8. Marine Ecology
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JMS 70 1 031-041 Eyh003 FINAL
PHYLOGENY AND HISTORICAL BIOGEOGRAPHY OF LIMPETS OF THE ORDER PATELLOGASTROPODA BASED ON MITOCHONDRIAL DNA SEQUENCES TOMOYUKI NAKANO AND TOMOWO OZAWA Department of Earth and Planetary Sciences, Nagoya University, Nagoya 464-8602,Japan (Received 29 March 2003; accepted 6June 2003) ABSTRACT Using new and previously published sequences of two mitochondrial genes (fragments of 12S and 16S ribosomal RNA; total 700 sites), we constructed a molecular phylogeny for 86 extant species, covering a major part of the order Patellogastropoda. There were 35 lottiid, one acmaeid, five nacellid and two patellid species from the western and northern Pacific; and 34 patellid, six nacellid and three lottiid species from the Atlantic, southern Africa, Antarctica and Australia. Emarginula foveolata fujitai (Fissurellidae) was used as the outgroup. In the resulting phylogenetic trees, the species fall into two major clades with high bootstrap support, designated here as (A) a clade of southern Tethyan origin consisting of superfamily Patelloidea and (B) a clade of tropical Tethyan origin consisting of the Acmaeoidea. Clades A and B were further divided into three and six subclades, respectively, which correspond with geographical distributions of species in the following genus or genera: (AÍ) north eastern Atlantic (Patella ); (A2) southern Africa and Australasia ( Scutellastra , Cymbula-and Helcion)', (A3) Antarctic, western Pacific, Australasia ( Nacella and Cellana); (BÍ) western to northwestern Pacific (.Patelloida); (B2) northern Pacific and northeastern Atlantic ( Lottia); (B3) northern Pacific (Lottia and Yayoiacmea); (B4) northwestern Pacific ( Nipponacmea); (B5) northern Pacific (Acmaea-’ânà Niveotectura) and (B6) northeastern Atlantic ( Tectura). Approximate divergence times were estimated using geo logical events and the fossil record to determine a reference date. -
Aus Dem Institut Für Polarökologie Der Christian-Albrechts-Universität Zu Kiel
Aus dem Institut für Polarökologie der Christian-Albrechts-Universität zu Kiel ECOLOGICAL STUDIES ON THE TROPICAL LIMPET Cellana testudinaria (Linnaeus, 1758): INFLUENCE OF ENVIRONMENTAL FACTORS ON THE ROCKY SHORE BENTHOS OF THE BIG KAI ISLAND, SOUTHEAST MOLLUCCAS, INDONESIA Dissertation zur Erlangung des Doktorgrades der Mathematisch-Naturwissenschaftlichen Fakultät der Christian-Albrechts-Universität zu Kiel Vorgelegt von Abraham Seumel Khouw KIEL 2003 „Gedruckt mit Unterstützung des Deutschen Akademischen Austauschdienstes“ Referent: Priv. –Doz. Dr. D. Piepenburg Korreferent: Prof. Dr. M. Spindler Tag der mündlichen Prüfung: 4 December 2003 Zum Druck genehmigt: Kiel, den ................................ Der Dekan ECOLOGICAL STUDIES ON THE TROPICAL LIMPET Cellana testudinaria (LINNAEUS, 1758) : INFLUENCE OF ENVIRONMENTAL FACTORS ON THE ROCKY SHORE BENTHOS OF THE BIG KAI ISLAND, SOUTHEAST MOLLUCCAS, INDONESIA Dedication This thesis is dedicated to : my beloved wife Costavina Litamahuputty and our son Winfield Bierhoff Khouw Da ist das Meer – so unendlich groß und weit, unzählbar sind die Tiere darin, große wie kleine Psalm 104, 25 Es lobe IHN Himmel und Erde, die Meere mit allem, was sich darin regt. Psalm 69, 35 © A.S.Khouw ACKNOWLEDGEMENTS ACKNOWLEDGEMENTS So many people have worked so hard and supported me so much to make this thesis possible. Words alone cannot express my appreciation. I am truly grateful for all their help and encouragement. First and foremost, I would like to thank Priv. Doz. Dr. D. Piepenburg who supervised me and to whom I will be eternally indebted for the surveillance, guidance, support and opportunity that he has given me. I hope that one day I will be able to return all his efforts to my students in Indonesia. -
