBK - INSUFFICIENTLY KNOWN - Taxa recently discovered or recognised in Great Britain which may prove to be more widespread in the future (although some recent discoveries may be placed in other categories if the group to which they belong is thought not to be under- recorded). PROVISIONAL RED DATA BOOK. pRDB - The prefix 'p' before any Red Data Book category implies that the grading is provisional. NATIONALLY SCARCE - Species which are thought to occur in between 16 and 100 10 km squares of the National Grid or, for less well recorded groups between eight and twenty vice-counties.
Maps Distribution maps have been produced using Dr Alan Morton’s DMAP software (version 7.0b). Unlike previous atlases, which omitted undated records from the maps, it has now been possible to include these undated records. Any records flagged on the YNU database as unconfirmed are not mapped, however. In addition, the database possesses a few peat sub-fossil records which are not considered for inclusion in the atlas. Nomenclature The YNU database is presently held in a copy of Recorder 6, which uses a new Coleoptera checklist based on Duff (2008) and this new list incorporates a very large number of taxon name changes since the checklist of Pope (1977); there is now a second edition of the latest Coleoptera checklist (Duff, 2012) which has not yet been incorporated fully into Recorder 6. Atlas of Coleoptera of Yorkshire. Part 5a – Staphylinidae: Aleocharinae Page | 1
Occasionally we receive records using “noddy names” (see Cooter, (1999) and Key (1999) for an excellent treatment of this subject), e.g. “Snail Hunter”, “Crucifix Beetle” and “Heath-glory ground beetle” (Cychrus caraboides, Panagaeus crux-major and Carabus nitens respectively). Such inventions were designed for use in political documents, and seem to have originated somewhere within Natural England (previously English Nature) or JNCC. For the purposes of serious science these terms are, in our opinion, either vague, ridiculous, confusing or unhelpful. We do not accept records using such names only, unless accompanied by specimens or other convincing evidence.
For the purposes of this Atlas the 2nd edition of the checklist by Duff (2012) will be used as the basis for nomenclature and arrangement of species. As this part of the Atlas goes to press there is new 3rd Edition of the British Checklist (Duff, 2018) which will be used in future sections of the Atlas.
Quality of the records
A regrettable aspect of many records within the YNU database is the lack of detail accompanying the record, especially of accurate grid references, lack of pertinent comments as to the habitat of the capture, its association with plants/animals, and sometimes, in the case of some modern records, a lack of care in determination, and the recording of the sex of the specimen. Old records, although sound, tend not to give much or indeed any of this detail. Old records on the database so rarely give habitat details but sometimes interpretation of the site name can give clues to the habitat type. Grid references could not be applied before 1930 anyway, and grid references do not figure in the paper archive of the YNU Coleoptera database until the late 1970s. Apparently it was not considered necessary to give that level of accuracy. Entering records into Recorder 6 requires a grid reference, and with older records I have added a grid reference where I believe this has been possible. If the site name is a vague one I have used a 10km reference, and where a 1km reference is deemed accurate enough I have included that. There can be little excuse with modern records not to include some accurate comment with the record, and at least a six-figure grid reference. Taxonomic knowledge and expertise have increased greatly since the days of Walsh, Bayford and Stainforth. Much good literature now exists for the taxonomist in the form of dichotomous keys, and when identifying specimens, especially with critical species pairs or groups, too much reliance on the outcome of keys can be placed on a determination, without apparently considering the habitat in which the specimen was taken, or the host plant/animal. Atlases of the national distributions of various orders of insect are now becoming available and should always be consulted. These often give a guide to the likelihood or otherwise of a determination, especially in scarce taxa. Museum collections containing correctly determined and labelled specimens are invaluable in accurate recording; museum collections are there for our use and should be consulted. If a species is ‘new’ to a vice-county or is very significant in some other respect, a second opinion should always be sought or an acknowledged specialist in the group consulted. A voucher should always be retained by the collector if possible. The recorder can only deal with the information he is given, and often has to beware of identifications made on photographic evidence only, and identifications of ‘difficult’ species by non-specialists. It takes many years to become competent in many areas of entomology, and the recorder often has to rely on the expertise of co-workers, especially when records covering large Orders are being documented. We have adopted a policy in our county recording whereby any received record of a species afforded Notable B or above, especially if a member of a ‘difficult’ beetle group, should be confirmed by a specialist in that group before the record is entered into the database as a correct record.
Ultimately, the responsibility for verification of records on the YNU database rests with the two authors. With modern records received from reputable coleopterists there is little problem. Some older records requiring vouchers for confirmation have been retained in the database but flagged as requiring confirmation.
Contributors
ABD – Tony Drane EJS – Eric Smith AG – Andy Godfrey EWA – Ted Aubrook AS – A Smith EWM – EW Morse ASL – Alan Lazenby FEK – Frank Kenington CES – CE Stott GS – G Simpson CJ – Colin Johnson HB – Harry Britten DAL – Derek Lott HHC – HH Corbett DH – David Heaver HRL – Horace Last DGH – David Hemingway JA – J Ashworth DM - Dave Maude JAO – John Owen EAN – EA Newbury JHD – JH Day EAW – EA Waterhouse JM – J Muona ECH – EC Horrell JRDL – JRD Lister Atlas of Coleoptera of Yorkshire. Part 5a – Staphylinidae: Aleocharinae Page | 2
KGP – Ken Payne RLF – RL Fowler KL – K Lawson RM – Robert Merritt LA - Len Auckland ROSC – ROS Clarke ME – Mick Eyre RRUK – RRU Kaufmann MGT – Mark Telfer RSK – Roger Key MH – Martin Hammond RW – Richard Wilson MLD – Mike Denton SAW – Alex Williams MLL – Martin Luff SGC – Steve Compton MLT – ML Thompson SE – S Ellis PH – Peter Hammond SLM – Seth Lister Mosley PJH – Peter Hodge TS – T Stainforth PK – Peter Kendall TW – T Wilkinson PR – Paul Richards WAE – Bill Ely PS – Peter Skidmore WCH – WC Hey RBA – Robert Angus WDH – Walter Douglas Hincks RCa – Ronald Carr WJF – William John Fordham RCl – Robin Clarke WJS – WJ Sanders RGB – Roger Booth WOS – Bill Steel RGL – Dick Loxton WRD – Bill Dolling RJH – Robin Hunt YNU – Yorkshire Naturalists' Union (various reports in RJM – Bob Marsh Naturalist) RL – R Lawson YU – York University
Atlas of Coleoptera of Yorkshire. Part 5a – Staphylinidae: Aleocharinae Page | 3
Subfamily ALEOCHARINAE Fleming, 1821
Tribe ACTOCHARINI Bernhauer & Schubert, 1911
ACTOCHARIS Sharp, 1870 readingii Sharp, 1870
NO RECORDS
Tribe ALEOCHARINI Fleming, 1821
ALEOCHARA Gravenhorst, 1802
Subgenus ALEOCHARA Gravenhorst, 1802 brevipennis Gravenhorst, 1806
Aleochara (Aleochara) brev ipennis Grav enhorst, 1806
3 WATSONIAN (3,0,1,0,1) There are five records, from 2 YORKSHIRE three localities on the YNU database: Bretton Park (SE2812), 17/07/1981 1 (MLD); Holmpton (TA3624), 21/09/1998 0 NY NZ NZ OV SD SE SE TA (FEK), in grass cuttings; 02/04/1999 (FEK), 9 on gorse; and 11/01/2000 (FEK) and 8 Nosterfield NR (SE2779), 18-19/07/2018 7 (MH det. MLD), in pitfall traps. The status of Nationally Scarce is rather misleading 6 as the species is found throughout the 5 British Isles where it appears to be 4 hygrophilous. 3 2 1
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curtula (Goeze, 1777)
Aleochara (Aleochara) curtula (Goeze, 1777)
3 WATSONIAN (12,14,61,18,12) Widely distributed and 2 YORKSHIRE common in Yorkshire. Found throughout the British Isles (with the exception of 1 northern Scotland), this is the most 0 NY NZ NZ OV SD SE SE TA common carrion-frequenting British 9 member of the genus. Most Yorkshire 8 specimens have been found in 7 association with carrion, but it has also been recorded from a grass pile, pitfall 6 traps under Calluna, and in horse and 5 cow dung. 4 3 2 1
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lata Gravenhorst, 1802
NO RECORDS
Subgenus BARYODMA Thomson, C.G., 1858 intricata Mannerheim, 1830
Aleochara (Baryodma) intricata Mannerheim, 1830
3 WATSONIAN (2,6,12,2,0) Widely distributed in Yorkshire, but not common. Yorkshire 2 YORKSHIRE specimens have been located in flood 1 refuse and the dung of horse, cow and 0 NY NZ NZ OV SD SE SE TA deer. 9 8 7 6 5 4 3 2 1
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Subgenus CERANOTA Stephens, 1839 ruficornis Gravenhorst, 1802
Aleochara (Ceranota) ruficornis Grav enhorst, 1802
3 WATSONIAN (6,8,16,3,1) Widely distributed in the 2 YORKSHIRE county, but not common. This Nationally Scarce species is found throughout 1 England and southern Scotland where it 0 NY NZ NZ OV SD SE SE TA has been recorded from a variety of 9 habitats (although woodland appears to 8 provide its main habitat). The few 7 Yorkshire records with collecting data show that they have been found in fungi 6 (including Agaricus campestris), and pitfall 5 traps. 4 3 2 1
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Subgenus COPROCHARA Mulsant & Rey, 1874 bilineata Gyllenhal, 1810
Aleochara (Coprochara) bilineata Gyllenhal, 1810
3 WATSONIAN (2,4,6,0,0) Despite the interest shown in the subgenus Coprochara by a number of 2 YORKSHIRE Yorkshire coleopterists and visiting 1 specialists, this species appears to be 0 NY NZ NZ OV SD SE SE TA genuinely rare in the county and, rather 9 worryingly, there is only a single post-2000 8 record. Yorkshire specimens have been found in dog dung (in sand dunes and an 7 inland garden) and pitfall traps. 6 5 4 3 2 1
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binotata Kraatz, 1858
Aleochara (Coprochara) binotata Kraatz, 1858
3 WATSONIAN (1,0,0,0,0) There is a single record on the 2 YORKSHIRE YNU database: Spurn (TA4212), 23/07/1963 (RCW). The single specimen, a 1 female, was originally described as A. 0 NY NZ NZ OV SD SE SE TA verna Say, 1836 (Welch, 1969), but its true 9 identity has now been correctly 8 determined and further details can be 7 found in Welch, 1990. This was the first British record and, although now found to 6 be more widely distributed in England and 5 Wales, it still remains a very rare species. 4 3 2 1
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bipustulata (Linnaeus, 1761)
Aleochara (Coprochara) bipustulata (Linnaeus, 1761)
3 WATSONIAN (28,8,50,9,1) The realisation that more 2 YORKSHIRE than one species was masquerading under this name (Welch, 1997) means that 1 without voucher specimens all records on 0 NY NZ NZ OV SD SE SE TA the YNU database are suspect. Dissection 9 of available and more recent specimens, 8 however, has indicated that the species is 7 common and widely distributed in the county. Yorkshire specimens have been 6 located in dung (dog, horse and sheep), 5 carrion, fungi, decaying vegetation, and in 4 the litter in beds of Phragmites and Carex 3 paniculata. 2
1
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verna Say, 1836
Aleochara (Coprochara) v erna Say, 1836
3 WATSONIAN (5,3,5,3,1) A revision of the subgenus 2 YORKSHIRE Coprochara (Welch, 1997) has indicated 1 that the species is far more widespread in
NY NZ NZ OV the British Isles than previously thought. 0 SD SE SE TA 9 Prior to this revision the species had only been recorded from the Hebrides (Inner 8 and Outer) and afforded the national 7 status pRDBK; this will obviously require 6 revision. Yorkshire specimens have been 5 located in flood refuse, dung (cow and 4 sheep), decaying vegetation, old fungi, 3 under seaweed, and in a dead fawn. 2
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Subgenus EMPLENOTA Casey, 1884
obscurella Gravenhorst, 1806
Aleochara (Emplenota) obscurella Grav enhorst, 1806
3 WATSONIAN (13,11,0,0,0) There are records from the full length of the Yorkshire coast, from 2 YORKSHIRE Saltburn (NZ6721) in the north to Spurn 1 (TA4212) in the south; also along the 0 NY NZ NZ OV SD SE SE TA Humber Estuary as far as Sunk Island 9 (TA2819). This is a littoral species which is 8 widely distributed in seaweed and carrion around the coasts of the British Isles. 7
6 5 4 3 2 1
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phycophila Allen, 1937
NO RECORDS
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Subgenus POLYSTOMOTA Casey, 1906
POLYSTOMA Stephens, 1835 non Zeder, 1800 grisea Kraatz, 1856
Aleochara (Polystomota) grisea Kraatz, 1856
3 WATSONIAN (9,4,0,0,0) There are records from four 2 YORKSHIRE localities on the YNU database: Redcar (NZ6024), date unknown (MLT & WJF det. 1 EAN); Saltburn (NZ6721), date unknown 0 NY NZ NZ OV SD SE SE TA (MLT); Spurn (TA4212), regularly between 9 1950 and 2016 (WOS, RCW etc.), mainly 8 from strandline seaweed on the Humber 7 shore but also from carrion; Kilnsea (TA4212), 04/05/1999 (WRD teste MLD). 6 This is a littoral species which is widely 5 distributed in seaweed and carrion around 4 the coasts of the British Isles. 3 2 1
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punctatella Motschulsky, 1858
Aleochara (Polystomota) punctatella Motschulsky, 1858
3 WATSONIAN (3,4,0,0,0) This is a littoral species which is 2 YORKSHIRE widely distributed around the coasts of the British Isles. 1
NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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Subgenus RHEOCHARA Mulsant & Rey, 1875 spadicea (Erichson, 1837)
Aleochara (Rheochara) spadicea (Erichson, 1837)
3 WATSONIAN (6,4,2,6,0) Widely distributed, especially 2 YORKSHIRE in the eastern half of the county, but not common. Rather worryingly, however, is 1 the fact that there are no post-2000 0 NY NZ NZ OV SD SE SE TA records. Although present throughout the 9 British Isles, most records are from 8 southern England. The species is usually 7 found in the subterranean nests of the Mole. The few Yorkshire records with 6 collecting data show they have been 5 found in pitfall traps and the nests of 4 moles and a social wasp. 3 2 1
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Subgenus XENOCHARA Mulsant & Rey, 1874 cuniculorum Kraatz, 1858
Aleochara (Xenochara) cuniculorum Kraatz, 1858
3 WATSONIAN (4,0,3,2,1) Widely distributed in the 2 YORKSHIRE county, but rare. This species is usually found in or in the vicinity of Rabbit 1 burrows, and it is possibly associated with 0 NY NZ NZ OV SD SE SE TA Rabbit dung. It has also been recorded 9 from the nests and burrows of Badger and 8 Sand Martin. Yorkshire specimens have 7 been located at the entrance to Rabbit burrows, in fungi, in pony dung, and by 6 sweeping vegetation. 5 4 3 2 1
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discipennis Mulsant & Rey, 1853
Aleochara (Xenochara) discipennis Mulsant & Rey, 1853
3 WATSONIAN (0,0,2,0,0) There are two records of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Roche Abbey (SK5489), 1 18/05/1986 (WAE det. CJ); Wharncliffe 0 NY NZ NZ OV SD SE SE TA Wood (SK2998), 02/05/2000 (EJS teste 9 MLD). 8 7 6 5 4 3 2 1
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fumata Gravenhorst, 1802
Aleochara (Xenochara) fumata Grav enhorst, 1802
3 WATSONIAN (0,0,3,0,0) There are three records on the YNU database: Newmillerdam (SE3315), 2 YORKSHIRE 15/07/1966 (EWA teste RCW); Hatfield 1 Moor (SE7005), 30/07/1991 (PS det. 0 NY NZ NZ OV SD SE SE TA MLD); Loxley Common (SK3090), 9 29/07/1993 (PR det. EJS). This species is 8 generally found in fungi, although there are records from carrion and nests of owls. 7 It is mainly confined to south-east 6 England, but there are a few records from 5 northern England and Scotland. It is 4 afforded the national status pRDBK. 3 2 1
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funebris Wollaston, 1864
Aleochara (Xenochara) funebris Wollaston, 1864
3 WATSONIAN (1,2,19,0,2) Widely distributed, especially 2 YORKSHIRE in the southern half of the county, but rare. The species was found repeatedly at 1 Blackmoorfoot Reservoir (SE0912) 0 NY NZ NZ OV SD SE SE TA between 1983 and 2007. Although it 9 occurs in a variety of habitats, it is most 8 often found in carrion. 7 6 5 4 3 2 1
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inconspicua Aubé, 1850
Aleochara (Xenochara) inconspicua Aubé, 1850
3 WATSONIAN (0,0,1,0,0) There is a single record on the 2 YORKSHIRE YNU database: Thorne Moor (SE7514), 24/10/1990 (DH det. CJ), in a water trap. 1 The larvae are parasitic in the puparia of 0 NY NZ NZ OV SD SE SE TA the Wheat Bulb Fly Delia coarctata. It has 9 only been recorded from England, where 8 it is very thinly distributed and has the 7 status pRDBK.
6 5 4 3 2 1
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kamila Likovský, 1984
Aleochara kamila Likov ský, 1984
3 WATSONIAN (1,1,1,3,0) There are six authentic records, 2 YORKSHIRE from five localities, of this Nationally Scarce species on the YNU database: 1 Knaresborough Ringing Station (SE3656) 0 NY NZ NZ OV SD SE SE TA (no longer extant), 03/1971 (EWA det. 9 MLD); Drop Clough (SE0413), 04/02/1987 8 (MLD); Willowgarth, Knottingley (SE5124), 7 10/05/1992 (RJH det. MLD teste CJ) and 23/04/1993 (RJH det. MLD); Wheldrake 6 Ings (SE7044), 13/02/1993 (MLD & RJM), 5 in flood refuse; May Moss (SE8796), 4 02/09/1996 (LA det. MLD). 3 2 Past confusion with A. diversa Sahlberg, J., 1876 renders the following records 1 unreliable without voucher specimens: 0 SD SE SE TA SJ SK SK TF Pannal Ash (SE3052), 04/1936 (RRUK); 9 Leeds (SE33?), 25/04/1948 (WJS det. 8 post 2000 WOS). pre 2000 Map produced for Yorkshire Naturalists' Union by DMAP 8/2/2019 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
lanuginosa Gravenhorst, 1802
Aleochara (Xenochara) lanuginosa Grav enhorst, 1802
3 WATSONIAN (15,24,110,23,17) Widely distributed and common in Yorkshire. This species can 2 YORKSHIRE virtually be guaranteed to be present in 1 most ungulate dung (especially that of 0 NY NZ NZ OV SD SE SE TA cow) and also in a number of other 9 habitats, including decaying vegetable 8 matter, fungi, and carrion.
7 6 5 4 3 2 1
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lygaea Kraatz, 1862
Aleochara (Xenochara) lygaea Kraatz, 1862
3 WATSONIAN (0,2,0,0,0) There are two records of this 2 YORKSHIRE RDB I species on the YNU database: Saltburn (NZ6721), 04/1898 (MLT); 1 Acklamb (NZ4917), 1925 (MLT). This 0 NY NZ NZ OV SD SE SE TA species is predominantly southern in 9 distribution, although there are old 8 records north to south-west Scotland. 7 6 5 4 3 2 1
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maculata Brisout de Barneville, 1863
NO RECORDS
moerens Gyllenhal, 1827
Aleochara moerens Gyllenhal, 1827
3 WATSONIAN (0,0,4,0,0) There are four records of this 2 YORKSHIRE Nationally Scarce species on the YNU database, all from the same locality: 1 Langsett (SE1900), 02/08/1985, 0 NY NZ NZ OV SD SE SE TA 02/09/1985 (EJS teste CJ) and 07 & 9 09/12/2007 (EJS), in gill fungi on the same 8 Sycamore on all occasions. 7 6 5 4 3 2 1
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moesta Gravenhorst, 1802
NO RECORDS
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sanguinea (Linnaeus, 1758)
NO RECORDS
sparsa Heer, 1839
Aleochara (Xenochara) sparsa Heer, 1839
3 WATSONIAN (5,8,5,6,2) There are authentic records 2 YORKSHIRE from a number of widely distributed localities in Yorkshire. In Yorkshire 1 specimens have been located in cow 0 NY NZ NZ OV SD SE SE TA dung, leaf litter, carrion, rot hole debris, 9 and pitfall traps. 8 7 Past confusion with identification renders the 16 pre-1960 Yorkshire 6 records unreliable without voucher 5 specimens. 4 3 2 1
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stichai Likovský, 1965
NO RECORDS
tristis Gravenhorst, 1806
NO RECORDS
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villosa Mannerheim, 1830
Aleochara (Xenochara) v illosa Mannerheim, 1830
3 WATSONIAN (0,2,0,0,0) The only records on the YNU 2 YORKSHIRE database involve numerous specimens from dovecotes in the Scarborough area 1 (TA08?) during 1930 (RLF & GBW). Most 0 NY NZ NZ OV SD SE SE TA specimens in British collections originate 9 from this source (Coleopterist 6: 41-42). 8 The species is found in a wide variety of 7 rotting vegetable matter and dung. It is widely scattered in England with 6 occasional records from Scotland and is 5 afforded the national status pRDBK. 4 3 2 1
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TINOTUS Sharp, 1883 202 morion (Gravenhorst, 1802)
Tinotus morion (Grav enhorst, 1802)
3 WATSONIAN (8,10,26,10,2) Widely distributed and 2 YORKSHIRE fairly common in Yorkshire. The species is usually found in moss and leaf litter. 1 Yorkshire specimens have been located in 0 NY NZ NZ OV SD SE SE TA dung (horse, cow, sheep and dog), carrion, 9 stable refuse, decomposing vegetable 8 matter, and litter in a Phragmites bed. 7 6 5 4 3 2 1
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Tribe ATHETINI Casey, 1910
Subtribe ATHETINA Casey, 1910
ACROTONA Thomson, C.G., 1859
aterrima (Gravenhorst, 1802)
Acrotona aterrima (Grav enhorst, 1802)
3 WATSONIAN (15,12,135,17,8) Widely distributed and 2 YORKSHIRE common in the county. Yorkshire specimens have been located in a wide 1 variety of decaying vegetable matter, 0 NY NZ NZ OV SD SE SE TA carrion, dung, stable refuse, and fungi 9 (including Pleurotus ostreatus). 8 7 6 5 4 3 2 1
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benicki (Allen, 1940)
Acrotona benicki (Allen, 1940)
3 WATSONIAN (0,1,0,0,0) There is a single record on the 2 YORKSHIRE YNU database: Wykeham Forest (SE9488), 05/08/2001 (LA det. MLD). The species is 1 probably associated with open grazing 0 NY NZ NZ OV SD SE SE TA land, where it has been found under horse 9 dung. 8 7 6 5 4 3 2 1
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convergens (Strand, 1958)
NO RECORDS
exigua (Erichson, 1837)
Acrotona exigua (Erichson, 1837)
3 WATSONIAN (0,0,1,0,0) There is a single record on the 2 YORKSHIRE YNU database: Holme Moss (SE0904), 07/04/1983 (CJ). 1
NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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muscorum (Brisout de Barneville, 1860)
Acrotona muscorum (Brisout de Barnev ille, 1860)
3 WATSONIAN (7,7,35,21,6) Widely distributed in 2 YORKSHIRE Yorkshire, but records show that it is mainly confined to a north-south central 1 band. The species is normally found in 0 NY NZ NZ OV SD SE SE TA decomposing plant litter. Yorkshire 9 specimens have been located in all types 8 of decaying vegetable matter, in fungi 7 (including Pleurotus ostreatus), in horse, cow and dog dung, and in carrion. 6 5 4 3 2 1
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obfuscata (Gravenhorst, 1802)
Acrotona obfuscata (Grav enhorst, 1802)
3 WATSONIAN (0,0,4,0,0) There are four records of this Nationally Scarce species on the YNU 2 YORKSHIRE database, all from Hillsborough (SK3288): 1 10/05/1989 (EJS det. CJ), 27/06/1989 (EJS 0 NY NZ NZ OV SD SE SE TA teste CJ) and 02/10/1990 (EJS), under 9 leaves from mud surface. This is a species 8 of wet habitats and is mainly confined to southern Britain, Yorkshire being at the 7 species’ northern limit. 6 5 4 3 2 1
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parens (Mulsant & Rey, 1852)
NO RECORDS
parvula (Mannerheim, 1830)
Acrotona parv ula (Mannerheim, 1830)
3 WATSONIAN (13,5,43,14,9) Widely distributed and 2 YORKSHIRE common in the county. The lack of records from most of VC61 may indicate under- 1 recording. The species is usually 0 NY NZ NZ OV SD SE SE TA associated with dung and decomposing 9 vegetable matter. Yorkshire specimens 8 have been located in flood refuse, grass 7 piles, stable refuse, horse, cow and sheep dung, carrion, and fungi (including 6 Pleurotus ostreatus). 5 4 3 2 1
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pseudotenera (Cameron, 1933)
NO RECORDS
pygmaea (Gravenhorst, 1802)
Acrotona pygmaea (Grav enhorst, 1802)
3 WATSONIAN (1,1,25,4,0) Widely distributed in the 2 YORKSHIRE county, but not common. Yorkshire specimens have mainly been located in 1 decaying vegetable matter and dead 0 NY NZ NZ OV SD SE SE TA wood. 9 8 7 6 5 4 3 2 1
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sylvicola (Kraatz, 1856)
NO RECORDS
troglodytes (Motschulsky, 1858)
NO RECORDS
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ADOTA Casey, 1909 maritima (Mannerheim, 1843)
Adota maritima (Mannerheim, 1843)
3 WATSONIAN (1,0,0,0,0) There is a single record on the YNU database: Bridlington (TA1968), 2 YORKSHIRE 16/05/2015 (RCa det. SAW), in tideline 1 seaweed. This North American species 0 NY NZ NZ OV SD SE SE TA was first recorded in Britain from 9 specimens found in north-east Scotland in 8 1968 (Easton, 1971), who described it as Atheta (Phyconoma) immigrans sp. n. The 7 species is still only known from very few 6 widely scattered localities, where it is 5 found among seaweed and other driftline 4 refuse on sandy shores. 3 2 1
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ALAOBIA Thomson, C.G., 1858 gagatina (Baudi, 1848)
Alaobia gagatina (Baudi, 1848)
3 WATSONIAN (3,11,9,3,1) This species is associated with fungi and, although widely distributed in 2 YORKSHIRE the county, is not common. The only 1 Yorkshire specimens with collecting data 0 NY NZ NZ OV SD SE SE TA show that it has been found in old fungi 9 and pitfall traps. 8 7 6 5 4 3 2 1
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hybrida (Sharp, 1869)
NO RECORDS
linderi (Brisout de Barneville, 1863)
NO RECORDS
pallidicornis (Thomson, C.G., 1856)
Alaobia pallidicornis (Thomson, C.G., 1856)
3 WATSONIAN (2,2,19,11,1) Widely distributed in Yorkshire, but not common. Yorkshire 2 YORKSHIRE specimens have been located in fungi 1 (including Polyporus squamosus), reedbed 0 NY NZ NZ OV SD SE SE TA and leaf litter, in moss, and under bark. 9 8 7 6 5 4 3 2 1
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scapularis (Sahlberg, C.R., 1831)
NO RECORDS
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sodalis (Erichson, 1837)
Alaobia sodalis (Erichson, 1837)
3 WATSONIAN (1,12,17,7,2) Widely distributed, but not 2 YORKSHIRE common, in the county. The lack of records from most of VC61 may be due to 1 under-recording. The only Yorkshire 0 NY NZ NZ OV SD SE SE TA records with collecting data show they 9 had been found under the bark of a pine 8 stump, in grass traps, and in the nest of a 7 Mole.
