DOCSLIB.ORG

New Specimens of Sparassodonta (Mammalia, Metatheria) From

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
New Specimens of Sparassodonta (Mammalia, Metatheria) From NEW SPECIMENS OF SPARASSODONTA (MAMMALIA, METATHERIA) FROM CHILE AND BOLIVIA by RUSSELL K. ENGELMAN Submitted in partial fulfillment of the requirements for the degree of Master of Science Department of Biology CASE WESTERN RESERVE UNIVERSITY January, 2019 CASE WESTERN RESERVE UNIVERSITY SCHOOL OF GRADUATE STUDIES We hereby approve the thesis/dissertation of Russell K. Engelman candidate for the degree of Master of Science*. Committee Chair Hillel J. Chiel Committee Member Darin A. Croft Committee Member Scott W. Simpson Committee Member Michael F. Benard Date of Defense July 20, 2018 *We also certify that written approval has been obtained for any proprietary material contained therein. ii TABLE OF CONTENTS NEW SPECIMENS OF SPARASSODONTA (MAMMALIA, METATHERIA) FROM CHILE AND BOLIVIA ....................................................................................................... i TABLE OF CONTENTS ................................................................................................... iii LIST OF TABLES ............................................................................................................. vi LIST OF FIGURES .......................................................................................................... vii ACKNOWLEDGEMENTS ................................................................................................ 1 LIST OF ABBREVIATIONS ............................................................................................. 4 ABSTRACT ........................................................................................................................ 7 1. INTRODUCTION ...................................................................................................... 8 2. AUSTRALOGALE LEPTOGNATHUS GEN. ET SP. NOV., A SECOND SPECIES OF SMALL SPARASSODONT (MAMMALIA, METATHERIA) FROM THE MIDDLE MIOCENE LOCALITY OF QUEBRADA HONDA, BOLIVIA .................... 13 2.1 INTRODUCTION .............................................................................................. 13 2.2 MATERIALS AND METHODS ....................................................................... 15 2.3 SYSTEMATIC PALEONTOLOGY .................................................................. 17 2.4 DESCRIPTION .................................................................................................. 19 2.5 DISCUSSION .................................................................................................... 30 2.5.1 The p3 locus in UATF-V-001900 ............................................................... 30 2.5.2 Systematic position of Australogale leptognathus ..................................... 34 2.5.3 Reduction of p3 in Metatherians ................................................................. 42 iii 2.5.4 Paleobiology of Australogale leptognathus ................................................ 44 2.5.5 The Sparassodont Fauna of Quebrada Honda and the Laventan SALMA . 47 3. A LATE EOCENE SPARASSODONT (MAMMALIA, METATHERIA, SPARASSODONTA) FROM THE LOCALITY OF LOS HELADOS (CENTRAL CHILE) ............................................................................................................................. 51 3.1 INTRODUCTION .............................................................................................. 51 3.2 MATERIALS AND METHODS ....................................................................... 53 3.3 SYSTEMATIC PALEONTOLOGY .................................................................. 53 3.4 DESCRIPTION .................................................................................................. 55 3.5 DISCUSSION .................................................................................................... 60 4. EOMAKHAIRA MOLOSSUS, A NEW BORHYAENOID SPARASSODONT (MAMMALIA, METATHERIA, SPARASSODONTA) FROM THE EARLY OLIGOCENE (?TINGUIRIRICAN) CACHAPOAL FAUNA, ANDEAN MAIN RANGE, CENTRAL CHILE ............................................................................................ 64 4.1 INTRODUCTION .............................................................................................. 64 4.2 MATERIALS AND METHODS ....................................................................... 68 4.3 SYSTEMATIC PALEONTOLOGY .................................................................. 