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Aspects of the Functional Morphology in the Cranial and Cervical Skeleton of the Sabre-Toothed Cat Paramachairodus Ogygia (Kaup, 1832) (Felidae
Blackwell Science, LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082The Lin- nean Society of London, 2005? 2005 1443 363377 Original Article FUNCTIONAL MORPHOLOGY OF P. OGYGIAM. J. SALESA ET AL. Zoological Journal of the Linnean Society, 2005, 144, 363–377. With 11 figures Aspects of the functional morphology in the cranial and cervical skeleton of the sabre-toothed cat Paramachairodus ogygia (Kaup, 1832) (Felidae, Machairodontinae) from the Late Miocene of Spain: Downloaded from https://academic.oup.com/zoolinnean/article-abstract/144/3/363/2627519 by guest on 18 May 2020 implications for the origins of the machairodont killing bite MANUEL J. SALESA1*, MAURICIO ANTÓN2, ALAN TURNER1 and JORGE MORALES2 1School of Biological & Earth Sciences, Byrom Street, Liverpool John Moores University, Liverpool, L3 3AF, UK 2Departamento de Palaeobiología, Museo Nacional de Ciencias Naturales-CSIC, José Gutiérrez Abascal, 2. 28006 Madrid, Spain Received January 2004; accepted for publication March 2005 The skull and cervical anatomy of the sabre-toothed felid Paramachairodus ogygia (Kaup, 1832) is described in this paper, with special attention paid to its functional morphology. Because of the scarcity of fossil remains, the anatomy of this felid has been very poorly known. However, the recently discovered Miocene carnivore trap of Batallones-1, near Madrid, Spain, has yielded almost complete skeletons of this animal, which is now one of the best known machairodontines. Consequently, the machairodont adaptations of this primitive sabre-toothed felid can be assessed for the first time. Some characters, such as the morphology of the mastoid area, reveal an intermediate state between that of felines and machairodontines, while others, such as the flattened upper canines and verticalized mandibular symphysis, show clear machairodont affinities. -
(Barbourofelinae, Nimravidae, Carnivora), from the Middle Miocene of China Suggests Barbourofelines Are Nimravids, Not Felids
UCLA UCLA Previously Published Works Title A new genus and species of sabretooth, Oriensmilus liupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids Permalink https://escholarship.org/uc/item/0g62362j Journal JOURNAL OF SYSTEMATIC PALAEONTOLOGY, 18(9) ISSN 1477-2019 Authors Wang, Xiaoming White, Stuart C Guan, Jian Publication Date 2020-05-02 DOI 10.1080/14772019.2019.1691066 Peer reviewed eScholarship.org Powered by the California Digital Library University of California Journal of Systematic Palaeontology ISSN: 1477-2019 (Print) 1478-0941 (Online) Journal homepage: https://www.tandfonline.com/loi/tjsp20 A new genus and species of sabretooth, Oriensmilus liupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids Xiaoming Wang, Stuart C. White & Jian Guan To cite this article: Xiaoming Wang, Stuart C. White & Jian Guan (2020): A new genus and species of sabretooth, Oriensmilusliupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids , Journal of Systematic Palaeontology, DOI: 10.1080/14772019.2019.1691066 To link to this article: https://doi.org/10.1080/14772019.2019.1691066 View supplementary material Published online: 08 Jan 2020. Submit your article to this journal View related articles View Crossmark data Full Terms & Conditions of access and use can be found at https://www.tandfonline.com/action/journalInformation?journalCode=tjsp20 Journal of Systematic Palaeontology, 2020 Vol. 0, No. 0, 1–21, http://dx.doi.org/10.1080/14772019.2019.1691066 A new genus and species of sabretooth, Oriensmilus liupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids a,bà c d Xiaoming Wang , Stuart C. -
