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Ent11 3 317 324 Bolshakov10rus.Pm6 Russian Entomol. J. 11 (3): 317324 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2002 Microlepidoptera Òóëüñêîé îáëàñòè. 10. Ìèííî-÷åõëèêîâûå è äëèííîóñûå ìîëè (Lepidoptera: Adeloidea: Incurvariidae, Adelidae) Microlepidoptera of Tula Area. 10. Incurvariid and Adelid moths (Lepidoptera: Adeloidea) Ë.Â. Áîëüøàêîâ L.V. Bolshakov Îáúåäèíåíèå Òóëüñêèé îáëàñòíîé èñòîðèêî-àðõèòåêòóðíûé è ëèòåðàòóðíûé ìóçåé (îòäåë ïðèðîäû), óë.Ñîâåòñêàÿ, 68, Òóëà 300000 Ðîññèÿ. Association Tula Regional Historical, Architectural and Literary Museum (Department of Nature), Sovetskaya Str. 68, Tula 300000 Russia. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: ìèííî-÷åõëèêîâûå è äëèííîóñûå ìîëè (Lepidoptera: Adeloidea: Incurvariidae, Adelidae), Òóëüñêàÿ îáëàñòü, ôàóíèñòèêà, ñïèñîê, çîíàëüíàÿ è áèîòîïè÷åñêàÿ ïðèóðî÷åííîñòü, ýêîëîãè÷åñ- êèå äàííûå. KEY WORDS: Incurvariid and Adelid moths (Lepidoptera: Adeloidea), Tula Area, faunistics, check-list, zonal and biotopic distribution, ecological data. ÐÅÇÞÌÅ: Íà îñíîâàíèè ìíîãîëåòíèõ èññëåäîâà-  ñïèñêå È.Â.Øìûòîâîé [1997] èñïîëüçîâàëàñü íèé â Òóëüñêîé îáëàñòè ïðåäñòàâëåíû ñïèñîê âèäîâ è ñèñòåìàòèêà è íîìåíêëàòóðà, ïðèíÿòàÿ â ðàáîòå ýêîëîãî-ôàóíèñòè÷åñêàÿ êëàññèôèêàöèÿ ìèííî-÷åõ- À.Ê. Çàãóëÿåâà [1978].  ïîñëåäóþùèõ ðàáîòàõ ïî ëèêîâûõ è äëèííîóñûõ ìîëåé (Lepidoptera: Adeloidea: Adeloidea (íàïðèìåð, [Razowski, 1978, 1987; Êóïðèÿ- Incurvariidae, Adelidae). Ñïèñîê âèäîâ íàñ÷èòûâàåò íîâ, 1995; Êîçëîâ, 1997]) ïîÿâëÿþòñÿ áîëåå èëè ìå- 20 âèäîâ, èç êîòîðûõ 6 ïðèâîäÿòñÿ âïåðâûå äëÿ òåð- íåå ñóùåñòâåííûå ñèñòåìàòè÷åñêèå èçìåíåíèÿ, à òàê- ðèòîðèè îáëàñòè.Adela cupriacella (Hbn., [1819]) âïåð- æå óòî÷íåíû ïðèîðèòåòíûå âèäîâûå íàçâàíèÿ è àâ- âûå ïðèâîäèòñÿ äëÿ Öåíòðà Åâðîïåéñêîé Ðîññèè. òîðñòâà ïåðâîîïèñàíèé. Îäíàêî îáùåïðèíÿòàÿ ñèñ- Ðàññìàòðèâàþòñÿ îñîáåííîñòè ýêîëîãèè âèäîâ. òåìà íàäñåìåéñòâà åùå íå âïîëíå