REVISED STRATIGRAPHY of the BEAUFORT GROUP in the SOUTHERN KAROO BASIN by B.R
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JVP 26(3) September 2006—ABSTRACTS
Neoceti Symposium, Saturday 8:45 acid-prepared osteolepiforms Medoevia and Gogonasus has offered strong support for BODY SIZE AND CRYPTIC TROPHIC SEPARATION OF GENERALIZED Jarvik’s interpretation, but Eusthenopteron itself has not been reexamined in detail. PIERCE-FEEDING CETACEANS: THE ROLE OF FEEDING DIVERSITY DUR- Uncertainty has persisted about the relationship between the large endoskeletal “fenestra ING THE RISE OF THE NEOCETI endochoanalis” and the apparently much smaller choana, and about the occlusion of upper ADAM, Peter, Univ. of California, Los Angeles, Los Angeles, CA; JETT, Kristin, Univ. of and lower jaw fangs relative to the choana. California, Davis, Davis, CA; OLSON, Joshua, Univ. of California, Los Angeles, Los A CT scan investigation of a large skull of Eusthenopteron, carried out in collaboration Angeles, CA with University of Texas and Parc de Miguasha, offers an opportunity to image and digital- Marine mammals with homodont dentition and relatively little specialization of the feeding ly “dissect” a complete three-dimensional snout region. We find that a choana is indeed apparatus are often categorized as generalist eaters of squid and fish. However, analyses of present, somewhat narrower but otherwise similar to that described by Jarvik. It does not many modern ecosystems reveal the importance of body size in determining trophic parti- receive the anterior coronoid fang, which bites mesial to the edge of the dermopalatine and tioning and diversity among predators. We established relationships between body sizes of is received by a pit in that bone. The fenestra endochoanalis is partly floored by the vomer extant cetaceans and their prey in order to infer prey size and potential trophic separation of and the dermopalatine, restricting the choana to the lateral part of the fenestra. -
On the Stratigraphic Range of the Dicynodont Taxon Emydops (Therapsida: Anomodontia) in the Karoo Basin, South Africa
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Wits Institutional Repository on DSPACE On the stratigraphic range of the dicynodont taxon Emydops (Therapsida: Anomodontia) in the Karoo Basin, South Africa Kenneth D. Angielczyk1*, Jörg Fröbisch2 & Roger M.H. Smith3 1Department of Earth Sciences, University of Bristol, Wills Memorial Building, Queens Road, BS8 1RJ, United Kingdom 2Department of Biology, University of Toronto at Mississauga, 3359 Mississauga Rd., Mississauga, ON, L5L 1C6, Canada 3Divison of Earth Sciences, South African Museum, P.O. Box 61, Cape Town, 8000 South Africa Received 19 May 2005. Accepted 8 June 2006 The dicynodont specimen SAM-PK-708 has been referred to the genera Pristerodon and Emydops by various authors, and was used to argue that the first appearance of Emydops was in the Tapinocephalus Assemblage Zone in the Karoo Basin of South Africa. However, the specimen never has been described in detail, and most discussions of its taxonomic affinities were based on limited data. Here we redescribe the specimen and compare it to several small dicynodont taxa from the Tapinocephalus and Pristerognathus assemblage zones. Although the specimen is poorly preserved, it possesses a unique combination of features that allows it to be assigned confidently to Emydops. The locality data associated with SAM-PK-708 are vague, but they allow the provenance of the specimen to be narrowed down to a relatively limited area southwest of the town of Beaufort West. Strata from the upper Tapinocephalus Assemblage Zone and the Pristerognathus Assemblage Zone crop out in this area, but we cannot state with certainty from which of these biostratigraphic divisions the specimen was collected. -
A New Mid-Permian Burnetiamorph Therapsid from the Main Karoo Basin of South Africa and a Phylogenetic Review of Burnetiamorpha
