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Cambios De Numeración Cambios de numeración Occasional papers from the Herbarium Greuter – 4 Código Internacional de Nomenclatura para algas, hongos y plantas (Código de Shenzhen) vii Traducción al español de la versión oficial en inglés autorizada por la International Association for Plant Taxonomy Publicado por: Stiftung Herbarium Greuter (Fundación Herbario Greuter) Englerallee 24B 14195 Berlín, Alemania © Stiftung Herbarium Greuter ISBN: 978-3-9820137-4-9 eISBN: 978-3-9820137-0-1 Impresión: Texto traducido por: Werner Greuter, Herbarium Mediterraneum, Università degli Studi di Palermo, Italy; and Botanischer Garten & Botanisches Museum Berlin-Dahlem, Freie Universität Berlin, Germany ([email protected]). Rosa Rankin Rodríguez, Jardín Botánico Nacional, Universidad de La Haba- na, Cuba ([email protected]). Con la colaboración de: Luis Alberto Parra Sánchez, Avda. Padre Claret 7, 5º G, 09400 Aranda de Duero, Burgos, España ([email protected]). Juan Bautista Martínez Laborde, Departamento de Biotecnología-Biología Vege- tal, Escuela Técnica Superior de Ingeniería Agronómica, Alimentaria y de Biosis- temas, Universidad Politécnica de Madrid, España (juanbau.martinez@ upm.es). Alina Freire Fierro, Apartado Postal Personal 17-17-793, Quito, Ecuador (ali- [email protected]). Renée Hersilia Fortunato, CONICET, Instituto de Recursos Biológicos, CIRN- INTA, N. Repetto y Los Reseros s/n, Hurlingham 1686, Facultad de Agronomía y Ciencias Agroalimentarias, Universidad de Morón, Prov. de Buenos Aires, Argen- tina ([email protected]). Comité editorial Código Internacional de Nomenclatura para algas, hongos y plantas (Código de Shenzhen) adoptado por el decimonoveno Congreso Internacional de Botánica Shenzhen, China, julio de 2017 Preparado y editado por NICHOLAS J. TURLAND, Presidente JOHN H. WIERSEMA, Secretario, y FRED R. BARRIE WERNER GREUTER, DAVID L. HAWKSWORTH PATRICK S. HERENDEEN SANDRA KNAPP WOLF-HENNING KUSBER DE-ZHU LI KAROL MARHOLD TOM W. MAY JOHN MCNEILL ANNA M. MONRO JEFFERSON PRADO MICHELLE J. PRICE GIDEON F. SMITH Miembros del Comité Editorial Edición en español a cargo de WERNER GREUTER y ROSA RANKIN RODRÍGUEZ Berlín, 2018 4–10 CCambios de numeración Edición original: Regnum Vegetabile Volumen 159 ISSN 0080-0694 Koeltz Scientific Books, 61479 Glashütten, Germany http://www.koeltz.com/product.aspx?pid=212180 © 2018, International Association for Plant Taxonomy (Europe) ISBN 978-3-946583-16-5 Ilustración de la carátula: La ilustración de la carátula fue dibujada por Pollyanna von Knorring (Estocolmo). El diseño se basa en el tema ginkgo que caracterizaba el XIX Congreso Botánico Internacional en Shenzhen, China, en julio de 2017 e incluye representantes de los tres grupos mencionados en el título del Código, además de los organismos fósiles. Las plantas están representadas por hojas de Ginkgo biloba L.; las algas, por los talos de la rodofícea marina antártica Ne- reoginkgo adiantifolia Kylin; los hongos, por el basidiomiceto Bartheletia para- doxa G. Arnaud ex Scheuer & al. formando manchas oscuras en las hojas caídas de G. biloba, con sus teliósporas y basidios; y los fósiles, por la reconstrucción de Ginkgo yimaensis Zhi-Yan Zhou & Bo-Le Zhang del Jurásico de China. Téngase en cuenta que las imágenes no están a escala. Comité editorial COMITÉ EDITORIAL PARA ESTE CÓDIGO Nicholas J. Turland, Botanischer Garten und Botanisches Museum Berlin, Freie Univer- sität Berlin, Königin-Luise-Str. 6–8, 14195 Berlin, Germany; [email protected] (Chair) John H. Wiersema, USDA, Agricultural Research Service, National Germplasm Re- sources Laboratory, Bldg. 003, BARC-West, Beltsville, Maryland 20705, U.S.A.; [email protected] (Secretary) Fred R. Barrie, Missouri Botanical Garden, P.O. Box 299, St. Louis, Missouri 63166- 0299, U.S.A.; address for correspondence: Department of Science and Education, The Field Museum, 1400 S. Lake Shore Drive, Chicago, Illinois 60605, U.S.A.; fbar- [email protected] Werner Greuter, Herbarium Mediterraneum, c/o Orto Botanico, Via Lincoln 2/A, 90123 Palermo, Italy; and Botanischer Garten und Botanisches Museum Berlin, Freie Universi- tät Berlin, Königin-Luise-Str. 6–8, 14195 Berlin, Germany; [email protected] David L. Hawksworth, Comparative Plant and Fungal Biology, Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3AE, U.K.; and Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, U.K.; and Jilin Agricultural Univer- sity, Changchun, Jilin 130118, China; [email protected] Patrick S. Herendeen, Chicago Botanic Garden, 1000 Lake Cook Road, Glencoe, Illinois 60022, U.S.A.; [email protected] Sandra Knapp, Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, U.K.; [email protected] Wolf-Henning Kusber, Botanischer Garten