1 the Proposed Relocation of Dwellers at Ingula Pumped Storage
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University of Birmingham Endocranial Anatomy
University of Birmingham Endocranial anatomy and life habits of the Early Triassic archosauriform Proterosuchus fergusi Brown, Emily; Butler, Richard; Ezcurra, Martin; Bhullar, Bhart-Anjan; Lautenschlager, Stephan DOI: 10.1111/pala.12454 License: Other (please specify with Rights Statement) Document Version Peer reviewed version Citation for published version (Harvard): Brown, E, Butler, R, Ezcurra, M, Bhullar, B-A & Lautenschlager, S 2020, 'Endocranial anatomy and life habits of the Early Triassic archosauriform Proterosuchus fergusi', Palaeontology, vol. 63, no. 2, pp. 255-282. https://doi.org/10.1111/pala.12454 Link to publication on Research at Birmingham portal Publisher Rights Statement: This is the peer reviewed version of the following article: Brown, E. E., Butler, R. J., Ezcurra, M. D., Bhullar, B. S. and Lautenschlager, S. (2019), Endocranial anatomy and life habits of the Early Triassic archosauriform Proterosuchus fergusi. Palaeontology, which has been published in final form at https://doi.org/10.1111/pala.12454. This article may be used for non-commercial purposes in accordance with Wiley Terms and Conditions for Use of Self-Archived Versions. General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. -
8. Archosaur Phylogeny and the Relationships of the Crocodylia
8. Archosaur phylogeny and the relationships of the Crocodylia MICHAEL J. BENTON Department of Geology, The Queen's University of Belfast, Belfast, UK JAMES M. CLARK* Department of Anatomy, University of Chicago, Chicago, Illinois, USA Abstract The Archosauria include the living crocodilians and birds, as well as the fossil dinosaurs, pterosaurs, and basal 'thecodontians'. Cladograms of the basal archosaurs and of the crocodylomorphs are given in this paper. There are three primitive archosaur groups, the Proterosuchidae, the Erythrosuchidae, and the Proterochampsidae, which fall outside the crown-group (crocodilian line plus bird line), and these have been defined as plesions to a restricted Archosauria by Gauthier. The Early Triassic Euparkeria may also fall outside this crown-group, or it may lie on the bird line. The crown-group of archosaurs divides into the Ornithosuchia (the 'bird line': Orn- ithosuchidae, Lagosuchidae, Pterosauria, Dinosauria) and the Croco- dylotarsi nov. (the 'crocodilian line': Phytosauridae, Crocodylo- morpha, Stagonolepididae, Rauisuchidae, and Poposauridae). The latter three families may form a clade (Pseudosuchia s.str.), or the Poposauridae may pair off with Crocodylomorpha. The Crocodylomorpha includes all crocodilians, as well as crocodi- lian-like Triassic and Jurassic terrestrial forms. The Crocodyliformes include the traditional 'Protosuchia', 'Mesosuchia', and Eusuchia, and they are defined by a large number of synapomorphies, particularly of the braincase and occipital regions. The 'protosuchians' (mainly Early *Present address: Department of Zoology, Storer Hall, University of California, Davis, Cali- fornia, USA. The Phylogeny and Classification of the Tetrapods, Volume 1: Amphibians, Reptiles, Birds (ed. M.J. Benton), Systematics Association Special Volume 35A . pp. 295-338. Clarendon Press, Oxford, 1988. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Tiago Rodrigues Simões