6 5 4 3 2 1
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subglabra (Sharp, 1869)
NO RECORDS
taxiceroides Munster, 1932
Alaobia taxiceroides Munster, 1932
3 WATSONIAN (0,1,5,0,0) There are records from six Yorkshire localities, all except one from 2 YORKSHIRE VC63. The only Yorkshire records with 1 collecting data show they originated from 0 NY NZ NZ OV SD SE SE TA the fungi Piptoporus betulinus and 9 Amanita muscaria. 8 7 6 5 4 3 2 1
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trinotata (Kraatz, 1856)
Alaobia trinotata (Kraatz, 1856)
3 WATSONIAN (29,18,121,23,5) Widely distributed and 2 YORKSHIRE common in the county. The species is usually found in decomposing plant 1 material and fungi. Yorkshire specimens 0 NY NZ NZ OV SD SE SE TA have been located in all types of vegetable 9 refuse, fungi (including Polyporus 8 squamosus), carrion, and dung. 7 6 5 4 3 2 1
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ALEVONOTA Thomson, C.G., 1858
egregia (Rye, 1876)
NO RECORDS gracilenta (Erichson, 1839)
NO RECORDS rufotestacea (Kraatz, 1856)
NO RECORDS
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ALIANTA Thomson, C.G., 1858 incana (Erichson, 1837)
Alianta incana (Erichson, 1837)
3 WATSONIAN (6,1,39,9,2) This species, which is found in 2 YORKSHIRE litter in damp places, is widely distributed in the southern half of the county. 1 Yorkshire specimens have mainly been 0 NY NZ NZ OV SD SE SE TA found in the litter in Typha, Glyceria and 9 Phragmites beds, occasionally in flood and 8 vegetable refuse. 7 6 5 4 3 2 1
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ALOCONOTA Thomson, C.G., 1858
Subgenus ALOCONOTA Thomson, C.G., 1858
cambrica (Wollaston, 1855)
Aloconota (Aloconota) cambrica (Wollaston, 1855)
3 WATSONIAN (0,3,0,1,1) There are five records on the YNU database: Hilla Green (SE9589), 2 YORKSHIRE 20/05/1925 (GBW); Whitby (NZ8911), 1 20/03/1935 (HB); Mulgrave Wood 0 NY NZ NZ OV SD SE SE TA (NZ8412), 20/03/1935 (HB); Queen Mary’s 9 Dub (SE3074), 04/09/1948 (EWA teste CJ); 8 Langton Bridge (SE2996), 25/07/2002 (RJM), in riverside shingle. 7
6 5 4 3 2 1
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currax (Kraatz, 1856)
Aloconota (Aloconota) currax (Kraatz, 1856)
3 WATSONIAN (2,9,4,0,0) Widely distributed, but not 2 YORKSHIRE common, in the county. The cluster of records from SE68 are from captures at 1 locations along the River Rye at Helmsley. 0 NY NZ NZ OV SD SE SE TA Most Yorkshire records stem from 9 specimens located in riverine shingle, only 8 occasionally in leaf litter. 7 6 5 4 3 2 1
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eichhoffi (Scriba, 1867)
NO RECORDS
gregaria (Erichson, 1839)
Aloconota (Aloconota) gregaria (Erichson, 1839)
3 WATSONIAN (58,27,133,36,7) This is a very common 2 YORKSHIRE and widely distributed species in the county. Yorkshire specimens have been 1 located in a wide variety of decomposing 0 NY NZ NZ OV SD SE SE TA vegetable matter, in fungi, carrion, dung, 9 shingle, under seaweed, and by sweeping 8 vegetation. 7 6 5 4 3 2 1
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insecta (Thomson, C.G., 1856)
Aloconota (Aloconota) insecta (Thomson, C.G., 1856)
3 WATSONIAN (3,2,13,3,2) Despite the national status of 2 YORKSHIRE ‘Common’ the species has only been recorded from 22 widely distributed 1 localities in the county. The few Yorkshire 0 NY NZ NZ OV SD SE SE TA records with collecting data have all 9 originated from flood refuse. 8 7 6 5 4 3 2 1
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mihoki Bernhauer, 1913
NO RECORDS
planifrons (Waterhouse, G.R., 1864)
NO RECORDS
subgrandis Brundin, 1954
NO RECORDS
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sulcifrons (Stephens, 1832)
Aloconota (Aloconota) sulcifrons (Stephens, 1832)
3 WATSONIAN (4,1,28,1,1) Widely distributed in the 2 YORKSHIRE county but, apparently, only common in VC63 (although this may reflect collecting 1 effort). The few Yorkshire records with 0 NY NZ NZ OV SD SE SE TA collecting data indicate they have been 9 located in ground litter, in a pile of 8 decaying weeds and under leaves on a 7 mud surface.
6 5 4 3 2 1
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Subgenus DISOPORA Thomson, C.G., 1859 coulsoni (Last, 1952)
NO RECORDS
languida (Erichson, 1837)
Aloconota (Disopora) languida (Erichson, 1837)
3 WATSONIAN (4,0,1,2,0) The names of the species belonging to the subgenus Disopora were, until resolved YORKSHIRE 2 by Last in 1952 (Ent. Monthly Mag. 88: 263- 1 264), misinterpreted by British workers.
NY NZ NZ OV Without voucher specimens any records prior 0 SD SE SE TA to this date are likely to be confused. As no 9 specimens have been located, the records from 8 Bubwith (SE7030) in 1915 (GBW) and 1918 (WJF), although likely correct because of the 7 authentic records below, remain in doubt. 6 5 There are seven authentic records of this Nationally Scarce species on the YNU database: 4 Bubwith (SE7030), 01/1919 (WJF teste RGB), 3 02/1919 (WJF teste MLD), 11/1919 (WJF teste 2 CJ), all from flood refuse; near Duffield (SE6936), 1930 (GBW det. HRL), in flood refuse; 1 Sharow Mires, Ripon (SE3370), 27/06/2002 SD SE SE TA 0 SJ SK SK TF (RJM teste MLD); Bishopthorpe (SE5948), 9 22/06/2009 (RJM); Potteric Carr (SE5900), 31/05/2013 (RJM teste MLD). 8 post 2000 Map produced for Yorkshire Naturalists' Union by DMAP 29/1/2019 pre 2000 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
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longicollis (Mulsant & Rey, 1852)
NO RECORDS
AMIDOBIA Thomson, C.G., 1858 talpa (Heer, 1841)
Amidobia talpa (Heer, 1841)
3 WATSONIAN (1,11,13,0,0) With the exception of a single record in VC61, the other Yorkshire 2 YORKSHIRE records are confined to VC63 and VC62. 1 This is a Nationally Scarce woodland 0 NY NZ NZ OV SD SE SE TA species which is associated with the wood 9 ants Formica rufa and F. lugubris and 8 possibly F. aquilonia. The Yorkshire records with collecting data confirm the 7 association with F. lugubris. 6 5 4 3 2 1
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AMISCHA Thomson, C.G., 1858 analis (Gravenhorst, 1802)
Amischa analis (Grav enhorst, 1802)
3 WATSONIAN (50,47,262,45,16) This is a very common 2 YORKSHIRE and widely distributed species in Yorkshire and has been recorded from a wide variety 1 of habitat types, including all types of 0 NY NZ NZ OV SD SE SE TA decaying vegetation, dung, and fungi. 9 8 7 6 5 4 3 2 1
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bifoveolata (Mannerheim, 1830)
Amischa bifov eolata (Mannerheim, 1830)
3 WATSONIAN (14,8,21,6,2) Widely, but thinly, distributed in the county. Yorkshire 2 YORKSHIRE specimens have been found in grass 1 heaps, an old straw bale, stable refuse, 0 NY NZ NZ OV SD SE SE TA flood refuse, moss, and sheep dung. 9 8 7 6 5 4 3 2 1
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decipiens (Sharp, 1869)
Amischa decipiens (Sharp, 1869)
3 WATSONIAN (10,6,39,7,4) Widely distributed in 2 YORKSHIRE Yorkshire, but records show that it is mainly confined to a north-south running 1 central band. It is found in a wide variety 0 NY NZ NZ OV SD SE SE TA of rotting organic material. Yorkshire 9 specimens have been recorded from 8 ground litter, an old straw heap, fungi, 7 moss, under bark, and by sweeping vegetation. 6 5 4 3 2 1
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forcipata Mulsant & Rey, 1873
Amischa forcipata Mulsant & Rey, 1873
3 WATSONIAN (5,0,5,1,0) This species, which is 2 YORKSHIRE associated with moss and plant litter etc, has only been recorded from 11 widely 1 distributed Yorkshire localities. The few 0 NY NZ NZ OV SD SE SE TA Yorkshire records with collecting data 9 indicate they were located in old straw 8 and by sweeping vegetation. 7 6 5 4 3 2 1
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nigrofusca (Stephens, 1829)
Amischa nigrofusca (Stephens, 1829)
3 WATSONIAN (1,1,12,3,2) Although the national 2 YORKSHIRE distribution of this species is clouded as it was previously considered a variety of A. 1 analis (Gravenhorst, 1802), it is still widely 0 NY NZ NZ OV SD SE SE TA distributed in the county. Yorkshire 9 specimens have been located in grass 8 cuttings, stable refuse, fungoid leaf litter, 7 and flood refuse.
6 5 4 3 2 1
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ANOPLETA Mulsant & Rey, 1874 corvina (Thomson, C.G., 1856)
Anopleta corv ina (Thomson, C.G., 1856)
3 WATSONIAN (0,4,8,0,2) Widely distributed in Yorkshire, 2 YORKSHIRE but not common. The few Yorkshire records with collecting data indicate they 1 were located in old fungi (including 0 NY NZ NZ OV SD SE SE TA Polyporus). 9 8 7 6 5 4 3 2 1
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kochi (Roubal, 1937)
NO RECORDS puberula (Sharp, 1869)
Anopleta puberula (Sharp, 1869)
3 WATSONIAN (0,0,1,0,0) There is a single record on the 2 YORKSHIRE YNU database: Brookhouse Dike (SK5088), 12/11/1989 (WAE det. CJ), in leaf litter. 1 The species is mainly restricted to 0 NY NZ NZ OV SD SE SE TA southern England, but there are 9 occasional records north to northern 8 Scotland. It is afforded the national status 7 pRDBK.
6 5 4 3 2 1
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soedermani (Bernhauer, 1931)
NO RECORDS
ATHETA Thomson, C.G., 1858 aeneicollis (Sharp, 1869)
Atheta aeneicollis (Sharp, 1869)
3 WATSONIAN (17,27,65,22,2) Widely distributed and common in the county. Although the 2 YORKSHIRE ecology appears to be unknown, Yorkshire 1 specimens have been located in fungi 0 NY NZ NZ OV SD SE SE TA (including Polyporus squamosus, 9 Laetiporus sulphureus and Agaricus 8 campestris), flood refuse, grass cuttings, stable refuse, carrion, and ungulate dung. 7
6 5 4 3 2 1
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aquatica (Thomson, C.G., 1852)
Atheta aquatica (Thomson, C.G., 1852)
3 WATSONIAN (1,8,37,5,1) Widely distributed in the 2 YORKSHIRE county, but not common. The cluster of records in SE11 (VC63) may simply reflect 1 recorder effort and, conversely, the lack of 0 NY NZ NZ OV SD SE SE TA records from a large area of VC61 may be 9 due to under-recording. It occurs in 8 decaying vegetable matter and fungi. 7 6 5 4 3 2 1
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aquatilis (Thomson, C.G., 1867)
Atheta aquatilis (Thomson, C.G., 1867)
3 WATSONIAN (1,3,32,1,2) Despite the national status of 2 YORKSHIRE ‘Nationally Scarce’, Yorkshire records indicate that it is widely distributed in 1 VC63 with occasional records from other 0 NY NZ NZ OV SD SE SE TA areas of the county. The species is usually 9 found near water, where it occurs in 8 rotting organic matter. Yorkshire 7 specimens have been found in fungi (including Pleurotus ostreatus), moss, 6 reed and Glyceria beds, carrion, sheep 5 dung, and ground litter under conifers. 4 3 2 1
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brunneipennis (Thomson, C.G., 1852)
Atheta brunneipennis (Thomson, C.G., 1852)
3 WATSONIAN (1,4,22,5,1) Widely distributed in 2 YORKSHIRE Yorkshire, but not common. The number of records from VC63 may simply reflect 1 recorder effort and, conversely, the lack of 0 NY NZ NZ OV SD SE SE TA records from a large area of VC61 may be 9 due to under-recording. Yorkshire 8 specimens have been found in ground 7 litter, grass cuttings, decaying fungi, carrion, and deer dung. 6
5 4 3 2 1
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castanoptera (Mannerheim, 1830)
Atheta castanoptera (Mannerheim, 1830)
3 WATSONIAN (2,25,125,24,4) Widely distributed and 2 YORKSHIRE common in Yorkshire but, rather surprisingly, is the near lack of records 1 from VC61. Yorkshire specimens have 0 NY NZ NZ OV SD SE SE TA been located in fungi (including Pleurotus 9 ostreatus and Polyporus squamosus), all 8 types of decomposing vegetable matter, 7 moss, carrion, and pitfall and water traps.
6 5 4 3 2 1
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ebenina (Mulsant & Rey, 1874)
NO RECORDS
graminicola (Gravenhorst, 1806)
Atheta graminicola (Grav enhorst, 1806)
3 WATSONIAN (39,29,235,55,16) Widely distributed and 2 YORKSHIRE common in the county. Yorkshire specimens have been found in beds of 1 Typha, Phragmites, Glyceria and Carex, 0 NY NZ NZ OV SD SE SE TA flood refuse, stable refuse, grass piles, 9 compost heaps, fungi (including Hirneola 8 auricula-judae), ungulate dung, and by 7 sweeping vegetation.
6 5 4 3 2 1
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heymesi Hubenthal, 1913
NO RECORDS
hypnorum (Kiesenwetter, 1850)
Atheta hypnorum (Kiesenwetter, 1850)
3 WATSONIAN (1,14,18,5,0) Widely distributed in Yorkshire, but not common. The 2 YORKSHIRE noticeable lack of records from VC65 and 1 most of VC61 may be due to under- 0 NY NZ NZ OV SD SE SE TA recording. The species occurs in rotting 9 vegetable matter and fungi. 8 7 6 5 4 3 2 1
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incognita (Sharp, 1869)
Atheta incognita (Sharp, 1869)
3 WATSONIAN (0,10,9,0,0) Although only reported from 2 YORKSHIRE six Yorkshire localities, this species appears to have two centres of population 1 in the county – in the eastern quarter of 0 NY NZ NZ OV SD SE SE TA VC62 and the southern half of VC63. The 9 only Yorkshire specimen with collecting 8 data was found in the litter of a deciduous 7 woodland.
6 5 4 3 2 1
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laevicauda Sahlberg, J., 1876
NO RECORDS
triangulum (Kraatz, 1856)
Atheta triangulum (Kraatz, 1856)
3 WATSONIAN (20,4,24,12,2) Widely distributed in 2 YORKSHIRE Yorkshire. The species is usually found in decaying vegetable matter, dung and 1 carrion. Yorkshire specimens have been 0 NY NZ NZ OV SD SE SE TA located in all types of decomposing plant 9 material, carrion, dung, under bark, and in 8 a Malaise trap. 7 6 5 4 3 2 1
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xanthopus (Thomson, C.G., 1856)
Atheta xanthopus (Thomson, C.G., 1856)
3 WATSONIAN (7,7,41,5,3) Widely distributed in 2 YORKSHIRE Yorkshire. Yorkshire specimens have been located in fungi (including Polyporus 1 squamosus and Agaricus campestris), 0 NY NZ NZ OV SD SE SE TA grass piles, old straw, moss, carrion, and 9 by sweeping vegetation. 8 7 6 5 4 3 2 1
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autumnalis (Erichson, 1839)
NO RECORDS basicornis (Mulsant & Rey, 1852)
Atheta basicornis (Mulsant & Rey, 1852)
3 WATSONIAN (0,0,1,3,0) There are four records, from 2 YORKSHIRE three localities, on the YNU database: Thorne Moor (SE7315), 27/04/1985 (RJM 1 teste CJ), in leaf litter; Askham Bog 0 NY NZ NZ OV SD SE SE TA (SE5648), 23/06/1996 (RGB & JAO) and 9 12/04/2003 (WRD teste MLD); Sharow 8 Mires, Ripon (SE3370), 14/07/2002 (RJM), 7 in fungoid debris under the bark of a fallen poplar. This Nationally Scarce species is 6 restricted to England and is associated 5 with wet habitats. 4 3 2 1
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boletophila (Thomson, C.G., 1856)
NO RECORDS
britanniae (Bernhauer & Scheerpeltz, 1926)
Atheta britanniae (Bernhauer & Scheerpeltz, 1926)
3 WATSONIAN (3,10,27,6,4) Widely distributed in the county, with the exception of a near 2 YORKSHIRE dearth of records for VC61. The species 1 occurs in dung, carrion, fungi and all types 0 NY NZ NZ OV SD SE SE TA of decomposing plant material. 9 8 7 6 5 4 3 2 1
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crassicornis (Fabricius, 1792)
Atheta crassicornis (Fabricius, 1792)
3 WATSONIAN (24,54,291,65,15) Widely distributed and 2 YORKSHIRE very common in Yorkshire. The lack of records from a large area of VC61 may be 1 due to under-recording. Yorkshire 0 NY NZ NZ OV SD SE SE TA specimens have been found in dung, 9 carrion, fungi (including Polyporus 8 squamosus, Pleurotus ostreatus, Coprinus 7 atramentarius, and Laetiporus sulphureus), and all types of decaying 6 plant material. 5 4 3 2 1
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diversa (Sharp, 1869)
Atheta div ersa (Sharp, 1869)
3 WATSONIAN (0,0,1,0,0) There is a single record on the 2 YORKSHIRE YNU database: Wessenden Head (SE0707), 16/06/1984 (MLD teste CJ), a 1 single male. Although widely distributed in 0 NY NZ NZ OV SD SE SE TA Scotland the above was the first English 9 record. 8 7 6 5 4 3 2 1
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divisa (Märkel, 1844)
Atheta div isa (Märkel, 1844)
3 WATSONIAN (1,0,1,4,0) There are six records, from five 2 YORKSHIRE localities, on the YNU database: Bubwith (SE7030), 02/1919 (CES); Queen Mary’s 1 Dub (SE3074), 04/09/1968 (CJ); Cadeby 0 NY NZ NZ OV SD SE SE TA (SE5100), 07/09/2001 (RJM teste DAL), in 9 garden refuse; Little Stainforth (SD8164), 8 06/08/2006 (RGB); Studley Royal 7 (SE2770), 12/06/2016 & 03/09/2016 (AG det. MLD). 6
5 4 3 2 1
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euryptera (Stephens, 1832)
Atheta euryptera (Stephens, 1832)
3 WATSONIAN (3,4,3,2,0) Despite the national status of 2 YORKSHIRE ‘Common’ the species has only been recorded from six Yorkshire localities: 1 Egton (NZ8106), date unknown (HB); 0 NY NZ NZ OV SD SE SE TA Whitby (NZ8911), date and recorder 9 unknown; Shirley Pool (SE5612), 10/1911 8 (recorder unknown); Thorne Moor 7 (SE7315), 27/04/1971 (CJ); Fairburn (SE4728), 11/07/1987 (MLD teste CJ); 6 Spurn (TA4212), 18/06/1990 (MLD). 5 4 3 2 1
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fungicola (Thomson, C.G., 1852)
Atheta fungicola (Thomson, C.G., 1852)
3 WATSONIAN (0,5,7,1,0) Widely distributed in the 2 YORKSHIRE county, especially in VCs 62 and 63, but far from common. The species is usually 1 found in fungi and carrion. The few 0 NY NZ NZ OV SD SE SE TA Yorkshire records with collecting data 9 indicate they were found in decaying 8 fungi, carrion, and ground litter under 7 logs.
6 5 4 3 2 1
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harwoodi (Williams, 1930)
Atheta harwoodi (Williams, 1930)
3 WATSONIAN (10,5,58,12,4) Widely distributed in 2 YORKSHIRE Yorkshire. Yorkshire specimens have been found in horse dung, carrion, stable 1 refuse, a garden compost heap, on a sap 0 NY NZ NZ OV SD SE SE TA run on an oak tree, and under dry 9 seaweed. 8 7 6 5 4 3 2 1
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intermedia (Thomson, C.G., 1852)
Atheta intermedia (Thomson, C.G., 1852)
3 WATSONIAN (1,1,3,3,0) This species, which is usually 2 YORKSHIRE found in dung, has only been recorded from eight widely distributed localities in 1 Yorkshire. Within Yorkshire, specimens 0 NY NZ NZ OV SD SE SE TA have been record from ungulate dung, 9 fungi (including Polyporus squamosus), 8 carrion, and grass clippings. 7 6 5 4 3 2 1
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liturata (Stephens, 1832)
Atheta liturata (Stephens, 1832)
3 WATSONIAN (1,7,8,11,4) Widely distributed in 2 YORKSHIRE Yorkshire, but not common. Usually found in fungi, Yorkshire specimens have been 1 found in a variety of species including 0 NY NZ NZ OV SD SE SE TA Polyporus squamosus, P. sulphureus, 9 Pleurotus ostreatus and Hypholoma 8 fasciculare. 7 6 5 4 3 2 1
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nidicola (Johansen, 1914)
Atheta nidicola (Johansen, 1914)
3 WATSONIAN (0,1,2,2,0) There are five records, from 2 YORKSHIRE four localities, on the YNU database: Dalby Forest (SE88), 18/10/1966 (GS teste EWA 1 & MLD), from a Sparrowhawk nest; 0 NY NZ NZ OV SD SE SE TA Askham Bog (SE5648), 20/04/1967 (CJ) 9 and 16/04/2005 (MLD); Blackbrook Wood 8 (SK2986), 05/01/1992 (EJS teste CJ); 7 Ewden Pond (SK2296), 31/03/1997 (EJS teste MLD). 6
5 4 3 2 1
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nigritula (Gravenhorst, 1802)
Atheta nigritula (Grav enhorst, 1802)
3 WATSONIAN (1,0,2,0,0) There are records from three localities on the YNU database: 2 YORKSHIRE Wharncliffe Wood (SK3195), 11/10/1992 1 (EJS teste CJ); Skipwith Common (SE6637), 0 NY NZ NZ OV SD SE SE TA 17/07/1993 (MLD); Rivelin (SK2987), 06- 9 10/1998 (EJS teste MLD), several 8 specimens in litter. This Nationally Scarce species is mainly restricted to southern 7 England, but there are scattered records 6 north to south-east Scotland. 5 4 3 2 1
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oblita (Erichson, 1839)
Atheta oblita (Erichson, 1839)
3 WATSONIAN (1,2,4,0,0) This species, which appears to 2 YORKSHIRE be widely distributed in England, has only been recorded from seven Yorkshire 1 localities. The only Yorkshire record with 0 NY NZ NZ OV SD SE SE TA collecting data indicates that specimens 9 were found in old fungi. 8 7 6 5 4 3 2 1
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paracrassicornis Brundin, 1954
Atheta paracrassicornis Brundin, 1954
3 WATSONIAN (0,0,0,1,0) A single male of this boreo- alpine species collected at Malham Tarn 2 YORKSHIRE (SD8967) on 13/09/1967 by CJ added the 1 species to the British list. A full account 0 NY NZ NZ OV SD SE SE TA can be found in Entomologist 101: 64-66. 9 8 There is a record on the NBN Gateway of a single male from Hatfield Forest, 7 Hertfordshire (TL5318) on 05/06/2008. 6 The determination is attributed to Tony 5 Drane (ABD), but he informs us that he has 4 never recorded the species and is looking 3 into how the discrepancy occurred.
