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On the Stratigraphic Range of the Dicynodont Taxon Emydops (Therapsida: Anomodontia) in the Karoo Basin, South Africa
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Wits Institutional Repository on DSPACE On the stratigraphic range of the dicynodont taxon Emydops (Therapsida: Anomodontia) in the Karoo Basin, South Africa Kenneth D. Angielczyk1*, Jörg Fröbisch2 & Roger M.H. Smith3 1Department of Earth Sciences, University of Bristol, Wills Memorial Building, Queens Road, BS8 1RJ, United Kingdom 2Department of Biology, University of Toronto at Mississauga, 3359 Mississauga Rd., Mississauga, ON, L5L 1C6, Canada 3Divison of Earth Sciences, South African Museum, P.O. Box 61, Cape Town, 8000 South Africa Received 19 May 2005. Accepted 8 June 2006 The dicynodont specimen SAM-PK-708 has been referred to the genera Pristerodon and Emydops by various authors, and was used to argue that the first appearance of Emydops was in the Tapinocephalus Assemblage Zone in the Karoo Basin of South Africa. However, the specimen never has been described in detail, and most discussions of its taxonomic affinities were based on limited data. Here we redescribe the specimen and compare it to several small dicynodont taxa from the Tapinocephalus and Pristerognathus assemblage zones. Although the specimen is poorly preserved, it possesses a unique combination of features that allows it to be assigned confidently to Emydops. The locality data associated with SAM-PK-708 are vague, but they allow the provenance of the specimen to be narrowed down to a relatively limited area southwest of the town of Beaufort West. Strata from the upper Tapinocephalus Assemblage Zone and the Pristerognathus Assemblage Zone crop out in this area, but we cannot state with certainty from which of these biostratigraphic divisions the specimen was collected. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Tiago Rodrigues Simões
Diapsid Phylogeny and the Origin and Early Evolution of Squamates by Tiago Rodrigues Simões A thesis submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy in SYSTEMATICS AND EVOLUTION Department of Biological Sciences University of Alberta © Tiago Rodrigues Simões, 2018 ABSTRACT Squamate reptiles comprise over 10,000 living species and hundreds of fossil species of lizards, snakes and amphisbaenians, with their origins dating back at least as far back as the Middle Jurassic. Despite this enormous diversity and a long evolutionary history, numerous fundamental questions remain to be answered regarding the early evolution and origin of this major clade of tetrapods. Such long-standing issues include identifying the oldest fossil squamate, when exactly did squamates originate, and why morphological and molecular analyses of squamate evolution have strong disagreements on fundamental aspects of the squamate tree of life. Additionally, despite much debate, there is no existing consensus over the composition of the Lepidosauromorpha (the clade that includes squamates and their sister taxon, the Rhynchocephalia), making the squamate origin problem part of a broader and more complex reptile phylogeny issue. In this thesis, I provide a series of taxonomic, phylogenetic, biogeographic and morpho-functional contributions to shed light on these problems. I describe a new taxon that overwhelms previous hypothesis of iguanian biogeography and evolution in Gondwana (Gueragama sulamericana). I re-describe and assess the functional morphology of some of the oldest known articulated lizards in the world (Eichstaettisaurus schroederi and Ardeosaurus digitatellus), providing clues to the ancestry of geckoes, and the early evolution of their scansorial behaviour. -
The Role of Fossils in Interpreting the Development of the Karoo Basin
Palaeon!. afr., 33,41-54 (1997) THE ROLE OF FOSSILS IN INTERPRETING THE DEVELOPMENT OF THE KAROO BASIN by P. J. Hancox· & B. S. Rubidge2 IGeology Department, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa 2Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa ABSTRACT The Permo-Carboniferous to Jurassic aged rocks oft1:J.e main Karoo Basin ofSouth Africa are world renowned for the wealth of synapsid reptile and early dinosaur fossils, which have allowed a ten-fold biostratigraphic subdivision ofthe Karoo Supergroup to be erected. The role offossils in interpreting the development of the Karoo Basin is not, however, restricted to biostratigraphic studies. Recent integrated sedimentological and palaeontological studies have helped in more precisely defming a number of problematical formational contacts within the Karoo Supergroup, as well as enhancing palaeoenvironmental reconstructions, and basin development models. KEYWORDS: Karoo Basin, Biostratigraphy, Palaeoenvironment, Basin Development. INTRODUCTION Invertebrate remains are important as indicators of The main Karoo Basin of South Africa preserves a facies genesis, including water temperature and salinity, retro-arc foreland basin fill (Cole 1992) deposited in as age indicators, and for their biostratigraphic potential. front of the actively rising Cape Fold Belt (CFB) in Fossil fish are relatively rare in the Karoo Supergroup, southwestern Gondwana. It is the deepest and but where present are useful indicators of gross stratigraphically most complete of several depositories palaeoenvironments (e.g. Keyser 1966) and also have of Permo-Carboniferous to Jurassic age in southern biostratigraphic potential (Jubb 1973; Bender et al. Africa and reflects changing depositional environments 1991). -