Diapsid Phylogeny and the Origin and Early Evolution of Squamates by Tiago Rodrigues Simões A thesis submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy in SYSTEMATICS AND EVOLUTION Department of Biological Sciences University of Alberta © Tiago Rodrigues Simões, 2018 ABSTRACT Squamate reptiles comprise over 10,000 living species and hundreds of fossil species of lizards, snakes and amphisbaenians, with their origins dating back at least as far back as the Middle Jurassic. Despite this enormous diversity and a long evolutionary history, numerous fundamental questions remain to be answered regarding the early evolution and origin of this major clade of tetrapods. Such long-standing issues include identifying the oldest fossil squamate, when exactly did squamates originate, and why morphological and molecular analyses of squamate evolution have strong disagreements on fundamental aspects of the squamate tree of life. Additionally, despite much debate, there is no existing consensus over the composition of the Lepidosauromorpha (the clade that includes squamates and their sister taxon, the Rhynchocephalia), making the squamate origin problem part of a broader and more complex reptile phylogeny issue. In this thesis, I provide a series of taxonomic, phylogenetic, biogeographic and morpho-functional contributions to shed light on these problems. I describe a new taxon that overwhelms previous hypothesis of iguanian biogeography and evolution in Gondwana (Gueragama sulamericana). I re-describe and assess the functional morphology of some of the oldest known articulated lizards in the world (Eichstaettisaurus schroederi and Ardeosaurus digitatellus), providing clues to the ancestry of geckoes, and the early evolution of their scansorial behaviour. -
Clippety Clop), Kwelera, East London, Great Kei Municipality, Eastern Cape
PALAEONTOLOGICAL ASSESSMENT: COMBINED FIELD ASSESSMENT AND DESKTOP STUDY Proposed development of Portion 3 of Farm 695 (Clippety Clop), Kwelera, East London, Great Kei Municipality, Eastern Cape. JOHN E. ALMOND (PhD, Cantab) Natura Viva cc, PO Box 12410 Mill Street, CAPE TOWN 8010, RSA. [email protected] October 2011 1. SUMMARY The proposed holiday housing development on Portion 3 of Farm 695 (Clippety Clop), Kwelera, East London, is situated on the northern banks of the tidal Kwelera River, some 20 km northeast of East London, Eastern Cape. The development footprint is largely underlain by Late Permian continental sediments of the Adelaide Subgroup (Lower Beaufort Group, c. 253-251 million years old). These rocks are overlain by Early Triassic sandstones of the Katberg Formation (Tarkastad Subgroup) that build the cliffs and higher ground to the northeast. South of the river the Beaufort Group sediments are intruded and baked by Early Jurassic igneous intrusions of the Karoo Dolerite Suite. The Balfour Formation fluvial sediments are potentially fossiliferous, having yielded elsewhere a wide range of terrestrial vertebrates (bones and teeth of pareiasaurs, therapsids, amphibians et al.), bivalves, trace fossils and vascular plants. The overall impact of this project on local palaeontological heritage is likely to be very minor, however, because the potentially fossiliferous Beaufort Group sediments here are (a) deeply weathered, (b) sparsely fossiliferous, (c) have probably been extensively baked by nearby dolerite intrusions, and (d) are mostly covered with a thick (> 3m) mantle of fossil-poor alluvium. No fossils were observed within good exposures of the Balfour Formation rocks at the coast and in excellent roadcuts inland. -
Osteohistology of Late Triassic Prozostrodontian Cynodonts from Brazil
Osteohistology of Late Triassic prozostrodontian cynodonts from Brazil Jennifer Botha-Brink1,2, Marina Bento Soares3 and Agustín G. Martinelli3 1 Department of Karoo Palaeontology, National Museum, Bloemfontein, South Africa 2 Department of Zoology and Entomology, University of the Free State, Bloemfontein, South Africa 3 Departamento de Paleontologia e Estratigrafia, Instituto de Geociências, Universidade Federal do Rio Grande do Sul, Porto Alegre, Brazil ABSTRACT The Prozostrodontia includes a group of Late Triassic-Early Cretaceous eucynodonts plus the clade Mammaliaformes, in which Mammalia is nested. Analysing their growth patterns is thus important for understanding the evolution of mammalian life histories. Obtaining material for osteohistological analysis is difficult due to the rare and delicate nature of most of the prozostrodontian taxa, much of which