2 1
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pilicornis (Thomson, C.G., 1852)
Atheta pilicornis (Thomson, C.G., 1852)
3 WATSONIAN (0,1,0,1,0) There are two records of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Saltburn (NZ6721), 10/1893 1 (MLT); Cayton Gill (SE2862), 05/05/2005 0 NY NZ NZ OV SD SE SE TA (MLD). The species is found in England and 9 southern Scotland where it is associated 8 with woodland. 7
6
5 4 3 2 1
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procera (Kraatz, 1856)
NO RECORDS
ravilla (Erichson, 1839)
Atheta rav illa (Erichson, 1839)
3 WATSONIAN (4,7,40,9,3) Widely distributed in the county, but not common. The lack of 2 YORKSHIRE records from most of VC61 may reflect 1 under-recording. Yorkshire specimens 0 NY NZ NZ OV SD SE SE TA have been found in carrion, dung, fungi 9 (including Polyporus squamosus and 8 Pleurotus ostreatus), in all types of 7 decaying plant material, and among old bones. 6 5 4 3 2 1
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strandiella (Brundin, 1954)
Atheta strandiella (Brundin, 1954)
3 WATSONIAN (1,0,0,0,0) There is a single record of this Nationally Scarce species on the YNU 2 YORKSHIRE database: Spurn (TA4212), 18/06/1990 1 (MLD det. CJ), a single male in tidal refuse 0 NY NZ NZ OV SD SE SE TA on the Humber shore at the Warren. Not 9 added to the British list until 1971 (Ent. 8 Monthly Mag. 107: 153-157), the species has now been recorded from parts of 7 England and northern Scotland. British 6 specimens have been recorded from 5 carrion, sheep dung, fen litter, moss, flood 4 refuse, tide line litter, and pitfall traps. 3
2 1
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vaga (Heer, 1839)
Atheta v aga (Heer, 1839)
3 WATSONIAN (4,9,44,13,1) Widely distributed in 2 YORKSHIRE Yorkshire but not common, apart from VC63 (although this may reflect 1 collecting effort). The species is usually 0 NY NZ NZ OV SD SE SE TA found in rotting fungi on trees and in bird 9 nests. Yorkshire specimens have been 8 located in old straw and grass heaps, in a 7 compost heap and other decaying vegetation, in fungi (including decaying 6 Polyporus), in carrion, and in the nests of 5 an owl and Rook. 4 3 2 1
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BADURA Mulsant & Rey, 1873 macrocera (Thomson, C.G., 1856)
Badura macrocera (Thomson, C.G., 1856)
3 WATSONIAN (3,3,15,6,2) Widely distributed in 2 YORKSHIRE Yorkshire, but not common. Yorkshire specimens have been located in ungulate 1 dung, fungi, moss, leaf litter, a manure 0 NY NZ NZ OV SD SE SE TA heap, and in the dry remains of a dead 9 Rabbit. 8 7 6 5 4 3 2 1
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puncticollis (Benick, 1938)
Badura puncticollis (Benick, 1938)
3 WATSONIAN (0,0,1,0,0) There is a single record on the 2 YORKSHIRE YNU database: Orange Wood, Blackmoorfoot Reservoir (SE1012), 1 19/06/2012 (MLD), a single female in 0 NY NZ NZ OV SD SE SE TA moss on a dead oak Quercus spp. This is a 9 very localised species and, with the 8 exceptions of records from Devon, 7 Yorkshire and Northumberland, it is confined to Scotland. 6
5 4 3 2 1
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BESSOBIA Thomson, C.G., 1858 excellens (Kraatz, 1856)
Bessobia excellens (Kraatz, 1856)
3 WATSONIAN (1,7,3,8,0) There are Yorkshire records 2 YORKSHIRE from 15 localities, mainly in the north and west of the county. The species mainly 1 occurs in fungi in autumn. The only 0 NY NZ NZ OV SD SE SE TA Yorkshire record with collecting data 9 indicates that specimens were located in a 8 grass heap. 7 6 5 4 3 2 1
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fungivora (Thomson, C.G., 1867)
Bessobia fungiv ora (Thomson, C.G., 1867)
3 WATSONIAN (3,7,4,6,0) Widely distributed in Yorkshire, but not common. It should be noted that 2 YORKSHIRE there is only a single post-2000 record. 1 The species is found in rotting fungi in 0 NY NZ NZ OV SD SE SE TA autumn. The only Yorkshire record with 9 collecting data shows that specimens 8 were found in pitfall traps set in a wet meadow. 7
6 5 4 3 2 1
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monticola (Thomson, C.G., 1852)
Bessobia monticola (Thomson, C.G., 1852)
3 WATSONIAN (1,3,4,3,0) Widely distributed in the 2 YORKSHIRE county, but not common. The only Yorkshire records with collecting data 1 show they had originated from a pile of 0 NY NZ NZ OV SD SE SE TA rotting grass, grass roots and in a rotting 9 Giant Puffball Calvatia gigantea. 8 7 6 5 4 3 2 1
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occulta (Erichson, 1837)
Bessobia occulta (Erichson, 1837)
3 WATSONIAN (0,3,3,0,1) There are Yorkshire records 2 YORKSHIRE from seven widely distributed localities. It should be noted that the most recent 1 record was in 1992. The species is found 0 NY NZ NZ OV SD SE SE TA in rotting fungi in autumn. The only 9 Yorkshire record with collecting data 8 indicates that specimens were found in a 7 dead gull on the coast. 6 5 4 3 2 1
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BOREOPHILIA Benick, 1973 eremita (Rye, 1866)
Boreophilia eremita (Rye, 1866)
3 WATSONIAN (4,10,18,5,6) Widely distributed in the 2 YORKSHIRE county - the cluster of records in VC63 may simply reflect recorder effort and, 1 conversely, the lack of records from the 0 NY NZ NZ OV SD SE SE TA eastern half of VC61 may be due to under- 9 recording. The species occurs in all types 8 of rotting vegetation, fungi, etc. Yorkshire 7 specimens have been located in fungi, litter, moss, flood refuse, pitfall traps, and 6 in seashore refuse. 5 4 3 2 1
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BRUNDINIA Tottenham, 1949
marina (Mulsant & Rey, 1853) Brundinia marina (Mulsant & Rey, 1853)
3 WATSONIAN (19,0,0,0,0) All Yorkshire localities for this 2 YORKSHIRE species are clustered around the mouth of the River Humber with records between 1 Spurn (TA4212) and Fisherman’s Channel, 0 NY NZ NZ OV SD SE SE TA Sunk Island (TA2819). 9 8 7 6 5 4 3 2 1
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meridionalis (Mulsant & Rey, 1853)
Brundinia meridionalis (Mulsant & Rey, 1853)
3 WATSONIAN (4,0,0,0,0) Most Yorkshire records of this 2 YORKSHIRE coastal species are centred around the lower reaches of the Humber Estuary: 1 Spurn (TA4212), 1952 (WOS); Fisherman’s 0 NY NZ NZ OV SD SE SE TA Channel, Sunk Island (TA2819), 9 17/08/1986 (RJM) and 27/04/1991 (MLD 8 & RJM); Beacon Ponds (TA4116), 7 21/06/1990 (MLD det. CJ); Speeton (TA1574), 12/07/2006 (MGT det. RJM). 6
5 4 3 2 1
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CADAVEROTA Yosii & Sawada, 1976 cadaverina (Brisout de Barneville, 1860)
Cadav erota cadav erina (Brisout de Barnev ille, 1860)
3 WATSONIAN (0,9,15,1,0) Widely distributed in 2 YORKSHIRE Yorkshire, but not common. The cluster of records in VC63 may simply reflect 1 recorder effort. The species is normally 0 NY NZ NZ OV SD SE SE TA found in decomposing organic material. 9 The few Yorkshire records with collecting 8 data show that the species has been 7 located in carrion, rotting fungi, and a pile of rotting grass. 6
5 4 3 2 1
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hansseni (Strand, 1943)
NO RECORDS
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CALLICERUS Gravenhorst, 1802 obscurus Gravenhorst, 1802
Callicerus obscurus Grav enhorst, 1802
3 WATSONIAN (4,3,6,1,0) Widely distributed in the 2 YORKSHIRE county, but not common. The only Yorkshire records with collecting data 1 show that they have been located in flood 0 NY NZ NZ OV SD SE SE TA refuse and pitfall traps. 9 8 7 6 5 4 3 2 1
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rigidicornis (Erichson, 1839)
Callicerus rigidicornis (Erichson, 1839)
3 WATSONIAN (1,1,16,2,0) Widely distributed in the 2 YORKSHIRE county, but not common. The abundance of records from VC63 may simply indicate 1 collecting effort. Yorkshire specimens 0 NY NZ NZ OV SD SE SE TA have been found under beach bark, in 9 straw bales, and in pitfall traps. 8 7 6 5 4 3 2 1
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CERITAXA Mulsant & Rey, 1873 dilaticornis (Kraatz, 1856)
NO RECORDS
pervagata (Benick, 1974)
NO RECORDS
testaceipes (Heer, 1839)
NO RECORDS
CHAETIDA Mulsant & Rey, 1874 longicornis (Gravenhorst, 1802)
Chaetida longicornis (Grav enhorst, 1802)
3 WATSONIAN (21,22,144,19,11) Widely distributed and common in Yorkshire. Yorkshire 2 YORKSHIRE specimens have been found in dung 1 (horse, cow, sheep and deer), carrion, 0 NY NZ NZ OV SD SE SE TA fungi (including Piptoporus betulinus and 9 Pleurotus ostreatus), and in all types of 8 decaying vegetable material. 7 6 5 4 3 2 1
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COPROTHASSA Thomson, C.G., 1859 melanaria (Mannerheim, 1830)
Coprothassa melanaria (Mannerheim, 1830)
3 WATSONIAN (2,2,0,0,0) There are four records, from three localities, on the YNU database: 2 YORKSHIRE Scarborough (TA0488), date unknown 1 (RL); Ravenscar (NZ9801), date unknown 0 NY NZ NZ OV SD SE SE TA (WJF); Spurn (TA4212), 06/1950 (WOS) 9 and 19/11/1990 (MLD), in sheep dung at 8 Chalk Bank.
7 6 5 4 3 2 1
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DADOBIA Thomson, C.G., 1858 immersa (Erichson, 1837)
Dadobia immersa (Erichson, 1837)
3 WATSONIAN (1,1,4,6,0) Widely distributed in Yorkshire, 2 YORKSHIRE but no common. The species is generally found under bark in woodland (both 1 deciduous and coniferous). The only 0 NY NZ NZ OV SD SE SE TA Yorkshire records with collecting data 9 show they were found under bark and in 8 hay litter. 7 6 5 4 3 2 1
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DALOTIA Casey, 1910 coriaria (Kraatz, 1856)
Dalotia coriaria (Kraatz, 1856)
3 WATSONIAN (6,2,45,13,0) Widely distributed in 2 YORKSHIRE Yorkshire, especially in the southern half of the county. Yorkshire specimens have 1 been found in fungi (including Polyporus 0 NY NZ NZ OV SD SE SE TA squamosus), a garden compost heap, 9 stable refuse, Carex litter, old straw, 8 woodchips, and on a sap run on an oak 7 tree.
6 5 4 3 2 1
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DATOMICRA Mulsant & Rey, 1874 canescens (Sharp, 1869)
Datomicra canescens (Sharp, 1869)
3 WATSONIAN (0,0,5,0,0) There are five records on the 2 YORKSHIRE YNU database: Eggborough Power Station (SE5724), 24/05/1985 (RJM det. CJ); 1 Melton Wood (SE5103), 01/05/1987 0 NY NZ NZ OV SD SE SE TA (RJM); Edge Moor, Blackmoorfoot 9 Reservoir (SE1012), 09/06/2007 (MLD), in 8 stable refuse; Yorkshire Wildlife Park 7 (SE6401), 30/07/2012 (RJM), in cattle dung; Thorpe Marsh (SE5809), 6 19/09/2016 (RJM), in deer dung. 5 4 3 2 1
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celata (Erichson, 1837)
Datomicra celata (Erichson, 1837)
3 WATSONIAN (24,16,223,26,12) Widely distributed and common in the county. Yorkshire 2 YORKSHIRE specimens have been located in flood 1 refuse, stable refuse, Juncus litter, 0 NY NZ NZ OV SD SE SE TA rotting vegetation, compost heaps, 9 moss, fungi (including Polyporus 8 squamosus and Pleurotus ostreatus), 7 and carrion.
6 5 4 3 2 1
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dadopora (Thomson, C.G., 1867)
Datomicra dadopora (Thomson, C.G., 1867)
3 WATSONIAN (4,5,46,12,4) Widely distributed in 2 YORKSHIRE Yorkshire, but not common. Yorkshire specimens have been located in horse and 1 sheep dung, fungi, moss, stable refuse, 0 NY NZ NZ OV SD SE SE TA carrion, decaying vegetable piles, and in a 9 Malaise trap. 8 7 6 5 4 3 2 1
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nigra (Kraatz, 1856)
Datomicra nigra (Kraatz, 1856)
3 WATSONIAN (34,12,152,35,11) Widely distributed and 2 YORKSHIRE common in the county. Yorkshire specimens have been found in a wide 1 variety of decaying organic matter, fungi, 0 NY NZ NZ OV SD SE SE TA sheep and cow dung, and also by 9 sweeping vegetation. 8 7 6 5 4 3 2 1
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sordidula (Erichson, 1837)
Datomicra sordidula (Erichson, 1837)
3 WATSONIAN (11,8,48,6,3) Widely distributed in Yorkshire, but not common. In Yorkshire 2 YORKSHIRE specimens have been located in fungi 1 (including Polyporus squamosus), dung 0 NY NZ NZ OV SD SE SE TA (cow and horse), carrion, a compost heap, 9 and in the litter in beds of Phragmites, 8 Glyceria and Typha.
7 6 5 4 3 2 1
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zosterae (Thomson, C.G., 1856) Datomicra zosterae (Thomson, C.G., 1856)
3 WATSONIAN (2,1,0,0,0) There are three authentic records on the YNU database: Aughton 2 YORKSHIRE Ings (SE6938), 25/04/1988 (RJM teste CJ), 1 in flood refuse; Wheldrake Ings (SE7043), 0 NY NZ NZ OV SD SE SE TA 07/09/1989(RJM teste CJ), in a Phragmites 9 bed; Howe Hill (SE8070), 19/07/2003 8 (ABD), a single female in the stump of a 7 Polyporus squamosus. This Nationally Scarce species is associated with wetlands 6 and is predominantly southern in 5 distribution, although there are records 4 from Scotland. 3 2 Past confusion with nomenclature renders the following records unreliable without 1 voucher specimens: Pannal Ash (SE3052), 0 SD SE SE TA SJ SK SK TF 1936-1939 (RRUK); Askham Bog (SE5748), 9 04/1943 (WDH). 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 16/11/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
DILACRA Thomson, C.G., 1858 luteipes (Erichson, 1837)
Dilacra luteipes (Erichson, 1837)
3 WATSONIAN (5,1,12,4,0) Widely distributed in the 2 YORKSHIRE county, but not common. The few Yorkshire records with collecting data 1 indicate that they are usually found in the 0 NY NZ NZ OV SD SE SE TA ground litter associated with wet habitats. 9 Specimens have also been located in such 8 habitats by sweeping vegetation. 7 6 5 4 3 2 1
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vilis (Erichson, 1837)
Dilacra v ilis (Erichson, 1837)
3 WATSONIAN (2,0,11,6,0) Yorkshire records are restricted to a dozen locations, all in the 2 YORKSHIRE southern half of the county. The few 1 Yorkshire records with collecting data 0 NY NZ NZ OV SD SE SE TA have been located in the ground litter 9 associated with wet habitats (including 8 Phragmites and Glyceria beds) and, on one occasion, in an old straw heap. 7
6 5 4 3 2 1
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DIMETROTA Mulsant & Rey, 1873 aeneipennis (Thomson, C.G., 1856)
Dimetrota aeneipennis (Thomson, C.G., 1856)
3 WATSONIAN (0,0,4,0,0) There are four records on the 2 YORKSHIRE YNU database: Abbey Brook (SK1792), 15/11/1987 (EJS teste CJ); Bradfield 1 (SK2792), 24/09/1988 (EJS teste CJ); 0 NY NZ NZ OV SD SE SE TA Broomhead Reservoir (SK2696), 24/09/88 9 (EJS); Strines (SK2291), 02/09/1989 (EJS 8 teste CJ). The species is generally 7 associated with deer dung.
6
5 4 3 2 1
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atramentaria (Gyllenhal, 1810)
Dimetrota atramentaria (Gyllenhal, 1810)
3 WATSONIAN (25,14,136,22,16) Widely distributed and very common in the county. Yorkshire 2 YORKSHIRE specimens have been found in association 1 with a wide variety of animal dung, 0 NY NZ NZ OV SD SE SE TA carrion, and all types of rotting vegetation. 9 8 7 6 5 4 3 2 1
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cauta (Erichson, 1837)
Dimetrota cauta (Erichson, 1837)
3 WATSONIAN (6,0,2,1,0) Despite the national status of 2 YORKSHIRE ‘Common’ there are records from only 1 eight Yorkshire localities, all of which are
NY NZ NZ OV in the southern half of the county. 0 SD SE SE TA 9 Although the ecology of this species is unknown, Yorkshire specimens have been 8 located in horse dung, reedbed litter, 7 flood refuse, and pitfall traps in a grazing 6 marsh. 5 4 3 2 1
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cinnamoptera (Thomson, C.G., 1856)
Dimetrota cinnamoptera (Thomson, C.G., 1856)
3 WATSONIAN (0,17,32,4,6) Widely distributed in the county, but not common. The cluster of 2 YORKSHIRE records in VC63 may simply reflect 1 recorder effort and, conversely, the lack of 0 NY NZ NZ OV SD SE SE TA records from VC61 may be due to under- 9 recording. In Yorkshire the species has 8 been located in carrion, dung (cow, horse and deer), fungi, and a variety of decaying 7 plant material. 6 5 4 3 2 1
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ischnocera (Thomson, C.G., 1870)
Dimetrota ischnocera (Thomson, C.G., 1870)
3 WATSONIAN (2,2,32,7,1) Widely distributed in the 2 YORKSHIRE county, but not common. The cluster of records in the Huddersfield and Doncaster 1 areas simply reflect recorder effort. 0 NY NZ NZ OV SD SE SE TA Yorkshire specimens have been found in 9 dung (cow and horse), fungi, under beech 8 bark, and in decaying vegetable matter. 7 6 5 4 3 2 1
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laevana (Mulsant & Rey, 1852)
Dimetrota laev ana (Mulsant & Rey, 1852)
3 WATSONIAN (0,5,10,1,2) Widely distributed in the 2 YORKSHIRE county, but not common. The species is usually associated with dung but Yorkshire 1 specimens, although usually associated 0 NY NZ NZ OV SD SE SE TA with this medium, have also been located 9 in old straw, carrion, and decaying fungi. 8 7 6 5 4 3 2 1
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marcida (Erichson, 1837)
Dimetrota marcida (Erichson, 1837)
3 WATSONIAN (0,3,59,2,1) Although widely distributed in the county, with the exception of the 2 YORKSHIRE western half of VC63, it appears to be 1 genuinely rare. The few Yorkshire records 0 NY NZ NZ OV SD SE SE TA with collecting data indicate they were 9 found in fungi, moss, leaf litter, and 8 decaying vegetation. 7 6 5 4 3 2 1
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nigripes (Thomson, C.G., 1856)
Dimetrota nigripes (Thomson, C.G., 1856)
3 WATSONIAN (8,3,48,5,7) Widely distributed and 2 YORKSHIRE common in Yorkshire. The lack of records from most of VC61 may reflect under- 1 recording. In Yorkshire specimens have 0 NY NZ NZ OV SD SE SE TA been located in dung (horse, sheep and 9 dog), ground litter in a Phragmites bed, 8 stable refuse, grass heaps, carrion, and in 7 a Malaise trap.
6 5 4 3 2 1
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setigera (Sharp, 1869)
Dimetrota setigera (Sharp, 1869)
3 WATSONIAN (4,3,34,3,1) Widely distributed in 2 YORKSHIRE Yorkshire, especially in the southern half of the County. The species is usually found 1 in association with rotting organic 0 NY NZ NZ OV SD SE SE TA material, dung and carrion. The few 9 Yorkshire records with collecting data 8 indicate they have been located in horse 7 dung, fungi, vegetable refuse, and pitfall traps. 6
5 4 3 2 1
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DINARAEA Thomson, C.G., 1858 aequata (Erichson, 1837)
Dinaraea aequata (Erichson, 1837)
3 WATSONIAN (16,18,86,23,1) Widely distributed 2 YORKSHIRE throughout most of Yorkshire. The lack of records from most of VC61 may reflect 1 under-recording. Although the species 0 NY NZ NZ OV SD SE SE TA usually occurs in rotting wood, Yorkshire 9 specimens have also been located in 8 decaying vegetation, fungi (including 7 Polyporus sulphureus), carrion, and by sweeping vegetation. 6
5 4 3 2 1
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angustula (Gyllenhal, 1810)
Dinaraea angustula (Gyllenhal, 1810)
3 WATSONIAN (22,2,54,3,1) Widely distributed in Yorkshire, but not as common as D. 2 YORKSHIRE aequata (Erichson, 1837), with most 1 records stemming from the southern half 0 NY NZ NZ OV SD SE SE TA of the county. In Yorkshire it has been 9 found in a wide variety of rotting organic 8 material.
7 6 5 4 3 2 1
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linearis (Gravenhorst, 1802)
Dinaraea linearis (Grav enhorst, 1802)
3 WATSONIAN (0,1,3,1,0) There are five records on the 2 YORKSHIRE YNU database: Shirley Pool (SE5612), 10/1911 (recorder unknown); Melton 1 Wood (SE5104), 14/07/1982 (RJM teste 0 NY NZ NZ OV SD SE SE TA CJ), in leaf litter; Askham Bog (SE5748), 9 21/03/1987 (RJM), in deep litter in willow 8 bog; Rawcliffe Meadows (SE5853), 7 22/02/1997 (MH det. MLD); Potteric Carr (SK5900), 2002 (EJS). 6
5 4 3 2 1
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DOCHMONOTA Thomson, C.G., 1859 clancula (Erichson, 1837)
Dochmonota clancula (Erichson, 1837)
3 WATSONIAN (0,0,19,6,0) Yorkshire records of this 2 YORKSHIRE Nationally Scarce species suggest that it is restricted to the south and centre of the 1 county. The species is confined to England 0 NY NZ NZ OV SD SE SE TA where it occurs in bogs, fens, and 9 marshes. All Yorkshire records with 8 collecting data show they were collected 7 from ground litter, including that of Phragmites and Juncus, in wet habitats. 6
5 4 3 2 1
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ENALODROMA Thomson, C.G., 1859 hepatica (Erichson, 1839)
Enalodroma hepatica (Erichson, 1839)
3 WATSONIAN (1,2,8,9,3) Widely distributed in the 2 YORKSHIRE county, but not common. Although the ecology of this species is unknown, in 1 Yorkshire it has been located in fungi and 0 NY NZ NZ OV SD SE SE TA all types of rotting vegetation. 9 8 7 6 5 4 3 2 1
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GEOSTIBA Thomson, C.G., 1858 circellaris (Gravenhorst, 1806)
Geostiba circellaris (Grav enhorst, 1806)
3 WATSONIAN (36,21,302,55,13) This is a very common 2 YORKSHIRE and widely distributed species in Yorkshire with records from a wide variety of rotting 1 organic material, also under seaweed on 0 NY NZ NZ OV SD SE SE TA the coast and in the nest of the ant 9 Formica lugubris. 8 7 6 5 4 3 2 1
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HALOBRECTA Thomson, C.G., 1858 algae (Hardy, 1851)
Halobrecta algae (Hardy, 1851)
3 WATSONIAN (3,0,0,0,0) There are three records on the 2 YORKSHIRE YNU database: Spurn (TA4212), 1947- 1953 (WDH); Welwick Salt Marsh 1 (TA3421), 10/06/1947 (WOS); Bridlington 0 NY NZ NZ OV SD SE SE TA (TA1968), 16/05/2015 (RCa det. SAW), in 9 tideline seaweed. 8 7 6 5 4 3 2 1
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algophila (Fenyes, 1909)
NO RECORDS
flavipes Thomson, C.G., 1861
Halobrecta flav ipes Thomson, C.G., 1861
3 WATSONIAN (4,3,0,0,0) This is a littoral species which is 2 YORKSHIRE mainly confined to the English and Welsh coast lines. It has been recorded from six 1 Yorkshire localities – along the full length 0 NY NZ NZ OV SD SE SE TA of the coast and at the mouth of the 9 Humber estuary. 8 7 6 5 4 3 2 1
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princeps (Sharp, 1869)
NO RECORDS
HYDROSMECTA Thomson, C.G., 1858 delicatissima (Bernhauer, 1908)
NO RECORDS
delicatula (Sharp, 1869)
Hydrosmecta delicatula (Sharp, 1869)
3 WATSONIAN (0,1,0,0,0) There is a single record on the YNU database: Scarborough (TA0488), 2 YORKSHIRE date unknown (RL). The species is 1 associated with fine shingle besides rivers 0 NY NZ NZ OV SD SE SE TA and streams and is afforded the national 9 status pRDBK. 8 7 6 5 4 3 2 1
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eximia (Sharp, 1869)
Hydrosmecta eximia (Sharp, 1869)
3 WATSONIAN (0,2,0,1,0) There are three records on the 2 YORKSHIRE YNU database: Scarborough (TA0488), date unknown (RL); Whitby (NZ8911), 1 date unknown (HB); East Keswick Fits 0 NY NZ NZ OV SD SE SE TA (SE3545), 03/08/2009 (AG det. MLD), in 9 riverine shingle. 8 7 6 5 4 3 2 1
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fragilis (Kraatz, 1854)
Hydrosmecta fragilis (Kraatz, 1854)
3 WATSONIAN (0,2,0,0,0) There are two records of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Scarborough (TA0488), date 1 unknown (RL); Saltburn (NZ6721), 0 NY NZ NZ OV SD SE SE TA 01/07/1901 (MLT). The species is 9 associated with fine sandy shingle besides 8 rivers and streams. 7 6 5 4 3 2 1
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longula (Heer, 1839)
NO RECORDS
subtilissima (Kraatz, 1854)
Hydrosmecta subtilissima (Kraatz, 1854)
3 WATSONIAN (0,0,0,1,0) There is a single record of this 2 YORKSHIRE Nationally Scarce species on the YNU database: East Keswick Fits (SE3545), 1 03/08/2009 (AG det. MLD), in riverine 0 NY NZ NZ OV SD SE SE TA shingle. The species is widespread in 9 northern England and parts of Scotland 8 where it occurs in the drier areas of sand 7 and fine shingle besides rivers and streams. 6
5 4 3 2 1
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LIOGLUTA Thomson, C.G., 1858 alpestris (Heer, 1839)
Liogluta alpestris (Heer, 1839)
3 WATSONIAN (2,34,15,2,3) Widely distributed in the 2 YORKSHIRE county, but not common. The only Yorkshire records with collecting data 1 show they were located in a grass pile and 0 NY NZ NZ OV SD SE SE TA in pitfall traps. 9 8 7 6 5 4 3 2 1
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granigera (Kiesenwetter, 1850)
Liogluta granigera (Kiesenwetter, 1850)
3 WATSONIAN (1,2,18,2,2) Yorkshire records show the 2 YORKSHIRE species to be widely distributed, especially in the south and west of the county. The 1 species is restricted to Scotland and 0 NY NZ NZ OV SD SE SE TA northern England. The few Yorkshire 9 records with collecting data indicate they 8 were located in leaf litter, vegetable 7 refuse and debris around a pond edge.