A New Late Permian Burnetiamorph from Zambia Confirms Exceptional
fevo-09-685244 June 19, 2021 Time: 17:19 # 1 ORIGINAL RESEARCH published: 24 June 2021 doi: 10.3389/fevo.2021.685244 A New Late Permian Burnetiamorph From Zambia Confirms Exceptional Levels of Endemism in Burnetiamorpha (Therapsida: Biarmosuchia) and an Updated Paleoenvironmental Interpretation of the Upper Madumabisa Mudstone Formation Edited by: 1 † 2 3,4† Mark Joseph MacDougall, Christian A. Sidor * , Neil J. Tabor and Roger M. H. Smith Museum of Natural History Berlin 1 Burke Museum and Department of Biology, University of Washington, Seattle, WA, United States, 2 Roy M. Huffington (MfN), Germany Department of Earth Sciences, Southern Methodist University, Dallas, TX, United States, 3 Evolutionary Studies Institute, Reviewed by: University of the Witwatersrand, Johannesburg, South Africa, 4 Iziko South African Museum, Cape Town, South Africa Sean P. Modesto, Cape Breton University, Canada Michael Oliver Day, A new burnetiamorph therapsid, Isengops luangwensis, gen. et sp. nov., is described Natural History Museum, on the basis of a partial skull from the upper Madumabisa Mudstone Formation of the United Kingdom Luangwa Basin of northeastern Zambia. Isengops is diagnosed by reduced palatal *Correspondence: Christian A. Sidor dentition, a ridge-like palatine-pterygoid boss, a palatal exposure of the jugal that [email protected] extends far anteriorly, a tall trigonal pyramid-shaped supraorbital boss, and a recess †ORCID: along the dorsal margin of the lateral temporal fenestra. The upper Madumabisa Christian A. Sidor Mudstone Formation was deposited in a rift basin with lithofacies characterized orcid.org/0000-0003-0742-4829 Roger M. H. Smith by unchannelized flow, periods of subaerial desiccation and non-deposition, and orcid.org/0000-0001-6806-1983 pedogenesis, and can be biostratigraphically tied to the upper Cistecephalus Assemblage Zone of South Africa, suggesting a Wuchiapingian age. -
Physical and Environmental Drivers of Paleozoic Tetrapod Dispersal Across Pangaea
ARTICLE https://doi.org/10.1038/s41467-018-07623-x OPEN Physical and environmental drivers of Paleozoic tetrapod dispersal across Pangaea Neil Brocklehurst1,2, Emma M. Dunne3, Daniel D. Cashmore3 &Jӧrg Frӧbisch2,4 The Carboniferous and Permian were crucial intervals in the establishment of terrestrial ecosystems, which occurred alongside substantial environmental and climate changes throughout the globe, as well as the final assembly of the supercontinent of Pangaea. The fl 1234567890():,; in uence of these changes on tetrapod biogeography is highly contentious, with some authors suggesting a cosmopolitan fauna resulting from a lack of barriers, and some iden- tifying provincialism. Here we carry out a detailed historical biogeographic analysis of late Paleozoic tetrapods to study the patterns of dispersal and vicariance. A likelihood-based approach to infer ancestral areas is combined with stochastic mapping to assess rates of vicariance and dispersal. Both the late Carboniferous and the end-Guadalupian are char- acterised by a decrease in dispersal and a vicariance peak in amniotes and amphibians. The first of these shifts is attributed to orogenic activity, the second to increasing climate heterogeneity. 1 Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK. 2 Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstraße 43, 10115 Berlin, Germany. 3 School of Geography, Earth and Environmental Sciences, University of Birmingham, Birmingham B15 2TT, UK. 4 Institut -
Taphonomy As an Aid to African Palaeontology*