comprises mostly of crania or sometimes even only teeth. Here we present a rare opportunity to observe the osteohistology of several postcranial elements of the basal prozostrodontid Prozostrodon brasiliensis, the tritheledontid Irajatherium hernandezi, and the brasilodontids Brasilodon quadrangularis and Brasilitherium riograndensis from the Late Triassic of Brazil (Santa Maria Supersequence). Prozostrodon and Irajatherium reveal similar growth patterns of rapid early growth with annual interruptions later in ontogeny. These interruptions are associated with wide zones of slow growing bone tissue. Brasilodon and Brasilitherium exhibit a mixture of woven-fibered bone tissue and slower growing parallel-fibered and lamellar bone. The slower growing bone tissues are present even during early ontogeny. The relatively slower growth in Brasilodon and Brasilitherium may be related to their small body size compared to Prozostrodon and Irajatherium. These brasilodontids also exhibit osteohistological similarities with the Late Triassic/Early Jurassic mammaliaform Morganucodon and the Late Cretaceous multituberculate mammals Kryptobaatar and Nemegtbaatar. -
Physical and Environmental Drivers of Paleozoic Tetrapod Dispersal Across Pangaea
ARTICLE https://doi.org/10.1038/s41467-018-07623-x OPEN Physical and environmental drivers of Paleozoic tetrapod dispersal across Pangaea Neil Brocklehurst1,2, Emma M. Dunne3, Daniel D. Cashmore3 &Jӧrg Frӧbisch2,4 The Carboniferous and Permian were crucial intervals in the establishment of terrestrial ecosystems, which occurred alongside substantial environmental and climate changes throughout the globe, as well as the final assembly of the supercontinent of Pangaea. The fl 1234567890():,; in uence of these changes on tetrapod biogeography is highly contentious, with some authors suggesting a cosmopolitan fauna resulting from a lack of barriers, and some iden- tifying provincialism. Here we carry out a detailed historical biogeographic analysis of late Paleozoic tetrapods to study the patterns of dispersal and vicariance. A likelihood-based approach to infer ancestral areas is combined with stochastic mapping to assess rates of vicariance and dispersal. Both the late Carboniferous and the end-Guadalupian are char- acterised by a decrease in dispersal and a vicariance peak in amniotes and amphibians. The first of these shifts is attributed to orogenic activity, the second to increasing climate heterogeneity. 1 Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK. 2 Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstraße 43, 10115 Berlin, Germany. 3 School of Geography, Earth and Environmental Sciences, University of Birmingham, Birmingham B15 2TT, UK. 4 Institut -
Stratigraphy, Sedimentary Facies and Diagenesis of the Ecca Group, Karoo Supergroup in the Eastern Cape, South Africa
STRATIGRAPHY, SEDIMENTARY FACIES AND DIAGENESIS OF THE ECCA GROUP, KAROO SUPERGROUP IN THE EASTERN CAPE, SOUTH AFRICA By Nonhlanhla Nyathi Dissertation submitted in fulfilment of the requirements for the degree of Master of Science In Geology FACULTY OF SCIENCE AND AGRICULTURE UNIVERSITY OF FORT HARE SUPERVISOR: PROFESSOR K. LIU CO-SUPERVISOR: PROFESSOR O. GWAVAVA APRIL 2014 DECLARATION I, Nonhlanhla Nyathi, hereby declare that the research described in this dissertation was carried out in the field and under the auspices of the Department of Geology, University of Fort Hare, under the supervision of Professor K. Liu and Prof. O. Gwavava. This dissertation and the accompanying photographs represent original work by the author, and have not been submitted, in whole or in part, to any other university for the purpose of a higher degree. Where reference has been made to the work of others, it has been dully acknowledged in the text. N. NYATHI Date signed: 12/04/2014 Place signed: Alice ACKNOWLEDGEMENTS The author is indebted to Professor K. Liu for his guidance, knowledge on all aspects of the project, constant supervision and help in doing field wok. Professor O. Gwavava is much appreciated for being my co-supervisor and helping me obtain financial support from the Goven Mbeki Research Development Centre. I am deeply grateful for the emotional support and encouragement from my parents and family. I express my profound gratitude to Mr Edwin Mutototwa and Mr Eric Madifor always taking time to read through my dissertation. The Rhodes University Geology laboratory technicians are thanked for their assistance in the making of thin sections. -