70 4.4 DESCRIPTION .................................................................................................. 73 4.4.1 Cranium....................................................................................................... 77 4.4.2 Mandible ..................................................................................................... 86 4.4.3 Dentition ..................................................................................................... 92 iv 4.5 PHYLOGENETIC ANALYSIS ....................................................................... 118 4.6 DISCUSSION .................................................................................................. 127 4.6.1 Paleobiology of SGOPV 3490 .................................................................. 127 4.6.2 Carnassial Rotation in Sparassodonts ....................................................... 135 4.6.3 Proborhyaenidae and the Origin of Thylacosmilids ................................. 140 5. FUTURE DIRECTIONS ........................................................................................ 151 SUPPLEMENTARY INFORMATION ......................................................................... 155 LITERATURE CITED ................................................................................................... 278 v LIST OF TABLES Table 2.1. Mandibular depth of UATF-V-001900 compared to other small Neogene sparassodonts. ................................................................................................................... 20 Table 2.2. Measurements of the dentition and mandibular ramus of UATF-V-001900. .. 22 Table 2.3. Proportions of m1-2 in Australogale leptognathus gen. et. sp. nov. compared to other small sparassodonts and the slightly larger Sipalocyon gracilis. ........................ 23 Table 2.4. Comparison of features among Australogale leptognathus gen. et sp. nov. and other very small to medium-sized South American tritubercular metatherians. .............. 37 Table 3.1. Anteroposterior lengths (in mm) of the lower dentition of SGOPV 6200. ...... 59 Table 3.2. Lower molar row anteroposterior length (Lm1-4), estimated body mass, and relative dentary depth at m4 (m4D) of SGOPV 6200 and species of Pharsophorus and Plesiofelis. ......................................................................................................................... 61 Table 4.1. Measurements of the holotype of Eomakhaira molossus (SGOPV 3490) in mm. Greatest dorsoventral height of maxilla was measured from the alveolar border of P3 to the dorsal border of the maxilla. ................................................................................... 79 Table 4.2. Dental measurements of the holotype of Eomakhaira molossus (SGOPV 3490) in mm. ............................................................................................................................... 93 Table 4.3. Upper canine proportions of SGOPV 3490 compared to other sparassodonts, focusing on canine shape (ratio of anteroposterior length/labiolingual width) and relative canine size. ........................................................................................................................ 94 Table 4.4. Morphometric values of the dentition used to infer dietary habits in Eomakhaira molossus. .................................................................................................... 133 Table 4.5. Angle of inward canting of the posterior upper molars in sparassodonts. ..... 137 vi LIST OF FIGURES Figure 2.1. Temporal and geographical location of Quebrada Honda. ............................. 14 Figure 2.2. UATF-V-001900, Australogale leptognathus gen. et sp. nov. (holotype), right dentary fragment in A, labial; B, occlusal; and C, lingual views. .................................... 19 Figure 2.3. Photograph (A) and line drawing (B) of m1-2 of UATF-V-001900 in occlusal view. .................................................................................................................................. 25 Figure 2.4. Line drawings of m1-2 of (A) Australogale leptognathus, UATF-V-001900; (B) the small sparassodont Pseudonotictis pusillus, MLP 11-26, and (C) the didelphoid Hesperocynus dolgopolae, FMNH P14469 (modified from Abello et al., 2015), scaled to the same anteroposterior length, showing the greater resemblance of Australogale to small sparassodonts than didelphoids. .............................................................................. 