6 5 4 3 2 1
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longiuscula (Gravenhorst, 1802)
Liogluta longiuscula (Grav enhorst, 1802)
3 WATSONIAN (6,9,18,2,1) Widely distributed in the county, but not common. The species is 2 YORKSHIRE found in moist ground litter, flood refuse, 1 etc. The only Yorkshire records with 0 NY NZ NZ OV SD SE SE TA collecting data indicate they were sieved 9 from plant litter and a grass heap. 8 7 6 5 4 3 2 1
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microptera Thomson, C.G., 1867
Liogluta microptera Thomson, C.G., 1867
3 WATSONIAN (1,4,72,13,3) Widely distributed in 2 YORKSHIRE Yorkshire, especially in the southern half of the county, but there is a lack of records 1 from most of VC61. In Yorkshire 0 NY NZ NZ OV SD SE SE TA specimens have been located in marsh 9 litter (including Phragmites and Carex 8 paniculata beds), fungi, moss, old hay, and 7 in pitfall traps set in woodland.
6 5 4 3 2 1
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pagana (Erichson, 1839)
Liogluta pagana (Erichson, 1839)
3 WATSONIAN (1,3,0,2,0) This Nationally Scarce species 2 YORKSHIRE has been recorded from six Yorkshire localities. There are no modern-day 1 records, however, the most recent being 0 NY NZ NZ OV SD SE SE TA from Malham Tarn (SD8967) in 10/1957. 9 The species probably occurs in a wide 8 variety of habitats as it has been recorded 7 from woodland, parkland, and marshes. The Yorkshire records with collecting data 6 show that it has been found in leaf litter, a 5 rotting grass pile, and with the ant Formica 4 lugubris. 3 2 1
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LYPROCORRHE Thomson, C.G., 1859 anceps (Erichson, 1837)
Lyprocorrhe anceps (Erichson, 1837)
3 WATSONIAN (0,6,5,1,0) This species, which is 2 YORKSHIRE myrmecophilous, appears to have two centres of population in Yorkshire – in the 1 eastern quarter of VC62 and the southern 0 NY NZ NZ OV SD SE SE TA half of VC63. All the Yorkshire records with 9 collecting data show they had originated 8 from the nests of the ant Formica lugubris 7 and, on single occasions, in cow dung, stable refuse and decaying plant debris. 6
5 4 3 2 1
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MICRODOTA Mulsant & Rey, 1873 aegra (Heer, 1841)
NO RECORDS
amicula (Stephens, 1832)
Microdota amicula (Stephens, 1832)
3 WATSONIAN (22,23,103,29,11) Widely distributed and common in the county. In Yorkshire it has 2 YORKSHIRE been found in fungi, (including Polyporus 1 squamosus and Pleurotus ostreatus), dung 0 NY NZ NZ OV SD SE SE TA (horse, cow and dog), under bark, carrion, 9 sweeping vegetation, pitfall traps in a wood, 8 and all types of rotting vegetation (including 7 tidal refuse).
6
5 4 3
2
1 SD SE SE TA 0 SJ SK SK TF 9
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atomaria (Kraatz, 1856)
NO RECORDS
atricolor (Sharp, 1869)
Microdota atricolor (Sharp, 1869)
3 WATSONIAN (4,4,14,3,1) Widely distributed in the 2 YORKSHIRE county, but not common. Although the ecology of this species is unknown, 1 Yorkshire specimens have been found in 0 NY NZ NZ OV SD SE SE TA fungi (including Piptoporus betulinus), 9 carrion, rotting vegetation, tidal refuse 8 and ungulate dung. 7 6 5 4 3 2 1
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benickiella (Brundin, 1948)
Microdota benickiella (Brundin, 1948)
3 WATSONIAN (0,0,2,2,0) There are four records of this Nationally Scarce species on the YNU 2 YORKSHIRE database: Studley Royal Park (SE2870), date 1 and recorder unknown; Rivelin (SK2987), 0 NY NZ NZ OV SD SE SE TA 05/01/1993 (EJS det. MLD); Askham Bog 9 (SE5648), 27/10/1997 (RGB); Fenwick 8 Churchyard (SE5916), 19/09/2016 (RJM), a single sieved from a compost heap. Although 7 generally associated with woodland there 6 are records from other habitats. 5 4 3 2 1
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boreella (Brundin, 1948)
Microdota boreella (Brundin, 1948)
3 WATSONIAN (0,0,2,1,0) There are three records on the 2 YORKSHIRE YNU database: Blackmoorfoot Reservoir (SE0912), 12/05/1990 (MLD teste CJ), in a 1 grass pile; Pighill Wood (SE0917), 0 NY NZ NZ OV SD SE SE TA 08/06/2006 (MLD), in decaying fungi; 9 Church Wood (SE2736), 27/04/2007 (MLD). 8 7 6 5 4 3 2 1
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excelsa (Bernhauer, 1911)
Microdota excelsa (Bernhauer, 1911)
3 WATSONIAN (0,0,3,0,0) There are three records of this Nationally Scarce species on the YNU 2 YORKSHIRE database: Langsett (SE2000), 13/06/1987 1 (EJS); Edge Moor, Blackmoorfoot Reservoir 0 NY NZ NZ OV SD SE SE TA (SE1012), 13/06/2012 (MLD), a single 9 female in horse dung; Dyson Wood 8 (SE1620), 11/05/2016 (MLD). This is a widely distributed species, but very rare. It 7 is found in broad-leaved woodland, pasture- 6 woodland and sallow carr where it has been 5 recorded from the dung of various 4 ungulates, carrion, fungi and grass cuttings. 3 2 1
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glabricula (Thomson, C.G., 1867)
Microdota glabricula (Thomson, C.G., 1867)
3 WATSONIAN (0,0,0,0,1) There is a single record of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Foxglove Covert (SE1596), 1 19/05/2008 (RJM), in rather dry cattle 0 NY NZ NZ OV SD SE SE TA dung. Predominantly southern in 9 distribution, where it has been located in 8 pasture-woodland, coniferous woodland 7 and possibly mixed woodland.
6
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indubia (Sharp, 1869)
Microdota indubia (Sharp, 1869)
3 WATSONIAN (5,6,38,13,2) Widely distributed and common in the county. The lack of records 2 YORKSHIRE from most of VC61 may reflect under- 1 recording. Although the ecology of this 0 NY NZ NZ OV SD SE SE TA species is unknown, Yorkshire specimens 9 have been found in fungi, rotting 8 vegetation, pitfall traps, dung, carrion, and 7 in the nest of a vole.
6 5 4 3 2 1
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inquinula (Gravenhorst, 1802)
NO RECORDS
liliputana (Brisout de Barneville, 1860)
Microdota liliputana (Brisout de Barnev ille, 1860)
3 WATSONIAN (0,0,2,0,0) There are two records on the 2 YORKSHIRE YNU database: Blackmoorfoot Reservoir (SE0912), 11/10/1987 (MLD), in rotting 1 fungi; Linthwaite (SE0914), 10/11/1990 0 NY NZ NZ OV SD SE SE TA (MLD det. CJ), on a sappy birch stump. 9 8 7 6 5 4 3 2 1
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minuscula (Brisout de Barneville, 1860)
Microdota minuscula (Brisout de Barnev ille, 1860)
3 WATSONIAN (0,0,1,0,0) There is a single record on the YNU database: Drop Clough (SE0413), 2 YORKSHIRE 10/01/1988 (DM det. CJ), in fungi. The 1 species is found in litter in marshes and 0 NY NZ NZ OV SD SE SE TA carrs. 9 8 7 6 5 4 3 2 1
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palleola (Erichson, 1837)
NO RECORDS
spatuloides (Benick, 1939)
Microdota spatuloides (Benick, 1939)
3 WATSONIAN (0,1,0,0,0) There is a single record on the 2 YORKSHIRE YNU database: Duncombe Park (SE6083), 10/09/1983 (RJM det. CJ), in carrion. The 1 species has only been recorded from 0 NY NZ NZ OV SD SE SE TA Scotland and Yorkshire, where it is usually 9 associated with woodland and pasture- 8 woodland. It is afforded the national 7 status pRDBK.
6 5 4 3 2 1
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subtilis (Scriba, 1866)
Microdota subtilis (Scriba, 1866)
3 WATSONIAN (0,0,2,3,0) There are five records on the 2 YORKSHIRE YNU database: Malham Tarn (SD8866), 09/1957 (WOS), in carrion; Askham Bog 1 (SE5648), 14/05/1969 (EWA teste MLD); 0 NY NZ NZ OV SD SE SE TA Bishop Wood (SE5634), 20/04/1984 9 (PJH), in a sappy birch stump; 8 Wharncliffe Wood (SK3195), 10/03/1988 7 (EJS); Eccleshall Wood (SK3282), 28/07/2001 (EJS teste MLD), in fungus on 6 a log. 5 4 3 2 1
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MOCYTA Mulsant & Rey, 1874 amplicollis (Mulsant & Rey, 1873)
Mocyta amplicollis (Mulsant & Rey, 1873)
3 WATSONIAN (2,2,13,2,2) Widely distributed in 2 YORKSHIRE Yorkshire but not common, with most records stemming from the southern half 1 of the county. Although the ecology of this 0 NY NZ NZ OV SD SE SE TA species is unknown, Yorkshire specimens 9 have been found in fungi and rotting 8 organic material. 7 6 5 4 3 2 1
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clientula (Erichson, 1839)
Mocyta clientula (Erichson, 1839)
3 WATSONIAN (2,5,16,4,0) Widely distributed in Yorkshire, but not common. The cluster of 2 YORKSHIRE records in the eastern half of VC63 almost 1 certainly reflects recorder effort. In 0 NY NZ NZ OV SD SE SE TA Yorkshire specimens have been located in 9 decomposing vegetable matter (mainly) 8 and fungi. 7 Past confusion with other members of this 6 difficult genus renders the following 5 records unreliable without voucher 4 specimens: Spurn (TA4212), 1947-1953 3 (WOS); Whitby (NZ9811), date unknown 2 (HB); Mulgrave Wood (NZ8412), date unknown (HB); Robin Hood’s Bay 1 (NZ9603), 28/06/1935 (recorder 0 SD SE SE TA SJ SK SK TF unknown); Goathland (SE8599), 9 14/09/1935 (HB). 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 17/7/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
fungi (Gravenhorst, 1806)
Mocyta fungi (Grav enhorst, 1806)
3 WATSONIAN (150,79,921,168,45) Widely distributed 2 YORKSHIRE and very common in the county. This is one of the commonest Aleocharinae in the 1 county and has been recorded from fungi 0 NY NZ NZ OV SD SE SE TA (including Polyporus squamosus), dung, 9 stable refuse, all types of decomposing 8 vegetable matter (including Phragmites, 7 Glyceria, Carex and Typha litter), and carrion. 6
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fussi (Bernhauer, 1908)
NO RECORDS
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orbata (Erichson, 1837)
Mocyta orbata (Erichson, 1837)
3 WATSONIAN (5,0,28,8,3) Widely distributed in Yorkshire, but not common. The cluster of 2 YORKSHIRE records in the eastern half of VC63 almost 1 certainly reflects recorder effort. In 0 NY NZ NZ OV SD SE SE TA Yorkshire specimens have been located in 9 all types of decaying vegetable matter, 8 horse dung, fungi, and by sweeping vegetation. 7
6 5 4 3 2 1
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orphana (Erichson, 1837)
Mocyta orphana (Erichson, 1837)
3 WATSONIAN (1,0,3,0,0) There are four records of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Wheldrake Ings (SE7043), 1 02/02/1985 (RGB det. JM); Rivelin 0 NY NZ NZ OV SD SE SE TA (SK2987), 09/03/1998 (EJS), in ground 9 litter and fungi; Wortley Top Forge 8 (SK2999), 05/09/2007 (EJS), sieved from 7 decaying plant debris; St James Churchyard, High Melton (SE5001), 6 11/05/2010 (RJM), sieved from grass 5 heaps. The species is mainly confined to 4 southern Britain, Yorkshire being at its 3 northern limit. 2 1
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MYCETOTA Ádám, 1987 fimorum (Brisout de Barneville, 1860)
NO RECORDS
laticollis (Stephens, 1832)
Mycetota laticollis (Stephens, 1832)
3 WATSONIAN (39,15,230,42,11) Widely distributed and common in the county. The species is 2 YORKSHIRE usually found in any type of decomposing 1 plant material. Yorkshire specimens have 0 NY NZ NZ OV SD SE SE TA been located in ground litter in Phragmites 9 and Typha beds, grass piles, stable refuse, 8 tide line debris, pitfall traps and the fungus 7 Pleurotus ostreatus.
6 5 4 3 2 1
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NEHEMITROPIA Lohse, 1971 lividipennis (Mannerheim, 1830)
Nehemitropia liv idipennis (Mannerheim, 1830)
3 WATSONIAN (16,13,53,14,1) Widely distributed and 2 YORKSHIRE common in the county. In Yorkshire specimens have been located in vegetable 1 refuse, grass heaps, old straw, fungi, and 0 NY NZ NZ OV SD SE SE TA cow dung. 9 8 7 6 5 4 3 2 1
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NEOHILARA Lohse, 1971 subterranea (Mulsant & Rey, 1853)
Neohilara subterranea (Mulsant & Rey, 1853)
3 WATSONIAN (0,0,2,0,0) There are two records on the YNU database: Blaxton Common (SE6901), 2 YORKSHIRE 08/07/1972 (EWA det. MLD), in the nest of 1 the ant Formica lugubris; Hatfield Moor 0 NY NZ NZ OV SD SE SE TA (SE6904), 25/07/2006 (AG det. MLD). 9 8 7 6 5 4 3 2 1
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NOTOTHECTA Thomson, C.G., 1858 confusa (Märkel, 1844)
Notothecta confusa (Märkel, 1844)
3 WATSONIAN (0,0,1,0,0) There is a single record of this 2 YORKSHIRE Nationally Scarce species which is generally associated with ants: Hall Dike 1 (SE1112), 17/12/1983 (DM det. MLD teste 0 NY NZ NZ OV SD SE SE TA CJ). 9 8 7 6 5 4 3 2 1
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flavipes (Gravenhorst, 1806)
Notothecta flav ipes (Grav enhorst, 1806)
3 WATSONIAN (0,6,14,0,0) The Yorkshire records are confined to VC63 and VC62. The species is 2 YORKSHIRE associated with the nests of ants and, with 1 the exception of specimens along the edge 0 NY NZ NZ OV SD SE SE TA of a tarn, the other Yorkshire records 9 indicate an association with the ant 8 Formica lugubris.
7 6 5 4 3 2 1
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OREOSTIBA Ganglbauer, 1895 tibialis (Heer, 1839)
Oreostiba tibialis (Heer, 1839)
3 WATSONIAN (0,0,0,8,4) This species is found on 2 YORKSHIRE mountains in northern England and Scotland. The Yorkshire distribution, from 1 ten localities, reflects the species’ upland 0 NY NZ NZ OV SD SE SE TA habitat requirements. Rather worrying is 9 the fact that are no post-2000 records. 8 7 6 5 4 3 2 1
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OUSIPALIA des Gozis, 1886 caesula (Erichson, 1839)
Ousipalia caesula (Erichson, 1839)
3 WATSONIAN (0,1,0,0,0) There is a single record on the 2 YORKSHIRE YNU database: Givendale (SE8885), 1925 (GBW det. HB). 1
NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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PACHNIDA Mulsant & Rey, 1874 nigella (Erichson, 1837)
Pachnida nigella (Erichson, 1837)
3 WATSONIAN (24,2,83,8,1) Most Yorkshire records are 2 YORKSHIRE confined to the southern half of the county. This is predominantly a southern 1 species which is found in marshy places. 0 NY NZ NZ OV SD SE SE TA The Yorkshire records with collecting data 9 reflect the habitat requirements of the 8 species, all records being from ground 7 litter in wet habitats (including Phragmites, Typha, Glyceria and 6 Sparganium beds). 5 4 3 2 1
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PACHYATHETA Munster, 1925 cribrata (Kraatz, 1856)
NO RECORDS
mortuorum (Thomson, C.G., 1867)
Pachyatheta mortuorum (Thomson, C.G., 1867)
3 WATSONIAN (0,0,0,1,0) There is a single record on the YNU database: Bishop Monkton Railway 2 YORKSHIRE Cutting (SE3165), 23/05-05/06/2010 (RW 1 det. RJM), from pitfall traps in grassland. 0 NY NZ NZ OV SD SE SE TA The national status of this species is 9 taxonomically clouded as Microdota 8 atricolor (Sharp, 1869), a widely 7 distributed species, was formerly known as mortuorum. 6 5 4 3 2 1
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PARAMEOTICA Ganglbauer, 1895 difficilis (Brisout de Barneville, 1860)
Parameotica difficilis (Brisout de Barnev ille, 1860)
3 WATSONIAN (17,1,1,0,1) The Yorkshire records of this 2 YORKSHIRE Nationally Scarce species indicate that the stronghold for the species is centred on 1 the Lower Derwent Valley. It is restricted 0 NY NZ NZ OV SD SE SE TA to England where it is found in wetlands 9 and marshes. In Yorkshire it has been 8 recorded from flood refuse, in vegetable 7 refuse, from a Typha bed, and by sweeping vegetation in a wet area. 6
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PARANOPLETA Brundin in Hansen, 1954 inhabilis (Kraatz, 1856) NO RECORDS
PHILHYGRA Mulsant & Rey, 1873 arctica (Thomson, C.G., 1856)
Philhygra arctica (Thomson, C.G., 1856)
3 WATSONIAN (0,8,2,8,5) The species is restricted to 2 YORKSHIRE mountains and high moorland in north Wales, northern England and Scotland. 1 The Yorkshire distribution, mainly from 0 NY NZ NZ OV SD SE SE TA two centres, reflects the habitat 9 requirements of the species. In Yorkshire 8 specimens have been located in flood 7 refuse, reed litter, and pitfall traps.
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britteni Joy, 1913
Philhygra britteni Joy, 1913
3 WATSONIAN (6,2,9,0,0) The 11 Yorkshire localities 2 YORKSHIRE which have yielded records are mainly confined to the southern half of the 1 county. This is a Nationally Scarce species 0 NY NZ NZ OV SD SE SE TA which is restricted to wet habitats. The 9 few Yorkshire records with collecting data 8 show that they have all been sieved from 7 flood refuse.
6 5 4 3 2 1
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debilis (Erichson, 1837)
Philhygra debilis (Erichson, 1837)
3 WATSONIAN (7,0,6,0,1) Recorded from 12 widely 2 YORKSHIRE distributed Yorkshire localities. It should be noted, however, that none of these are 1 post-2000. All the VC64 records stem from 0 NY NZ NZ OV SD SE SE TA various sites in the Lower Derwent Valley 9 with specimens being located in flood 8 refuse. 7 6 5 4 3 2 1
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deformis (Kraatz, 1856)
Philhygra deformis (Kraatz, 1856)
3 WATSONIAN (0,1,0,0,1) There are two records of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Scarborough (TA0488), date 1 unknown (RL); Semerwater (SD9287), 0 NY NZ NZ OV SD SE SE TA 28/08/1988 (RJM), in flood refuse. The 9 species usually occurs in wetland or wet 8 areas in woodland. It is considered 7 possible that it is associated with small mammals’ nests and runs. 6
5 4 3 2 1
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elongatula (Gravenhorst, 1802)
Philhygra elongatula (Grav enhorst, 1802)
3 WATSONIAN (17,22,54,28,14) Widely distributed and 2 YORKSHIRE common in Yorkshire. This is the commonest Philhygra species in the 1 county. In Yorkshire specimens have been 0 NY NZ NZ OV SD SE SE TA found in a wide variety of habitats, 9 including vegetable, stable, flood and old 8 hay refuse; in wet moss; and Phragmites, 7 Typha, Carex and Glyceria litter.
6 5 4 3 2 1
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fallaciosa (Sharp, 1869)
Philhygra fallaciosa (Sharp, 1869)
3 WATSONIAN (1,2,2,0,0) There are five records on the 2 YORKSHIRE YNU database: Danby High Moor (NZ7103), 27/05/1987 (MLD); Egton High 1 Moor (NZ7702), 03/09/1987 (RJM); 0 NY NZ NZ OV SD SE SE TA Skipwith Common (SE6637), 27/09/1989 9 (RBA det. MLD); May Moss (SE8796), 8 05/05/1996 (LA teste MLD); Thorne Moor 7 (SE7315), 10/1990 (DH det. MLD).
6 5 4 3 2 1
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gyllenhalii (Thomson, C.G., 1856)
Philhygra gyllenhalii (Thomson, C.G., 1856)
3 WATSONIAN (5,1,17,3,0) Despite the national status of 2 YORKSHIRE ‘Common’ the species shows a rather restricted distribution in the county, with 1 records stemming from the south and 0 NY NZ NZ OV SD SE SE TA centre. In Yorkshire specimens have been 9 located in Juncus litter, in pitfall traps 8 along the edge of a reed bed, and in moss. 7 6 5 4 3 2 1
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hygrobia (Thomson, C.G., 1856)
Philhygra hygrobia (Thomson, C.G., 1856)
3 WATSONIAN (7,1,16,3,0) Widely distributed in 2 YORKSHIRE Yorkshire, especially in the southern half of the county. It will be seen that most 1 post-2000 records, however, are confined 0 NY NZ NZ OV SD SE SE TA to a narrow north-south running central 9 band. This Nationally Scarce species is 8 associated with a wide variety of wet 7 habitats. In Yorkshire specimens have been found in decaying vegetable matter, 6 reedbed litter, old hay, and moss. 5 4 3 2 1
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hygrotopora (Kraatz, 1856)
Philhygra hygrotopora (Kraatz, 1856)
3 WATSONIAN (2,5,14,3,3) Widely distributed in 2 YORKSHIRE Yorkshire, but not common. Yorkshire specimens have been located in wet moss 1 behind a waterfall, in leaf litter, flood 0 NY NZ NZ OV SD SE SE TA refuse, and by sweeping vegetation. 9 8 7 6 5 4 3 2 1
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luridipennis (Mannerheim, 1830)
Philhygra luridipennis (Mannerheim, 1830)
3 WATSONIAN (2,3,7,3,0) Widely distributed in the 2 YORKSHIRE county, but not common. It should be noted that there is only a single post-2000 1 record. Most Yorkshire specimens have 0 NY NZ NZ OV SD SE SE TA been located in flood refuse. 9 8 7 6 5 4 3 2 1
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malleus (Joy, 1913)
Philhygra malleus (Joy, 1913)
3 WATSONIAN (6,5,31,4,2) Widely distributed, especially 2 YORKSHIRE in the southern half of the county. In Yorkshire most specimens have originated 1 from flood refuse, only occasionally in 0 NY NZ NZ OV SD SE SE TA reed beds, leaf litter around a pond edge, 9 and rotting vegetation. 8 7 6 5 4 3 2 1
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melanocera (Thomson, C.G., 1856)
Philhygra melanocera (Thomson, C.G., 1856)
3 WATSONIAN (6,4,32,5,0) Widely distributed, especially 2 YORKSHIRE in the southern half of the county. Yorkshire specimens have mainly 1 originated from flood refuse and grass 0 NY NZ NZ OV SD SE SE TA piles. 9 8 7 6 5 4 3 2 1
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obtusangula (Joy, 1913)
Philhygra obtusangula (Joy, 1913)
3 WATSONIAN (1,0,15,2,2) Widely distributed in the 2 YORKSHIRE western half of the county. The concentration of records in the western 1 half of VC63 may simply reflect recorder 0 NY NZ NZ OV SD SE SE TA effort. The few Yorkshire records with 9 collecting data show they have been 8 located in flood refuse, Juncus litter, and 7 decaying fungi.