Hyaenodontidae (Creodonta, Mammalia) and the Position of Systematics in Evolutionary Biology
Hyaenodontidae (Creodonta, Mammalia) and the Position of Systematics in Evolutionary Biology by Paul David Polly B.A. (University of Texas at Austin) 1987 A dissertation submitted in partial satisfaction of the requirements for the degree of Doctor of Philosophy in Paleontology in the GRADUATE DIVISION of the UNIVERSITY of CALIFORNIA at BERKELEY Committee in charge: Professor William A. Clemens, Chair Professor Kevin Padian Professor James L. Patton Professor F. Clark Howell 1993 Hyaenodontidae (Creodonta, Mammalia) and the Position of Systematics in Evolutionary Biology © 1993 by Paul David Polly To P. Reid Hamilton, in memory. iii TABLE OF CONTENTS Introduction ix Acknowledgments xi Chapter One--Revolution and Evolution in Taxonomy: Mammalian Classification Before and After Darwin 1 Introduction 2 The Beginning of Modern Taxonomy: Linnaeus and his Predecessors 5 Cuvier's Classification 10 Owen's Classification 18 Post-Darwinian Taxonomy: Revolution and Evolution in Classification 24 Kovalevskii's Classification 25 Huxley's Classification 28 Cope's Classification 33 Early 20th Century Taxonomy 42 Simpson and the Evolutionary Synthesis 46 A Box Model of Classification 48 The Content of Simpson's 1945 Classification 50 Conclusion 52 Acknowledgments 56 Bibliography 56 Figures 69 Chapter Two: Hyaenodontidae (Creodonta, Mammalia) from the Early Eocene Four Mile Fauna and Their Biostratigraphic Implications 78 Abstract 79 Introduction 79 Materials and Methods 80 iv Systematic Paleontology 80 The Four Mile Fauna and Wasatchian Biostratigraphic Zonation 84 Conclusion 86 Acknowledgments 86 Bibliography 86 Figures 87 Chapter Three: A New Genus Eurotherium (Creodonta, Mammalia) in Reference to Taxonomic Problems with Some Eocene Hyaenodontids from Eurasia (With B. Lange-Badré) 89 Résumé 90 Abstract 90 Version française abrégéé 90 Introduction 93 Acknowledgments 96 Bibliography 96 Table 3.1: Original and Current Usages of Genera and Species 99 Table 3.2: Species Currently Included in Genera Discussed in Text 101 Chapter Four: The skeleton of Gazinocyon vulpeculus n. -
7 X 11 Long.P65
Cambridge University Press 978-0-521-73586-5 - Carnivoran Evolution: New Views on Phylogeny, Form, and Function Edited by Anjali Goswami and Anthony Friscia Index More information Index Page numbers in bold type refers to figures. Acinonyx jubatus, 226, 304 Arctoidea, 12, 27, 30, 31, 40, 94, 104 Actiocyon, 115, 125 Artiodactyla, 312 active replacement, 311 Atilix paludinosis, 9 Adelphailurus, 411 Ailuridae, 11, 30, 92–126, 231, 304 badgers, see Mustelidae diagnosis, 116 bamboo, 12, 94 Ailurinae, 92, 116–20 Barbourofelinae, 10, 34, 295, 301 Ailuropoda, 15, 20, 31, 94, 104, 107, 226 basicranium, 37, 65, 107, 155, 157, 158, 159 Ailurus, 117, 126 Bassaricyon, 11 Ailurus fulgens, 3, 30, 93, 126 Bassaricyon lasius, 377 alisphenoid canal, 82, 107, 110 Bassariscus, 231, 238 allometry, 43, 165, 168 Bassariscus astutus, 388 multiphasic, 172–6, 178–85 Bathygale, 32 postcranial, 411–59 bear-dogs, see Amphicyonidae Alopecocyon, 112, 113, 115, 122, 125 behaviour, 411, 454 Alopex lagopus, 392 Bergmann’s Rule, 396 American lion, see Panthera leo cf. atrox biogeography, 225–39, 247–65, 361 Amphictis, 30, 92, 109, 112, 115, 116, biomechanics 123–124 cranial, 466–81 Amphicynodon, 109 postcranial, 450–9 Amphicynopsis, 297 bite force, 466–81 Amphicyon major, 299 body size, 39–43, 226, 248, 249, 269, Amphicyonidae, 17, 34, 41, 142, 193, 295 270, 301, 314, 325, 330, 412, 413 Ancient DNA, 25 bone cracking, 8, 16, 19, 289, 304 Andrewsarchus, 304 brain size, 39, 43–7 Aonyx, 236 lions, 168 aquatic species, 4, see Pinnipedia Buxolestes, 271 ArcGIS, 379 Arctictis, -