Editors' choice A new mid-Permian burnetiamorph therapsid from the Main Karoo Basin of South Africa and a phylogenetic review of Burnetiamorpha MICHAEL O. DAY, BRUCE S. RUBIDGE, and FERNANDO ABDALA Day, M.O., Rubidge, B.S., and Abdala, F. 2016. A new mid-Permian burnetiamorph therapsid from the Main Karoo Basin of South Africa and a phylogenetic review of Burnetiamorpha. Acta Palaeontologica Polonica 61 (4): 701–719. Discoveries of burnetiamorph therapsids in the last decade and a half have increased their known diversity but they remain a minor constituent of middle–late Permian tetrapod faunas. In the Main Karoo Basin of South Africa, from where the clade is traditionally best known, specimens have been reported from all of the Permian biozones except the Eodicynodon and Pristerognathus assemblage zones. Although the addition of new taxa has provided more evidence for burnetiamorph synapomorphies, phylogenetic hypotheses for the clade remain incongruent with their appearances in the stratigraphic column. Here we describe a new burnetiamorph specimen (BP/1/7098) from the Pristerognathus Assemblage Zone and review the phylogeny of the Burnetiamorpha through a comprehensive comparison of known material. Phylogenetic analysis suggests that BP/1/7098 is closely related to the Russian species Niuksenitia sukhonensis. Remarkably, the supposed mid-Permian burnetiids Bullacephalus and Pachydectes are not recovered as burnetiids and in most cases are not burnetiamorphs at all, instead representing an earlier-diverging clade of biarmosuchians that are characterised by their large size, dentigerous transverse process of the pterygoid and exclusion of the jugal from the lat- eral temporal fenestra. The evolution of pachyostosis therefore appears to have occurred independently in these genera. -
South Africa's Coalfields — a 2014 Perspective
International Journal of Coal Geology 132 (2014) 170–254 Contents lists available at ScienceDirect International Journal of Coal Geology journal homepage: www.elsevier.com/locate/ijcoalgeo South Africa's coalfields — A 2014 perspective P. John Hancox a,⁎,AnnetteE.Götzb,c a University of the Witwatersrand, School of Geosciences and Evolutionary Studies Institute, Private Bag 3, 2050 Wits, South Africa b University of Pretoria, Department of Geology, Private Bag X20, Hatfield, 0028 Pretoria, South Africa c Kazan Federal University, 18 Kremlyovskaya St., Kazan 420008, Republic of Tatarstan, Russian Federation article info abstract Article history: For well over a century and a half coal has played a vital role in South Africa's economy and currently bituminous Received 7 April 2014 coal is the primary energy source for domestic electricity generation, as well as being the feedstock for the Received in revised form 22 June 2014 production of a substantial percentage of the country's liquid fuels. It furthermore provides a considerable source Accepted 22 June 2014 of foreign revenue from exports. Available online 28 June 2014 Based on geographic considerations, and variations in the sedimentation, origin, formation, distribution and quality of the coals, 19 coalfields are generally recognised in South Africa. This paper provides an updated review Keywords: Gondwana coal of their exploration and exploitation histories, general geology, coal seam nomenclature and coal qualities. With- Permian in the various coalfields autocyclic variability is the norm rather than the exception, whereas allocyclic variability Triassic is much less so, and allows for the correlation of genetically related sequences. During the mid-Jurassic break up Coalfield of Gondwana most of the coal-bearing successions were intruded by dolerite. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Meandering in the Main Karoo Basin, Eastern Cape, South Africa FIELD TRIP LEADERS: Emese Bordy & Goonie Marsh