Variability of the Parietal Foramen and the Evolution of the Pineal Eye in South African Permo-Triassic Eutheriodont Therapsids
The sixth sense in mammalian forerunners: Variability of the parietal foramen and the evolution of the pineal eye in South African Permo-Triassic eutheriodont therapsids JULIEN BENOIT, FERNANDO ABDALA, PAUL R. MANGER, and BRUCE S. RUBIDGE Benoit, J., Abdala, F., Manger, P.R., and Rubidge, B.S. 2016. The sixth sense in mammalian forerunners: Variability of the parietal foramen and the evolution of the pineal eye in South African Permo-Triassic eutheriodont therapsids. Acta Palaeontologica Polonica 61 (4): 777–789. In some extant ectotherms, the third eye (or pineal eye) is a photosensitive organ located in the parietal foramen on the midline of the skull roof. The pineal eye sends information regarding exposure to sunlight to the pineal complex, a region of the brain devoted to the regulation of body temperature, reproductive synchrony, and biological rhythms. The parietal foramen is absent in mammals but present in most of the closest extinct relatives of mammals, the Therapsida. A broad ranging survey of the occurrence and size of the parietal foramen in different South African therapsid taxa demonstrates that through time the parietal foramen tends, in a convergent manner, to become smaller and is absent more frequently in eutherocephalians (Akidnognathiidae, Whaitsiidae, and Baurioidea) and non-mammaliaform eucynodonts. Among the latter, the Probainognathia, the lineage leading to mammaliaforms, are the only one to achieve the complete loss of the parietal foramen. These results suggest a gradual and convergent loss of the photoreceptive function of the pineal organ and degeneration of the third eye. Given the role of the pineal organ to achieve fine-tuned thermoregulation in ecto- therms (i.e., “cold-blooded” vertebrates), the gradual loss of the parietal foramen through time in the Karoo stratigraphic succession may be correlated with the transition from a mesothermic metabolism to a high metabolic rate (endothermy) in mammalian ancestry. -
An Examination of Non-Mammalian Cynodont Cranial Endocasts
.6 \\0 \. \ '3 ~ -=t-5 c. I BLOEMfONTeiN . OUIUOTEEK. LI£lRARY fÏËiliiiËÊKSEÏ'~lPi·AAriM~GëN-D'ËR University Free State 1111111111111111111111\111\111\111111111111111 IIIII ~II IIIII IIIII III 11\1 ,::~:EN O~lSTANDJGHED£ UIT DIE 34300004543215 universiteitVrY'stáat ~~:::[~:!'VERWYDER WORD NIE 'n"'= '.."mc*.,._....,_.,p j----------------------~ AN EXAMINATION OF NON-MAMMALIAN CYNODONT CRANIAL ENDOCASTS by DEWALD DU PLESSIS Submitted in fulfilment of the requirements for the degree MAGISTER SCiENTlAE in the Department of Zoology and Entomo.logy Faculty of Natural and Agricultural Sciences University of the Free State Bloemfontein Study leaders: Dr. J. Botha-Brink Mr. H.J.B. Butler September 2010 DECLARATION I, the undersigned, hereby declare that the work contained in this dissertation is my own original work and that I have not previously in its entirety or in part submitted it at any university for a degree. I further more cede copyright of the dissertation in favour of the University of the Free State. Signature: " Date: " . ii DEDICATION To the King of kings, the only living God, the Lord and my Saviour, Jesus Christ. All of the praise, all of the honour, all of the glorification belongs to You. I thank you for Your gifts, blessings and inspiration. Thank you for the three years during which I was blessed with the privilege to work with these animals. Everything that I am, is grace from Your hands. Without You I am nothing. iii ACKNOWLEDGEMENTS The present study was completed at the University of the Free State, Bloemfontein, in conjunction with the National Museum in Bloemfontein and the Nuclear Energy Corporation of South Africa (Neesa), with the support of a grant from the National Research Foundation. -