28 Figure 2.5. X-ray (A) and micro-CT (B) images of UATF-V-001900, showing the lack of a replacement tooth below p3 (second tooth from right) and the fully formed anterior root of m3 (at left). ................................................................................................................... 31 Figure 2.6. Strict consensus tree of 8 most parsimonious trees showing the relationship of Australogale leptognathus among sparassodonts and other metatherians. ....................... 36 Figure 3.1. Map of Chile (left) and central Chile (inset box) showing the location of Los Helados (LH, at arrow) and other selected Paleogene Chilean fossil mammal localities of the
Load more

Recommended publications
  • Aspects of the Functional Morphology in the Cranial and Cervical Skeleton of the Sabre-Toothed Cat Paramachairodus Ogygia (Kaup, 1832) (Felidae
    Blackwell Science, LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082The Lin- nean Society of London, 2005? 2005 1443 363377 Original Article FUNCTIONAL MORPHOLOGY OF P. OGYGIAM. J. SALESA ET AL. Zoological Journal of the Linnean Society, 2005, 144, 363–377. With 11 figures Aspects of the functional morphology in the cranial and cervical skeleton of the sabre-toothed cat Paramachairodus ogygia (Kaup, 1832) (Felidae, Machairodontinae) from the Late Miocene of Spain: Downloaded from https://academic.oup.com/zoolinnean/article-abstract/144/3/363/2627519 by guest on 18 May 2020 implications for the origins of the machairodont killing bite MANUEL J. SALESA1*, MAURICIO ANTÓN2, ALAN TURNER1 and JORGE MORALES2 1School of Biological & Earth Sciences, Byrom Street, Liverpool John Moores University, Liverpool, L3 3AF, UK 2Departamento de Palaeobiología, Museo Nacional de Ciencias Naturales-CSIC, José Gutiérrez Abascal, 2. 28006 Madrid, Spain Received January 2004; accepted for publication March 2005 The skull and cervical anatomy of the sabre-toothed felid Paramachairodus ogygia (Kaup, 1832) is described in this paper, with special attention paid to its functional morphology. Because of the scarcity of fossil remains, the anatomy of this felid has been very poorly known. However, the recently discovered Miocene carnivore trap of Batallones-1, near Madrid, Spain, has yielded almost complete skeletons of this animal, which is now one of the best known machairodontines. Consequently, the machairodont adaptations of this primitive sabre-toothed felid can be assessed for the first time. Some characters, such as the morphology of the mastoid area, reveal an intermediate state between that of felines and machairodontines, while others, such as the flattened upper canines and verticalized mandibular symphysis, show clear machairodont affinities.
    [Show full text]
  • Chronostratigraphy of the Mammal-Bearing Paleocene of South America 51
    Thierry SEMPERE biblioteca Y. Joirriiol ofSoiiih Ainorirari Euirli Sciriin~r.Hit. 111. No. 1, pp. 49-70, 1997 Pergamon Q 1‘197 PublisIlcd hy Elscvicr Scicncc Ltd All rights rescrvcd. Printed in Grcnt nrilsin PII: S0895-9811(97)00005-9 0895-9X 11/97 t I7.ol) t o.(x) -. ‘Inshute qfI Human Origins, 1288 9th Street, Berkeley, California 94710, USA ’Orstom, 13 rue Geoffroy l’Angevin, 75004 Paris, France 3Department of Geosciences, The University of Arizona, Tucson, Arizona 85721, USA Absfract - Land mammal faunas of Paleocene age in the southern Andean basin of Bolivia and NW Argentina are calibrated by regional sequence stratigraphy and rnagnetostratigraphy. The local fauna from Tiupampa in Bolivia is -59.0 Ma, and is thus early Late Paleocene in age. Taxa from the lower part of the Lumbrera Formation in NW Argentina (long regarded as Early Eocene) are between -58.0-55.5 Ma, and thus Late Paleocene in age. A reassessment of the ages of local faunas from lhe Rfo Chico Formation in the San Jorge basin, Patagonia, southern Argentina, shows that lhe local fauna from the Banco Negro Infeiior is -60.0 Ma, mak- ing this the most ancient Cenozoic mammal fauna in South,America. Critical reevaluation the ltaboraí fauna and associated or All geology in SE Brazil favors lhe interpretation that it accumulated during a sea-level lowsland between -$8.2-56.5 Ma. known South American Paleocene land inammal faunas are thus between 60.0 and 55.5 Ma (i.e. Late Paleocene) and are here assigned to the Riochican Land Maminal Age, with four subages (from oldest to youngest: Peligrian, Tiupampian, Ilaboraian, Riochican S.S.).