ñëîæèëàñü. Áîëü- øèíñòâî îòå÷åñòâåííûõ àâòîðîâ íå ïðèíèìàþò íåêî- ABSTRACT: On the basis of the long-term research òîðûå íîâøåñòâà, ïðåäëîæåííûå â 197080-å ãîäû in Tula Area the list of species of Incurvariid and Adelid ðÿäîì çàïàäíîåâðîïåéñêèõ àâòîðîâ (îñâåùåííûå É. moths (Lepidoptera: Adeloidea) is introduced. The list Ðàçîâñêèì [Razowski, 1987]), íî âûãëÿäåâøèå íå î÷åíü includes 20 species, 6 of them are reported from the Area åñòåñòâåííî. Ñ ó÷åòîì ïåðå÷èñëåííûõ ðàáîò íàìè (â for the first time. Adela cupriacella (Hbn., [1819]) are ðàáî÷åì ïîðÿäêå) â ñèñòåìàòèêó áàçîâîãî ñïèñêà ðå- recorded in the Center of European Russia for the first ãèîíàëüíîé ôàóíû ââîäÿòñÿ íåêîòîðûå èçìåíåíèÿ. time. Pecularities of ecology are discussed. Íàì ïðåäñòàâëÿåòñÿ, ÷òî ïåðåâîä ðîäà Lampronia Stph., 1829 â íåàðêòè÷åñêóþ ãðóïïó Prodoxidae (èëè Ôàóíà è ýêîëîãèÿ ìîëåîáðàçíûõ ÷åøóåêðûëûõ Prodoxinae) íå îïðàâäàí (ïðåæäå âñåãî, â ñâÿçè ñ íàäñåìåéñòâà Adeloidea â Åâðîïåéñêîé ÷àñòè è äðó- îñîáåííîñòÿìè áèîëîãèè è ñòðîåíèÿ ãåíèòàëèé âè- ãèõ ðåãèîíàõ Ðîññèè èçó÷åíû î÷åíü ôðàãìåíòàðíî. äîâ [Davis, 1967; Çàãóëÿåâ, 1978; Razovski, 1978]). Cïèñîê ñåìåéñòâ Incurvariidae è Adelidae Òóëüñêîé Ýòîò ðîä ìû îñòàâëÿåì â ñåìåéñòâå Incurvariidae. è Êàëóæñêîé îáëàñòåé [Øìûòîâà, 1997], ôàêòè÷åñ- Ïîäñåìåéñòâî Nematopogoninae ñ ðîäîì Nematopo- êè íàñ÷èòûâàþùèé 19 âèäîâ, îãðàíè÷èâàåòñÿ ñâåäå- gon Zell., 1839 ïåðåâîäèòñÿ ïî ñîâîêóïíîñòè ïðè- íèÿìè î ìåñòàõ ñáîðîâ è ñðîêàõ ëåòà áàáî÷åê. Ïðè çíàêîâ â ñåìåéñòâî Adelidae.  ïîäñåìåéñòâå Adeli- ýòîì äëÿ Òóëüñêîé îáëàñòè ïðèâîäèëîñü 14 âèäîâ (ñ nae, íàðÿäó ñ ðîäîì Adela Latr., 1796, íàìè ñîõðàíÿ- îøèáî÷íûì îïðåäåëåíèåì "Nemotois" fasciellus (F., åòñÿ õîðîøî îáîñîáëåííûé ïî âíåøíèì ïðèçíàêàì 1775) è îòäåëüíûìè íåòî÷íîñòÿìè, êàñàþùèìèñÿ è ýêîëîãî-ýòîëîãè÷åñêèì îñîáåííîñòÿì (õîòÿ è íå íîìåíêëàòóðû, ìåñòîíàõîæäåíèé, ñðîêîâ ëåòà). Áî- âûäåëÿþùèéñÿ ïî ñòðîåíèþ ãåíèòàëèé) ðîä Cauchas ëåå ïîäðîáíûé ðåãèîíàëüíûé îáçîð ýòîé íåáîëü- Zell., 1839. Òàêñîí Nemophora Ill. et Hoffm., 1798 øîé, íî î÷åíü îáîñîáëåííîé ãðóïïû, àêòóàëåí â (âûäåëåííûé ïî æèëêîâàíèþ çàäíèõ êðûëüåâ) ìû îáùåýíòîìîëîãè÷åñêîì è ñîçîáèîëîãè÷åñêîì àñ- ðàññìàòðèâàåì êàê ñáîðíûé ïîäðîä â ðîäå Adela. ïåêòàõ.  