Mammals from the Earliest Uintan (Middle Eocene) Turtle Bluff Member, Bridger Formation, Southwestern Wyoming, USA, Part 1: Primates and Rodentia
Palaeontologia Electronica palaeo-electronica.org Mammals from the earliest Uintan (middle Eocene) Turtle Bluff Member, Bridger Formation, southwestern Wyoming, USA, Part 1: Primates and Rodentia Thomas S. Kelly and Paul C. Murphey ABSTRACT The Turtle Bluff Member (TBM) is the stratotype section for the earliest Uintan bio- chron, Ui1a, of the middle Eocene Uintan North American Land Mammal age. For more than a century, the TBM had yielded only a few fragmentary specimens. As the result of many years of field work, numerous mammal fossils have now been recov- ered and provide an unprecedented opportunity to better define this poorly known interval. This is the first in a series of papers that provide detailed descriptions and tax- onomic revisions of the fauna of the TBM. Here we document the occurrence of the fol- lowing taxa in the TBM: Uintasorex parvulus; Microsyops annectans; Notharctus robustior; Omomys carteri; Trogolemur myodes; Washakius insignis; Thisbemys corru- gatus; Microparamys minutus; Microparamys sp.; Sciuravus nitidus; Tillomys senex; Tillomys? parvidens; Taxymys lucaris; Pauromys sp., cf. P. perdit us ; three informal sci- uravid species (sp. A, B and C); cf. Pareumys sp.; Metanoiamys sp.; and Elymys? emryi new species. Except for the previously described Hemiacodon engardae, all of the primates from the TBM are holdover taxa from the earlier Bridgerian Land Mammal age, whereas the rodents exhibit a modest diversification during the earliest Uintan. Elymys? emryi and four additional informal rodent species (Microparamys sp., sci- uravid sp. A, cf. Pareumys sp., and Metanoiamys sp.) make their appearances in the TBM and, as such, can be added to the list of index species characterizing biochron Ui1a. -
Diplomarbeit
DIPLOMARBEIT KILLING BEHAVIOR IN SMILODON FATALIS (MAMMALIA, CARNIVORA, FELIDAE) BASED ON FUNCTIONAL ANATOMY AND BODY PROPORTIONS OF THE FRONT- AND HIND LIMBS BY JENS-UWE SCHMIEDER GEOLOGISCHES INSTITUT DER EBERHARDT-KARLS-UNIVERSITÄT TÜBINGEN JULY 2000 Jens Schmieder Page II 02.07.2003 Hiermit versichere ich, die vorliegende Arbeit selbständig und nur unter Zuhilfenahme erlaubter und angegebener Hilfsmittel angefertigt zu haben. Tübingen, August 2000. (Jens Schmieder) Jens Schmieder Page III 02.07.2003 Abstract Elongated canines exclusively evolved in carnivores, which are able to stabilize their victims with their anterior extremities. It was shown that power and agility of the front limbs are strongly correlated with the development of sabers. Limb- and skull proportions of the extinct cat Smilodon fatalis were therefore compared with those of six extant species of large felids and those of Canis lupus. Furthermore, differences in hunting behavior and locomotory capabilities were analyzed. Ratios of limb segment lengths have been shown to relate to functional and locomotory differences (e.g., cursoriality) in both extinct and extant felines. S. fatalis is equipped with relatively short and sturdy limbs. Moreover, it possessed a great angle of inclination of the olecranon fossa relative to the long axis of the humerus, in addition to a wide and laterally oriented radial notch. The radial head was more circular than in any other extant cat member. Additionally, the Teres major muscle inserts further away from the shoulder joint and the joints are more powerfully built and demonstrate a great amount of strength and flexibility. It is very likely that Smilodon preyed on the large contemporary megafauna because of its overall more powerful anatomy compared to that of modern felines. -
University of Michigan University Library
CONTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN VOL.31, NO. 2, PP. 43-78 AU~US~15,2003 SMALL LIMNOCYONINES (HYmNODONTIDAE, MAMMALIA) FROM THE BRIDGERIAN MIDDLE EOCENE OF WYOMING: THINOCYON, PROLIMNOCYON, AND IMDODON, NEW GENUS MICHAEL MORLO AND GREGG F. GUNNELL MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN ANN ARBOR CONTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY Philip D. Gingerich, Director This series of contributions from the Museum of Paleontology is a medium for publication of papers based chiefly on collections in the Museum. When the number of pages issued is sufficient to make a volume, a title page plus a table of contents will be sent to libraries on the Museum's mailing list. This will be sent to individuals on request. A list of the separate issues may also be obtained by request. Correspondence should be directed to the Publications Secretary, Museum of Paleontology, The University of Michigan, 1109 Geddes Road, Ann Arbor, Michigan 48 109-1079 ([email protected]). VOLS. 2-3 1: Parts of volumes may be obtained if available. Price lists are available upon inquiry. Text and illustrations 02003 by the Museum of Paleontology, University of Michigan SMALL LIMNOCYONINES (HYAENODONTIDAE, MAMMALIA) FROM THE BRIDGERIAN MIDDLE EOCENE OF WYOMING: THINOCYON, PROLIMNOCYON, AND IRIDODON, NEW GENUS MICHAEL MORLO1 AND GREGG F. GUNNELL~ Abstract - The fossil record of small Limnocyoninae from the Bridgerian land- mammal age is reviewed and new specimens are described. Among 120 examined specimens of Thinocyon, 96 represent the type species, T velox, with the other 24 representing T medius. The relatively large sample size of I: velox demonstrates the great size and morphological variability of this species. -
The Role of Clade Competition in the Diversification of North American Canids
The role of clade competition in the diversification of North American canids Daniele Silvestroa,b,c,1, Alexandre Antonellia,d, Nicolas Salaminb,c, and Tiago B. Quentale,1 aDepartment of Biological and Environmental Sciences, University of Gothenburg, 413 19 Gothenburg, Sweden; bDepartment of Ecology and Evolution, University of Lausanne, 1015 Lausanne, Switzerland; cSwiss Institute of Bioinformatics, Quartier Sorge, 1015 Lausanne, Switzerland; dGothenburg Botanical Garden, 413 19 Gothenburg, Sweden; and eDepartamento de Ecologia, Universidade de São Paulo, 05508-900, São Paulo, SP, Brazil, 05508-900 Edited by Mike Foote, The University of Chicago, Chicago, IL, and accepted by the Editorial Board May 30, 2015 (received for review February 10, 2015) The history of biodiversity is characterized by a continual replace- The subfamilies within the dog family Canidae show the char- ment of branches in the tree of life. The rise and demise of these acteristic sequential clade replacement repeatedly seen in the branches (clades) are ultimately determined by changes in speci- deep history of biodiversity. Canids comprise three clades: the ation and extinction rates, often interpreted as a response to Hesperocyoninae and Borophaginae subfamilies, which are ex- varying abiotic and biotic factors. However, understanding the tinct, and the subfamily Caninae, which includes extinct and relative importance of these factors remains a major challenge in living species. Their diversification dynamics have been causally evolutionary biology. Here we analyze the rich North American linked to the evolution of intrinsic properties, including increase fossil record of the dog family Canidae and of other carnivores to in body size and an exclusively carnivorous diet (hypercarnivory) tease apart the roles of competition, body size evolution, and (22–24). -
Genus/Species Skull Ht Lt Wt Stage Range Abacinonyx See Acinonyx Abathomodon See Speothos A