Palaeont. afr., 24 (1981 ) PRESIDENTIAL ADDRESS: TAPHONOMY AS AN AID TO AFRICAN PALAEONTOLOGY* by C.K. Brain Transvaal Museum, P.O. Box 413, Pretoria 0001 SUMMARY Palaeontology has its roots in both the earth and life sciences. Its usefulness to geology comes from the light which the understanding of fossils may throw on the stratigraphic re lationships of sediments, or the presence of economic deposits such as coal or oil. In biology, the study of fossils has the same objectives as does the study of living animals or plants and such objectives are generally reached in a series of steps which may be set out as follows: STEP I. Discovering what forms of life are, or were, to be found in a particular place at a particular time. Each form is allocated a name and is fitted into a system of classification. These contributions are made by the taxonomist or the systematist. STEP 2. Gaining afuller understanding ofeach described taxon as a living entity. Here the input is from the anatomist, developmental biologist, genetIcIst, physi ologist or ethologist and the information gained is likely to modify earlier decisions taken on the systematic position of the forms involved. STEP 3. Understanding the position ofeach form in the living community or ecosystem. This step is usually taken by a population biologist or ecologist. Hopefully, any competent neo- or palaeobiologist (I use the latter term deliberately in this context in preference to "palaeontologist") should be able to contribute to more than one of the steps outlined above. Although the taxonomic and systematic steps have traditionally been taken in museums or related institutions, it is encouraging to see that some of the steps subsequent to these very basic classificatory ones are now also being taken by museum biol ogists. -
Proceedings of the 18Th Biennial Conference of the Palaeontological Society of Southern Africa Johannesburg, 11–14 July 2014
Proceedings of the 18th Biennial Conference of the Palaeontological Society of Southern Africa Johannesburg, 11–14 July 2014 Table of Contents Letter of Welcome· · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · 63 Programme · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · 64 · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · 66 Hand, K.P., Bringing Two Worlds Together: How Earth’s Past and Present Help Us Search for Life on Other Planets · · · · · · · 66 · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · 67 Erwin, D.H., Major Evolutionary Transitions in Early Life: A Public Goods Approach · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · 67 Lelliott, A.D., A Survey of Visitors’ Experiences of Human Origins at the Cradle of Humankind, South Africa· · · · · · · · · · · · · · 68 Looy, C., The End-Permian Biotic Crisis: Why Plants Matter · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · · 69 Reed, K., Hominin Evolution and Habitat: The Importance of Analytical Scale · · · · · · · · · · · · · · · · · · · · · · · · · · · · · -
Stratigraphy, Sedimentary Facies and Diagenesis of the Ecca Group, Karoo Supergroup in the Eastern Cape, South Africa
STRATIGRAPHY, SEDIMENTARY FACIES AND DIAGENESIS OF THE ECCA GROUP, KAROO SUPERGROUP IN THE EASTERN CAPE, SOUTH AFRICA By Nonhlanhla Nyathi Dissertation submitted in fulfilment of the requirements for the degree of Master of Science In Geology FACULTY OF SCIENCE AND AGRICULTURE UNIVERSITY OF FORT HARE SUPERVISOR: PROFESSOR K. LIU CO-SUPERVISOR: PROFESSOR O. GWAVAVA APRIL 2014 DECLARATION I, Nonhlanhla Nyathi, hereby declare that the research described in this dissertation was carried out in the field and under the auspices of the Department of Geology, University of Fort Hare, under the supervision of Professor K. Liu and Prof. O. Gwavava. This dissertation and the accompanying photographs represent original work by the author, and have not been submitted, in whole or in part, to any other university for the purpose of a higher degree. Where reference has been made to the work of others, it has been dully acknowledged in the text. N. NYATHI Date signed: 12/04/2014 Place signed: Alice ACKNOWLEDGEMENTS The author is indebted to Professor K. Liu for his guidance, knowledge on all aspects of the project, constant supervision and help in doing field wok. Professor O. Gwavava is much appreciated for being my co-supervisor and helping me obtain financial support from the Goven Mbeki Research Development Centre. I am deeply grateful for the emotional support and encouragement from my parents and family. I express my profound gratitude to Mr Edwin Mutototwa and Mr Eric Madifor always taking time to read through my dissertation. The Rhodes University Geology laboratory technicians are thanked for their assistance in the making of thin sections. -
Comparing Middle Permian and Early Triassic Environments: Mud Aggregates As a Proxy for Climate Change in the Karoo Basin, South Africa