6 5 4 3 2 1
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palustris (Kiesenwetter, 1844)
Philhygra palustris (Kiesenwetter, 1844)
3 WATSONIAN (6,4,7,1,0) Although widely distributed in 2 YORKSHIRE Yorkshire there are only records from 12 localities. The few Yorkshire records with 1 collecting data show they have been 0 NY NZ NZ OV SD SE SE TA located in flood refuse, vegetable refuse, 9 and in Glyceria beds. 8 7 6 5 4 3 2 1
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parca (Mulsant & Rey, 1873)
NO RECORDS
scotica (Elliman, 1909)
NO RECORDS
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terminalis (Gravenhorst, 1806)
Philhygra terminalis (Grav enhorst, 1806)
3 WATSONIAN (4,0,0,0,0) There are four records, from two localities, on the YNU database: 2 YORKSHIRE Bubwith (SE7030), 1919 (WJF) and 1933 1 (GBW); Wheldrake Ings (SE7044), 0 NY NZ NZ OV SD SE SE TA 11/03/1995 and 25/10/1996 (MLD), both 9 in flood refuse. The species is confined to 8 very few counties in England, Yorkshire being at the species’ northern limit. It is 7 restricted to the banks of rivers and 6 ponds, and other marshy habitats and is 5 afforded the national status pRDBK. The 4 statement that the Bubwith captures in 3 1919 were described as new to science (Naturalist 125:73) was given in error; 2 they were new to Britain. 1 SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 17/7/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
volans (Scriba, 1859)
Philhygra v olans (Scriba, 1859)
3 WATSONIAN (3,1,13,3,1) Widely distributed in 2 YORKSHIRE Yorkshire but not common, with most records stemming from the southern half 1 of the county. Yorkshire specimens have 0 NY NZ NZ OV SD SE SE TA been located in flood refuse, Carex litter, 9 Typha beds, Molinia fen, fungi, and in the 8 heartwood of a red-rotted oak. 7 6 5 4 3 2 1
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PLATARAEA Thomson, C.G., 1858 brunnea (Fabricius, 1798)
Plataraea brunnea (Fabricius, 1798)
3 WATSONIAN (12,6,33,1,0) Widely distributed in the 2 YORKSHIRE county, but not common. The lack of records from VC65 may be due to under- 1 ecording, but the lack of records from 0 NY NZ NZ OV SD SE SE TA most of VC64 is more difficult to explain. 9 Although generally found under wild 8 conditions it can also be synanthropic and 7 has been located indoors at Crosland Moor, Huddersfield (SE1114) on a number 6 of occasions. Other Yorkshire specimens 5 have been found in flood refuse, pitfall 4 traps, a Malaise trap, and by sweeping 3 vegetation. 2 1
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PYCNOTA Mulsant & Rey, 1874 paradoxa (Mulsant & Rey, 1861)
Pycnota paradoxa (Mulsant & Rey, 1861)
3 WATSONIAN (3,0,0,1,0) There are four records, from 2 YORKSHIRE three localities, of this Nationally Scarce species on the YNU database: Roundhay 1 Park (SE3338), 01/03/1910 (EWM); 0 NY NZ NZ OV SD SE SE TA Bubwith (SE7030), 04/1914 and 08/1916 9 (WJF), in moles’ nests; North Duffield 8 Carrs (SE6936), 08/1916 (WJF). The 7 species is confined to England where it is generally associated with moles’ nest. 6
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RHAGOCNEME Munster, 1923 subsinuata (Erichson, 1839)
Rhagocneme subsinuata (Erichson, 1839)
3 WATSONIAN (1,4,15,2,0) Widely distributed in Yorkshire, but rare. The only locality with 2 YORKSHIRE multiple records is Blackmoorfoot 1 Reservoir (SE0912 & SE1012) where it has 0 NY NZ NZ OV SD SE SE TA been repeatedly recorded since 1999. 9 Although the ecology of the species is 8 unknown, Yorkshire specimens have been 7 located in fungi, moss, decaying vegetable matter, and stable refuse. 6 5 4 3 2 1
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SCHISTOGLOSSA Kraatz, 1856 aubei (Brisout de Barneville, 1860)
NO RECORDS bergvalli Palm, 1968
NO RECORDS
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curtipennis (Sharp, 1869)
Schistoglossa curtipennis (Sharp, 1869)
3 WATSONIAN (0,0,6,2,0) There are records from eight 2 YORKSHIRE Yorkshire localities, most of which are in VC63, although this may simply reflect 1 recorder effort. As undissected specimens 0 NY NZ NZ OV SD SE SE TA or females of this species may include S. 9 bergvalli Palm, 1968, the records from 8 Yedingham (SE8979), 07/09/1931 (GBW) 7 and Langsett (SE2101), 16/03/1989 (EJS) are removed from the Yorkshire list. 6 Authenticated Yorkshire specimens have 5 been located in flood refuse and reed 4 litter. 3 2 1
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gemina (Erichson, 1837)
Schistoglossa gemina (Erichson, 1837)
3 WATSONIAN (3,0,3,2,0) Very rare with records from 2 YORKSHIRE eight Yorkshire localities, indicating that it is confined to a narrow north-south 1 central band. This is a Nationally Scarce 0 NY NZ NZ OV SD SE SE TA species which is generally associated with 9 wetlands. Yorkshire specimens have been 8 located in grass tussocks in wet habitats, 7 marsh litter, and Carex paniculata litter.
6 5 4 3 2 1
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viduata (Erichson, 1837)
NO RECORDS
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THINOBAENA Thomson, C.G., 1859 vestita (Gravenhorst, 1806)
Thinobaena v estita (Grav enhorst, 1806)
3 WATSONIAN (9,6,0,0,0) This species is confined to the 2 YORKSHIRE shoreline where it is found in litter. The inland record, which involved a single 1 female at Aughton Ings (SE6938) on 0 NY NZ NZ OV SD SE SE TA 14/05/1988 (MLD det. CJ), is most 9 unusual. 8 7 6 5 4 3 2 1
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TRAUMOECIA Mulsant & Rey, 1874 picipes (Thomson, C.G., 1856)
NO RECORDS
TRICHIUSA Casey, 1893 immigrata Lohse, 1984
Trichiusa immigrata Lohse, 1984
3 WATSONIAN (10,0,39,0,1) This immigrant species was first discovered in Britain during 1992 2 YORKSHIRE when specimens were found in Kent (Heal, 1 1993) and by the end of 1994 it had spread 0 NY NZ NZ OV SD SE SE TA north to Leicestershire (Lott, 1995). The 9 species then continued its northward 8 march and was first recorded in Yorkshire at Elstronwick (TA2332) on 27/05/1996 7 (WRD det. MLD). Since this time it has 6 continued to spread throughout the 5 county and also extended its range further 4 north; the first Scottish record came with 3 the finding of a specimen on Raasay on 08/09/2003 (Moore, 2004 & 2012). 2 Yorkshire specimens have been located in 1 all types of decomposing vegetable 0 SD SE SE TA SJ SK SK TF matter, including that containing dung. 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 17/7/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 Atlas of Coleoptera of Yorkshire. Part 5a – Staphylinidae: Aleocharinae Page | 97
XENOTA Mulsant & Rey, 1874 myrmecobia (Kraatz, 1856)
NO RECORDS
Subtribe THAMIARAEINA Fenyes, 1921
THAMIARAEA Thomson, C.G., 1858 cinnamomea (Gravenhorst, 1802)
Thamiaraea cinnamomea (Grav enhorst, 1802)
3 WATSONIAN (0,0,1,0,0) Very rare in Yorkshire with a 2 YORKSHIRE single record on the YNU database: Orange Wood, Blackmoorfoot Reservoir (SE1012), 1 11/11/1985 (MLD teste CJ), a single under 0 NY NZ NZ OV SD SE SE TA bark. The species is usually associated with 9 sap runs on oak Quercus spp. which have 8 been burrowed by the larvae of the Goat 7 Moth Cossus cossus. The Goat Moth is now very scarce in Yorkshire (Sutton & 6 Beaumont, 1989) with no records from the 5 Huddersfield area. 4 3 2 1
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hospita (Märkel, 1844)
Thamiaraea hospita (Märkel, 1844)
3 WATSONIAN (0,1,0,0,0) There is a single record of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Scarborough (TA0488), date and 1 recorder unknown. This species is found in 0 NY NZ NZ OV SD SE SE TA broad-leaved and pasture woodland and is 9 restricted to southern England, with 8 scattered records further north (although it 7 has not been recorded from Scotland).
6 5 4 3 2 1
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Tribe AUTALIINI Thomson, C.G., 1859
AUTALIA Leach, 1819 impressa (Olivier, 1795)
Autalia impressa (Oliv ier, 1795)
3 WATSONIAN (4,15,120,16,3) Widely distributed and common in Yorkshire. The lack of records 2 YORKSHIRE from most of VC61 may indicate under- 1 recording. This species is usually found in 0 NY NZ NZ OV SD SE SE TA decaying gill fungi. Yorkshire specimens 9 have been found in a wide variety of fungi 8 (including Piptoporus betulinus, Polyporus squamosus, Pleurotus ostreatus, Grifola 7 frondosa and Phallus sp.), in piles of 6 decaying vegetable matter (including a 5 garden compost heap), in leaf litter, in moss, 4 and in pitfall traps. 3 2 1
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longicornis Scheerpeltz, 1947
Autalia longicornis Scheerpeltz, 1947
3 WATSONIAN (4,2,31,3,3) It was not until 1983 that this 2 YORKSHIRE species, which had been masqueraded under the name A. impressa (Olivier, 1 1795), was described as new to Britain 0 NY NZ NZ OV SD SE SE TA (Ent. Monthly Mag. 119: 223-225). The 9 species has now been found to be widely 8 distributed but not common in Yorkshire. 7 The cluster of records in the Doncaster (VC63) area may simply reflect recorder 6 effort. Yorkshire specimens have been 5 located in rotting grass, dung (horse and 4 sheep), fungi (including Polyporus 3 squamosus and Grifola frondosa), and by 2 sweeping vegetation.
1
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puncticollis Sharp, 1864
Autalia puncticollis Sharp, 1864
3 WATSONIAN (0,0,0,2,1) Despite the national status of 2 YORKSHIRE ‘Common’ there are only three records on the YNU database: Malham (SD8967), 1 06/1954 & 09/1957 (WOS), in dung and 0 NY NZ NZ OV SD SE SE TA rotting grass; Rowton Beck (NY9126), 9 20/06/1981 (PS). 8 7 6 5 4 3 2 1
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rivularis (Gravenhorst, 1802)
Autalia riv ularis (Grav enhorst, 1802)
3 WATSONIAN (19,16,148,25,9) Widely distributed and 2 YORKSHIRE common in Yorkshire, especially in the southern half of the county. This species is 1 usually found in herbivore dung and 0 NY NZ NZ OV SD SE SE TA decaying plant material. Most Yorkshire 9 specimens with collecting data have been 8 located in the dung of horse, pony, cow, 7 sheep, or deer. Others have been found in compost and grass heaps, in stable refuse, 6 in fungi, in moss, and in carrion. 5 4 3 2 1
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Tribe DEINOPSINI Sharp, 1883
DEINOPSIS Matthews, A.H., 1838 erosa (Stephens, 1832)
Deinopsis erosa (Stephens, 1832)
3 WATSONIAN (4,0,13,3,0) The records indicate that the species is confined to the southern half of 2 YORKSHIRE the county. The cluster of records in the 1 eastern half of VC63 may simply indicate 0 NY NZ NZ OV SD SE SE TA recorder effort. The species is found in 9 damp places, mainly in woodland. In 8 Yorkshire specimens have been found in a Phragmites bed, in marsh litter, in moss, 7 and by sweep netting along the edge of a 6 pond. 5 4 3 2 1
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Tribe DIGLOTTINI Jakobson, 1909
DIGLOTTA Champion, G.C., 1899 mersa (Haliday, 1837)
Diglotta mersa (Haliday, 1837)
3 WATSONIAN (3,1,0,0,0) There are four records, from 2 YORKSHIRE two localities, on the YNU database: Spurn (TA4212), 21/06/1947 (EWA teste MLD), 1 1948-1952 (WOS) and 22/07/1963 (RCW), 0 NY NZ NZ OV SD SE SE TA all under stones below the high water 9 mark; Coatham Sands (NZ5725), 1995 8 (ME), in an area of slack and sand dunes. 7 This species is found under stones on sand between the tides. 6
5 4 3 2 1
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sinuaticollis (Mulsant & Rey, 1871)
NO RECORDS
Tribe FALAGRIINI Mulsant & Rey, 1873
BOHEMIELLINA Machulka, 1941 flavipennis (Cameron, 1920)
NO RECORDS
BORBOROPORA Kraatz in Kraatz & Fuss, 1862 kraatzii Fuss in Kraatz & Fuss, 1862
NO RECORDS
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CORDALIA Jacobs, 1925 obscura (Gravenhorst, 1802)
Cordalia obscura (Grav enhorst, 1802)
3 WATSONIAN (0,2,2,1,0) There are five records on the 2 YORKSHIRE YNU database: York district (SE55?), date unknown (WDH); Cloughton (TA0194), 1 1914 (WDH); Whitby (NZ8911), 0 NY NZ NZ OV SD SE SE TA 29/04/1035 (HB); Mag Wood, 9 Huddersfield (SE1313), 27/11/1983 (DM 8 det. MLD teste CJ); Cawood (SE5737), 7 10/05/1990 (MLD).
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FALAGRIA Leach, 1819 caesa Erichson, 1837
Falagria caesa Erichson, 1837
3 WATSONIAN (0,5,15,1,0) Widely distributed in 2 YORKSHIRE Yorkshire, especially in the southern half of the county, but not common. The few 1 Yorkshire specimens with collecting data 0 NY NZ NZ OV SD SE SE TA show they have been found in 9 decomposing vegetable matter, stable 8 refuse, and fungi. 7 6 5 4 3 2 1
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sulcatula (Gravenhorst, 1806)
Falagria sulcatula (Grav enhorst, 1806)
3 WATSONIAN (0,0,4,0,0) There are four records of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Royd House Wood (SE1613), 1 25/11/1984 (DM det. MLD); 0 NY NZ NZ OV SD SE SE TA Blackmoorfoot Reservoir (SE0912), 9 09/09/1985 (MLD); Dale Dike Reservoir 8 (SK2491), 02/04/1988 (EJS); Bradfield Dale 7 (SK2491), 28/12/1988 (EJS). The species is confined to England where it is associated 6 with wetlands, both fresh and brackish. 5 4 3 2 1
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FALAGRIOMA Casey, 1906 thoracica (Stephens, 1832)
Falagrioma thoracica (Stephens, 1832)
3 WATSONIAN (0,1,2,0,0) There are three records on 2 YORKSHIRE the YNU database: Saltburn (NZ6721), date unknown (MLT); Wentbridge 1 (SE4917), 1879 (recorder unknown); 0 NY NZ NZ OV SD SE SE TA Bradley Gate Wood (SE1619), 9 06/06/2006 (MLD). 8 7 6 5 4 3 2 1
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MYRMECOCEPHALUS MacLeay, 1873 concinnus (Erichson, 1839)
Myrmecocephalus concinnus (Erichson, 1839)
3 WATSONIAN (0,0,1,0,0) There is a single record on the 2 YORKSHIRE YNU database: Melton Wood (SE5104), 15/09/2014 (RJM), a single specimen by 1 sieving leaf litter and moss. 0 NY NZ NZ OV SD SE SE TA 9 8 7 6 5 4 3 2 1
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MYRMECOPORA Saulcy, 1865
Subgenus XENUSA Mulsant & Rey, 1875 brevipes Butler, 1909
NO RECORDS oweni Assing, 1997
NO RECORDS
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sulcata (Kiesenwetter, 1850)
Myrmecopora (Xenusa) sulcata (Kiesenwetter, 1850)
3 WATSONIAN (3,2,0,0,0) There are five records, from 2 YORKSHIRE four localities, on the YNU database: Scarborough (TA0488), date unknown 1 (TW); Saltburn (NZ6721), 1896 (MLT); 0 NY NZ NZ OV SD SE SE TA Spurn (TA4212), 1919 (GBW) and 1952 9 (WOS), on the Humber shore; Filey 8 (TA1281), 21/05/2005 (RJM det. MLD), in 7 strand line debris.
6 5 4 3 2 1
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uvida (Erichson, 1840)
Myrmecopora (Xenusa) uv ida (Erichson, 1840)
3 WATSONIAN (0,1,0,0,0) There is a single record on the
2 YORKSHIRE YNU database: Saltburn (NZ6721), 05/1896 (MLT). 1
NY NZ NZ OV 0 SD SE SE TA 9 8
7 6 5 4 3 2 1
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Tribe GYMNUSINI Heer, 1839
GYMNUSA Gravenhorst, 1806 brevicollis (Paykull, 1800)
Gymnusa brev icollis (Paykull, 1800)
3 WATSONIAN (1,8,24,3,3) Widely distributed but never common. The species is generally found in 2 YORKSHIRE moss in marshy places. The Yorkshire 1 records with collecting data show they 0 NY NZ NZ OV SD SE SE TA have been found in Sphagnum, in 9 vegetable refuse, at the roots of heather, 8 by sweeping vegetation, and in pitfall and 7 water traps.
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variegata Kiesenwetter, 1845
Gymnusa v ariegata Kiesenwetter, 1845
3 WATSONIAN (1,4,10,0,2) Widely distributed but rarely located. The cluster of records in the 2 YORKSHIRE western half of VC63 may simply reflect 1 recorder effort. This Nationally Scarce 0 NY NZ NZ OV SD SE SE TA species is confined to marshy areas. 9 Yorkshire specimens have been located in 8 Sphagnum and in wet pond litter.
7 6 5 4 3 2 1
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Tribe HOMALOTINI Heer, 1839
Subtribe BOLITOCHARINA Thomson, C.G., 1859
BOLITOCHARA Mannerheim, 1830 bella Märkel, 1844 Bolitochara bella Märkel, 1844
3 WATSONIAN (3,0,1,1,0) There are five records, from 2 YORKSHIRE three localities, on the YNU database: Malham Tarn (SD8967), 09/1957 (WOS), in 1 Polystictus versicolor; Lepton Great Wood 0 NY NZ NZ OV SD SE SE TA (SE1914), 07/07/1985 (DM det. MLD teste 9 CJ); Brook Farm, Elstronwick (TA2332), 8 06/05/2001 (WRD teste MLD) and 7 25/05/2002 & 22/09/2013 (WRD), all specimens were associated with Ash. The 6 species is found in fungi and fungoid 5 wood. 4 3 2 1
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lucida (Gravenhorst, 1802)
Bolitochara lucida (Grav enhorst, 1802)
3 WATSONIAN (0,7,13,7,0) Widely distributed in 2 YORKSHIRE Yorkshire, but not common. The species is found in fungi and fungoid wood. In 1 Yorkshire specimens have been located in 0 NY NZ NZ OV SD SE SE TA rotting fungi, under bark, and in woodland 9 leaf litter. 8 7 6 5 4 3 2 1
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mulsanti Sharp, 1875
Bolitochara mulsanti Sharp, 1875
3 WATSONIAN (0,3,6,0,1) There are Yorkshire records 2 YORKSHIRE from eight widely distributed localities, but only two of these are post-2000. This 1 Nationally Scarce species is northern in 0 NY NZ NZ OV SD SE SE TA distribution, Yorkshire being near to its 9 southern limit. It is found in decaying 8 fungus and under fungus infected bark in 7 broad-leaved and coniferous woodlands, and pasture-woodland. There is a record 6 on the YNU database from Kildale Wood 5 (NZ60) in June 1917 (MLT) which requires 4 confirmation (and therefore not mapped). 3 2 1
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obliqua Erichson, 1837
Bolitochara obliqua Erichson, 1837
3 WATSONIAN (9,22,124,18,4) Widely distributed and common in the county. The lack of records 2 YORKSHIRE from most of VC61 may indicate under- 1 recording. The species is usually found in 0 NY NZ NZ OV SD SE SE TA fungi. Yorkshire specimens have been 9 located in a variety of fungi (including 8 Polyporus squamosus), by beating fungoid 7 infested sticks, under bark, in moss, in ground and leaf litter, in flood refuse, and 6 under stones. 5 4 3 2 1
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pulchra (Gravenhorst, 1806)
Bolitochara pulchra (Grav enhorst, 1806)
3 WATSONIAN (0,1,1,0,0) There are two records of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Doncaster district (SE50?), date 1 unknown (HHC); Kildale (NZ6109), 0 NY NZ NZ OV SD SE SE TA 07/1917 (MLT), in a rotten branch. This is 9 a woodland species that is more widely 8 distributed in Scotland than further south. 7 It has been recorded from several fungus species including Piptoporus betulinus, 6 and under rotten beech bark. 5 4 3 2 1
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EURYUSA Erichson, 1837 optabilis Heer, 1839
NO RECORDS sinuata Erichson, 1837
NO RECORDS
HETEROTA Mulsant & Rey, 1874 plumbea (Waterhouse, G.R., 1858)
Heterota plumbea (Waterhouse, G.R., 1858)
3 WATSONIAN (1,0,0,0,0) There is a single record on the YNU database: Spurn (TA4212), 2 YORKSHIRE 21/10/1984 (MLD). This Nationally Scarce 1 coastal species is confined to the English 0 NY NZ NZ OV SD SE SE TA and Welsh coasts, Yorkshire being on its 9 northern limit along the eastern seaboard. 8 7 6 5 4 3 2 1
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LEPTUSA Kraatz, 1856 fumida (Erichson, 1839)
Leptusa fumida (Erichson, 1839)
3 WATSONIAN (10,29,177,33,5) Widely distributed and 2 YORKSHIRE common in Yorkshire, especially in VC63 (although this may simply reflect recorder 1 effort). The species occurs under bark and 0 NY NZ NZ OV SD SE SE TA in rotting wood. In Yorkshire specimens 9 have been located in fungi (including 8 Polyporus squamosus and Piptoporus), 7 under the bark of sycamore, beech and pine, in moss, ground litter under conifers, 6 and by sweeping vegetation. 5 4 3 2 1
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norvegica Strand, A., 1941
Leptusa norv egica Strand, A., 1941
3 WATSONIAN (0,3,2,0,0) There are five records of this Nationally Scarce species on the YNU 2 YORKSHIRE database: Langdale End (SE9491), 1 10/1979 (RSK det. MLD); Troutdale 0 NY NZ NZ OV SD SE SE TA (SE9288), 20/02/1982 (RSK det. MLD); 9 Elland Park Wood (SE1122), 09/09/1987 8 (MLD teste CJ); Hagg Wood Marsh (SE8390), 10/06/1989 (MLD); Orange 7 Wood, Blackmoorfoot Reservoir (SE1012), 6 25/06/2013 (MLD), under deciduous bark. 5 This species is found under bark (both 4 deciduous and evergreen) and is mainly 3 confined to Scotland, but there are scattered records throughout the rest of 2 Britain. 1 SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 22/7/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
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pulchella (Mannerheim, 1830)
Leptusa pulchella (Mannerheim, 1830)
3 WATSONIAN (1,7,13,3,2) Widely distributed in 2 YORKSHIRE Yorkshire, but not common. It is usually found under conifer bark. The few 1 Yorkshire records with collecting data 0 NY NZ NZ OV SD SE SE TA indicate that it has been found under the 9 bark of conifers and sycamore, and in leaf 8 litter under conifers. 7 6 5 4 3 2 1
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ruficollis (Erichson, 1839)
Leptusa ruficollis (Erichson, 1839)
3 WATSONIAN (1,11,33,6,1) Widely distributed and 2 YORKSHIRE common in Yorkshire, especially in VC63 (although this may simply reflect recorder 1 effort). The species is to be found in fungi 0 NY NZ NZ OV SD SE SE TA or under the bark of trees. In Yorkshire it 9 has been located in fungi (including 8 Piptoporus), in dead wood, moss, under 7 beech bark, and in the rotted heartwood of an oak. 6
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PSEUDOMICRODOTA Machulka, 1935 paganettii (Bernhauer, 1909)
NO RECORDS
PSEUDOPASILIA Ganglbauer, 1895 testacea (Brisout de Barneville, 1863)
NO RECORDS
RHOPALOCERINA Reitter, 1909 clavigera (Scriba, 1859)
NO RECORDS
TACHYUSIDA Mulsant & Rey, 1871 gracilis (Erichson, 1837)
NO RECORDS
THECTUROTA Casey, 1893 marchii (Dodero, 1922)
Thecturota marchii (Dodero, 1922)
3 WATSONIAN (0,0,1,0,0) There is a single record of this naturalised species on the YNU database: 2 YORKSHIRE Slaithwaite Tip (SE0713) (no longer extant), 1 15/04/1948 (EWA teste CJ). 0 NY NZ NZ OV SD SE SE TA 9 8 7 6 5 4 3 2 1
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williamsi (Bernhauer, 1936)
NO RECORDS
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Subtribe GYROPHAENINA Kraatz, 1856
AGARICOCHARA Kraatz, 1856 latissima (Stephens, 1832)
Agaricochara latissima (Stephens, 1832)
3 WATSONIAN (0,6,50,7,2) Widely distributed in 2 YORKSHIRE Yorkshire, especially in VC63 (although this may simply reflect recorder effort). In 1 Yorkshire specimens have been located in 0 NY NZ NZ OV SD SE SE TA fungi (including Piptoporus), carrion, 9 rotting vegetation, and cow dung. 8 7 6 5 4 3 2 1
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BRACHIDA Mulsant & Rey, 1871 exigua (Heer, 1839)
NO RECORDS
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ENCEPHALUS Stephens, 1832 complicans Stephens, 1832
Encephalus complicans Stephens, 1832
3 WATSONIAN (5,3,18,9,2) Widely distributed in Yorkshire, but not common. In Yorkshire 2 YORKSHIRE specimens have mainly been found by 1 sieving ground litter, but they have also 0 NY NZ NZ OV SD SE SE TA been located in moss, vegetable refuse, 9 and fungi. 8 7 6 5 4 3 2 1
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GYROPHAENA Mannerheim, 1830 affinis Mannerheim, 1830
Gyrophaena affinis Mannerheim, 1830
3 WATSONIAN (7,5,17,3,0) Widely distributed in the county, but not common. Most Yorkshire 2 YORKSHIRE specimens have been recorded from fungi 1 (including Polyporus squamosus, P. 0 NY NZ NZ OV SD SE SE TA sulphureus, Pleurotus ostreatus, 9 Laetiporus sulphureus, and Marasmius 8 ramealis), only occasionally from moss, ground litter and by sweeping vegetation. 7
6 5 4 3 2 1
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bihamata Thomson, C.G., 1867
Gyrophaena bihamata Thomson, C.G., 1867
3 WATSONIAN (2,0,8,1,0) There are records from eight 2 YORKSHIRE Yorkshire localities, all in the southern half of the county. This species is found in 1 fungi, particularly bracket fungi on trees, 0 NY NZ NZ OV SD SE SE TA often in large numbers. 9 8 7 6 5 4 3 2 1
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congrua Erichson, 1837
NO RECORDS
fasciata (Marsham, 1802)
Gyrophaena fasciata (Marsham, 1802)
3 WATSONIAN (1,6,22,10,0) Widely distributed in the 2 YORKSHIRE county, but not common. This species is associated with damaged or decaying 1 fungi. The Yorkshire records with 0 NY NZ NZ OV SD SE SE TA collecting data show that it is mainly found 9 in association with fungi (including 8 Polyporus squamosus), only occasionally 7 from moss and leaf litter.