Umsobomvu Wind Energy Facility Near Middelburg, Pixley Ka Seme & Chris Hani District Municipalities, Northern and Eastern Ca
1 Palaeontological specialist assessment: combined desktop and field-based study Umsobomvu Wind Energy Facility near Middelburg, Pixley ka Seme & Chris Hani District Municipalities, Northern and Eastern Cape John E. Almond PhD (Cantab.) Natura Viva cc, PO Box 12410 Mill Street, Cape Town 8010, RSA [email protected] April 2015 EXECUTIVE SUMMARY InnoWind (Pty) Ltd is proposing to develop a wind energy facility (WEF) of up to 140 MW generation capacity in the mountainous Agter-Renosterberg region of the eastern Karoo, situated some 25 km to the northwest of Middelburg, on the border between the Northern Cape and Eastern Cape Provinces. Land parcels concerned in the proposed WEF total over 18 200 hectares and include 60/1, 133/RE, 133/1, 133/3, 61/0, 61/2, 61/4, 61/6, 135/1, 135/RE, 3/RE, 3/2, 3/3, 3/4, 3/7, 3/8, 120/RE and 136/RE, falling under the Pixley ka Seme & Chris Hani District Municipalities. The electrical energy generated by the WEF will be fed directly into 400 kV power lines from a small on-site substation. The proposed Umsobomvu WEF study area is largely underlain by continental (fluvial, lacustrine) sediments of the Beaufort Group (Karoo Supergroup). These include (1) latest Permian to earliest Triassic rocks forming the uppermost portion of the Adelaide Subgroup (equivalents of the Balfour Formation of the eastern Main Karoo Basin) that crop out in low-lying, hilly terrain around the periphery of the Klein-Renosterberg massif, as well as (2) Early Triassic sediments of the Katberg Formation (Tarkastad Group) that build the Klein-Renosterberg escarpment and large parts of the upland plateau. -
Early Triassic) of Southern Brazil
Palaeont. afr., 38, 93-98 (2002) FIRST RECORD OF NON-MAMMALIAN CYNODONTS (THERAPSIDA) IN THE SANGA DO CABRAL FORMATION (EARLY TRIASSIC) OF SOUTHERN BRAZIL by Fernando Abdala1, Sergio Dias-da-Silva2 and Juan C. Cisneros3 1Museu de Ciencias e Tecnologia, Pontij(cia Universidade Cat6lica do Rio Grande do Sui, Porto Alegre, RS, Brazil (Current address, Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Private Bag 3, WITS 2050, Johannesburg, South Africa). Email: abdalaf@ geosciences. wits.ac.za 2Instituto de Geociencias, Universidade Federal do Rio Grande do Sui, Campus do Vale, Bento Gonc;alvez, Porto Alegre (91540-000), RS, Brazil 3Museo de Historia Natural de El Salvador, Final C. Los Viveros, Col. Nicaragua, San Salvador, El Salvador. ABSTRACT New material from the Sanga do Cabral Formation in southern Brazil is described, including one distal and three proximal portions of femora, and a distal portion of a humerus. Comparative studies indicate affinities of the proximal femora and the distal humerus with non-mammalian cynodonts, whereas the distal femur cannot be attributed with certainty to this group. These postcranial elements are the first record of non-mammal ian cynodonts for the Sang a do Cabral Formation and demonstrate the presence of small- to medium-sized representatives of this group in the Early Triassic fauna of Brazil. KEYWORDS: Triassic, non-mammalian cynodonts, Sanga do Cabral Formation, Brazil. INTRODUCTION 1981). Isolated and mostly fragmentary fossil remains Permian and Triassic outcrops of the Parana Basin in of small size are found within the intraformational southern Brazil have produced abundant remains of conglomerates, rendering taxonomic identification continental tetrapods (Barberena et al.