Colby College Digital Commons @ Colby Honors Theses Student Research 2011 Comparing Middle Permian and Early Triassic Environments: Mud Aggregates as a Proxy for Climate Change in the Karoo Basin, South Africa Bryce Pludow Colby College Follow this and additional works at: https://digitalcommons.colby.edu/honorstheses Part of the Geology Commons Colby College theses are protected by copyright. They may be viewed or downloaded from this site for the purposes of research and scholarship. Reproduction or distribution for commercial purposes is prohibited without written permission of the author. Recommended Citation Pludow, Bryce, "Comparing Middle Permian and Early Triassic Environments: Mud Aggregates as a Proxy for Climate Change in the Karoo Basin, South Africa" (2011). Honors Theses. Paper 622. https://digitalcommons.colby.edu/honorstheses/622 This Honors Thesis (Open Access) is brought to you for free and open access by the Student Research at Digital Commons @ Colby. It has been accepted for inclusion in Honors Theses by an authorized administrator of Digital Commons @ Colby. COMPARING MIDDLE PERMIAN AND EARLY TRIASSIC ENVIRONMENTS: MUD AGGREGATES AS A PROXY FOR CLIMATE CHANGE IN THE KAROO BASIN, SOUTH AFRICA B. Amelia Pludow „11 A Thesis Submitted to the Faculty of the Geology Department of Colby College in Fulfillment of the Requirements for Honors in Geology Waterville, Maine May, 2011 COMPARING MIDDLE PERMIAN AND EARLY TRIASSIC ENVIRONMENTS: MUD AGGREGATES AS A PROXY FOR CLIMATE CHANGE IN THE KAROO BASIN, SOUTH AFRICA Except where reference is made to the work of others, the work described in this thesis is my own or was done in collaboration with my advisory committee B. -
Ska Core and Phase 1 Development Area, Great Karoo, Northern Cape - Palaeontological Heritage
1 SKA CORE AND PHASE 1 DEVELOPMENT AREA, GREAT KAROO, NORTHERN CAPE - PALAEONTOLOGICAL HERITAGE John E. Almond PhD (Cantab.) Natura Viva cc, PO Box 12410 Mill Street, Cape Town 8010, RSA [email protected] March 2016 EXECUTIVE SUMMARY The SKA core and Phase 1 study area in the Northern Cape is situated on the north-western margins of the Main Karoo Basin. The region is underlain by marine / lacustrine to terrestrial sediments of the Karoo Supergroup (Ecca and Lower Beaufort Groups) of Early to Middle Permian age. The fossil record of the Ecca Group here is dominated by a wide range of non-marine, subaqueously-generated trace fossils. A key trace fossil locality in the Tierberg Formation (Farm Brassefontein) occurs c. 65 km southwest of Brandvlei. Well-preserved mesosaurid reptiles and bony fish are recorded from the Whitehill Formation but are rare, while the Whitehill mudrocks are extensively intruded by dolerite in this region. Classic outcrops of storm-dominated shelf sediments of the Waterford Formation (previously Carnarvon Formation) are located in the Kareeberge of the Williston – Carnarvon area. These shelf successions contain a distinctive trace fossil assemblage as well as fairly common blocks of petrified driftwood. Middle Permian continental sediments of the Lower Beaufort Group (Abrahamskraal Formation) between Williston and Fraserburg have yielded a small number of vertebrate fossils of the Tapinocephalus Assemblage Zone including therapsids, pareiasaur John E. Almond (2016) Natura Viva cc 2 reptiles and bony fish. Important trace fossil assemblages, including tetrapod trackways, are associated with sandstone palaeosurfaces such as the well-known Gansfontein site near Fraserburg. -
Remagnetizations in Late Permian and Early Triassic Rocks from Southern Africa and Their Implications for Pangeareconstructions
412 Earth and Planetary Science Letters, 79 (1986) 412-418 Elsevier Science Publishers B.V., Amsterdam - Printed in The Netherlands 151 Remagnetizations in Late Permian and Early Triassic rocks from southern Africa and their implications for Pangea reconstructions Martha M. Ballard ‘, Rob Van der Voo * and I.W. HZlbich 2 ’ Department of Geological Sciences, University of Michigan, Am Arbor, MI 48109 (U.S.A.) ’ Department of Geology, University of Stellenbosch, Stellenbosclr (Republic of South Africa) Received March 31,1985; revised version received June 24, 1986 -A paleomagnetic study of late Paleozoic and early Mesozoic sedimentary rocks from southern Africa suggests wide-spread remagnetization of these rocks. Samples of the Mofdiahogolo Formation in Botswana and of the Lower Beaufort Group in South Africa were treated using thermal, alternating field and chemical demagnetization. The Mofdiahogolo redbeds show a univectoral decay of the remanence revealing a characteristic direction of D = 340°, I = - 58O, k = 64, a9s = 12O. The Lower Beaufort sandstones, using thermal and alternating field demagnetization, show a very similar direction of D = 337O, I = -63”, k = 91, ags = 6O. A fold test on the Beaufort rocks is negative indicating that this magnetization is secondary and acquired after the Permo-Triassic Cape Belt folding event. Previous studies have reported similar directions in the Upper Beaufort redbeds as well as in the Kenyan Maji ya Chumvi Formation of Early Triassic age. The poles of these studies have been used in testing the validity of the various Pangea reconstructions for the Late Permian and the Early Triassic. Our results suggest that these poles may also be based on remagnetized data and that their use to document the position of Gondwana in Pangea reconstructions should be treated with caution.