Differences in Sterol Composition of Gonads of the Lottiid Limpets
Journal of Oleo Science Copyright ©2011 by Japan Oil Chemists’ Society J. Oleo Sci. 60, (10) 501-504 (2011) NOTE Differences in Sterol Composition of Gonads of the Lottiid Limpets Nipponacmea concinna and Nipponacmea fuscoviridis from Northeastern Japan Hideki Kawashima1* , Masao Ohnishi2 and Satoshi Ogawa3 1 Bioscience Laboratory, Miyako College, Iwate Prefectural University (1-5-1 Kanan, Miyako, Iwate 027-0039, JAPAN) 2 Department of Food Science, Obihiro University of Agriculture and Veterinary Medicine (2-11 Inada-cho, Obihiro, Hokkaido 080-8555, JAPAN) 3 Department of Chemistry and Bioengineering, Faculty of Engineering, Iwate University (4-3-5 Morioka, Iwate 020-8551, JAPAN) Abstract: This is the first report on the sterol composition in Nipponacmea concinna and Nipponacmea fuscoviridis, 2 dominant species of lottiid limpets. There were signifi cant differences in sterol composition between male and female gonads of the limpets. Previous studies have shown that zymostenol and zymosterol are major lipid components of male gonads of the nacellid limpets Cellana grata and Cellana toreuma. In contrast, in this study, only trace amounts of zymosterol were detected in male gonads of N. fuscoviridis. Key words: gastropod, limpet, Nipponacmea concinna, Nipponacmea fuscoviridis, sterol composition 1 INTRODUCTION gonads(the oocytes and sperm)of N. fuscoviridis13)are Limpets, which are among the most common mollusks of currently being used in a developmental study on fertiliza- the class Gastropoda living in most intertidal rock shores tion in invertebrate marine organisms, very little is known worldwide, are model invertebrate animals for ecological about the composition and distribution of sterols in these 2 and physiological research1, 2)as well as for environmental limpet species. -
DNA Barcoding Reveal Patterns of Species Diversity Among
www.nature.com/scientificreports OPEN DNA barcoding reveal patterns of species diversity among northwestern Pacific molluscs Received: 04 April 2016 Shao’e Sun, Qi Li, Lingfeng Kong, Hong Yu, Xiaodong Zheng, Ruihai Yu, Lina Dai, Yan Sun, Accepted: 25 August 2016 Jun Chen, Jun Liu, Lehai Ni, Yanwei Feng, Zhenzhen Yu, Shanmei Zou & Jiping Lin Published: 19 September 2016 This study represents the first comprehensive molecular assessment of northwestern Pacific molluscs. In total, 2801 DNA barcodes belonging to 569 species from China, Japan and Korea were analyzed. An overlap between intra- and interspecific genetic distances was present in 71 species. We tested the efficacy of this library by simulating a sequence-based specimen identification scenario using Best Match (BM), Best Close Match (BCM) and All Species Barcode (ASB) criteria with three threshold values. BM approach returned 89.15% true identifications (95.27% when excluding singletons). The highest success rate of congruent identifications was obtained with BCM at 0.053 threshold. The analysis of our barcode library together with public data resulted in 582 Barcode Index Numbers (BINs), 72.2% of which was found to be concordantly with morphology-based identifications. The discrepancies were divided in two groups: sequences from different species clustered in a single BIN and conspecific sequences divided in one more BINs. In Neighbour-Joining phenogram, 2,320 (83.0%) queries fromed 355 (62.4%) species-specific barcode clusters allowing their successful identification. 33 species showed paraphyletic and haplotype sharing. 62 cases are represented by deeply diverged lineages. This study suggest an increased species diversity in this region, highlighting taxonomic revision and conservation strategy for the cryptic complexes. -