POST 10 Meandering in the main Karoo Basin, Eastern Cape, South Africa FIELD TRIP LEADERS: Emese Bordy & Goonie Marsh This comprehensive five day trip will take us through, via an unique route in the shortest travel time and distance, a geo-traverse through over 400 million years of South African geological history. The main focus of the field trip is the sedimentary fill of the southern main Karoo Basin, including the nature of some of the major Karoo intrusive and volcanic complexes of the Eastern Cape. Field Trip Leaders: Emese Bordy and Goonie Marsh Start: Port Elizabeth End: Port Elizabeth Dates: 3-8 September 2016 ITINERARY SUGGESTION OF FLIGHT BOOKING FROM CAPE TOWN TO PORT ELIZABETH: SOUTH AFRICAN AIRWAYS FLIGHT 1803 DEPARTING CAPE TOWN AT 07h00, ARRIVING PORT ELIZABETH AT 08h15 Day 1 3 September 2016, Saturday Arrival at Port Elizabeth Airport and transfer to Grahamstown Overnight at the Graham Hotel in Grahamstown BB via Addo Elephant Park Day 2 4 September 2016, Sunday Stop 1 : Overview of 400 million years of South African geological history though the rocks and landscape of Grahamstown, Eastern Cape. Topics covered: Cape and Karoo systems, formation of the main Karoo Basin and Cape Fold Belt, Gondwana breakup, Cenozoic sea level changes. Location : 1820 Settlers' Monument Features to be seen : Relationship of the geology and geomorphology Cape Fold Belt - large-scale geological and geomorphological features: Witteberg quartzite ridges with some mudstone lenses to the south (cut by the N2 national road) & to the north (called Botha’s Ridge) running across the horizon. City bowl overlying the E-W running contact between soft Witteberg mudstones to the south and Dwyka tillites to the north Flat peneplain underlain by hard silcretes (with Joza/ King’s Flat township on it), visible along the northeastern horizon. -
The Sauropodomorph Biostratigraphy of the Elliot Formation of Southern Africa: Tracking the Evolution of Sauropodomorpha Across the Triassic–Jurassic Boundary
Editors' choice The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary BLAIR W. MCPHEE, EMESE M. BORDY, LARA SCISCIO, and JONAH N. CHOINIERE McPhee, B.W., Bordy, E.M., Sciscio, L., and Choiniere, J.N. 2017. The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary. Acta Palaeontologica Polonica 62 (3): 441–465. The latest Triassic is notable for coinciding with the dramatic decline of many previously dominant groups, followed by the rapid radiation of Dinosauria in the Early Jurassic. Among the most common terrestrial vertebrates from this time, sauropodomorph dinosaurs provide an important insight into the changing dynamics of the biota across the Triassic–Jurassic boundary. The Elliot Formation of South Africa and Lesotho preserves the richest assemblage of sauropodomorphs known from this age, and is a key index assemblage for biostratigraphic correlations with other simi- larly-aged global terrestrial deposits. Past assessments of Elliot Formation biostratigraphy were hampered by an overly simplistic biozonation scheme which divided it into a lower “Euskelosaurus” Range Zone and an upper Massospondylus Range Zone. Here we revise the zonation of the Elliot Formation by: (i) synthesizing the last three decades’ worth of fossil discoveries, taxonomic revision, and lithostratigraphic investigation; and (ii) systematically reappraising the strati- graphic provenance of important fossil locations. We then use our revised stratigraphic information in conjunction with phylogenetic character data to assess morphological disparity between Late Triassic and Early Jurassic sauropodomorph taxa. Our results demonstrate that the Early Jurassic upper Elliot Formation is considerably more taxonomically and morphologically diverse than previously thought. -
Tiago Rodrigues Simões
Diapsid Phylogeny and the Origin and Early Evolution of Squamates by Tiago Rodrigues Simões A thesis submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy in SYSTEMATICS AND EVOLUTION Department of Biological Sciences University of Alberta © Tiago Rodrigues Simões, 2018 ABSTRACT Squamate reptiles comprise over 10,000 living species and hundreds of fossil species of lizards, snakes and amphisbaenians, with their origins dating back at least as far back as the Middle Jurassic. Despite this enormous diversity and a long evolutionary history, numerous fundamental questions remain to be answered regarding the early evolution and origin of this major clade of tetrapods. Such long-standing issues include identifying the oldest fossil squamate, when exactly did squamates originate, and why morphological and molecular analyses of squamate evolution have strong disagreements on fundamental aspects of the squamate tree of life. Additionally, despite much debate, there is no existing consensus over the composition