Genus/Species Skull Ht Lt Wt Stage Range Abacinonyx see Acinonyx Abathomodon see Speothos A. fossilis see Icticyon pacivorus? Pleistocene Brazil Abelia U.Miocene Europe Absonodaphoenus see Pseudarctos L.Miocene USA A. bathygenus see Cynelos caroniavorus Acarictis L.Eocene W USA cf. A. ryani Wasatchian Colorado(US) A. ryani Wasatchian Wyoming, Colorado(US) Acinomyx see Acinonyx Acinonyx M.Pliocene-Recent Europe,Asia,Africa,N America A. aicha 2.3 m U.Pliocene Morocco A. brachygnathus Pliocene India A. expectata see Miracinonyx expectatus? Or Felis expectata? A. intermedius M.Pleistocene A. jubatus living Cheetah M.Pliocene-Recent Algeria,Europe,India,China A. pardinensis 91 cm 3 m 60 kg Astian-Biharian Italy,India,China,Germany,France A. sp. L.Pleistocene Tanzania,Ethiopia A. sp. Cf. Inexpectatus Blancan-Irvingtonian California(US) A. studeri see Miracinonyx studeri Blancan Texas(US) A. trumani see Miracinonyx trumani Rancholabrean Wyoming,Nevada(US) Acionyx possibly Acinonyx? A. cf. Crassidens Hadar(Ethiopia) Acrophoca 1.5 m U.Miocene-L.Pliocene Peru,Chile A. longirostris U.Miocene-L.Pliocene Peru A. sp. U.Miocene-L.Pliocene Chile Actiocyon M-U.Miocene W USA A. leardi Clarendonian California(US) A. sp. M.Miocene Oregon(US) Adcrocuta 82 cm 1.5 m U.Miocene Europe,Asia A. advena A. eximia 80 cm 1.5 m Vallesian-Turolian Europe(widespread),Asia(widespread) Adelphailurus U.Miocene-L.Pliocene W USA, Mexico,Europe A. kansensis Hemphillian Arizona,Kansas(US),Chihuahua(Mexico) Adelpharctos M.Oligocene Europe Adilophontes L.Miocene W USA A. brachykolos Arikareean Wyoming(US) Adracodon probably Adracon Eocene France A. quercyi probably Adracon quercyi Eocene France Adracon U.Eocene-L.Oligocene France A. -
Revista Ilicitana De Paleontología Y Mineralogía Núm. 30 2010
Cidaris Revista Ilicitana de Paleontología y Mineralogía Núm. 30 2010 VIII Encuentro de Jóvenes Investigadores en Paleontología VOLUMEN DE ACTAS GRUPO CULTURAL PALEONTOLÓGICO DE ELCHE NUEVOS HALLAZGOS DE SANSANOSMILUS JOURDANI (FILHOL 1883) (CARNIVORA: BARBOUROFELIDAE) DEL MIOCENO MEDIO DE LA SERIE ESTRATIGRÁFICA DEL ABOCADOR DE CAN MATA (CUENCA DEL VALLÈS-PENEDÈS, BARCELONA, ESPAÑA) NEW FINDS OF SANSANOSMILUS JOURDANI (FILHOL 1883) (CARNIVORA: BARBOUROFELIDAE) FROM THE MIDDLE MIOCENE STRATIGRAPHIC SERIES OF ABOCADOR DE CAN MATA (VALLÈS- PENEDÈS BASIN, BARCELONA, SPAIN) Josep M. Robles1,2, David M. Alba2, Raül Carmona1, Cheyenn Rotgers1,2, Jordi Galindo2, Jordi Balaguer1, Sergio Almécija2 y Salvador Moyà-Solà3 1FOSSILIA Serveis Paleontològics i Geològics, S.L. c/ Jaume I 87, 1er 5a Sant Celoni, Barcelona (Spain). E-mail: josep. [email protected], [email protected], [email protected], [email protected] 2Institut Català de Paleontologia, Universitat Autònoma de Barcelona. Edifi ci ICP, Campus de la UAB s/n, 08193 Cerdanyola del Vallès, Barcelona (Spain). E-mail: [email protected], [email protected], [email protected] 3ICREA en el Institut Català de Paleontologia y la Unitat d’Antropologia Biològica (Dept. BABVE), Universitat Autònoma de Barcelona. Edifi ci ICP, Campus de la UAB s/n, 08193 Cerdanyola del Vallès, Barcelona (Spain). E-mail: [email protected] RESUMEN La escasez de restos fósiles del carnívoro dientes-de-sable Sansanosmilus jourdani y otros barbourofélidos euroasiáticos ha difi cultado la clarifi cación de su estatus taxonómico. En este trabajo, se dan a conocer los nuevos hallazgos de esta especie, a partir de varias localidades del Aragoniense superior (Mioceno medio) de la serie estratigráfi ca del Abocador de Can Mata (els Hostalets de Pierola, cuenca del Vallès-Penedès, Barcelona, España). -
A Survey of Cenozoic Mammal Baramins