    [Show full text]
  • Petrology and Geochemistry of Volcanic Rocks Behind the Cenozoic Arc Front in the Andean Cordillera, Central Chile (33°50'S) Andean Geology, Vol
    Andean Geology ISSN: 0718-7092 [email protected] Servicio Nacional de Geología y Minería Chile Muñoz, Marcia; Fuentes, Francisco; Vergara, Mario; Aguirre, Luis; Olov Nyström, Jan; Féraud, Gilbert; Demant, Alain Abanico East Formation: petrology and geochemistry of volcanic rocks behind the Cenozoic arc front in the Andean Cordillera, central Chile (33°50'S) Andean Geology, vol. 33, núm. 1, enero, 2006, pp. 109-140 Servicio Nacional de Geología y Minería Santiago, Chile Available in: http://www.redalyc.org/articulo.oa?id=173918422005 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative Abanico East Formation: petrology and geochemistry of volcanic rocks behind the Cenozoic arc front in the Andean Cordillera, central Chile (33°50'S) Marcia Muñoz Departamento de Geología, Universidad de Chile, Casilla 13518, Correo 21, Santiago, Chile [email protected] Francisco Fuentes [email protected] Mario Vergara [email protected] Luis Aguirre [email protected] Jan Olov Nyström Swedish Museum of Natural History, SE-10405 Stockholm, Sweden [email protected] Gilbert Féraud UMR Géosciences Azur, CNRS-UNSA, Université de Nice- Sophia Antipolis, 06108 Nice Cedex 02, France [email protected] Alain Demant Laboratoire de Pétrologie Magmatique Université Aix-Marseille III, 13397 Marseille Cedex 20, France [email protected] ABSTRACT The stratigraphy, chemistry and age of rocks assigned to the eastern portion of the Abanico Formation exposed along the El Volcán river valley, Principal Cordillera east of Santiago (30º50'S/70º12'-70º5'W), are reported and discussed.
    [Show full text]
  • Southward-Directed Subduction of the Farallon–Aluk Spreading Ridge and Its Impact on Subduction Mechanics and Andean Arc Magmatism: Insights From
    feart-08-00121 May 7, 2020 Time: 11:30 # 1 ORIGINAL RESEARCH published: 08 May 2020 doi: 10.3389/feart.2020.00121 Southward-Directed Subduction of the Farallon–Aluk Spreading Ridge and Its Impact on Subduction Mechanics and Andean Arc Magmatism: Insights From Edited by: Marina Manea, Geochemical and Seismic National Autonomous University of Mexico, Mexico Tomographic Data Reviewed by: 1,2 1,2 1,2 1,2 Luca Ferrari, Sofía B. Iannelli *, Lucía Fernández Paz , Vanesa D. Litvak , Guido Gianni , Geosciences Center, National Lucas M. Fennell1,2, Javiera González3, Friedrich Lucassen4, Simone Kasemann4, Autonomous University of Mexico, Verónica Oliveros3 and Andrés Folguera1,2 Mexico 1 2 Jiashun Hu, Departamento de Ciencias Geológicas, Universidad de Buenos Aires, Buenos Aires, Argentina, Instituto de Estudios 3 California Institute of Technology, Andinos ‘Don Pablo Groeber’, CONICET- Universidad de Buenos Aires, Buenos Aires, Argentina, Departamento 4 United States de Ciencias de la Tierra, Universidad de Concepción, Concepción, Chile, MARUM - Center for Marine Environmental Sciences and Faculty of Geosciences, University of Bremen, Bremen, Germany *Correspondence: Sofía B. Iannelli sofi[email protected] Since the initial proposal of the past existence of a southward-directed mid-ocean ridge–subduction interaction in the Andes during Late Cretaceous–Paleogene times, Specialty section: This article was submitted to several studies have been devoted to uncover the tectonomagmatic evidence of this Structural Geology and Tectonics, process. The collision of a spreading ridge against a subduction margin provokes a section of the journal important tectonomagmatic changes, including, between them, variations in arc-related Frontiers in Earth Science magmatic activity and in the plate-margin stress regime.