Appendix 13.5 Marine Ecology Field Survey Result Excluding CWD
Expansion of Hong Kong International Airport into a Three-Runway System Environmental Impact Assessment Report Appendix 13.5 Marine Ecology Field Survey Result Excluding CWD 308875/ENL/ENL/03/07/C March 2014 P:\Hong Kong\ENL\PROJECTS\308875 3rd runway\03 Deliverables\07 Final EIA Report\Appendices\Ch 13 Marine Ecology\Appendix 13.5 Marine Ecological Field Survey Result (Excluding CWD).doc 1 Expansion of Hong Kong International Airport into a Three-Runway System Environmental Impact Assessment Report 1. Baseline Conditons of Subtidal Shore and Coral Communities The marine waters in the North Western Water Control Zone support both subtidal hard and soft bottom assemblages. The locations with coral communities of conservation importance recorded are shown in the habitat maps Drawing No. MCL/P132/EIA/13-014 to Drawing No. MCL/P132/EIA/13-020 . 1.1 Subtidal Hard Bottom Assemblages Spot dive surveys and rapid ecological assessments (REAs) at 16 coral survey points for hard bottom coral were undertaken between August 2012and September 2013. Dates for the hard-bottom and soft-bottom coral spot-check dive surveys and REAs are given in Table 1.1. Table 1-1: Dates for coral-check dive surveys and REAs (hard-bottom and soft-bottom) Hard-bottom Soft-bottom Location Date Location Date D1 9 Aug 2012 C1 10 May 2013 D2 9 Aug 2012 C2 14 May 2013 D3 9 Aug 2012 C3 14 May 2013 D4 9 Aug 2012 C4 9 May 2013 D5 9 Aug 2012 C5 9 May 2013 D6 9 Aug 2012 C6 24 May 2013 D7 9 Aug 2012 C7 24 May 2013 D8 9 Aug 2012 C8 10 May 2013 D9 2 Sept 2013 C9 10 May 2013 D10 31 Jul 2013 C10 10 May 2013 D11 31 Jul 2013 C11 24 May 2013 D12 4 Aug 2013 C12 10 May 2013 D13 4 Aug 2013 C13 14 May 2013 D14 31 Jul 2013 C14 14 May 2013 D15 11 Sept 2013 C15 21 May 2013 D16 2 Sept 2013 C16 21 May 2013 - C17 9 May 2013 - C18 9 May 2013 - C19 21 May 2013 - SC2 21 May 2013 - SC10 15 May 2013 - SC12 10 May 2013 Detailed findings of the hard bottom coral dive surveys are provided in Annex A1. -
Larval Dispersal Dampens Population Fluctuation and Shapes the Interspecific Spatial Distribution Patterns of Rocky Title Intertidal Gastropods
Larval dispersal dampens population fluctuation and shapes the interspecific spatial distribution patterns of rocky Title intertidal gastropods Sahara, Ryosuke; Fukaya, Keiichi; Okuda, Takehiro; Hori, Masakazu; Yamamoto, Tomoko; Nakaoka, Masahiro; Noda, Author(s) Takashi Ecography, 38, 1-9 Citation https://doi.org/10.1111/ecog.01354 Issue Date 2015 Doc URL http://hdl.handle.net/2115/62537 Rights The definitive version is available at www.blackwell-synergy.com Type article (author version) File Information 150723ecography.pdf Instructions for use Hokkaido University Collection of Scholarly and Academic Papers : HUSCAP Larval dispersal dampens population fluctuation and shapes the interspecific spatial distribution patterns of rocky intertidal gastropods Ryosuke Sahara1, Keiichi Fukaya2, Takehiro Okuda3, Masakazu Hori4, Tomoko Yamamoto5, Masahiro Nakaoka6, and Takashi Noda1* 1Faculty of Environmental Science, Hokkaido University, N10W5, Kita-ku, Sapporo, Hokkaido 060-0810 Japan 2The Institute of Statistical Mathematics, 10-3 Midoricho, Tachikawa, Tokyo 190-8562 Japan 3National Research Institute of Far Seas Fisheries, Fisheries Research Agency, 2-12-4, Fukura, Kanazawa-ku, Yokohama 236-8648 Japan 4National Research Institute of Fisheries and Environment of Inland Sea, Fisheries Research Agency, Maruishi 2-17-5, Hatsukaichi, Hiroshima 739-0452 Japan 5Faculty of Fisheries, Kagoshima