of the Lepidosauromorpha (the clade that includes squamates and their sister taxon, the Rhynchocephalia), making the squamate origin problem part of a broader and more complex reptile phylogeny issue. In this thesis, I provide a series of taxonomic, phylogenetic, biogeographic and morpho-functional contributions to shed light on these problems. I describe a new taxon that overwhelms previous hypothesis of iguanian biogeography and evolution in Gondwana (Gueragama sulamericana). I re-describe and assess the functional morphology of some of the oldest known articulated lizards in the world (Eichstaettisaurus schroederi and Ardeosaurus digitatellus), providing clues to the ancestry of geckoes, and the early evolution of their scansorial behaviour. -
The Role of Fossils in Interpreting the Development of the Karoo Basin
Palaeon!. afr., 33,41-54 (1997) THE ROLE OF FOSSILS IN INTERPRETING THE DEVELOPMENT OF THE KAROO BASIN by P. J. Hancox· & B. S. Rubidge2 IGeology Department, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa 2Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa ABSTRACT The Permo-Carboniferous to Jurassic aged rocks oft1:J.e main Karoo Basin ofSouth Africa are world renowned for the wealth of synapsid reptile and early dinosaur fossils, which have allowed a ten-fold biostratigraphic subdivision ofthe Karoo Supergroup to be erected. The role offossils in interpreting the development of the Karoo Basin is not, however, restricted to biostratigraphic studies. Recent integrated sedimentological and palaeontological studies have helped in more precisely defming a number of problematical formational contacts within the Karoo Supergroup, as well as enhancing palaeoenvironmental reconstructions, and basin development models. KEYWORDS: Karoo Basin, Biostratigraphy, Palaeoenvironment, Basin Development. INTRODUCTION Invertebrate remains are important as indicators of The main Karoo Basin of South Africa preserves a facies genesis, including water temperature and salinity, retro-arc foreland basin fill (Cole 1992) deposited in as age indicators, and for their biostratigraphic potential. front of the actively rising Cape Fold Belt (CFB) in Fossil fish are relatively rare in the Karoo Supergroup, southwestern Gondwana. It is the deepest and but where present are useful indicators of gross stratigraphically most complete of several depositories palaeoenvironments (e.g. Keyser 1966) and also have of Permo-Carboniferous to Jurassic age in southern biostratigraphic potential (Jubb 1973; Bender et al. Africa and reflects changing depositional environments 1991). -
A New Late Permian Burnetiamorph from Zambia Confirms Exceptional
fevo-09-685244 June 19, 2021 Time: 17:19 # 1 ORIGINAL RESEARCH published: 24 June 2021 doi: 10.3389/fevo.2021.685244 A New Late Permian Burnetiamorph From Zambia Confirms Exceptional Levels of Endemism in Burnetiamorpha (Therapsida: Biarmosuchia) and an Updated Paleoenvironmental Interpretation of the Upper Madumabisa Mudstone Formation Edited by: 1 † 2 3,4† Mark Joseph MacDougall, Christian A. Sidor * , Neil J. Tabor and Roger M. H. Smith Museum of Natural History Berlin 1 Burke Museum and Department of Biology, University of Washington, Seattle, WA, United States, 2 Roy M. Huffington (MfN), Germany Department of Earth Sciences, Southern Methodist University, Dallas, TX, United States, 3 Evolutionary Studies Institute, Reviewed by: University of the Witwatersrand, Johannesburg, South Africa, 4 Iziko South African Museum, Cape Town, South Africa Sean P. Modesto, Cape Breton University, Canada Michael Oliver Day, A new burnetiamorph therapsid, Isengops luangwensis, gen. et sp. nov., is described Natural History Museum, on the basis of a partial skull from the upper Madumabisa Mudstone Formation of the United Kingdom Luangwa Basin of northeastern Zambia. Isengops is diagnosed by reduced palatal *Correspondence: Christian A. Sidor dentition, a ridge-like palatine-pterygoid boss, a palatal exposure of the jugal that [email protected] extends far anteriorly, a tall trigonal pyramid-shaped supraorbital boss, and a recess †ORCID: along the dorsal margin of the lateral temporal fenestra. The upper Madumabisa Christian A. Sidor Mudstone Formation was deposited in a rift basin with lithofacies characterized orcid.org/0000-0003-0742-4829 Roger M. H. Smith by unchannelized flow, periods of subaerial desiccation and non-deposition, and orcid.org/0000-0001-6806-1983 pedogenesis, and can be biostratigraphically tied to the upper Cistecephalus Assemblage Zone of South Africa, suggesting a Wuchiapingian age. -