    [Show full text]
  • Exhumation and Uplift of the Western Main Cordillera Between 33° and 34°5
    6th International Symposium on Andean Geodynamics (ISAG 2005, Barcelona), Extended Abstracts: 273-276 Exhumation and uplift of the western Main Cordillera between 33° and 34°5 Andrés Fock" Reynaldo Charrier 2, Marcelo Fadas 3, Victor Maksaev 4, Mark Fanning 5, & Pamela Alvarez 6 1 Departamento de Geologia, Universidad de Chile, Santiago, Chile ([email protected]); 2 Departamento de Geologia, Universidad de Chile, Santiago, Chile ([email protected]); 3 Departamento de Geologia, Universidad de Chile, Santiago, Chile, and LMTG-IRD, Université Paul Sabatier, Toulouse, France ([email protected]); 4 Departamento de Geologia, Universidad de Chile, Santiago, Chile ([email protected]); 5 Research School of Earth Sciences, Australian National University, Camberra, Australia ([email protected]); 6 SIPETROL, Santiago, Chile ([email protected]) KEYWORDS: Central Chile, Andes, Apatite fission-track ages, Neogene mountain building INTRODUCTION ln this paper we discuss the control of major structures in the exhumation of Cenozoic rocks and surface uplift in the Andean Main Cordillera based on new apatite fission tracks age analysis and the study of geological cross -sections. Fig. 1: Principal morphostructural Units. The box shows de Study Region The study area is segmented in the following N-S oriented morphostructural units (Fig. 1), from west to east: Coastal Cordillera, Central Depression, Western and Eastern Main Cordil1era (WMC and EMC, respectively), and Frontal Cordillera. The Coastal Cordillera consists mainly of volcanic and sedirnentary Mesozoic rocks, the Aptian - Albian Las Chilcas and Maastrichtian La Valle formations and Late Mesozoic intrusive bodies (Sellés and Gana, 2001), which are in contact with Cenozoic rocks through west vergent thrust faults located in the Central Depression and the western border of the WMC.
    [Show full text]
  • A Dated Phylogeny of Marsupials Using a Molecular Supermatrix and Multiple Fossil Constraints
    Journal of Mammalogy, 89(1):175–189, 2008 A DATED PHYLOGENY OF MARSUPIALS USING A MOLECULAR SUPERMATRIX AND MULTIPLE FOSSIL CONSTRAINTS ROBIN M. D. BECK* School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales 2052, Australia Downloaded from https://academic.oup.com/jmammal/article/89/1/175/1020874 by guest on 25 September 2021 Phylogenetic relationships within marsupials were investigated based on a 20.1-kilobase molecular supermatrix comprising 7 nuclear and 15 mitochondrial genes analyzed using both maximum likelihood and Bayesian approaches and 3 different partitioning strategies. The study revealed that base composition bias in the 3rd codon positions of mitochondrial genes misled even the partitioned maximum-likelihood analyses, whereas Bayesian analyses were less affected. After correcting for base composition bias, monophyly of the currently recognized marsupial orders, of Australidelphia, and of a clade comprising Dasyuromorphia, Notoryctes,and Peramelemorphia, were supported strongly by both Bayesian posterior probabilities and maximum-likelihood bootstrap values. Monophyly of the Australasian marsupials, of Notoryctes þ Dasyuromorphia, and of Caenolestes þ Australidelphia were less well supported. Within Diprotodontia, Burramyidae þ Phalangeridae received relatively strong support. Divergence dates calculated using a Bayesian relaxed molecular clock and multiple age constraints suggested at least 3 independent dispersals of marsupials from North to South America during the Late Cretaceous or early Paleocene. Within the Australasian clade, the macropodine radiation, the divergence of phascogaline and dasyurine dasyurids, and the divergence of perameline and peroryctine peramelemorphians all coincided with periods of significant environmental change during the Miocene. An analysis of ‘‘unrepresented basal branch lengths’’ suggests that the fossil record is particularly poor for didelphids and most groups within the Australasian radiation.