íàñòîÿùåé ðàáîòå ïðåäñòàâëåíû ðåçóëüòà- Òàêñîí Nemotois Hbn., [1825] (âûäåëåííûé ïî ñòðî- òû èññëåäîâàíèé, îñíîâàííûõ íà îáðàáîòàííîì àâ- åíèþ ãëàç ñàìöîâ) ìîã áû èìåòü êàê ìèíèìóì ïîä- òîðîì ìàòåðèàëå ïî Òóëüñêîé îáëàñòè, ëèòåðàòóð- ðîäîâîé ñòàòóñ, íî ýòî íàçâàíèå ÿâëÿåòñÿ îáúåêòèâ- íûõ äàííûõ è îðèãèíàëüíîé ìåòîäèêå. íûì ñèíîíèìîì Nemophora, ò.å. íåïðèãîäíî. Äåé- 318 Ë.Â.Áîëüøàêîâ Ðèñ.1. Êàðòà ìåñò ñáîðà ìàòåðèàëà (îáîçíà÷åíû öèôðàìè) è áèîãåîãðàôè÷åñêèõ âûäåëîâ (îáîçíà÷åíû áóêâàìè) Òóëüñêîé îáëàñòè. Ëåñíàÿ çîíà: À ïîäçîíà õâîéíî-øèðîêîëèñòâåííûõ ëåñîâ; Á ïîäçîíà øèðîêîëèñòâåííûõ ëåñîâ (Á1 ëåñà Îêñêî- Óïèíñêîãî âîäîðàçäåëà, Á2 îñòðîâíûå ëåñà). Ëåñîñòåïíàÿ çîíà:  ïîäçîíà ñåâåðíîé ëåñîñòåïè ñåâåðíîãî âàðèàíòà; à ïîäçîíà ñåâåðíîé ëåñîñòåïè þæíîãî âàðèàíòà (òèïè÷íîé ëåñîñòåïè). Ìåñòîíàõîæäåíèÿ ðàñøèôðîâàíû â òåêñòå. Fig. 1. Map of localities (numbers) and biogeographical units (litters) of Tula Area. Forest zone: À subzone of coniferous- broadleaved forests; Á subzone of broadleaved forests (Á1 forests of Oka-Upa watershed, Á2 forest islands); Forest-steppe zone:  subzone of northern forest-steppe of northern kind; à subzone of northern forest-steppe of southern kind (typical forest- steppe). Localities are given in the text. ñòâèòåëüíî, èçìåí÷èâîñòü æèëêîâàíèÿ íàáëþäàåò- âûì. Îí õðàíèòñÿ â îñíîâíîì â ôîíäàõ Îáúåäèíå- ñÿ è â ðÿäå äðóãèõ ðîäîâ ðàññìàòðèâàåìîé ãðóïïû, íèÿ Òóëüñêèé îáëàñòíîé èñòîðèêî-àðõèòåêòóðíûé îñîáåííî Nematopogon è Cauchas. è ëèòåðàòóðíûé ìóçåé è â ëè÷íûõ êîëëåêöèÿõ,  ðåçóëüòàòå ýíòîìîëîãè÷åñêèõ èññëåäîâàíèé, íåêîòîðûå ýêçåìïëÿðû ïåðåäàíû â Çîîëîãè÷åñêèé îñíîâíûå àñïåêòû êîòîðûõ ðàññìàòðèâàëèñü ðàíåå èíñòèòóò ÐÀÍ. Òàêñîíîìè÷åñêàÿ èäåíòèôèêàöèÿ [Ñâèðèäîâ, Áîëüøàêîâ, 1997; Áîëüøàêîâ, 1998, ïðîâåäåíà ïðåèìóùåñòâåííî Ë.