The Proceedings of the International Conference on Creationism Volume 8 Print Reference: Pages 217-221 Article 43 2018 A Survey of Cenozoic Mammal Baramins C Thompson Core Academy of Science Todd Charles Wood Core Academy of Science Follow this and additional works at: https://digitalcommons.cedarville.edu/icc_proceedings DigitalCommons@Cedarville provides a publication platform for fully open access journals, which means that all articles are available on the Internet to all users immediately upon publication. However, the opinions and sentiments expressed by the authors of articles published in our journals do not necessarily indicate the endorsement or reflect the views of DigitalCommons@Cedarville, the Centennial Library, or Cedarville University and its employees. The authors are solely responsible for the content of their work. Please address questions to [email protected]. Browse the contents of this volume of The Proceedings of the International Conference on Creationism. Recommended Citation Thompson, C., and T.C. Wood. 2018. A survey of Cenozic mammal baramins. In Proceedings of the Eighth International Conference on Creationism, ed. J.H. Whitmore, pp. 217–221. Pittsburgh, Pennsylvania: Creation Science Fellowship. Thompson, C., and T.C. Wood. 2018. A survey of Cenozoic mammal baramins. In Proceedings of the Eighth International Conference on Creationism, ed. J.H. Whitmore, pp. 217–221, A1-A83 (appendix). Pittsburgh, Pennsylvania: Creation Science Fellowship. A SURVEY OF CENOZOIC MAMMAL BARAMINS C. Thompson, Core Academy of Science, P.O. Box 1076, Dayton, TN 37321, [email protected] Todd Charles Wood, Core Academy of Science, P.O. Box 1076, Dayton, TN 37321, [email protected] ABSTRACT To expand the sample of statistical baraminology studies, we identified 80 datasets sampled from 29 mammalian orders, from which we performed 82 separate analyses. -
Contributions from the Museum of Paleontology the University of Michigan
CONTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN VOL. 28, NO. 7, w. 141-180 September 30, 1991 SYSTEMATICS AND EVOLUTION OF LATE PALEOCENE AND EARLY EOCENE OXYAENIDAE (MAMMALIA, CREODONTA) IN THE CLARKS FORK BASIN, WYOMING GREGG F. GUNNELL and PHILIP D. GINGERICH MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN ANN ARBOR CONTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY Philip D. Gingerich, Director This series of contributions from the Museum of Paleontology is a medium for publication of papers based chiefly on collections in the Museum. When he number of pages issued is sufficient to make a volume, a title page and a table of contents will be sent to libraries on the mailing list, and to individuals on reqcest. A list of the separate issues may also be obtained by request. Correspondence should be directed to the Museum of Paleontology, The University of Michigan, Ann Arbor, Michigan 48109-1079. VOLS. 2-27. Parts of volumes may be obtained if available. Price lists are available upon inquiry. SYSTEMATICS AND EVOLUTION OF LATE PALEOCENE AND EARLY EOCENE OXYAENIDAE (MAMMALIA, CREODONTA) IN THE CLARKS FORK BASIN, WYOMING BY GREGG F. GUNNELL and PHILIP D. GINGERICH Abstract.- Oxyaenidae is a family of archaic carnivorous mammals included in the mammalian order Creodonta. The major oxyaenid radiation occurred in North America during the late Paleocene and early Eocene, while smaller radiations occurred in Europe and Asia in the Eocene. Three subfamilies are represented in early Cenozoic sediments in the Clarks Fork Basin of north- western Wyoming: Tytthaeninae (new), Palaeonictinae, and Oxyaeninae. Tytthaenines are represented by one genus: Tiffanian-Clarkforkian Tytthaena; palaeonictines are represented by two genera: Tiffanian-Clarkforkian Dip- salodon and Clarkforkian-Wasatchian Palaeonictis; oxyaenines are represented by two genera: Clarkforkianarly Wasatchian Dipsalidictis, and Wasatchian Oxyaena.