    [Show full text]
  • A Phylogeny and Timescale for Marsupial Evolution Based on Sequences for Five Nuclear Genes
    J Mammal Evol DOI 10.1007/s10914-007-9062-6 ORIGINAL PAPER A Phylogeny and Timescale for Marsupial Evolution Based on Sequences for Five Nuclear Genes Robert W. Meredith & Michael Westerman & Judd A. Case & Mark S. Springer # Springer Science + Business Media, LLC 2007 Abstract Even though marsupials are taxonomically less diverse than placentals, they exhibit comparable morphological and ecological diversity. However, much of their fossil record is thought to be missing, particularly for the Australasian groups. The more than 330 living species of marsupials are grouped into three American (Didelphimorphia, Microbiotheria, and Paucituberculata) and four Australasian (Dasyuromorphia, Diprotodontia, Notoryctemorphia, and Peramelemorphia) orders. Interordinal relationships have been investigated using a wide range of methods that have often yielded contradictory results. Much of the controversy has focused on the placement of Dromiciops gliroides (Microbiotheria). Studies either support a sister-taxon relationship to a monophyletic Australasian clade or a nested position within the Australasian radiation. Familial relationships within the Diprotodontia have also proved difficult to resolve. Here, we examine higher-level marsupial relationships using a nuclear multigene molecular data set representing all living orders. Protein-coding portions of ApoB, BRCA1, IRBP, Rag1, and vWF were analyzed using maximum parsimony, maximum likelihood, and Bayesian methods. Two different Bayesian relaxed molecular clock methods were employed to construct a timescale for marsupial evolution and estimate the unrepresented basal branch length (UBBL). Maximum likelihood and Bayesian results suggest that the root of the marsupial tree is between Didelphimorphia and all other marsupials. All methods provide strong support for the monophyly of Australidelphia. Within Australidelphia, Dromiciops is the sister-taxon to a monophyletic Australasian clade.
    [Show full text]
  • (Barbourofelinae, Nimravidae, Carnivora), from the Middle Miocene of China Suggests Barbourofelines Are Nimravids, Not Felids
    UCLA UCLA Previously Published Works Title A new genus and species of sabretooth, Oriensmilus liupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids Permalink https://escholarship.org/uc/item/0g62362j Journal JOURNAL OF SYSTEMATIC PALAEONTOLOGY, 18(9) ISSN 1477-2019 Authors Wang, Xiaoming White, Stuart C Guan, Jian Publication Date 2020-05-02 DOI 10.1080/14772019.2019.1691066 Peer reviewed eScholarship.org Powered by the California Digital Library University of California Journal of Systematic Palaeontology ISSN: 1477-2019 (Print) 1478-0941 (Online) Journal homepage: https://www.tandfonline.com/loi/tjsp20 A new genus and species of sabretooth, Oriensmilus liupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids Xiaoming Wang, Stuart C. White & Jian Guan To cite this article: Xiaoming Wang, Stuart C. White & Jian Guan (2020): A new genus and species of sabretooth, Oriensmilusliupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids , Journal of Systematic Palaeontology, DOI: 10.1080/14772019.2019.1691066 To link to this article: https://doi.org/10.1080/14772019.2019.1691066 View supplementary material Published online: 08 Jan 2020. Submit your article to this journal View related articles View Crossmark data Full Terms & Conditions of access and use can be found at https://www.tandfonline.com/action/journalInformation?journalCode=tjsp20 Journal of Systematic Palaeontology, 2020 Vol. 0, No. 0, 1–21, http://dx.doi.org/10.1080/14772019.2019.1691066 A new genus and species of sabretooth, Oriensmilus liupanensis (Barbourofelinae, Nimravidae, Carnivora), from the middle Miocene of China suggests barbourofelines are nimravids, not felids a,bà c d Xiaoming Wang , Stuart C.