University, Shimoarata 4-50-20, Kagoshima, Kagoshima 890-0056 Japan 6Akkeshi Marine Station, Field Science Centre for the Northern Biosphere, Hokkaido University, Aikappu, Akkeshi, Hokkaido 088-1113 Japan *Corresponding author: Takashi NODA; email: [email protected] 1 Abstract Many marine benthic invertebrates pass through a planktonic larval stage whereas others spend their entire lifetimes in benthic habitats. Recent studies indicate that non-planktonic species show relatively greater fine-scale patchiness than do planktonic species, but the underlying mechanisms remain unknown. -
407072.F1.Pdf
Supplementary Table 1 Fragment Primer Primer sequence 28S rRNA small 28SF 5'-GAC CCG TCT TGA AGC ACG-3' 28SR 5'-CCA CAG CGC CAG TTC TGC TTA C-3' 28S rRNA large 28SF2 5'-ACC TAT TCT CAA ACT TTA AAT GG-3' 28SR2 5'-GAC TTC CCT TAC CTA CAT-3' 18S rRNA part A 18Sa2.0 5'-ATG GTT GCA AAG CTG AAA C-3' 18S9R 5'-GAT CCT TCC GCA GGT TCA CCT AC-3' 18S rRNA part B 18S1F 5'-TAC CTG GTT GAT CCT GCC AGT AG-3' 18S5R 5'-CTT GGC AAA TGC TTT CGC-3' 18S rRNA part C 18S3F 5'-GTT CGA TTC CGG AGA GGG A-3' 18Sbi 5'-GAG TCT CGT TCG TTA TCG GA-3' 16S rRNA 16Sa 5'-CGC CTG TTT ATC AAA AAC AT-3' 16Sb 5'-CTC CGG TTT GAA CTC AGA TCA-3' H3 H3aF 5'-ATG GCT CGT ACC AAG CAG ACV GC-3' H3aR 5'-ATA TCC TTR GGC ATR ATR GTG AC-3' PCR conditions Initial step 94°C 6min Denaturation 94°C 1min Annealing 50°C 1min Elongation 72°C 1min 30sec Final elongation 72°C 6min Cycles 45 Supplementary Table 2 Species 18S 28S1 28S2 28S3 28S4 H3 COI 16S Outgroups Annelida Eunice pennata AY040684 AY340391 DQ779731 AF321418 Tubifex tubifex GQ355437 GQ355465 GQ355465 EF179544 EU117545 Brachiopoda Eohemithiris grayi AF025936 AY839242 AY839242 AY839242 Lingula adamsi U08329 Terebratalia transversa FJ196115 AF342802 AF342802 AF342802 AF342802 FJ196085 Terebratulina retusa U08324 AY839244 AY839244 AY839244 DQ779768 AF334238 Cycliophora Symbion americanus EF142068 EF142087 EF142087 EF140778 EF140771 Symbion pandora Y14811 AY218133 AY218084 Entoprocta Barentsia gracilis FJ196109 AY210456 AY210456 AY210456 AY210456 FJ196079 Loxosomella murmanica AY218100 DQ279950 DQ279950 DQ279950 DQ279950 AY218150 Pedicellina -
Characterization and Quantification of Biochemical Components of Selected Molluscs Species from the Kerala Coast
Characterization and quantification of biochemical components of selected Molluscs species from the Kerala coast Thesis submitted to Cochin University of Science and Technology in partial fulfilment of the requirements for the degree of Doctor of Philosophy in Environmental Chemistry Under the Faculty of Marine Sciences by Ragi A. S. Reg. No. 4224 Department of Chemical Oceanography School of Marine Sciences Cochin University of Science and Technology Kochi – 682016 November, 2017 Characterization and quantification of biochemical components of selected Molluscs species from the Kerala coast Ph. D. Thesis under the Faculty of Marine Sciences Author Ragi A. S. Research Scholar Department of Chemical Oceanography School of Marine Sciences Cochin University of Science and Technology Kochi - 682016 Email: [email protected] Supervising Guide Dr. S. Muraleedharan Nair Professor Department of Chemical Oceanography School of Marine Sciences Cochin University of Science and Technology Kochi - 682016 Email: [email protected] Department of Chemical Oceanography School of Marine Sciences Cochin University of Science and Technology Kochi - 682016 November, 2017 DEDICATION To my Parents, teachers, husband, friends and beloved son for always supporting, helping and standing by me. COCHIN UNIVERSITY OF SCIENCE AND TECHNOLOGY DEPARTMENT OF CHEMICAL OCEANOGRAPHY Fine Arts Avenue Kochi - 682016, India Tel: 91-484-2383131(O) Dr. S. Muraleedharan Nair 91-484-2863402(O) Fax: 91-848-2374164 Professor E-mail: [email protected] This is to certify that the thesis entitled “Characterization and quantification of biochemical components of selected Molluscs species from the Kerala coast” is an authentic record of the research work carried out by Mrs. Ragi A. S. -