und Botanisches Museum Berlin, Freie Uni- versität Berlin, Königin-Luise-Str. 6–8, 14195 Berlin, Germany; [email protected] De-Zhu Li, Kunming Institute of Botany, Chinese Academy of Sciences, 132 Lanhei Road, Heilongtan, Kunming, Yunnan 650201, P. R. China; [email protected] Karol Marhold, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, 845 23 Bratislava, Slovak Republic; and Department of Botany, Facul- ty of Science, Charles University, Benátská 2, 128 01 Praha, Czech Republic; ka- [email protected] Tom W. May, Royal Botanic Gardens Victoria, 100 Birdwood Avenue, Melbourne, Victo- ria 3004, Australia; [email protected] John McNeill, Royal Botanic Garden, Edinburgh, 20A Inverleith Row, Edinburgh EH3 5LR, U.K.; and Royal Ontario Museum, Toronto, Canada; [email protected] Anna M. Monro, Australian National Herbarium, Centre for Australian National Biodi- versity Research, GPO Box 1700, Canberra ACT 2601, Australia; anna.monro@envi- ronment.gov.au Jefferson Prado, Instituto de Botânica, Av. Miguel Estéfano 3687, CEP 04301-902, São Paulo, SP, Brazil; [email protected] Michelle J. Price, Conservatoire et Jardin botaniques de la Ville de Genève (CJBG), che- min de l’Impératrice 1, 1292 Chambésy, Genève, Switzerland; [email protected] Gideon F. Smith, Department of Botany, P.O. Box 77000, Nelson Mandela University, Port Elizabeth, 6031 South Africa; and Centre for Functional Ecology, Departamento de Ciências da Vida, Universidade de Coimbra, 3001-455 Coimbra, Portugal; smithgi- [email protected] v vi Foto delaSección CONTENIDO La Sección de Nomenclatura del XIX Congreso Internacional de Botánica, Shenzhen, China, 20 de julio de 2017, con el Directorio de Nomenclatura sentado en la primera fila (de izquierda a derecha): Li Zhang, Yun-Fei Deng (Relatores), Anna M. Monro (asistente de los Relatores), John H. Wiersema (Vicepresidente), Nicholas J. Turland (Ponente General), Sandra Knapp (Presidenta), Renée H. Fortunato, John McNeill, Werner Greuter, Gideón F. Smith y Karen L. Wilson (Vicepresidentes). (Foto: Li Zhang, por cortesía de la IAPT) Contenido CONTENIDO Introducción de los traductores ix Prefacio xiv Clave para los cambios de numeración de Artículos, Notas y Recomendaciones xxxix Fechas importantes en el Código lii Preámbulo (Pre. 1–14) 1 División I Principios (I–VI) 4 División II Reglas y Recomendaciones (Art. 1–62) 5 Capítulo I Los taxones y sus rangos (Art. 1–5) 5 Capítulo II Estatus, tipificación y prioridad de los nombres (Art. 6–15) 9 Sección 1 Definiciones de estatus (Art. 6) 9 Sección 2 Tipificación (Art. 7–10) 15 Sección 3 Prioridad (Art. 11–12) 40 Sección 4 Limitación del principio de prioridad (Art. 13–15) 49 Capítulo III Nomenclatura de los taxones según su rango (Art. 16–28) 58 Sección 1 Nombres de los taxones de rango superior a familia (Art. 16– 17) 58 Sección 2 Nombres de las familias, subfamilias, tribus y subtribus (Art. 18–19) 61 Sección 3 Nombres de los géneros y de las subdivisiones de género (Art. 20–22) 67 Sección 4 Nombres de las especies (Art. 23) 74 Sección 5 Nombres de los taxones de rango inferior a especie (taxones infraespecíficos) (Art. 24–27) 80 Sección 6 Nombres de los organismos cultivados (Art. 28) 85 Capítulo IV Publicación efectiva (Art. 29–31) 87 Sección 1 Condiciones de la publicación efectiva (Art. 29–30) 87 Sección 2 Fechas de la publicación efectiva (Art. 31) 96 Capítulo V Publicación válida de los nombres (Art. 32–45) 98 Sección 1 Disposiciones generales (Art. 32–37) 98 Sección 2 Nombres de taxones nuevos (Art. 38–40) 110 Sección 3 Combinaciones nuevas, nombres en rango nuevo, nombres de reemplazo (Art. 41) 122 Sección 4 Nombres en grupos particulares (Art. 42–45) 130 Capítulo VI Citación (Art. 46–50) 136 Sección 1 Citas de autor (Art. 46–50) 136 vii Contenido Sección 2 Recomendaciones generales sobre la citación (Rec. 50A–G) 151 Capítulo VII Rechazo de nombres (Art. 51–58) 154 Capítulo [VIII] Nombres de los hongos con ciclo de vida pleomórfico (Art. F.8) 169 Capítulo IX Ortografía y género gramatical de los nombres (Art. 60–62)170 Sección 1 Ortografía (Art. 60–61) 170 Sección 2 Género gramatical (Art. 62) 187 Capítulo F Nombres de organismos que se consideran hongos (Art. F.1–9) 190 Sección 1 Limitación al principio de la prioridad (Art. F.1–3) 192 – Punto de partida nomenclatural (Art. F.1) 192 – Nombres protegidos (Art. F.2) 192 – Nombres sancionados (Art. F.3) 193 Sección 2 Publicación válida y tipificación de los nombres (Art. F.4–5) 196 – Términos que denotan rango mal ubicados (Art. F.4) 196 – Registro de nombres y actos nomenclaturales ( Art. F.5) 197 Sección 3 Rechazo de los nombres (Art. F.6–7) 199 Sección 4 Nombres de hongos con ciclo de vida pleomórfico (Art. F.8) 200 Sección 5
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