    [Show full text]
  • Análisis Litoestratigráfico De La Formación Cerro Azul (Mioceno Superior) En La Provincia De La Pampa
    Revista de la Asociación Geológica Argentina 67 (2): 257 - 265 (2010) 257 ANÁLISIS LITOESTRATIGRÁFICO DE LA FORMACIÓN CERRO AZUL (MIOCENO SUPERIOR) EN LA PROVINCIA DE LA PAMPA Graciela VISCONTI1, Ricardo N. MELCHOR1,2, Claudia I. MONTALVO1, Aldo M. UMAZANO1,2 y Elena E. DE ELORRIAGA1 1 Universidad Nacional de La Pampa, Facultad de Ciencias Exactas y Naturales. E-mail: [email protected] 2 INCITAP (CONICET-UNLPam) RESUMEN La Formación Cerro Azul fue definida en 1980 para incluir a las sedimentitas continentales pliocenas (limolitas arenosas y are- niscas limosas) que afloran de manera discontinua en casi todo el ámbito de la provincia de La Pampa. No obstante, varias in- vestigaciones paleontológicas realizadas a partir de la segunda mitad de la década del 80’ han permitido ubicar geocronológi- camente a la unidad en el intervalo 10 Ma a 5,8-5,7 Ma. El objetivo del trabajo es realizar un análisis de las características li- toestratigráficas de la Formación Cerro Azul de acuerdo al Código Argentino de Estratigrafía. Se propone un lectoestratoti- po para la unidad, consistente en el perfil de Algarrobo del Águila y un perfil auxiliar en cerro El Morro. También se estable- cieron las relaciones estratigráficas con otras formaciones. Se interpreta un paleoambiente depositacional de llanura, donde alter- nan depósitos de loess con numerosos paleosuelos, detectándose escasos depósitos lacustres en la base y pocos cursos fluviales. Palabras clave: Formación Cerro Azul, Mioceno, Huayqueriense, La Pampa. ABSTRACT: Lithostratigrafic analysis of Cerro Azul Formation (Upper Miocene), La Pampa. Cerro Azul Formation was defined in 1980 to include the Pliocene continental sedimentary rocks (sandy silt and silty sand) that appear in discontinuous outcrops in al- most all La Pampa province.
    [Show full text]
  • The Record of Miocene Impacts in the Argentine Pampas
    Meteoritics & Planetary Science 41, Nr 5, 749–771 (2006) Abstract available online at http://meteoritics.org The record of Miocene impacts in the Argentine Pampas Peter H. SCHULTZ1*, Marcelo Z¡RATE2, Willis E. HAMES3, R. Scott HARRIS1, T. E. BUNCH4, Christian KOEBERL5, Paul RENNE6, and James WITTKE7 1Department of Geological Sciences, Brown University, Providence, Rhode Island 02912–1846, USA 2Facultad de Ciencias Exactas y Naturales, Universidad Nacional de La Pampa, Avda Uruguay 151, 6300 Santa Rosa, La Pampa, Argentina 3Department of Geology, Auburn University, Auburn, Alabama 36849, USA 4Department of Geology, Northern Arizona University, Flagstaff, Arizona 86011, USA 5Department of Geological Sciences, University of Vienna, Althanstrasse 14, A-1090 Vienna, Austria 6Berkeley Geochronology Center, 2455 Ridge Road, Berkeley, California 94709, USA 7Department of Geology, Northern Arizona University, Flagstaff, Arizona 86011, USA *Corresponding author. E-mail: [email protected] (Received 02 March 2005; revision accepted 14 December 2005) Abstract–Argentine Pampean sediments represent a nearly continuous record of deposition since the late Miocene (∼10 Ma). Previous studies described five localized concentrations of vesicular impact glasses from the Holocene to late Pliocene. Two more occurrences from the late Miocene are reported here: one near Chasicó (CH) with an 40Ar/39Ar age of 9.24 ± 0.09 Ma, and the other near Bahía Blanca (BB) with an age of 5.28 ± 0.04 Ma. In contrast with andesitic and dacitic impact glasses from other localities in the Pampas, the CH and BB glasses are more mafic. They also exhibit higher degrees of melting with relatively few xenoycrysts but extensive quench crystals. In addition to evidence for extreme heating (>1700 °C), shock features are observed (e.g., planar deformation features [PDFs] and diaplectic quartz and feldspar) in impact glasses from both deposits.