Â. Áîëüøàêîâûì 1999à,á, 2000, 2001], íà òåððèòîðèè Òóëüñêîé îá- ïî ðÿäó ñïåöèàëüíûõ ðàáîò [Wojtusiak, 1972; Çàãó- ëàñòè îáíàðóæåíî 4 âèäà Incurvariidae è 16 âèäîâ ëÿåâ, 1978; Razowski, 1978; Êîçëîâ, 1997]. Ìíîãèå Adelidae, àííîòèðîâàííûé ñïèñîê è ýêîëîãî-ôàó- ýêçåìïëÿðû ðàíåå îïðåäåëÿëèñü È.Â. Øìûòîâîé íèñòè÷åñêèé àíàëèç êîòîðûõ ïðåäñòàâëåíû íèæå. (Êàëóæñêèé îáúåäèíåííûé êðàåâåä÷åñêèé ìóçåé) Ïðè ýòîì 1 âèä Incurvariidae è 5 âèäîâ Adelidae â õîäå ðàáîòû íàä ñïèñêîì âèäîâ Òóëüñêîé è Êà- âïåðâûå ïðèâîäÿòñÿ äëÿ òåððèòîðèè îáëàñòè, à 1 ëóæñêîé îáëàñòåé [1997]. Íåêîòîðûå ñëîæíûå ýê- âèä Adelidae âïåðâûå äëÿ Öåíòðà Åâðîïåéñêîé çåìïëÿðû ñàìîê áûëè ëþáåçíî ïðîñìîòðåíû Ì.Â. Ðîññèè. Êîçëîâûì (Óíèâåðñèòåò Òóðêó, Ôèíëÿíäèÿ). Ðàç- Èññëåäîâàííûé ìàòåðèàë ñîáðàí ñ 1977 ïî 2001 íîñòîðîííþþ ïîìîùü àâòîðó îêàçûâàëè òàêæå À.Ê. ãã. ïðåèìóùåñòâåííî Ë.Â. Áîëüøàêîâûì, ÷àñòè÷- Çàãóëÿåâ è À.Ë. Ëüâîâñêèé (Çîîëîãè÷åñêèé èíñòè- íî Ñ.À. Àíäðååâûì, Â.Í. è Í.Â. Êðûëîâûìè, òóò ÐÀÍ), À.Â. Ñâèðèäîâ è Å.Ì. Àíòîíîâà (Çîîëî- À.Ô. Ëàêîìîâûì, Ä.À. Ñàôðîíîâûì, Ñ.À. Ðÿáî- ãè÷åñêèé ìóçåé ÌÃÓ). Microlepidoptera Òóëüñêîé îáëàñòè. 10. 319 Àâòîð âûðàæàåò ñåðäå÷íóþ áëàãîäàðíîñòü ïåðå- äîâ, ðàñïðîñòðàíåííûõ ïîâñåìåñòíî. Ïåðå÷èñëÿ- ÷èñëåííûì âûøå ñïåöèàëèñòàì, êîëëåãàì è èññëå- åì 59 ìåñòîíàõîæäåíèé, óïîìèíàåìûõ â íàñòîÿ- äîâàòåëÿì-ýíòóçèàñòàì. ùåé ðàáîòå: Èç 20 âèäîâ íàøåãî ñïèñêà 1 ãîëàðêòè÷åñêèé 3 ×åêàëèí, 4 Ñåëþêîâî, 5 Ðæàâåö, 6 (îäíàêî âîçìîæíî, ÷òî íà ñåâåðîàìåðèêàíñêèé êîí- Ïåñî÷åíñêèé, 7 Êàìûøåíêà, 9 Ôóðñîâî, 13 òèíåíò îí çàâåçåí), 4 òðàíñïàëåàðêòè÷åñêèå, 4 Íèêîëî-Ãàñòóíü, 13à Ìàòþõèíñêèé, 17 Îêîðîêîâî, çàïàäíî-öåíòðàëüíîïàëåàðêòè÷åñêèå è 11 çàïàä- 17à Ãîâîðåíêè, 17á Ìèõàéëîâêà (Ñóâîðîâñêèé ð-í), íîïàëåàðêòè÷åñêèå (ñêîðåå âñåãî, íåêîòîðûå èç íèõ 18à Ïëàòîâî, 18á Áàòüêîâî, 20 ×åðåïåòü, 21 Áîëîòî (Ñóâîðîâñêèé ð-í), 23 Âàðóøèöû, 24 Ìåä- ñî âðåìåíåì îáíàðóæàòñÿ, ïî êðàéíåé ìåðå, íà þãå