Similarity of Rocky Intertidal Assemblages Along the Pacific Coast of Japan : Effects of Spatial Scales and Geographic Title Distance
Similarity of rocky intertidal assemblages along the Pacific coast of Japan : effects of spatial scales and geographic Title distance Author(s) Nakaoka, M.; Ito, N.; Yamamoto, T.; Okuda, T.; Noda, T. Ecological Research, 21(3), 425-435 Citation https://doi.org/10.1007/s11284-005-0138-6 Issue Date 2006-05 Doc URL http://hdl.handle.net/2115/13476 Rights The original publication is available at www.springerlink.com Type article (author version) File Information Nakaoka_et_al_2nd_ver.pdf Instructions for use Hokkaido University Collection of Scholarly and Academic Papers : HUSCAP Nakaoka et al. 1 Masahiro Nakaoka, Norihiko Ito, Tomoko Yamamoto, Takehiro Okuda, Takashi 2 Noda 3 4 Similarity of rocky intertidal assemblages along the Pacific 5 Coast of Japan: Effects of spatial scales and geographic 6 distance 7 8 9 10 M. Nakaoka (Corresponding author), N. Ito 11 Graduate School of Science and Technology, Chiba University, Inage, Chiba 263-8522, 12 Japan 13 E-mail: [email protected] 14 Tel. +81-43-2903929 15 Fax +81-43-2903929 16 17 T. Yamamoto 18 Faculty of Fisheries Sciences, Kagoshima University, Kagoshima, Japan 19 20 T. Okuda, T. Noda 21 Graduate School of Fisheries Sciences, Hokkaido University, Hakodate, Japan 22 23 24 1 Nakaoka et al. 24 Abstract Factors and processes affecting community structures operate at various 25 spatial and temporal scales. We analyzed how similarities of rocky intertidal 26 assemblages vary at different spatial scales using a nested, hierarchical sampling design. 27 Intertidal assemblages consisting of algae, sessile animals and mobile animals were 28 censused on five rock walls at each of five shores chosen within each of six regions 29 along the Pacific coat of Japan encompassing 1800 km coastlines. -
Agreement No. CE32/2011 (CE) Planning and Engineering Study On
Project: Agreement No. CE32/2011 (CE) Planning and Engineering Study on the Remaining Development in Tung Chung Appendix 9.12 - Full list of Intertidal Faunal Species recorded in Tung Chung Bay Page 1 of 8 Protection status / No. Division/ Phylum Class Order Family Genus/Species Level of concern 1 Cnidaria Anthozoa Actiniaria Haliplanellidae Haliplanella lineata 2 Cnidaria Scyphomedusae Semaeostomeae Cyaneidae Cyanea nozakii 3 Annelida Polychaeta Phyllodocida Nereidae Dendronereis sp. 4 Annelida Polychaeta Phyllodocida Polynoidae Harmothoe sp. 5 Annelida Polychaeta Sabellida Serpulidae Hydroides sp. 6 Sipuncula - - - Unidentified Peanut worm* 7 Mollusca Bivalvia Arcoida Arcidae Barbatia virescens 8 Mollusca Bivalvia Arcoida Arcidae Scapharca cornea 9 Mollusca Bivalvia Mytiloida Mytilidae Modiolus philippinarum 10 Mollusca Bivalvia Mytiloida Mytilidae Perna viridis 11 Mollusca Bivalvia Mytiloida Mytilidae Septifer virgatus 12 Mollusca Bivalvia Ostreoida Ostreidae Saccostrea cucullata 13 Mollusca Bivalvia Pholadomyoida Laternulidae Laternula truncata 14 Mollusca Bivalvia Pterioida Isognomonidae Isognomon sp. 15 Mollusca Bivalvia Pterioida Pteriidae Pinctada sp. 16 Mollusca Bivalvia Veneroida Mesodesmatidae Caecella chinensis 17 Mollusca Bivalvia Veneroida Psammobiidae Soletellina diphos 18 Mollusca Bivalvia Veneroida Solenidae Solen sp. 19 Mollusca Bivalvia Veneroida Veneridae Anomalocardia flexuosa 20 Mollusca Bivalvia Veneroida Veneridae Anomalocardia squamosa 21 Mollusca Bivalvia Veneroida Veneridae Cyclina sinensis 22 Mollusca Bivalvia -
