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Insecta, Coleoptera, Carabidae, Pseudomorphinae)

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Insecta, Coleoptera, Carabidae, Pseudomorphinae) DOI: 10.18195/issn.0312-3162.24(2).2008.151-180 Records of the Western Australian l'vIuseum 24: 151-180 (2008). New and rare pseudomorphine species from Western Australia (Insecta, Coleoptera, Carabidae, Pseudomorphinae) Martin Baehr Zoologische Staatssammlung, Mi.inchhausenstr. 21, 0-81247 Mi.inchen, Germany. Email: martin.baehr(iDzsm.mwn.de Abstract - Eight new species of the carabid genera Sphallomorpha Westwood and Adelotopus Hope are described from VVestern Australia: Sphallomorpha parinterioris, S. permutata, S. szitoi, S. t7avorufil, S. c(ypeosetosa, AdeIotopus pi/barae, A. basalis and A. minutus. Records from Western Australia of the following rare or poorly documented species are also dealt with: Sphallomorpha froggatti (Macleay), S. faIIax (Westwood), S. polysetosa Baehr, S. guttigera (Newman), S. maClJ1ata (Newman), S. uniformis Baehr, S. flavopicea Baehr, Adelotopus brevipennis Macleay, A. puncticolIis angustemaculatus Baehr, A. rubiginosus Newman, A. houstoni Baehr, A. adustus Baehr, A. coriaceus Baehr, A. paroensis Castelnau, Cainogenion ipsoides occiclentale Baehr, and Paussotropus cylindricus (Chaudoir). For SphaIlomorpha t7avopicea Baehr and Aclelotopus houstoni Baehr the male genitalia are described and figured for the first time. All new species and those to which new taxonomic information is added, are inserted in the respective general keys. The Western Australian distributions of most mentioned species are figured in maps. Keywords: taxonomy, new species, SphaIIomorpha, Adelotopus, Cainogenion, Paussotropus INTRODUCTION dry country with but limited or scattered tree A recent collecting trip through the interior of growth pseudomorphine beetles can be rather Western Australia and visits to Western Australian neglected may be responsible. Fortunately, Western institutions and collections revealed a number of Australian authorities (Western Australian new and rare species of the pseudomorphine genera Museum, Department of Conservation and Land 5phallomorpha Westwood, Adelotopus Hope, Management, Western Australian Department of Cainogenion Notman, and Paussotropus Westwood Agriculture) during the past years conducted a that are described or mentioned in the present paper. number of surveys in certain parts of Western During the visit also several common and well Australia which revealed, inter alia, a number of documented species were collected or identified interesting pseudomorphine beetles. However, as (5phallomorpha castelnaui Reiche, 5. hydroporoides these beetles can best be sampled mainly by use of Westwood, 5. suturalis Germar, Adelotopus laevis specialized methods (pulling down bark into an Macleay, A. gyrinoides Hope) which are not umbrella, fogging bark), this group certainly is mentioned further in this paper. For information underrepresented in the samples from the previous about these species vide the revisions of the Goldfields and Wheat Belt surveys and the present Australian Pseudomorphinae (Baehr 1992, 1997). Pilbara survey. In spite of the previous revisions which included The scope of the present paper hence is not only almost all material available in collections when to describe the new species and to make available they were written, the pseudomorphine fauna of additional information on species diversity and Western Australia, in particular that of the interior, distribution, but also to draw the attention of probably is less well known than the faunas of most collectors to this beetle group and to the other parts of Australia. This deficiency may be arboricolous and corticolous beetle fauna of simply due to the large size of the country which Western Australia in generat and not only to the makes reasonable or even systematic sampling of beetle fauna but also the verv rich fauna of other the fauna almost impossible, but also the invertebrates that live on trees, even in the semiarid impression of putative collectors that in a largely and arid interior. 8th Supplement to the "Revision of the I'seudomorphinae of the Australian Region" 152 M. Baehr MATERIALS AND METHODS proeps, proepisternal setae, longitudinally and Whereas the specimens collected during the transversally on proepisternum (either side) ongoing Pilbara survey conducted by the marg, marginal setae, along margin of elytra (either Department of Environment and Conservation, side) Perth, were sampled by glycol pitfall trapping, the st VI, setae on posterior border of sternum VI, the material collected during my travel was either penultimate visible sternite (either side) sampled by pulling down sheets of loose bark from cS st VII, setae of male sternum VII, the terminal a number of bark-shedding eucalypts (Salmon visible sternite (either side) Gum, River Red Gum, Redwood), or by Pyrethrum 2 st VII; setae of female sternum VII, the terminal fogging the rough-barked trunks of a variety of visible sternite (either side) eucalypt and non-eucalypt tree~ (diverse Mallee, Ja~rah, Tuart, Red Tingle, Grevillea, Mulga, and other Acacias). SYSTEMATICS Dissecting methods and style and format of the descriptions exactly correspond to those in my Family Carabidae LatreiIIe revisions (Baehr 1992, 1997) and the following Subfamily Pseudomorphinae Newman supplements (Baehr 1993a, 1993b, 1994b, 2002, 2004, 2005, 2006, 2007) which also can be used to gain Genus Sphallomorpha Westwood, 1837 additional information about the mentioned genera, their morphology, distribution, and habits. Sphallomorpha Westwood, 1837: 414; Baehr, 1992: Complete references for previously described 15. species can be taken from the revisions and the Type species supplements and are not included in the literature. Sphallomorpha