    [Show full text]
  • The Fauna from the Tyrannosaurus Rex Excavation, Frenchman Formation (Late Maastrichtian), Saskatchewan
    The Fauna from the Tyrannosaurus rex Excavation, Frenchman Formation (Late Maastrichtian), Saskatchewan Tim T. Tokaryk 1 and Harold N. Bryant 2 Tokaryk, T.T. and Bryant, H.N. (2004): The fauna from the Tyrannosaurus rex excavation, Frenchman Formation (Late Maastrichtian), Saskatchewan; in Summary of Investigations 2004, Volume 1, Saskatchewan Geological Survey, Sask. Industry Resources, Misc. Rep. 2004-4.1, CD-ROM, Paper A-18, 12p. Abstract The quarry that contained the partial skeleton of the Tyrannosaurus rex, familiarly known as “Scotty,” has yielded a diverse faunal and floral assemblage. The site is located in the Frenchman River valley in southwestern Saskatchewan and dates from approximately 65 million years, at the end of the Cretaceous Period. The faunal assemblage from the quarry is reviewed and the floral assemblage is summarized. Together, these assemblages provide some insight into the biological community that lived in southwestern Saskatchewan during the latest Cretaceous. Keywords: Frenchman Formation, Maastrichtian, Late Cretaceous, southwestern Saskatchewan, Tyrannosaurus rex. 1. Introduction a) Geological Setting The Frenchman Formation, of latest Maastrichtian age, is extensively exposed in southwestern Saskatchewan (Figure 1; Fraser et al., 1935; Furnival, 1950). The lithostratigraphic units in the formation consist largely of fluvial sandstones and greenish grey to green claystones. Outcrops of the Frenchman Formation are widely distributed in the Frenchman River valley, southeast of Eastend. Chambery Coulee, on the north side of the valley, includes Royal Saskatchewan Museum (RSM) locality 72F07-0022 (precise locality data on file with the RSM), the site that contained the disarticulated skeleton of a Tyrannosaurus rex. McIver (2002) subdivided the stratigraphic sequence at this locality into “lower” and “upper” beds.
    [Show full text]
  • University of Florida Thesis Or Dissertation Formatting
    UNDERSTANDING CARNIVORAN ECOMORPHOLOGY THROUGH DEEP TIME, WITH A CASE STUDY DURING THE CAT-GAP OF FLORIDA By SHARON ELIZABETH HOLTE A DISSERTATION PRESENTED TO THE GRADUATE SCHOOL OF THE UNIVERSITY OF FLORIDA IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY UNIVERSITY OF FLORIDA 2018 © 2018 Sharon Elizabeth Holte To Dr. Larry, thank you ACKNOWLEDGMENTS I would like to thank my family for encouraging me to pursue my interests. They have always believed in me and never doubted that I would reach my goals. I am eternally grateful to my mentors, Dr. Jim Mead and the late Dr. Larry Agenbroad, who have shaped me as a paleontologist and have provided me to the strength and knowledge to continue to grow as a scientist. I would like to thank my colleagues from the Florida Museum of Natural History who provided insight and open discussion on my research. In particular, I would like to thank Dr. Aldo Rincon for his help in researching procyonids. I am so grateful to Dr. Anne-Claire Fabre; without her understanding of R and knowledge of 3D morphometrics this project would have been an immense struggle. I would also to thank Rachel Short for the late-night work sessions and discussions. I am extremely grateful to my advisor Dr. David Steadman for his comments, feedback, and guidance through my time here at the University of Florida. I also thank my committee, Dr. Bruce MacFadden, Dr. Jon Bloch, Dr. Elizabeth Screaton, for their feedback and encouragement. I am grateful to the geosciences department at East Tennessee State University, the American Museum of Natural History, and the Museum of Comparative Zoology at Harvard for the loans of specimens.
    [Show full text]