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New and Rare Species of the Pseudomorphine Genera Sphallomorpha Westwood, Adelotopus Hope, and Pseudomorpha Kirby Mainly from Far Northern Territory, Australia

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New and Rare Species of the Pseudomorphine Genera Sphallomorpha Westwood, Adelotopus Hope, and Pseudomorpha Kirby Mainly from Far Northern Territory, Australia ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de SPIXIANA 32 2 231–253 München, November 2009 ISSN 0341–8391 New and rare species of the pseudomorphine genera Sphallomorpha Westwood, Adelotopus Hope, and Pseudomorpha Kirby mainly from far Northern Territory, Australia (Coleoptera, Carabidae, Pseudomorphinae) 12th supplement to the “Revision of the Pseudomorphinae of the Australian Region” Martin Baehr Baehr, M. (2009): New and rare species of the pseudomorphine genera Sphallo- morpha Westwood, Adelotopus Hope, and Pseudomorpha Kirby mainly from far Northern Territory, Australia (Coleoptera, Carabidae, Pseudomorphinae). 12th supplement to the “Revision of the Pseudomorphinae of the Australian Region”. – Spixiana 32/2: 231-253 Five new species and one new subspecies of the pseudomorphine genera Sphallo- morpha Westwood, 1837 and Adelotopus Hope, 1834 from Australia are described: Sphallomorpha elseyi, S. laevior, S. triangularis, Adelotopus rufozonatus roperi, and A. frater, all from far Northern Territory, and S. aenigmatica from southern Western Australia. Records from various parts of Australia of a number of rare, ill docu- mented, or in the region so far unknown pseudomorphine species of the mentioned genera and of the genus Pseudomorpha Kirby, 1825 are also dealt with. All new spe- cies and subspecies are inserted in the respective general keys. Martin Baehr, Zoologische Staatssammlung, Münchhausenstr. 21, D-81247 München, Germany; e-mail: [email protected] Introduction 1867), S. biplagiata (Castelnau, 1867), S. nitiduloides Guérin,1844, S. amabilis (Castelnau, 1867), Adelotopus A recent collecting trip through far northern parts politus Castelnau, 1867, A. haemorrhoidalis Erichson, of Northern Territory and Western Australia and 1842, A. bimaculatus angustior Baehr, 1997, A. laticollis visits at certain Australian institutions and collections Baehr, 1997, A. g. gyrinoides Hope, 1834, A. d. dubius revealed a number of new and rare species of the Baehr, 1997, A. montorum Baehr, 1997, A. murrayanus pseudomorphine genera Sphallomorpha Westwood, Baehr, 1997, A. rufoguttatus Blackburn, 1893, A. maci- 1837, Adelotopus Hope, 1834, and Pseudomorpha lentus Baehr, 1997, A. paroensis Castelnau, 1867, Cai- Kirby, 1825 that are described or mentioned in nogenion ephippiatum (Newman, 1856), C. obscurum the present paper. During the visit also several (Castelnau, 1867), and C. tropicum Baehr, 1997. For common and well documented species were col- information about these species vide the revisions lected or identified that are not mentioned further of the Australian Pseudomorphinae and some of in this paper, i.e. Sphallomorpha froggatti (Macleay, the supplements (Baehr 1992, 1993a, 1994a, 1997, 1888), S. denisonensis (Castelnau, 1867), S. difficilis 2002, 2005, 2008). (Blackburn,1901), S. dubia (Castelnau, 1867), S. lae- In spite of the previous revisions and the many vigata (Castelnau, 1867), S. laevis (Castelnau, 1867), supplements which included the majority of the ma- S. polita (Macleay, 1871), S. guttigera (Newman, 1842), terial available in collections when they were written, S. castelnaui (Reiche, 1868), S. bicolor (Castelnau, the pseudomorphine fauna of large parts of interior 231 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de and far northern Australia is not yet satisfactorily bah, Terminalia, Satinash, certain Acacias, Grevilleas, recorded. New species are continuously detected and Casuarinas), or the rough-barked lower trunks of and of a great number of species few or even single bark shedding eucalypts (Red River Gum, White Gum). specimens were ever recorded, or the species are so A few specimens also were collected at black light in far known only from a single locality. This deficiency the night. In most specimens from other institutions the sampling