decipiens Westwood, 1837, by For the benefit of the reader, abbreviations of the monotypy. chaetotaxy which is particularly important for identification of Sphallomorpha species, are repeated below. Remarks The habitus photographs were obtained with a With about 145 described species this is the digital camera using ProgRes Capture Basic and largest genus of the subfamily Pseudomorphinae AutoMontage, and were subsequently prepared and, with respect to the great number of with Corel Photo Paint 11. plesiomorphic character states, it is the most basal Specimens are lodged in the following genus and altogether the adelphotaxon of all other institutions: DEC, Department of Environment and genera (Baehr 1994a). It's basal status is also Conservation, Woodvale, Western Australia' CBM demonstrated by the ovipary of all species, whereas working collection of M. Baehr in Zool~gisch~ all other genera of Pseudomorphinae (except Staatssammlung, Mtinchen; WADA, Western Cryptocephalomorpha the reproduction mode of Australian Department of Agriculture, South Perth; which is still unknown) are ovoviviparous. The WAM, Western Australian Museum, Perth. overwhelming majority of the species occur in The following abbrevations are used for Australia, with a few species recorded from New Guinea. In Australia, species of chaetoatxy: Sphallomorpha occur in most environments, provided some tree supraorb, supraorbital seta (either side) preorb, preorbital seta (either side) growth is present, because the species generally occur on trees and prefer bark-shedding eucalypts, c1yp, c1ypeal seta (either side) labr, labral setae (common) where during daytime they hide under the bark. ment.med, medial mental setae, at base of mental Several species have been recorded from rainforest excision or mental tooth (common) ecosystems, but the majority occur in open forest ment.lat, lateral mental setae, on wings of mentum and even in rather dry country. (either side) gloss, glossal setae, on ventral rim of apex of glossa (either side) Sphallomorpha froggatti (Madeay, 1888) gut, gular setae, inside of gular suture (either side) Silphomorpha froggatti Macleay, 1888: 457; postorb, postorbital setae, posteriorly of eye on a Sphallomorpha froggatti, Baehr, 1992: 63; Baehr, conspicuous rim (either side) 2002: 102. suborb, suborbital setae, below eye, laterally of gular suture (either side) New material examined pron.ant, anterior pronotal setae, near anterior angle Australia: Western Australia: 1 0, site WA06/ of pronotum (either side) 134, 2 km S. of crossing of Tom Price Road and pron.post, posterior pronotal setae, near posterior Paraburdoo Road, 22.95025°S, 117.34708°E, 396 m, angle of pronotum (either side) 17 February 2006, M. Baehr (WAM 67920). Pseudomorphine species from Western Australia 153 Remarks Etymology The new specimen is the first recent record of S, The name refers to the close relationship of this Froggatti from Western Australia south of the Great species to SphalIol11orpha interioris Baehr from Sandy Desert. This comrnon species was previously central Northern Territory, known from north-eastern Queensland from about I{ockhampton through northern tropical Northern Diagnosis Territorv to the Kimberley Division in northern- A species of the mastcrsii species-group (Baehr most Western Australia, 1(92) with coarsely punctate eIytral striae and fairly 'nl(' single specimen was fogged from the rough coarse punctations of intervals, and with slightly bark on the trunk of a Mulga Acacia (Acacia aneura), bent-down apex of the aedeagus, Distinguished from the most similar S, interioris Baehr by the more asymmetrical shape and less distinctly bent Sphallomorpha fallax (Westwood, 1837) apex of aedeagus, and the elongate female Figure 16 stvlomere 2, Silphol11orpha hlllax Westwood, 1837: 426, Description Spha/lol11orpha [a/l<l\ (Westwood): Baehr, 1992: 79; Measurements: length:
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    RADAR Research Archive and Digital Asset Repository Copyright © and Moral Rights for this thesis are retained by the author and/or other copyright owners. A copy can be downloaded for personal non-commercial research or study, without prior permission or charge. This thesis cannot be reproduced or quoted extensively from without first obtaining permission in writing from the copyright holder(s). The content must not be changed in any way or sold commercially in any format or medium without the formal permission of the copyright holders. Note if anything has been removed from thesis. Map on p233 When referring to this work, the full bibliographic details must be given as follows: Hogan, J. E. (2012) Taxonomy, Systematics and Biogeography of the Scaritinae (Insecta, Coleoptera, Carabidae). PhD Thesis. Oxford Brookes University. WWW.BROOKES.AC.UK/GO/RADAR Taxonomy, Systematics and Biogeography of the Scaritinae (Insecta, Coleoptera, Carabidae) James Edward Hogan Thesis submitted in partial fulfilment of the requirements of the award of Doctor of Philosophy Oxford Brookes University December 2012 Abstract Scaritinae are a subfamily of ground beetles (Carabidae), containing about 1900 species and 125 genera. They share a distinctive body shape linked to a burrowing lifestyle. The diversity of Scaritinae is concentrated in the tropics and warmer regions of the southern hemisphere, particularly Southern Africa, Madagascar and Australia. The evolutionary history (phylogeny) of scaritines is unknown, leading to conflicting classification schemes and uncertainty over the definition of genera, especially in one sub- group of scaritines, the tribe Scaritini. In particular, it is unclear whether Scaritinae are descended from a common ancestor (monophyletic) or whether they are an artificial group defined by convergent adaptations to burrowing.
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