6 5 4 3 2 1
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gentilis Erichson, 1839
Gyrophaena gentilis Erichson, 1839
3 WATSONIAN (6,16,38,13,2) Widely distributed in Yorkshire. Most Yorkshire specimens have 2 YORKSHIRE been found in association with fungi 1 (including Polyporus squamosus and 0 NY NZ NZ OV SD SE SE TA Pleurotus ostreatus), only occasionally in 9 decomposing vegetable material, the 8 rotted heartwood of oak, and carrion.
7 6 5 4 3 2 1
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hanseni Strand, A., 1946
NO RECORDS
joyi Wendeler, 1924
Gyrophaena j oyi Wendeler, 1924
3 WATSONIAN (1,0,0,2,0) There are three records of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Willow Garth, Ripon (SE3173), 1 24/06/2002 (RJM teste MLD), sieved from 0 NY NZ NZ OV SD SE SE TA old fungi under willow bark; Sharow 9 Mires, Ripon (SE3370), 06/08/2002 (RJM 8 teste MLD), in fungi under bark of poplar; 7 Humber Bridge CP (TA0126), 01/06/2003 (SE det. MLL). The species is associated 6 with marshy habitats. 5 4 3 2 1
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joyioides Wüsthoff, 1937
Gyrophaena j oyioides Wüsthoff, 1937
3 WATSONIAN (0,0,7,5,0) There are records from 11 2 YORKSHIRE Yorkshire localities, all in the southern half of the county. This Nationally Scarce 1 species is mainly restricted to southern 0 NY NZ NZ OV SD SE SE TA England where it is found in fungi in 9 woodland, including pasture-woodland. 8 All Yorkshire specimens have been located 7 in fungi (including Piptoporus betulinus).
6 5 4 3 2 1
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lucidula Erichson, 1837
NO RECORDS
manca Erichson, 1839
Gyrophaena manca Erichson, 1839
3 WATSONIAN (3,8,32,14,1) The species’ wide 2 YORKSHIRE distribution throughout England (including Yorkshire) strongly suggests 1 that the status of ‘Nationally Scarce’ 0 NY NZ NZ OV SD SE SE TA should be reduced to no more than ‘Local’. 9 The species occurs in fungi, particularly 8 that attached to trees. In Yorkshire 7 specimens have been located in fungi (including Polyporus squamosus, P. 6 sulphureus and Laetiporus sulphureus), 5 moss and leaf litter. 4 3 2 1
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minima Erichson, 1837
Gyrophaena minima Erichson, 1837
3 WATSONIAN (1,6,5,5,1) Widely distributed in the 2 YORKSHIRE county, but not common. The few Yorkshire records with collecting data 1 show they have been found in fungi 0 NY NZ NZ OV SD SE SE TA (including Polyporus) and in vegetable 9 refuse. 8 7
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munsteri Strand, A., 1935
Gyrophaena munsteri Strand, A., 1935
3 WATSONIAN (0,1,0,1,0) There are two records on the YNU database: Cayton Bay (TA0785), 2 YORKSHIRE 18/06/1985 (RBA det. MLD); Hackfall 1 Wood (SE2377), 16/06/1996 (RJM det. 0 NY NZ NZ OV SD SE SE TA MLD), in vegetable refuse by the river. The 9 species occurs in fungi in woodland and is 8 afforded the national status pRDBK. 7 6 5 4 3 2 1
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nana (Paykull, 1800)
Gyrophaena nana (Paykull, 1800)
3 WATSONIAN (0,0,1,1,1) There are three authentic records 2 YORKSHIRE on the YNU database: Bolton Percy (SE5341), 22/05/1943 (WOS), in Polyporus squamosus; 1 Torne Bridge (SE6703), 25/09/1969 (EWA 0 NY NZ NZ OV SD SE SE TA teste MLD); Cover Banks, Middleham 9 (SE1387), 22/08/1992 (RJM), in fungi under 8 bark. 7 Past confusion with other members of the 6 genus renders the following records suspect 5 without voucher specimens: Skipwith 4 Common (SE6637), 15/07/1916 (WJF); 3 Allerthorpe Common (SE7647), 16/09/1923 2 (WJF).
1
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poweri Crotch, 1867
Gyrophaena poweri Crotch, 1867
3 WATSONIAN (1,0,0,0,0) There is a single record on the 2 YORKSHIRE YNU database: Thorpe Estate, Rudston (TA1167), 19/08/1989 (MLD). The species 1 is found in fungi on trees and is 0 NY NZ NZ OV SD SE SE TA predominantly southern in distribution. It 9 is afforded the national status pRDBK. 8 7 6 5 4 3 2 1
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pseudonana Strand, A., 1939
NO RECORDS pulchella Heer, 1839
NO RECORDS
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rousi Dvorák, 1966
NO RECORDS
strictula Erichson, 1839
Gyrophaena strictula Erichson, 1839
3 WATSONIAN (0,0,4,0,0) There are four authentic records 2 YORKSHIRE on the YNU database: Hallfield Dore (SK2980), 01/04/1985 (EJS teste MLD); 1 Booth Wood, Wortley (SK3198), 0 NY NZ NZ OV SD SE SE TA 09/04/1985 (EJS); Damflask (SK2790), 9 15/04/1995 (EJS); Wharncliffe Wood 8 (SE3195), 04/04/1998 (EJS). This Nationally 7 Scarce species is confined to England and south-west Scotland where it is associated 6 with the hard fungus Daedalaea quercina 5 on oak. 4 3 Past confusion with other members of the 2 genus renders the record from Scarborough (TA0488) in 1868 (ECH) 1 unreliable without voucher specimens. 0 SD SE SE TA SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 16/11/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
transversalis Strand, 1939
NO RECORDS
williamsi Strand, A., 1935
Gyrophaena williamsi Strand, A., 1935
3 WATSONIAN (0,0,1,1,0) There are two records of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Harlow Carr Gardens (SE2954), 1 24/09/1948 (EWA teste MLD); Bottom’s 0 NY NZ NZ OV SD SE SE TA Mill Wood (SE1307), 20/06/1985 (DM det. 9 MLD). The species is predominantly 8 southern in distribution, the records above 7 being the most northerly.
6 5 4 3 2 1
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Subtribe HOMALOTINA Heer, 1839
ANOMOGNATHUS Solier, 1849 cuspidatus (Erichson, 1839)
Anomognathus cuspidatus (Erichson, 1839)
3 WATSONIAN (0,5,7,0,1) Despite the national status of 2 YORKSHIRE ‘Common’ the species is only very thinly distributed in Yorkshire. The species is 1 found under bark. 0 NY NZ NZ OV SD SE SE TA 9 8 7 6 5 4 3 2 1
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CYPHEA Fauvel, 1863 curtula (Erichson, 1837)
NO RECORDS
HOMALOTA Mannerheim, 1830 plana (Gyllenhal, 1810)
Homalota plana (Gyllenhal, 1810)
3 WATSONIAN (4,8,16,7,1) The species is widely, but 2 YORKSHIRE thinly, distributed in the county. Although usually found under bark, 1 Yorkshire specimens have also been 0 NY NZ NZ OV SD SE SE TA located in fungi (including Polystictus 9 versicolor and Laetiporus sulphureus) 8 and in a Glyceria bed. 7
6 5 4 3 2 1
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Subtribe SILUSINA Fenyes, 1918
SILUSA Erichson, 1837 rubiginosa Erichson, 1837
NO RECORDS
Tribe HYGRONOMINI Thomson, C.G., 1859
HYGRONOMA Erichson, 1837 dimidiata (Gravenhorst, 1806)
Hygronoma dimidiata (Grav enhorst, 1806)
3 WATSONIAN (26,7,80,15,5) Widely distributed in 2 YORKSHIRE Yorkshire, especially in VC63. It is generally associated with leaf litter and 1 moss in rich marshland, although it will 0 NY NZ NZ OV SD SE SE TA overwinter in reed stems and leaf 9 sheaths. In Yorkshire specimens have 8 been located in litter in the beds of Typha, 7 Phragmites, Carex, Glyceria and Sparganium. 6
5 4 3 2 1
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Tribe HYPOCYPHTINI Laporte, 1835
CYPHA Leach, 1819 apicalis (Brisout de Barneville, 1863)
NO RECORDS aprilis (Rey, 1882)
NO RECORDS
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discoidea (Erichson, 1839)
Cypha discoidea (Erichson, 1839)
3 WATSONIAN (4,0,1,0,1) There are six records, from five 2 YORKSHIRE localities, of this Nationally Scarce species on the YNU database: Hornsea (TA1948), 1 date and recorder unknown & 26/10/1996 0 NY NZ NZ OV SD SE SE TA (RJM), in ground litter; Fowlsyke Flash 9 (SK5298), 01/05/2000 (RJM), in reed bed 8 litter; Pulfin Bog (TA0544), 20/04/2011 7 (RJM); Hollym Carrs (TA3224), 08/09/2014 (RJM); Great Shunner Fell (SD8597), 6 10/06/2015 (RW det. RJM), at the edge of 5 a Sphagnum pool. The species is widely 4 distributed in England (mainly southern) 3 and is confined to fens, marshes and 2 estuaries where it is associated with plant litter. 1
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laeviuscula (Mannerheim, 1830)
Cypha laev iuscula (Mannerheim, 1830)
3 WATSONIAN (8,7,13,7,3) Widely distributed in 2 YORKSHIRE Yorkshire, but not common. Yorkshire specimens have been located in grass piles, 1 stable refuse and in the fungus Hypholoma 0 NY NZ NZ OV SD SE SE TA fasciculare. 9 8 7 6 5 4 3 2 1
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longicornis (Paykull, 1800)
Cypha longicornis (Paykull, 1800)
3 WATSONIAN (32,25,94,14,2) Widely distributed and 2 YORKSHIRE common in Yorkshire, especially in the southern half of the county. The species 1 occurs under damp conditions in 0 NY NZ NZ OV SD SE SE TA vegetable refuse and, in Yorkshire, has 9 been recorded from grass cuttings, 8 compost heaps, stable refuse, and the 7 litter in Phragmites and Carex paniculata beds. 6
5 4 3 2 1
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pulicaria (Erichson, 1839)
Cypha pulicaria (Erichson, 1839)
3 WATSONIAN (5,3,4,1,0) Widely distributed in the 2 YORKSHIRE county, but not common. This Nationally Scarce species is associated with bogs and 1 marshes. 0 NY NZ NZ OV SD SE SE TA 9 8 7 6 5 4 3 2 1
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punctum (Motschulsky, 1857)
Cypha punctum (Motschulsky, 1857)
3 WATSONIAN (1,2,0,0,0) There are three records on the 2 YORKSHIRE YNU database: Mulgrave Wood (NZ8513), 1934 (recorder unknown); Dunnington 1 Common (SE6750), 1971 (recorder 0 NY NZ NZ OV SD SE SE TA unknown); Ashberry Pastures (SE5685), 9 29/09/1972 (EWA), several at grass roots. 8 The species is usually found among moss, 7 litter and decaying vegetable matter.
6 5 4 3 2 1
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seminulum (Erichson, 1839)
Cypha seminulum (Erichson, 1839)
3 WATSONIAN (0,1,0,1,0) There are two authentic records on the YNU database: Duncombe 2 YORKSHIRE Park (SE6083), 10/06/1980 (CJ); Studley 1 (SE2870), 19/06/1989 (CJ). The species is 0 NY NZ NZ OV SD SE SE TA found in woodland where it has been 9 located in rotting wood, under bark, in 8 fungi and in moss. It is afforded the national status pRDBK. 7
6 Past confusion with nomenclature renders 5 the following records unreliable without 4 voucher specimens: Saltburn (NZ6721), 3 08/1896 (MLT); Cornelian Bay (TA0686), 09/1926 (recorder unknown). 2
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tarsalis (Luze, 1902)
NO RECORDS
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HOLOBUS Solier, 1849 flavicornis (Lacordaire, 1835)
NO RECORDS
OLIGOTA Mannerheim, 1830 apicata (Erichson, 1837)
Oligota apicata Erichson, 1837
3 WATSONIAN (3,8,7,3,2) Widely distributed but rarely encountered in Yorkshire. This Nationally 2 YORKSHIRE Scarce species is confined to England and 1 south-east Scotland. It is found in 0 NY NZ NZ OV SD SE SE TA woodland and pasture-woodland where it 9 is possibly a predator of Cis (Ciidae) larvae 8 and mites. Yorkshire specimens have been located in fungi, under bark, and in 7 woodland litter. 6 5 4 3 2 1
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granaria Erichson, 1837
NO RECORDS
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inflata (Mannerheim, 1830)
Oligota inflata (Mannerheim, 1830)
3 WATSONIAN (4,13,18,3,0) Widely distributed in the 2 YORKSHIRE county, but not common. The species occurs in haystacks, vegetable refuse, 1 moss, etc. In Yorkshire specimens have 0 NY NZ NZ OV SD SE SE TA been found in grass and straw heaps, a 9 garden compost heap, Carex litter, 8 fungoid leaf litter, horse dung, and in a 7 Robin’s nest.
6 5 4 3 2 1
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parva Kraatz, 1862
Oligota parv a Kraatz, 1862
3 WATSONIAN (12,1,24,0,0) This species, which is 2 YORKSHIRE associated with mouldy grass and manure heaps, although widely distributed in the 1 eastern half of the county, is not common. 0 NY NZ NZ OV SD SE SE TA In Yorkshire specimens have been located 9 in all types of rotting vegetation and, 8 occasionally, fungi. 7 6 5 4 3 2 1
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picipes (Stephens, 1832)
Oligota picipes (Stephens, 1832)
3 WATSONIAN (1,2,2,0,0) There are five authentic records on the YNU database: Stoney 2 YORKSHIRE Royd (SE3202), 25/01/1987 (EJS teste CJ); 1 Staxton (TA0279), 05/06/1996 (LA det. 0 NY NZ NZ OV SD SE SE TA MLD); Rawcliffe Meadows (SE5852), 9 14/05/1997 (MLD); Blenkins Farm, 8 Owstwick (TA2632), 15/11/2002 (WRD det. MLD); Blackmoorfoot Reservoir 7 (SE1212), 26/05/2005 (MLD), in grass 6 cuttings. 5 4 Past confusion with nomenclature renders 3 the following records unreliable without voucher specimens: Gawthorpe (SE2722), 2 29/03/1948 (EWA); Spurn (TA4212), 1 20/07/1948 & 13/08/1948 (WOS); 0 SD SE SE TA SJ SK SK TF Saltergate (SE8595), 10/04/1937 (HB). 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 16/11/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
pumilio Kiesenwetter, 1858
Oligota pumilio Kiesenwetter, 1858
3 WATSONIAN (0,0,1,0,0) There is a single authentic 2 YORKSHIRE record on the YNU database: Torne Bridge (SE6703), 25/09/1969 (EWA). 1
0 NY NZ NZ OV SD SE SE TA Past confusion with O. pusillima 9 (Gravenhorst, 1806) renders the following 8 records unreliable without voucher 7 specimens: Scarborough (TA0488), date unknown (KL); Malham Tarn (SD8967), 6 09/1957 (WOS). 5 4 3 2 1
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punctulata Heer, 1839
Oligota punctulata Heer, 1839
3 WATSONIAN (7,1,44,12,3) Widely distributed, 2 YORKSHIRE especially in the south and centre of the county. All records on the YNU database 1 have been reappraised since the revision 0 NY NZ NZ OV SD SE SE TA of the genus in Ent. Monthly Mag. 106: 54- 9 62. Yorkshire specimens have been 8 located in haystacks, grass cuttings, 7 compost heaps, Carex litter, fungi, and stable refuse. 6
5 4 3 2 1
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pusillima (Gravenhorst, 1806)
Oligota pusillima (Grav enhorst, 1806)
3 WATSONIAN (6,2,8,3,0) Widely distributed in the county, but not common. It is found in 2 YORKSHIRE manure heaps and grass piles, 1 occasionally in rotting fungi and carrion. 0 NY NZ NZ OV SD SE SE TA 9 Past confusion with O. pumilio 8 Kiesenwetter, 1858 renders the following record unreliable without voucher 7 specimens: Spurn (TA4212), 1947-1953 6 (WOS). 5 4 3 2 1
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Tribe LOMECHUSINI Fleming, 1821
Subtribe LOMECHUSINA Fleming, 1821
LOMECHUSA Gravenhorst, 1806 emarginata (Paykull, 1789)
Lomechusa emarginata (Paykull, 1789)
3 WATSONIAN (0,0,7,0,0) There are six records, from five 2 YORKSHIRE localities, on the YNU database: Rossington (SK6398), 10/05/1908 (HHC); Dalton Bank, 1 Huddersfield (SE1718), 1910 and 1916 0 NY NZ NZ OV SD SE SE TA (SLM); Doncaster (SE50?), 1913 (HHC); 9 Huddersfield (SE11?), 02/03/1917 (SLM) 8 (this record may refer to Dalton Bank); 7 Anston Stones Wood (SK5383), 19/05/1985 (WAE teste MLD). This Nationally Scarce 6 species is restricted to England, where it 5 occurs in the nests of a number of ant 4 species. 3 2 1
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paradoxah Gravenhorst, 1806
e r Lomechusa paradoxa Grav enhorst, 1806 e 3a WATSONIAN (0,1,0,0,0) There is a single record on the 2r YORKSHIRE YNU database: Castle Howard (SE7270), 17/09/2003 (ABD), beaten from oak. The 1e s species occurs in grassland and sandy 0 NY NZ NZ OV e SD SE SE TA places, often on the coast. It is afforded 9v the national status RDBK. 8e 7n 6r e 5 c 4o 3r 2d 1s , 0 SD SE SE TA f SJ SK SK TF 9r 8o post 31/12/1999 pre 31/12/1999 m Map produced for Yorkshire Naturalists' Union by DMAP 22/7/2018 f 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 i v e
l
o Atlasc of Coleoptera of Yorkshire. Part 5a – Staphylinidae: Aleocharinae a Page | 131 l i t i e
LOMECHUSOIDES Tottenham, 1939 strumosus (Fabricius, 1775)
NO RECORDS
Subtribe MYRMEDONIINA Thomson, C.G., 1867
DRUSILLA Leach, 1819 canaliculata (Fabricius, 1787)
Drusilla canaliculata (Fabricius, 1787)
3 WATSONIAN (38,37,291,18,8) Widely distributed and 2 YORKSHIRE common in the county. Yorkshire specimens have been located in ant nests’, 1 fungi, moss, carrion, litter in Phragmites 0 NY NZ NZ OV SD SE SE TA and Glyceria beds, in a saltmarsh, and in 9 flood refuse. 8 7 6 5 4 3 2 1
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MYRMOECIA Mulsant & Rey, 1874 plicata (Erichson, 1837)
NO RECORDS
PELLA Stephens, 1835 313 cognata (Märkel, 1842)
NO RECORDS funesta (Gravenhorst, 1806)
NO RECORDS
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humeralis (Gravenhorst, 1802)
Pella humeralis (Grav enhorst, 1802)
3 WATSONIAN (1,3,6,0,1) Widely distributed in Yorkshire, but not common. It will be noted that 2 YORKSHIRE there are no post-2000 records. Indeed, 1 the last record was as long ago as April 0 NY NZ NZ OV SD SE SE TA 1988. This species is associated with ants 9 and all Yorkshire records refer to 8 specimens from the nests of the ant Formica lugubris. 7 6 5 4 3 2 1
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laticollis (Märkel, 1845)
NO RECORDS
limbata (Paykull, 1789)
Pella limbata (Paykull, 1789)
3 WATSONIAN (1,0,2,0,0) There are three records on the YNU database: Eastrington (SE7830), 2 YORKSHIRE 01/06/2003 (SE det. MLL); Brodsworth 1 Community Woodland (SE5206), 0 NY NZ NZ OV SD SE SE TA 04/05/2016 (RJM), sieved from litter in a 9 reed bed; Sprotbrough (SE5301), 8 21/06/2016 (RJM), by general sweeping. 7 The species lives in the nests of ants of the genus Lasius. 6 5 4 3 2 1
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lugens (Gravenhorst, 1802)
NO RECORDS
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ZYRAS Stephens, 1835 collaris (Märkel, 1842)
NO RECORDS haworthi Stephens, 1835
NO RECORDS
Tribe MYLLAENINI Ganglbauer, 1895
MYLLAENA Erichson, 1837 brevicornis (Matthews, A.H., 1838)
Myllaena brev icornis (Matthews, A.H., 1838)
3 WATSONIAN (3,12,30,12,10) Widely distributed and 2 YORKSHIRE common. This is the commonest Myllaena species in the county and has 1 been located in flood refuse, decaying 0 NY NZ NZ OV SD SE SE TA plant debris, reedbed litter, grass roots, 9 and in moss. 8 7 6 5 4 3 2 1
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dubia (Gravenhorst, 1806)
Myllaena dubia (Grav enhorst, 1806)
3 WATSONIAN (10,5,46,4,3) Widely distributed and 2 YORKSHIRE common in the county. The cluster of records in the southern half of VC63 may 1 simply reflect recorder effort. This is a 0 NY NZ NZ OV SD SE SE TA very fast moving species which is found in 9 moss and plant litter. 8 7 6 5 4 3 2 1
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Myllaena elongata (Matthews, A.H., 1838)
3 WATSONIAN (1,3,3,0,0) Very rare with records from six 2 YORKSHIRE widely distributed localities. This Nationally Scarce species is found in damp 1 places where it has been located with 0 NY NZ NZ OV SD SE SE TA Bledius (Staphylinidae) and Heterocerus 9 (Heteroceridae). 8 7 6 5 4 3 2 1
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fowleri Matthews, A., 1883
NO RECORDS
gracilicornis Fairmaire & Brisout de Barneville, 1859
NO RECORDS
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gracilis (Matthews, A.H., 1838)
Myllaena gracilis (Matthews, A.H., 1838)
3 WATSONIAN (2,0,3,2,0) There are records from seven 2 YORKSHIRE Yorkshire localities. Rather worryingly, however, is the fact that there are no post- 1 2000 record. Although widely distributed 0 NY NZ NZ OV SD SE SE TA in England, it is chiefly southern. It is found 9 in wet habitats. The few Yorkshire records 8 with collecting data indicate they have 7 been located in a Phragmites bed, in vegetable refuse by a pond, and in a 6 grazed field. 5 4 3 2 1
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infuscata Kraatz, 1853
Myllaena infuscata Kraatz, 1853
3 WATSONIAN (10,4,16,12,1) Widely distributed with 2 YORKSHIRE most records stemming from the south and east of the county. Yorkshire 1 specimens have been found in Phragmites 0 NY NZ NZ OV SD SE SE TA and Carex litter, in a Glyceria bed, in old 9 lawn mowings, in vegetable refuse, and in 8 moss. 7 6 5 4 3 2 1
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intermedia Erichson, 1837
Myllaena intermedia Erichson, 1837
3 WATSONIAN (4,0,18,1,3) Despite the national status of 2 YORKSHIRE ‘Common’ the Yorkshire records indicate that it is confined to a narrow central 1 band. It is found in wet habitats. The only 0 NY NZ NZ OV SD SE SE TA Yorkshire specimens with collecting data 9 were found in a Typha bed and by 8 sweeping vegetation around a pond. 7 6 5 4 3 2 1
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kraatzi Sharp, 1871
Myllaena kraatzi Sharp, 1871
3 WATSONIAN (1,8,9,0,1) Very rare with records from a 2 YORKSHIRE number of widely scattered localities, especially in the south and north of the 1 county. This Nationally Scarce species is 0 NY NZ NZ OV SD SE SE TA found in bogs and marshes. In Yorkshire 9 specimens have been located in reedbed 8 and ground litter and in flood refuse. 7 6 5 4 3 2 1
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masoni Matthews, A., 1883
NO RECORDS
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minuta (Gravenhorst, 1806)
Myllaena minuta (Grav enhorst, 1806)
3 WATSONIAN (4,1,12,12,3) Despite the national status of 2 YORKSHIRE ‘Common’ the Yorkshire records are confined to a north- south central band. 1 This is a very fast moving species which is 0 NY NZ NZ OV SD SE SE TA found in moss and leaf litter. Yorkshire 9 specimens have been found in reed beds 8 (both Phragmites and Typha), in Glyceria 7 beds, in moss, in ground litter near water, and in a dead bird by the water’s edge. 6
5 4 3 2 1
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Tribe OXYPODINI Thomson, C.G., 1859
Subtribe DINARDINA Mulsant & Rey, 1873
DINARDA Leach, 1819 dentata (Gravenhorst, 1806)
NO RECORDS
hagensi Wasmann, 1889
NO RECORDS
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maerkeli Kiesenwetter, 1843
Dinarda maerkeli Kiesenwetter, 1843
3 WATSONIAN (1,7,2,0,0) There are Yorkshire records 2 YORKSHIRE from eight widely distributed localities but only one of these is post-2000. This 1 species is usually found in association with 0 NY NZ NZ OV SD SE SE TA ants’ nests and the only Yorkshire record 9 with collecting data indicates that it was 8 found in the nest of the ant Formica 7 lugubris.