Ecca Group) of South Africa: a Preliminary Review of Palaeoniscoid Fishes and Taphonomy
Records of the Western Australian Museum Supplement No. 57: 175-181 (1999). The Permian Whitehill Formation (Ecca Group) of South Africa: a preliminary review of palaeoniscoid fishes and taphonomy Fiona J. Evans 1 and Patrick A. Bender2 I Department of Zoology, University of Stellenbosch, Private Bag Xl, Matieland, 7602, South Africa; email: [email protected] 2 Museum of the Council for Geoscience, Private Bag Xl12, Pretoria, 0001, South Africa Abstract - The Permian (Artinskian) Whitehill Formation (Ecca Group) in South Africa and southern Namibia was deposited as part of a large stratified inland sea which extended from the southern margins of South Africa, and southern Namibia, northern Namibia (Aba-Huab Formation, Huab Basin) to the Iratl Formation of the Parana Basin in Brazil and parts of Uruguay. A common biota exists between these two continents, namely mesosaurid reptiles, pygocephalomorph crustacea, wood, palaeoniscoid fish and insects. The Whitehill Formation also contains plant stem fragments, palynomorphs, coprolites, a cephalochordate, rare sponge spicules and traces of arthropods and fish. The good preservation, particularly of the palaeoniscoid material, from the Whitehill Formation contrasts strongly with the disarticulated scales and spines from northern Namibia and Brazil. Recent collections in the Whitehill Formation near Calvinia and Louriesfontein (Northern Cape Province) of South Africa have demonstrated the necessity for a review of the palaeoniscoid taxa present in these units. Possible new species have also been collected, including a deep-bodied form. INTRODUCTION The extensive stratified inland sea in which the Whitehill Formation (Ecca Group, Karoo Supergroup; - - - - - Waterford Fm Figure 1) was deposited had limited or no access to ------------ the ocean according to reconstructions by Oelofsen Fort Brown Fm and Araujo (1987) and Pickford (1995). -
Taphonomy As an Aid to African Palaeontology*
Palaeont. afr., 24 (1981 ) PRESIDENTIAL ADDRESS: TAPHONOMY AS AN AID TO AFRICAN PALAEONTOLOGY* by C.K. Brain Transvaal Museum, P.O. Box 413, Pretoria 0001 SUMMARY Palaeontology has its roots in both the earth and life sciences. Its usefulness to geology comes from the light which the understanding of fossils may throw on the stratigraphic re lationships of sediments, or the presence of economic deposits such as coal or oil. In biology, the study of fossils has the same objectives as does the study of living animals or plants and such objectives are generally reached in a series of steps which may be set out as follows: STEP I. Discovering what forms of life are, or were, to be found in a particular place at a particular time. Each form is allocated a name and is fitted into a system of classification. These contributions are made by the taxonomist or the systematist. STEP 2. Gaining afuller understanding ofeach described taxon as a living entity. Here the input is from the anatomist, developmental biologist, genetIcIst, physi ologist or ethologist and the information gained is likely to modify earlier decisions taken on the systematic position of the forms involved. STEP 3. Understanding the position ofeach form in the living community or ecosystem. This step is usually taken by a population biologist or ecologist. Hopefully, any competent neo- or palaeobiologist (I use the latter term deliberately in this context in preference to "palaeontologist") should be able to contribute to more than one of the steps outlined above. Although the taxonomic and systematic steps have traditionally been taken in museums or related institutions, it is encouraging to see that some of the steps subsequent to these very basic classificatory ones are now also being taken by museum biol ogists.