âåæêà, 26 Ïîïîâêà (×åðíñêèé ð-í), 35 Ãëèíèùè, 35à Çàïàäíîé Ñèáèðè). Äîëãîòíûå ñîñòàâëÿþùèå àðåà- Îëåíèíêà, 44 Íîâîå Õàíèíî, 44à Íîâîàëåêñàíä- ëîâ 16 âèäîâ ìîæíî îõàðàêòåðèçîâàòü, êàê òåìïå- ðîâêà, 47 Áåðíèêè, 51 Òóëà, 54 Êîñàÿ Ãîðà, 55 ðàòíûå (ñ èçâåñòíûìè ñóæåíèÿìè â êîíòèíåíòàëü- Ñêóðàòîâñêèé (Ëåíèíñêèé ð-í), 56 ßñíàÿ Ïîëÿíà, 62 íûõ ðåãèîíàõ Ñåâåðíîé Àçèè), à 4 âèäîâ êàê Èíøèíñêèé, 64 Ñàäêè, 64à Ñâîáîäà, 64á Êðþêîâ- ñóááîðåàëüíûå. Ýòè ñâåäåíèÿ âðÿä ëè ìîæíî ñ÷è- êà (Ùåêèíñêèé ð-í), 72 Äàëìàòîâêà, 73 Ðåâÿêèíî, 74 òàòü îêîí÷àòåëüíûìè â ñâÿçè ñî ñëàáîé èçó÷åííîñ- Ñòðóíèíî, 75 Øåìåòîâî, 77â Õîâàíñêîå, 82 òüþ äàííîé ãðóïïû ìèêðî÷åøóåêðûëûõ âî ìíîãèõ Øóëüãèíî, 90 Âåëåãîæ, 91 Ïîëåíîâî, 93 Ïðèîê- ðåãèîíàõ óìåðåííîé Àçèè. Âåñüìà âîçìîæíî, ÷òî ïî ñêàÿ, 93à Ëàíüøèíñêèé, 96 Õîðîøåâêà, 97 Âîñåì- ñêîå, 103 Äåäèëîâñêèå Âûñåëêè, 103à Ïîòåòèíî, òåððèòîðèè íàøåãî ðåãèîíà ïðîõîäÿò ñåâåðíûå ãðà- 103á Ôèëàòîâî, 103â Äüÿêîâî, 107 Ðàññâåò, 114 íèöû àðåàëîâ íåêîòîðûõ ñóááîðåàëüíûõ âèäîâ. Îò- Êàìûíèíî, 115 Ïëàâñê, 116 Àãðîëåñ, 125 ìåòèì, ÷òî â Òóëüñêîé îáëàñòè îáíàðóæåíî îêîëî Íîâîìîñêîâñê, 135 Ùó÷üå, 143 Âîäÿíîå Ïîëå, 145 80% âèäîâ Adeloidea, óêàçàííûõ â öèòèðóåìîé ëèòå- Òàòèíêè, 146 Êóëèêîâêà, 149 Ãðèáîåäîâî, 157 ðàòóðå äëÿ Öåíòðà Åâðîïåéñêîé Ðîññèè. Âÿçîâî, 158 Øèëîâî.  ïðåäëàãàåìîì ñïèñêå îáùàÿ ïîðÿäêîâàÿ íóìåðàöèÿ. Çíàêîì * îòìå÷åí âèä, âïåðâûå ïðèâî- Ñïèñîê âèäîâ äèìûé äëÿ Öåíòðà Åâðîïåéñêîé Ðîññèè. Ñèñòåìà è íîìåíêëàòóðà ðàññìîòðåíû âûøå. Èç ñèíîíèìîâ ÑÅÌÅÉÑÒÂÎ INCURVARIIDAE ÌÈÍÍÎ-×ÅÕ- ïðèâîäÿòñÿ òîëüêî ñàìûå ðàñïðîñòðàíåííûå. Äëÿ ËÈÊÎÂÛÅ ÌÎËÈ âèäîâ, ðàíåå óêàçàííûõ äëÿ Òóëüñêîé îáëàñòè, äàíû ññûëêè íà ïóáëèêàöèþ È.Â. Øìûòîâîé [1997]. 1. Incurvaria oehlmanniella (Hbn., 1796) Àðåàëîãè÷åñêèå õàðàêòåðèñòèêè âèäîâ (ïî íî- [Øìûòîâà, 1997] ìåíêëàòóðå Ê.Á. Ãîðîäêîâà [1984]) äàþòñÿ íà îñíî- Òðàíñåâðàçèàòñêèé òåìïåðàòíûé âèä. âàíèè êàê âûøåóïîìÿíóòûõ èñòî÷íèêîâ, òàê è íî- Ìàòåðèàë:
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