6 5 4 3 2 1
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pygmaea Wasmann, 1894
NO RECORDS
HOMOEUSA Kraatz, 1856 acuminata (Märkel, 1842)
NO RECORDS
Subtribe MEOTICINA Seevers, 1978
MEOTICA Mulsant & Rey, 1873 anglica Benick in Muona, 1991
NO RECORDS
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exilis (Knoch in Gravenhorst, 1806)
Meotica exilis (Knoch in Grav enhorst, 1806)
3 WATSONIAN (3,0,1,2,1) There are seven records, from 2 YORKSHIRE five localities, on the YNU database: West Bretton (SE2815), 02/02/1949 (EWA); 1 Spurn (TA4212), 20/08/1949 (WOS), in 0 NY NZ NZ OV SD SE SE TA reed debris; Thorganby Ings (SE6942), 9 09/05/1971 and 09/06/1071 (EWA teste 8 CJ); Askham Bog (SE5648), 23/06/1996 7 (RGB & JAO) and 10/04/1999 (RJM), sieved from ground litter and moss; Hall 6 Garth Ponds, Ripon (SE3174), 09/09/2002 5 (RJM), sieved from litter in reed bed. 4 3 2 1
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exillima Sharp, 1915
Meotica exillima Sharp, 1915
3 WATSONIAN (2,1,3,1,0) Despite the national status of 2 YORKSHIRE ‘Local’ there are only records from six localities on the YNU database. The 1 species is associated with moss in damp 0 NY NZ NZ OV SD SE SE TA places. The only Yorkshire records with 9 collecting data show they have been 8 found in ground litter and Carex litter. 7 6 5 4 3 2 1
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filiformis (Motschulsky, 1860)
Meotica filiformis (Motschulsky, 1860)
3 WATSONIAN (0,2,9,2,0) Widely distributed in the 2 YORKSHIRE county. The lack of records from VC61 and VC65 may indicate under-recording. The 1 only Yorkshire records with collecting data 0 NY NZ NZ OV SD SE SE TA show they have been found in Carex 9 paniculata litter. 8 7 6 5 4 3 2 1
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pallens (Redtenbacher, 1849)
NO RECORDS
Subtribe OXYPODINA Thomson, C.G., 1859
AMAROCHARA Thomson, C.G., 1858 bonnairei (Fauvel, 1865)
NO RECORDS forticornis (Lacordaire, 1835)
NO RECORDS
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umbrosa (Erichson, 1837)
Amarochara umbrosa (Erichson, 1837)
3 WATSONIAN (0,2,1,0,0) There are three records on the 2 YORKSHIRE YNU database: Saltburn (NZ6721), 1896 (MLT); Netherby Dale (SE9084), 1 28/05/1988 (WAE det. CJ); Sheffield 0 NY NZ NZ OV SD SE SE TA (SK3689), 27/07/2001 (EJS det. MLD). This 9 is a Nationally Scarce species which is 8 associated with the subterranean runs of 7 small mammals.
6 5 4 3 2 1
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CALODERA Mannerheim, 1830
aethiops (Gravenhorst, 1802)
Calodera aethiops (Grav enhorst, 1802)
3 WATSONIAN (19,3,16,6,2) Widely distributed in Yorkshire, especially in the southern and 2 YORKSHIRE central area of the county. The species is 1 usually found on the ground amongst low 0 NY NZ NZ OV SD SE SE TA vegetation and plant debris. Yorkshire 9 specimens have been located in Typha 8 beds, Juncus litter, flood refuse, leaf litter, old grass mowings, and pitfall traps. 7
6 5 4 3 2 1
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nigrita Mannerheim, 1830
Calodera nigrita Mannerheim, 1830
3 WATSONIAN (2,2,3,1,1) With records from only eight 2 YORKSHIRE Yorkshire localities this species lives up to its status of Nationally Scarce. Rather 1 worryingly, however, is the fact that there 0 NY NZ NZ OV SD SE SE TA is only a single post-2000 record. The 9 species is confined to England where it is 8 found in wetlands and marshy habitats. In 7 Yorkshire specimens have been located in flood refuse, grass tussocks, leaf litter, and 6 in damp vegetable refuse. 5 4 3 2 1
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protensa Mannerheim, 1830
Calodera protensa Mannerheim, 1830
3 WATSONIAN (0,0,0,2,0) There are two records on the 2 YORKSHIRE YNU database, both from a wet meadow at Cawood (SE5737): 05/1990 (KGP det. 1 CJ), a single female in a pitfall trap; 0 NY NZ NZ OV SD SE SE TA 05/1992 (KGP det. MLD), a single male in a 9 pitfall trap. Confined to wet habitats, this 8 species has the national status of RDB I as 7 it has only been recorded from Essex, Oxfordshire and Yorkshire. The Yorkshire 6 records are the first since the 1920s, when 5 the species was recorded from Yarnton, 4 Oxfordshire. 3 2 1
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riparia Erichson, 1837
Calodera riparia Erichson, 1837
3 WATSONIAN (2,2,12,3,0) What few Yorkshire records 2 YORKSHIRE there are of this Nationally Scarce species are mainly from the southern half of the 1 county. The species is associated with wet 0 NY NZ NZ OV SD SE SE TA habitats, including marshes, fens and river 9 shingle. 8 7 6 5 4 3 2 1
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rubens (Erichson, 1837)
NO RECORDS
rufescens Kraatz, 1856
NO RECORDS
uliginosa Erichson, 1837
NO RECORDS
COUSYA Mulsant & Rey, 1875 320 defecta Mulsant & Rey, 1875
NO RECORDS
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longitarsis (Thomson, C.G., 1867)
Cousya longitarsis (Thomson, C.G., 1867)
3 WATSONIAN (0,12,0,0,0) There are 12 records from five 2 YORKSHIRE localities on the YNU database, all from the North York Moors: Danby Low Moor 1 (NZ7111), 05, 06, 08, 09 & 10/1987 and 08 0 NY NZ NZ OV SD SE SE TA & 10/1988 (all YU det. MLD); Egton High 9 Moor (NZ7702), 03/09/1987 (YU det. 8 RJM); Levisham Moor (SE8391), 7 03/09/1987 (YU det. MLD); Kildale Moor (NZ6211), 07/07/1987 & 14/09/1987 (YU 6 det. RJM); Spaunton Moor (SE7293), 5 05/10/1987 (YU det. MLD). This Nationally 4 Scarce species is restricted to north Wales, 3 northern England and northern Scotland 2 where it is found at the tops of mountains and on high moorland. 1
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nigrata (Fairmaire & Laboulbène, 1856)
NO RECORDS
nitidiventris (Fagel, 1958)
Cousya nitidiv entris (Fagel, 1958)
3 WATSONIAN (0,0,1,0,0) There is a single record on the 2 YORKSHIRE YNU database: Ringinglow (SK2984), 06/11/1964 (RCl & ROSC det. SAW). The 1 single specimen added the species to the 0 NY NZ NZ OV SD SE SE TA British list (Proc. Br. Ent. Nat. Hist. Soc. 12: 9 46-48). This is a little known species which 8 has only been recorded from six English 7 vice counties, the record above being the most northerly. It is afforded the national 6 status pRDBK. 5 4 3 2 1
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CRATARAEA Thomson, C.G., 1858 suturalis (Mannerheim, 1830)
Crataraea suturalis (Mannerheim, 1830)
3 WATSONIAN (1,1,5,4,1) This distinct species, although 2 YORKSHIRE widely distributed in Yorkshire, appears to be genuinely rare. Yorkshire specimens 1 have been found in old straw, in litter 0 NY NZ NZ OV SD SE SE TA under Alders in a swamp, and in a grass 9 heap. 8 7 6 5 4 3 2 1
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DEXIOGYIA Thomson, C.G., 1858 corticina (Erichson, 1837)
NO RECORDS
HAPLOGLOSSA Kraatz, 1856 gentilis (Märkel, 1844)
Haploglossa gentilis (Märkel, 1844)
3 WATSONIAN (1,0,0,0,0) There is a single record on the 2 YORKSHIRE YNU database: Spurn (TA4212), 08/06/1950 (WOS), in stable refuse from a 1 barn which was thought to have housed 0 NY NZ NZ OV SD SE SE TA an owl’s nest. This species has been 9 recorded from owls’ nests and also the 8 nests of the ant Lasius fuliginosus. 7 6 5 4 3 2 1
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marginalis (Gravenhorst, 1806)
NO RECORDS
nidicola (Fairmaire, 1852)
Haploglossa nidicola (Fairmaire, 1852)
3 WATSONIAN (2,1,2,2,1) The species is known from eight 2 YORKSHIRE widely distributed localities. Its limited distribution is due to the fact that it is 1 associated with the nests of Sand Martins. 0 NY NZ NZ OV SD SE SE TA 9 8 7 6 5 4 3 2 1
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picipennis (Gyllenhal, 1827)
NO RECORDS
villosula (Stephens, 1832)
Haploglossa v illosula (Stephens, 1832)
3 WATSONIAN (0,9,1,2,1) There are records from 13 2 YORKSHIRE widely scattered Yorkshire localities. This species is found in passerine birds’ nests, 1 and it also occurs on flowers. With the 0 NY NZ NZ OV SD SE SE TA exception of a record of a single in a 9 Robin’s nest, all other Yorkshire records 8 with collecting data refer to individuals 7 which have been swept from vegetation.
6 5 4 3 2 1
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HYGROPORA Kraatz, 1856 cunctans (Erichson, 1837)
NO RECORDS
ILYOBATES Kraatz, 1856 bennetti Donisthorpe, 1914
Ilyobates bennetti Donisthorpe, 1914
3 WATSONIAN (4,0,2,2,0) There are eight authentic 2 YORKSHIRE Yorkshire records from five localities: Wheldrake Ings (SE7044), 15/05/1979 1 (CJ); Bubwith Bridge (SE7036), 0 NY NZ NZ OV SD SE SE TA 24/04/1985 (PK det. RJM), 13/05/1986 (PK 9 det. MLD) and 11/08/1993 (MLD), in flood 8 refuse; Shirley Pool (SE5612), 19/07/1986 7 (RJM teste CJ); Roche Abbey (SK5489), 06/05/2000 (EJS teste MLD); Hopewell 6 Farm (SE3758), 11 and 24/06/2000 (MLD), 5 in pitfall traps. This Nationally Scarce 4 species appears to be restricted to 3 England (although past confusion with I. 2 nigricollis (Paykull, 1800) clouds the known distribution). 1
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nigricollis (Paykull, 1800)
Ilyobates nigricollis (Paykull, 1800)
3 WATSONIAN (0,0,2,1,0) There are three authentic 2 YORKSHIRE records on the YNU database: Askham Bog (SE5648), 03/05/1965 (EWA & CJ); Little 1 Don Valley (SK1899), date unknown (EJS); 0 NY NZ NZ OV SD SE SE TA Langsett (SE2101), 17/06/1990 (EJS det. 9 CJ). The records from Scarborough 8 (TA0488) on an unknown date (TW) and 7 Roundhay Park (SE3338) on 01/05/1910 (EWM) are probably referable to I. 6 bennetti Donisthorpe, 1914. Being 5 restricted to very few counties, the 4 species is afforded the national status 3 pRDBK. 2
1
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propinquus (Aubé, 1850)
Ilyobates propinquus (Aubé, 1850)
3 WATSONIAN (2,0,5,0,0) There are records from three 2 YORKSHIRE localities on the YNU database: Manor Farm, Eddlethorpe (SE7766), 2000-01 1 (RGL det. MLD), in pitfall traps; Rossington 0 NY NZ NZ OV SD SE SE TA Bridge (SK6399), 05/1985 (RJM), in litter 9 under Carex paniculata; Walton NR, 8 Wakefield (SE3618), 27/4/2011- 7 18/06/2011 (MLD), numerous specimens in pitfall traps along a pond edge. This 6 Nationally Scarce species is generally 5 found in sandy places, including river 4 banks and sand pits, and is predominantly 3 southern in distribution, Yorkshire being 2 on the species’ northern limit.
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ISCHNOGLOSSA Kraatz, 1856 obscura Wunderle, 1990
NO RECORDS prolixa (Gravenhorst, 1802)
Ischnoglossa prolixa (Grav enhorst, 1802)
3 WATSONIAN (0,0,10,2,0) The realisation that two 2 YORKSHIRE species were masquerading under this name (Entomologist’s Record and Journal 1 of Variation 106: 241-244) has cast doubt 0 NY NZ NZ OV SD SE SE TA on previous records. The map, therefore, 9 only includes extant specimens which 8 have been re-examined and more recent 7 captures. The species is usually found under bark. 6
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turcica Wunderle, 1992
Ischnoglossa turcica Wunderle, 1992
3 WATSONIAN (0,0,1,0,0) There is a single record on the 2 YORKSHIRE YNU database: Ringinglow (SK2883), 07/04/1991 (EJS det. CJ), a single female 1 under bark. This species was described as 0 NY NZ NZ OV SD SE SE TA new to science by Wunderle (1992) and 9 new to Britain by Owen (1994). The few 8 records which have accumulated since this 7 time suggest that it is predominantly southern in distribution, the Yorkshire 6 record above being the furthest north. 5 4 3 2 1
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MNIUSA Mulsant & Rey, 1875 incrassata (Mulsant & Rey, 1852)
Mniusa incrassata (Mulsant & Rey, 1852)
3 WATSONIAN (1,8,20,9,5) Widely distributed in the 2 YORKSHIRE county but, rather surprisingly, there is a lack of records from most of VC61. It is also 1 rather worrying that most records are pre- 0 NY NZ NZ OV SD SE SE TA 2000. The species is found among dead 9 leaves etc. In Yorkshire specimens have 8 been located in flood refuse, pitfall traps, 7 leaf litter, grass traps, moss, and vegetable refuse. 6
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OCALEA Erichson, 1837 badia Erichson, 1837
Ocalea badia Erichson, 1837
3 WATSONIAN (0,3,1,1,0) There are five records on the YNU database: Scarborough (TA0488), 2 YORKSHIRE date unknown (CES); Forge Valley 1 (SE9886), 09/1926 (recorder unknown); 0 NY NZ NZ OV SD SE SE TA Tadcaster (SE4843), 09/10/1969 (EWA 9 teste CJ); Upper Windleden Reservoir 8 (SE1501), 05/04/1985 (DM det. MLD teste CJ); Raincliffe Woods (SE9988), 7 08/01/1986 (MLD det. CJ). 6 5 4 3 2 1
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latipennis Sharp, 1870
Ocalea latipennis Sharp, 1870
3 WATSONIAN (0,0,1,1,0) There are two authentic records on the YNU database: Langsett 2 YORKSHIRE (SE1900), 07/04/1985 (DM det. MLD teste 1 CJ); Willow Garth (SE3173), 13/08/2002 0 NY NZ NZ OV SD SE SE TA (RJM det. MLD), in flood refuse. The 9 species is found in sand and shingle on the 8 banks of rivers. 7 Due to past confusion with O. rivularis 6 Miller, L., 1851 the following records are 5 suspect without voucher specimens: 4 Scarborough (TA0488), date unknown 3 (WCH); Raincliffe Woods (SE9988), date 2 unknown (GBW); Bingley (SE1139), 10/1908 (recorder unknown). 1 SD SE SE TA 0 SJ SK SK TF 9 8 post 2000 Map produced for Yorkshire Naturalists' Union by DMAP 29/1/2019 pre 2000 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
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picata (Stephens, 1832)
Ocalea picata (Stephens, 1832)
3 WATSONIAN (3,23,53,19,8) Widely distributed and 2 YORKSHIRE common in the county, although records in VC61 are limited to the western 1 extremity. The species is typically found in 0 NY NZ NZ OV SD SE SE TA damp places in moss etc. Yorkshire 9 specimens have been found in Carex, 8 Phragmites, Glyceria maxima and leaf 7 litter, flood refuse, under the bark of old logs, in decaying fungi, and in pitfall traps. 6
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rivularis Miller, L., 1851
NO RECORDS
OCYUSA Kraatz, 1856 maura (Erichson, 1837)
Ocyusa maura (Erichson, 1837)
3 WATSONIAN (11,2,75,10,1) This species of marshy 2 YORKSHIRE habitats, although widely distributed, is 1 commonest in the southern half of the
NY NZ NZ OV county. 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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picina (Aubé, 1850)
Ocyusa picina (Aubé, 1850)
3 WATSONIAN (1,1,5,3,0) There are Yorkshire records 2 YORKSHIRE from only seven widely distributed 1 localities, only two of which are post-
NY NZ NZ OV 2000, indicating that this is but a rare 0 SD SE SE TA 9 species in the county. 8 7 6 5 4 3 2 1
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OXYPODA Mannerheim, 1830 acuminata (Stephens, 1832)
Oxypoda acuminata (Stephens, 1832)
3 WATSONIAN (5,7,24,4,3) Widely distributed in the 2 YORKSHIRE county but, rather worryingly, is the fact that most records are pre-2000. The 1 species is usually found in vegetable 0 NY NZ NZ OV SD SE SE TA refuse. The only Yorkshire record with 9 collecting data refers to specimens from 8 stable refuse. 7 6 5 4 3 2 1
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alternans (Gravenhorst, 1802)
Oxypoda alternans (Grav enhorst, 1802)
3 WATSONIAN (8,23,104,20,3) Widely distributed and 2 YORKSHIRE common, especially in the southern half of the county. The species is generally found 1 in decaying fungi, vegetable refuse and 0 NY NZ NZ OV SD SE SE TA dung. Yorkshire specimens have been 9 located in a variety of fungi (including 8 Polyporus sulphureus, Piptoporus 7 betulinus, Pleurotus ostreatus, Grifola frondosa and Russula nigricans), in grass 6 piles, old hay debris, flood refuse, moss, 5 and in water and pitfall traps. 4 3 2 1
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annularis Mannerheim, 1830
Oxypoda annularis Mannerheim, 1830
3 WATSONIAN (2,6,22,5,0) Widely distributed in the 2 YORKSHIRE county but, rather worryingly, is the fact that most records are pre-2000. The 1 species is found in any decaying organic 0 NY NZ NZ OV SD SE SE TA matter. The few Yorkshire records with 9 collecting data indicate they have been 8 located in a garden compost heap, in 7 stable refuse, in a pile of rotting grass, and along the edge of a saltmarsh. 6
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brachyptera (Stephens, 1832)
Oxypoda brachyptera (Stephens, 1832)
3 WATSONIAN (9,0,3,1,0) Very locally distributed in Yorkshire, with records from only ten 2 YORKSHIRE centrally placed localities. The few 1 Yorkshire records with collecting data 0 NY NZ NZ OV SD SE SE TA indicate they have been found in Glyceria 9 and Carex litter and in flood refuse along 8 the strand line. 7 6 5 4 3 2 1
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brevicornis (Stephens, 1832)
Oxypoda brev icornis (Stephens, 1832)
3 WATSONIAN (28,21,79,12,4) Common and widely 2 YORKSHIRE distributed in the county. The species is usually found in vegetable refuse. 1 Yorkshire specimens have been found in 0 NY NZ NZ OV SD SE SE TA flood refuse, old hay debris, old fungi, a 9 garden compost heap, by sweeping 8 vegetation, and in Phragmites, Typha and 7 Iris pseudacorus litter.
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carbonaria (Heer, 1841)
Oxypoda carbonaria (Heer, 1841)
3 WATSONIAN (2,5,18,2,0) Widely distributed but rarely 2 YORKSHIRE located. The cluster of records in the western half of VC63 may simply reflect 1 recorder effort. The few Yorkshire records 0 NY NZ NZ OV SD SE SE TA with collecting data indicate they have 9 been located in vegetable refuse, in a pile 8 of rotting turnips in a garden, and at the 7 base of coastal sand dunes.
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elongatula Aubé, 1850
Oxypoda elongatula Aubé, 1850
3 WATSONIAN (26,18,258,46,7) Widely distributed and 2 YORKSHIRE common, especially in the southern half of the county. Yorkshire specimens have 1 been collected from a wide variety of 0 NY NZ NZ OV SD SE SE TA habitats, including vegetable, stable, flood 9 and old hay refuse; decaying fungi; in wet 8 moss; and Phragmites, Typha, Carex and 7 Glyceria litter.
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exoleta Erichson, 1839
Oxypoda exoleta Erichson, 1839
3 WATSONIAN (6,1,4,0,0) This Nationally Scarce species 2 YORKSHIRE which is usually found in sandy and heathy habitats has been recorded from nine 1 Yorkshire localities, all in the southern half 0 NY NZ NZ OV SD SE SE TA of the county. Yorkshire specimens have 9 been located in ground litter, in damp 8 sand under moss, in a garden compost 7 heap, and in pitfall traps set along a pond edge. 6
5 Past confusion with other members of the 4 genus renders the records from Spurn 3 (TA4212) in 1950, 1951 and 1952 (all WOS) 2 unreliable without voucher specimens.
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ferruginea Erichson, 1839
NO RECORDS
flavicornis Kraatz, 1856
Oxypoda flav icornis Kraatz, 1856
3 WATSONIAN (3,0,10,1,0) There are Yorkshire records from 11 localities, all in the southern half 2 YORKSHIRE of the county. This is a Nationally Scarce 1 species which is found throughout 0 NY NZ NZ OV SD SE SE TA England and also south-east Scotland. The 9 few Yorkshire records with collecting data 8 indicate they have been located in decaying straw and sieved from leaf litter. 7
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formiceticola Märkel, 1841
Oxypoda formiceticola Märkel, 1841
3 WATSONIAN (0,2,14,0,0) With the exception of two records in VC62 (Harewood Dale in 1924 2 YORKSHIRE and Hellwath Beck in 1936) all other 1 records are from VC63. As this is an ants’ 0 NY NZ NZ OV SD SE SE TA nest associate, however, it is possible that 9 the species may be under-recorded. All 8 the Yorkshire records with collecting data show they had originated from the nests 7 of the ant Formica lugubris. 6 5 4 3 2 1
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haemorrhoa (Mannerheim, 1830)
Oxypoda haemorrhoa (Mannerheim, 1830)
3 WATSONIAN (28,15,155,18,9) A widely distributed and common species in the county. The lack of 2 YORKSHIRE records in the northern half of VC62 and 1 the western halves of VC64 and VC65 may 0 NY NZ NZ OV SD SE SE TA simply be a reflection of under-recording. 9 The species occurs in decaying fungi, 8 vegetable refuse etc. Yorkshire specimens have been collected from a wide variety of 7 habitats, including all types of decaying 6 vegetation, horse dung, by sweeping 5 vegetation, in ground litter under Alnus 4 and conifers, in Phragmites and Carex 3 litter, under bark, and from the nests of the ant Formica lugubris. 2
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induta Mulsant & Rey, 1861
Oxypoda induta Mulsant & Rey, 1861
3 WATSONIAN (1,2,1,0,0) There are four records on the 2 YORKSHIRE YNU database: Whitby (NZ8911), 1935 (HB); Skelder (NZ8409), 15/04/1935 (HB); 1 Brook Farm, Elstronwick (TA2332), 2/1999 0 NY NZ NZ OV SD SE SE TA (WRD det. MLD); Whiteley Wood 9 (SK3185), 22/10/1999 (EJS teste MLD). 8 7 6 5 4 3 2 1
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islandica Kraatz, 1857
Oxypoda islandica Kraatz, 1857
3 WATSONIAN (0,0,0,3,0) There are three records, from two localities, of this Nationally Scarce 2 YORKSHIRE species on the YNU database: Malham 1 Tarn (SD9867), 08/1955 and 1957 (WOS), 0 NY NZ NZ OV SD SE SE TA in Larch litter; Penyghent (SD8373), 9 06/07/1967 (CJ). The species is confined to 8 moorland and mountains in northern England. 7
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lentula Erichson, 1837
Oxypoda lentula Erichson, 1837
3 WATSONIAN (15,1,7,0,0) With the exception of a single VC62 record (Saltburn in 1898) all other 2 YORKSHIRE Yorkshire records are from the south and 1 west of the county. Yorkshire specimens 0 NY NZ NZ OV SD SE SE TA have been found in flood refuse, reedbed 9 litter, a Glyceria bed, and in a pile of old 8 straw. 7 6 5 4 3 2 1
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longipes Mulsant & Rey, 1861
Oxypoda longipes Mulsant & Rey, 1861
3 WATSONIAN (2,0,1,0,0) There are three records, from two localities, on the YNU database: 2 YORKSHIRE Bubwith (SE7030), 1916 and 1918 (WJF); 1 Mag Wood, Huddersfield (SE1313), 0 NY NZ NZ OV SD SE SE TA 27/11/1983 (DM det. MLD teste CJ). 9 8 7 6 5 4 3 2 1
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lurida Wollaston, 1857
NO RECORDS mutata Sharp, 1871
NO RECORDS nigricornis Motschulsky, 1860
Oxypoda nigricornis Motschulsky, 1860
3 WATSONIAN (0,0,3,1,0) There are four records on the 2 YORKSHIRE YNU database: Malham Tarn (SD8967), 13/09/1967 (CJ); Brun Moor (SE0109), 1 14/09/1985 (DM det. MLD teste CJ); Drop 0 NY NZ NZ OV SD SE SE TA Clough (SE0413), 26/09/1987 (DM det. 9 MLD); Cartledge Flat (SK2093), 8 12/06/1988 (EJS). This is a Nationally 7 Scarce montane species which has been recorded from the Midlands, northern 6 England and north-east Scotland. 5 Yorkshire specimens have been located in 4 a rotting grass pile and sheep dung. 3 2 1
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NO RECORDS
opaca (Gravenhorst, 1802)
Oxypoda opaca (Grav enhorst, 1802)
3 WATSONIAN (18,11,71,18,1) A widely distributed and 2 YORKSHIRE common species, especially in the southern half of the county. The species is 1 found in any decaying organic matter. 0 NY NZ NZ OV SD SE SE TA Although the majority of Yorkshire records 9 show this affinity to decaying organic 8 matter, specimens have also been located 7 in horse and sheep dung and fungi under the bark of a poplar. 6
5 4 3 2 1
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praecox Erichson, 1839
NO RECORDS
procerula Mannerheim, 1830
Oxypoda procerula Mannerheim, 1830
3 WATSONIAN (1,3,26,6,3) Widely distributed but not common in the county. The species is 2 YORKSHIRE found in litter and moss in wet places. 1 Although most Yorkshire specimens have 0 NY NZ NZ OV SD SE SE TA been located in such habitats, they have 9 also been found in an old dung heap and 8 by sweeping lagoon-side vegetation.