Acdsee Print
FAR EASTERN BRANCH OF THE RUSSIAN ACADEMY OF SCIENCES Institute of Marine Biology FAR EASTERN NATIONAL UNIVERSITY Educational Center for Basic Marine Biota Research FAR EAST MALACOLOGICAL SOCIETY - « » ABSTRACTS OF THE CONFERENCE MOLLUSKS OF THE NORTHEASTERN ASIA AND NORTHERN PACIFIC: BIODIVERSITY, ECOLOGY, BIOGEOGRAPHY AND FAUNAL HISTORY October 4 6, 2004, Vladivostok, Russia - : , , 4 6 2004 ., , Vladivostok Dalnauka 2004 594 « - : , , - », 4 6 2004 ., , / . - . : , 2004. 170 . Abstracts of the Conference Mollusks of the Northeastern Asia and North- ern Pacific: Biodiversity, Ecology, Biogeography and Faunal History , October 4 6, 2004, Vladivostok, Russia / Comp. by O. Ya. Semenikhina. Vladivostok: Dal- nauka, 2004. 170 p. « » Conference was organized with financial support of the Far Eastern Branch of the Russian Academy of Sciences and Research and Educational Center for Basic Marine Biota Research of the Far Eastern National University ISBN © , 2004 © , 2004 Abstracts of the conference «Mollusks of the Northeastern Asia and Northern Pacific: Biodiversity, Ecology, Biogeography and Faunal History». Vladivostok, October 4 6, 2004 CONTENTS Konstantin A. Lutaenko. Tenth anniversary of the Russian Far East Malaco- logical Society . ... 11 Dmitry O. Alexeyev. Recent status of the systematics of the family Loligi- nidae (Cephalopoda) .. 13 Svetlana I. Andreeva, Nikolay I. Andreev. Speciation by means of acclimati- zation in bivalve molluscs Syndosmya segmentum Recluz of the Aral Sea ... 14 Alim P. Anisimov, Natalya P. Tokmakova, Irina A. Kirsanova, Natalya E. Zumchenko, Evgenia V. Tabakova, Aleksandra A. Brovar, Natalya A. Galimulina. To the problem of somatic polyploidy in mollusks: Estimation of DNA content in somatic cells nuclei of bivalves . ... 17 Maxim V. Astakhov, Anastasya G. Rodina. Bivalvia: To the question of the dependence of some growth characteristics from sexual belonging of indivi- duals . -
The Shell Structure of the Recent Patellogastropoda (Mollusca: Gastropoda)
Paleontological Research, vol. 9, no. 2, pp. 143–168, June 30, 2005 6 by the Palaeontological Society of Japan The shell structure of the Recent Patellogastropoda (Mollusca: Gastropoda) TAKESHI FUCHIGAMI1 AND TAKENORI SASAKI2 1Department of Earth and Planetary Sciences, Faculty of Science, University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033 2The University Museum, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033 ([email protected]) Received March 25, 2005; Revised manuscript accepted March 28, 2005 Abstract. The shell microstructure of 44 species belonging to 19 genera and 5 families of Patellogas- tropoda was observed by scanning electron microscopy on the basis of material mainly from the Northwest Pacific. As a result, 17 microstructures of prismatic, crossed, and lamellar structures were recognized. The comparison among species revealed 20 shell structure groups which are defined by microstructures and shell layer arrangement. The relations between taxa and shell structural composition indicate that the Recent patellogastropods generally have distinctive and stable shell structures at the genus level. This high level of consistency provides a firm basis for the application of shell structural characters to identify fossil pa- tellogastropods. However, the evolutionary process of microstructures and homology across different shell layers are mostly ambiguous in the absence of robust phylogeny and undoubted positional criteria for comparison. More studies from phylogenetic, ontogenetic and mineralogical viewpoints