7
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recondita Kraatz, 1856
NO RECORDS
soror Thomson, C.G., 1855
NO RECORDS
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spectabilis Märkel, 1844
Oxypoda spectabilis Märkel, 1844
3 WATSONIAN (1,7,2,2,0) There are Yorkshire records 2 YORKSHIRE from ten widely distributed localities. This Nationally Scarce species is found 1 throughout the British Isles in a wide 0 NY NZ NZ OV SD SE SE TA variety of habitats. 9 8 7 6 5 4 3 2 1
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tarda Sharp, 1871
NO RECORDS
tirolensis Gredler, 1863
NO RECORDS
vittata Märkel, 1842
Oxypoda v ittata Märkel, 1842
3 WATSONIAN (0,0,1,0,0) There is a single record on the YNU database: Limb Valley (SK3083), 2 YORKSHIRE 23/12/1987, (ASL teste EJS & CJ), a single 1 male in a Longworth mammal trap. The 0 NY NZ NZ OV SD SE SE TA species appears to be confined to England 9 and Wales, the record above being the 8 furthest north.
7 6 5 4 3 2 1
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PHLOEOPORA Erichson, 1837 concolor Kraatz, 1856
NO RECORDS
corticalis (Gravenhorst, 1802)
Phloeopora corticalis (Grav enhorst, 1802)
3 WATSONIAN (1,2,8,1,1) Widely distributed in the county, 2 YORKSHIRE but rare. This Nationally Scarce species is restricted to England where it is associated 1 with woodland, wood-edge and scrub. The 0 NY NZ NZ OV SD SE SE TA only Yorkshire specimens with collecting data 9 have been located under bark. 8 7 6 5 4 3 2 1
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nitidiventris Fauvel, 1904
NO RECORDS
scribae Eppelsheim, 1884
NO RECORDS
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testacea (Mannerheim, 1830)
Phloeopora testacea (Mannerheim, 1830)
3 WATSONIAN (3,5,29,5,0) This species, which is found under bark, is widely distributed, especially in 2 YORKSHIRE the southern half of the county. The lack of 1 records from the western and eastern parts 0 NY NZ NZ OV SD SE SE TA of the county may be due to under-recording. 9 The few Yorkshire records with collecting 8 data show they have been found under bark (both deciduous and evergreen), and in old 7 Piptoporus fungi. 6 5 4 3 2 1
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STICHOGLOSSA Fairmaire & Laboulbène, 1856 semirufa (Erichson, 1839)
NO RECORDS
TETRALAUCOPORA Bernhauer, 1928 337 longitarsis (Erichson, 1837)
Tetralaucopora longitarsis (Erichson, 1837)
3 WATSONIAN (11,11,33,6,5) This species, which lives 2 YORKSHIRE on mud at the side of water, especially rivers, is widely distributed in the 1 county. The lack of records from the 0 NY NZ NZ OV SD SE SE TA hilly areas in the west is intriguing, 9 however. The Yorkshire specimens 8 with collecting data indicate they have 7 been located in pitfall traps, in an old straw heap, in a Glyceria bed, under 6 stones, by sweeping vegetation, and in 5 dry seaweed. 4 3 2 1
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rubicunda (Erichson, 1837)
Tetralaucopora rubicunda (Erichson, 1837)
3 WATSONIAN (0,2,0,0,1) There are three records of this Nationally Scarce species on the 2 YORKSHIRE YNU database: Scarborough (TA0488), 1 date unknown (RL); Duncombe Park NY NZ NZ OV 0 SD SE SE TA (SE6083), 18/06/1983 (RJM), in river 9 shingle; Raydale Beck, Semerwater 8 (SD9287), 03/09/1985 (RSK det. MLD). The species is usually found on the 7 banks of rivers and streams. 6 5 4 3 2 1
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THIASOPHILA Kraatz, 1856 angulata (Erichson, 1837)
Thiasophila angulata (Erichson, 1837)
3 WATSONIAN (0,4,11,0,0) There are records from 14 2 YORKSHIRE localities on the YNU database. Interestingly, the records are 1 concentrated in the western half of VC63 0 NY NZ NZ OV SD SE SE TA and a cluster of four sites in eastern VC62. 9 The species is associated with ants’ nests, 8 particularly Formica rufa. With the 7 exception of an individual found under the bark of a deciduous tree, all the Yorkshire 6 records with collecting data show they 5 had originated from the nests of the ant 4 Formica lugubris. 3 2 1
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inquilina (Märkel, 1842)
NO RECORDS
Subtribe TACHYUSINA Thomson, C.G., 1859
BRACHYUSA Mulsant & Rey, 1874 concolor (Erichson, 1839)
NO RECORDS
DACRILA Mulsant & Rey, 1874 fallax (Kraatz, 1856)
Dacrila fallax (Kraatz, 1856)
3 WATSONIAN (5,0,5,0,0) This Nationally Scarce species is known from ten Yorkshire localities, all in 2 YORKSHIRE VC63 and the southern half of VC62. It is 1 confined to wetlands and mainly occurs in 0 NY NZ NZ OV SD SE SE TA southern Britain, Yorkshire being at its 9 northern limit. Yorkshire specimens have 8 been swept from vegetation near a pond, but most have been located in ground 7 litter, including that of a Phragmites bed 6 and in a saltmarsh. 5 4 3 2 1
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pruinosa (Kraatz, 1856)
NO RECORDS
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DASYGNYPETA Lohse, 1974 velata (Erichson, 1837)
Dasygnypeta v elata (Erichson, 1837)
3 WATSONIAN (2,0,0,0,0) There are two records of this 2 YORKSHIRE Nationally Scarce species on the YNU database, both from Wheldrake Ings 1 (SE7044): 29/07/1992 and 22/08/1992 0 NY NZ NZ OV SD SE SE TA (MLD det. CJ). The species is associated 9 with watery habitats and confined to 8 southern Britain, Yorkshire being on its 7 northern limit.
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GNYPETA Thomson, C.G., 1858 caerulea (Sahlberg, C.R., 1831)
Gnypeta caerulea (Sahlberg, C.R., 1831)
3 WATSONIAN (0,0,0,5,4) The nine Yorkshire records show this species to be confined to the 2 YORKSHIRE western halves of VC64 and VC65. It is a 1 Nationally Scarce species and is confined 0 NY NZ NZ OV SD SE SE TA to the margins of rivers. The few 9 Yorkshire records with collecting data are 8 from moss at the sides of streams or under waterfalls. 7
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carbonaria (Mannerheim, 1830)
Gnypeta carbonaria (Mannerheim, 1830)
3 WATSONIAN (8,6,20,4,3) Even when excluding the early th 2 YORKSHIRE 20 century records (which may have included G. rubrior Tottenham, 1939), the 1 species is widely distributed in the county. 0 NY NZ NZ OV SD SE SE TA The species is found on the muddy banks 9 of rivers and ponds. Yorkshire specimens 8 have been located along the edges of 7 ponds, in vegetable refuse near water, in wet trickles on sea cliffs, in reedbed litter, 6 and in fungi. 5 4 3 2 1
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ripicola (Kiesenwetter, 1844)
Gnypeta ripicola (Kiesenwetter, 1844)
3 WATSONIAN (0,1,3,0,0) There are four records of this Nationally Scarce species on the YNU 2 YORKSHIRE database: Margery Wood (SE2709), 1 12/07/1989 (EJS teste CJ); Duncombe Park 0 NY NZ NZ OV SD SE SE TA (SE6083), 26/06/1993 (RJM teste CJ); 9 Damflask (SK2790), 15/04/1995 (EJS teste 8 MLD); Pebley Pond, Barlborough (SK4978), 18/03/2000 (RM det. EJS). This 7 species is confined to habitats near water, 6 particularly the banks of rivers, streams 5 and ponds. 4 3 2 1
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rubrior Tottenham, 1939
Gnypeta rubrior Tottenham, 1939
3 WATSONIAN (3,2,14,0,0) Recorded from 16 Yorkshire 2 YORKSHIRE localities, mainly in VC63, but also on or near the coast. It is found on the muddy 1 banks of rivers. The few Yorkshire records 0 NY NZ NZ OV SD SE SE TA with collecting data reveal that it has been 9 found in ground litter in a reed bed, on the 8 mud in a saltmarsh, and along the muddy 7 margin of a brackish pond.
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ISCHNOPODA Stephens, 1835 leucopus (Marsham, 1802)
Ischnopoda leucopus (Marsham, 1802)
3 WATSONIAN (1,9,5,2,0) Widely, but thinly, distributed in the county. Given the species’ habitat 2 YORKSHIRE requirements of wet silt, sand and shingle 1 at the side of clean moving water, it is 0 NY NZ NZ OV SD SE SE TA surprising that there are no records from 9 the eastern half of VC61 and western 8 halves of VC64 or VC65.
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scitula (Erichson, 1837)
NO RECORDS
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umbratica (Erichson, 1837)
Ischnopoda umbratica (Erichson, 1837)
3 WATSONIAN (0,2,1,0,0) There are three records on the 2 YORKSHIRE YNU database: Ravenscar (NZ9801), date and recorder unknown; Redcar (NZ6024), 1 13/10/1913 (WJF); Wheatley (SE5905), 0 NY NZ NZ OV SD SE SE TA 15/05-02/07/2008 (MGT det. RJM). 9 8 7 6 5 4 3 2 1
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TACHYUSA Erichson, 1837 concinna Heer, 1839
Tachyusa concinna Heer, 1839
3 WATSONIAN (0,1,1,0,0) There are two records of this Nationally Scarce species on the YNU 2 YORKSHIRE database: Green Wood, Denby Dale 1 (SE2108), 15/05/1948 (EWA det. HB teste 0 NY NZ NZ OV SD SE SE TA MLD); Cayton Bay (TA0785), 13/07/2006 9 (MGT det. RJM teste MLD). This species is 8 found in reed beds and river and lake margins in southern Britain, Yorkshire 7 being its northern limit. 6 5 4 3 2 1
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constricta Erichson, 1837
Tachyusa constricta (Erichson, 1837)
3 WATSONIAN (0,6,1,6,0) Widely scattered but not 2 YORKSHIRE common in the county. The few Yorkshire records with collecting data show they 1 have been located on damp mud and in 0 NY NZ NZ OV SD SE SE TA shingle along river banks. 9 8 7 6 5 4 3 2 1
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objecta (Mulsant & Rey, 1870)
NO RECORDS
THINONOMA Thomson, C.G., 1859 340 atra (Gravenhorst, 1806)
Thinonoma atra (Grav enhorst, 1806)
3 WATSONIAN (18,4,40,6,1) Widely distributed in 2 YORKSHIRE Yorkshire, especially in the southern half of the county. The species is found on 1 sandy riverbanks, where it can be seen 0 NY NZ NZ OV SD SE SE TA running rapidly over bare sand in sunny 9 conditions. Yorkshire specimens have 8 been located in a wide variety of wet 7 habitats, including Typha and Phragmites beds, but they have also been swept from 6 vegetation in a woodland clearing. 5 4 3 2 1
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Tribe PHYTOSINI Thomson, C.G., 1867
ARENA Fauvel, 1862 tabida (Kiesenwetter, 1850)
Arena tabida (Kiesenwetter, 1850)
3 WATSONIAN (1,0,0,0,0) There is a single record on the 2 YORKSHIRE YNU database: Spurn (TA3910), 21/07/1963 (RCW & WOS), under 1 driftwood on the shore. This is a coastal 0 NY NZ NZ OV SD SE SE TA species which appears to be confined to 9 the English and Welsh coasts. It is afforded 8 the national status pRDBK. 7 6 5 4 3 2 1
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PHYTOSUS Curtis, 1838 balticus Kraatz, 1859
Phytosus balticus Kraatz, 1859
3 WATSONIAN (2,4,0,0,0) There are six records, from four localities, on the YNU database: 2 YORKSHIRE Scarborough (TA0488), 1880 (WCH) and 1 1909 (recorder unknown); Eston 0 NY NZ NZ OV SD SE SE TA (NZ5919), 1909 (recorder unknown); 9 Saltburn (NZ6721), 1909 (recorder 8 unknown); Spurn (TA4212), 07/1952 7 (WOS) and 23/07/1963 (RCW), under tidal refuse on the Humber shore. This species 6 is generally found under felts of green 5 algae on the seashore. 4 3 2 1
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nigriventris (Chevrolat, 1843)
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spinifer Curtis, 1838
Phytosus spinifer Curtis, 1838
3 WATSONIAN (1,2,0,0,0) There are three records on the 2 YORKSHIRE YNU database: Scarborough (TA0488), 31/08/1899 (recorder unknown); Saltburn 1 (NZ6721), 09/1902 (MLT), common in 0 NY NZ NZ OV SD SE SE TA dead gulls on the coast; Spurn (TA4212), 9 14/07/1952 (WOS). 8 7 6 5 4 3 2 1
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Tribe PLACUSINI Mulsant & Rey, 1871
PLACUSA Erichson, 1837 complanata Erichson, 1839
NO RECORDS
depressa Mäklin, 1845
Placusa depressa Mäklin, 1845
3 WATSONIAN (0,1,3,0,0) There are four records of this Nationally Scarce species on the YNU 2 YORKSHIRE database: Stocksmoor Common (SE2714), 1 16/09/1971 (EWA teste MLD); Wharncliffe 0 NY NZ NZ OV SD SE SE TA Wood (SK2996), 01/11/1987 (EJS teste CJ); 9 Langsett Quarry (SE2000), 11/03/1990 8 (EJS); Rawcliffe Meadows (SE5754), 07/06/1997 (MLD), a single under pine 7 bark. This is predominantly a northern 6 species, although there are records from 5 south-east England. 4 3 2 1
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pumilio (Gravenhorst, 1802)
NO RECORDS
tachyporoides (Waltl, 1838)
Placusa tachyporoides (Waltl, 1838)
3 WATSONIAN (0,0,0,1,0) There is a single record of this 2 YORKSHIRE Nationally Scarce species on the YNU database: Bishop Wood (SE5634), 1 26/10/1985 (PK det. RJM teste PH). The 0 NY NZ NZ OV SD SE SE TA species is restricted to England where it is 9 found under bark in deciduous woodland 8 and pasture-woodland. 7 6 5 4 3 2 1
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UNCONFIRMED, DOUBTFUL AND ERRONEOUSLY RECORDED SPECIES
In the early 20th century it was not fully appreciated that this taxonomically difficult group required genitalic dissection for positive identification. As a consequence, there are a good number of records on the YNU database which, without specimens for dissection and confirmation, cannot be included on the Yorkshire List. The following species have been dismissed for the reasons outlined below.
Aleochara (Aleochara) lata Gravenhorst, 1802 The records of this species from Cat Babbleton (SE9974), 22/06/1985 (WAE); Rainsborough Park (SK3999), 04/07/1985 (WAE); Sandbeck Park (SK5691), 04/07/1991 (WAE), documented as new for Yorkshire (Naturalist 111: 96), have proved to be A. curtula (Goeze, 1777). Although corrected (Naturalist 113: 151), the earlier record was, unfortunately, perpetuated (Coleopterist 6: 36). As there are no further records the species is removed from the Yorkshire list.
Aleochara (Xenochara) moesta Gravenhorst, 1802 There are five unconfirmed/erroneous records on the YNU database: Byland (SE5579), 14/09/1892 (MLT); Sunnydale, near Bradford (SE1043), 1916 (JA); Holme on Spalding Moor (SE8138), 17/05/1924 (TS); Arnecliffe Wood (NZ4601), 09/1925 (MLT); Adel Dam (SE2741), 17/11/1963 (JHF). Past confusion with nomenclature renders the first four records unreliable without voucher specimens. The Adel specimen (in Leeds City Museum) has been re-identified as A. lanuginosa Gravenhorst, 1802 (det. MLD). The species has not been added to the Yorkshire list.
Alaobia hybrida (Sharp, 1869) There are two unconfirmed records on the YNU database: Sleights (NZ8707), date unknown (HB); East Row (NZ8616), 19/03/1936 (HB). Past confusion with other members of the genus renders these records unreliable without voucher specimens. The species has not been added to the Yorkshire list.
Alevonota rufotestacea (Kraatz, 1856) There are two unconfirmed records on the YNU database: Studley (SE2870), date unknown (EAW); Saltburn Wood (NZ6720), 05/1905 (MLT). Past confusion with other members of the genus renders these records unreliable without voucher specimens. The species has not been added to the Yorkshire list.
Microdota inquinula (Gravenhorst, 1802) There is a single unconfirmed record on the YNU database: Skelder (NZ8409), 20/03/1936 (HB). Past confusion with nomenclature renders this record unreliable without a voucher specimen. The species has not been added to the Yorkshire list.
Mocyta gilvicollis (Scheerpeltz, 1949) There are three records on the YNU database: Thorne Moor (SE7315), 15/03/1969 (CJ); Wydale Hall (SE9283), 09/04/1984 (LA det. MLD); Runswick Bay (NZ2817), 09/06/1990 (MLD det. CJ). There is much confusion regarding the identification of species within the Mocyta group and, as there are no published British records of this species, it has not been included in the most recent checklist (Duff, 2012).
Schistoglossa aubei (Brisout de Barneville, 1860) Specimens from Bubwith (SE7030) in 02/1919 (JHD), originally standing as this species on the YNU database, are extant in the Carlisle Museum and have proved to be Philhygra melanocera (Thomson, C.G., 1856) (Naturalist 125: 75). The species has not been added to the Yorkshire list.
Gyrophaena congrua Erichson, 1837 There are four unconfirmed records on the YNU database: Saltburn Wood (NZ6720), 1895 (recorder unknown); Fylingdales Moor (SE99), 24/09/1930 (WJF); West Bretton (SE2914), 07/10/1948 (EWA); Buttercrambe (SE7358), 06/08/1951 (AS). Past confusion with other members of the genus renders these records unreliable without voucher specimens. The species has not been added to the Yorkshire list.
Gyrophaena lucidula Erichson, 1837 There are two unconfirmed records on the YNU database: Scarborough (TA0488), date and recorder unknown; Saltburn (NZ6721), 06/1896 (recorder unknown). Past confusion with other members of the genus renders these records unreliable without voucher specimens. The species has not been added to the Yorkshire list.
Myllaena graeca Kraatz, 1858 The only British record involved two specimens which were found at Scarborough (TA0488) in the late 19th century (TW). This species has a Mediterranean distribution and it is doubtful that its occurrence in the British Isles is anything more than “adventitious” (Entomologist’s Monthly Magazine 104: 183-187). Even though one of the specimens is extant in the Matthews Collection at Bolton Museum, the species is not included in the latest beetle checklist (Duff, 2012).
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Dinarda dentata (Gravenhorst, 1806) Past confusion with the commoner D. maerkeli Kiesenwetter, 1843 renders the following record suspect without voucher specimens: Hellwath Beck (SE9599), 26/09/1937 (HB). This record was stated to be the only authentic Yorkshire record in Denton (2003) and the opportunity to correct this error is hereby taken. The species has not been added to the Yorkshire list.
Meotica anglica Benick in Muona, 1991 There is a single unconfirmed record on the YNU database: Scarborough (TA0488), date unknown (RL). Past confusion with other members of the genus renders this record unreliable without a voucher specimen. The species has not been added to the Yorkshire list.
Calodera uliginosa Erichson, 1837 Specimens collected at West Bretton (SE2914) on 23/03/1948 were originally described as this species (Naturalist 74: 151). The specimens are in the Manchester University Museum collection and have been re-identified as C. riparia Erichson, 1837 (CJ pers. comm.). The species has not been added to the Yorkshire list.
Ocalea rivularis Miller, L. 1851. Past confusion with O. latipennis Sharp, 1870 renders the record from Raincliffe Woods (SE9988) on an unknown date (GBW) unreliable without a voucher specimen. The species has not been added to the Yorkshire list.
Dexiogyia corticina (Erichson, 1837) There is a single unconfirmed record on the YNU database: Filey (TA1280), 1914 (ECH). This record is unreliable without a voucher specimen and the species has not been added to the Yorkshire list.
References
Cooter, J. 1999. On ‘English’ names for British Coleoptera. Coleopterist 8 (1): 43-45. Denton, M.L. 2003. The Aleocharinae (Coleoptera: Staphylinidae) of Yorkshire. Naturalist 128: 103-125. Duff, A.G. (ed.). 2008. Checklist of the Beetles of the British Isles, 2008 edition. A.G. Duff publishing, Wells, Somerset. Duff, A.G. (ed.). 2012. Checklist of Beetles of the British Isles, 2nd edition. Pemberley Books. Duff, A.G., (ed.). 2018. Checklist of the Beetles of the British Isles, 3rd edition. Pemberley Books. Easton, A.M. 1971. A new British Atheta (Col., Staphylinidae) in a new subgenus. Entomologist’s Monthly Magazine 107: 24- 26. Heal, N.F. 1993. Trichiusa immigrata Lohse (Staphylinidae) – first record for Britain. Coleopterist 2: 18. Hyman, P.S., and Parsons, M.S. 1992. A review of the scarce and threatened Coleoptera of Great Britain, part 1. JNCC, Peterborough. Hyman, P.S. and Parsons, M.S. 1994. A review of the scarce and threatened Coleoptera of Great Britain, part 2. JNCC, Peterborough. Key, R.S. 1999. More on ‘noddy names’ for British beetles. Coleopterist 8 (2): 94. Lott, D.A. 1995. Trichiusa immigrata Lohse (Staphylinidae) in Leicestershire. Coleopterist 4: 14. Moore, R.R. 2004. Trichiusa immigrata Lohse (Staphylinidae) new to Scotland. Coleopterist 13: 34. Moore, R.R. 2012. The Beetles of the Isle of Raasay in the Inner Hebrides. Privately published. Owen, J.A. 1994. Ischnoglossa turcica Wunderle (Staphylinidae) new to Britain. Coleopterist 3: 65. Pope, R.D. 1977. Kloet & Hincks. A Check List of British Insects. Part 3: Coleoptera and Strepsiptera. Second revised edition. Handbks. Ident. Brit. Insects 11(3), pp. xiv+105. Royal Entomological Society, London. Shirt, D.B. (ed). 1987. British Red Data Books: 2. Insects. Nature Conservancy Council, Peterborough. Sutton, S.L. & Beaumont, H.E. (eds.). 1989. Butterflies and Moths of Yorkshire. Yorkshire Naturalists’ Union, Leeds. Welch, R.C. 1969. Aleochara verna Say (Col., Staphylinidae) new to Britain. Entomologist 102: 207-209. Welch, R.C. 1990. Aleochara binotata Kr., not A. verna Say (Col., Staphylinidae), a British insect. Entomologist’s Record and Journal of Variation 107: 225-226. Welch, R.C. 1997. The British species of the genus Aleochara Gravenhorst (Staphylinidae). Coleopterist 6: 1-45. Wunderle, P. 1992. Eine neue Art der Gattung Ischnoglossa Kraatz 1856 aus Turkie (Coleoptera. Staphylinidae. Aleocharinae). Entomol. Blatter. 88: 49-52.
Errors and omissions The authors would be pleased to receive feedback regarding any errors or omissions noted by users of this Atlas.
M L Denton FRES & R J Marsh FRES
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