methods were not mentioned on the labels, may be simply due to the large size of the country but I guess that they likewise generally were collected or the remoteness of many parts of Australia which from (under) bark of trees. makes reasonable or even systematic sampling of Altogether 32 pseudomorphine species in about 280 the fauna almost impossible. Moreover, generally specimens were collected during the five weeks travel pseudomorphine species can be only sampled using through northern parts of Northern Territory and West- specialized methods, either by use of an umbrella, ern Australia, though the bulk of specimens belongs to or by fogging tree trunks, because they occur under the three common northern Australian species Sphallo- bark or in deep cracks in the bark, of various trees. morpha laevis (Castelnau), S. froggatti (Macleay), and To get a reasonable overview of the fauna, these S. denisonensis (Castelnau). In the course of the travel, at methods must be carried out systematically at a great 90 localities about 500 trees were fogged and another number of localities, and at an even greater number 500 trees were sampled by pulling down stripes of bark. of trees, because pseudomorphines apparently tend Dissecting methods and style and format of the to concentrate on single trees, or they occur very descriptions exactly correspond to those in my revisions localized in certain areas. Hence collecting is a spe- (Baehr 1992, 1997) and the following supplements cialized and time-consuming task and seasonality (Baehr 1993a,b, 1994a, 2002, 2004, 2005, 2006, 2007, 2008, probably renders systematic sampling even more in press) which also can be used to gain additional in- difficult. However, when conditions are favourable formation about the mentioned genera, their morphol- and sampling methods and efforts are adequate, a ogy, distribution, and habits. Complete references for surprising great number and diversity of species previously described species can be taken from the revi- can be found on a single tree, which, for example, sions and the supplements and are not included in the was demonstrated on a few large Red River Gums citations of literature. in northern Australia that I tried to collect system- For the benefit of the reader, abbreviations of the chetotaxy which is particularly important for identifica- atically several years ago (Baehr 1990). “Rarity” of a tion of Sphallomorpha species, are repeated below. species thus does not automatically mean that it is The habitus photographs were obtained with a rare in nature, because this denotation as well may digital camera using ProgRes Capture Basic and Auto- mean “rarely collected” on whatever reasons: e.g. Montage and subsequently were worked with Corel seasonality, or geographically scattered occurrence, Photo Paint 11. or special habitats, of the species, but also inadequate All citations of label data exactly conform with the sampling because of too little collecting efforts, or labels, including any abbreviations, dates and any other ineffective or inappropriate sampling methods. information. The labels of all specimens collected during The scope of the present paper hence is not only the mentioned trip, including those of the types of the to describe the new species and to make available new species, are printed. additional information on species diversity, distri- The holotypes of the newly described species are bution, and habits, but also to draw the attention of located at Museum and Art Gallery of the Northern Territory, Darwin, and Australian Museum, Sydney. future collectors towards this very specialized and peculiar beetle group and to the arboricolous and corticolous beetle fauna of the Far North of Australia Scientific names of the mentioned tree species in general, and not only to the beetle fauna but also the very rich fauna of other invertebrates that live Red River Gum – Eucalyptus camaldulensis Dehnh. on or under bark of trees (see Baehr 1990). Ghost Gum – E. papuana F. Muell. White Gum – E. alba Reinw. ex Blume Darwin Stringybark – E. tetrodonta F. Muell. Material and Methods Darwin Whoolybutt – E. miniata Cunn. ex Schauer Smooth-stemmed Bloodwood – E. bleeseri Blakely The specimens collected during my travel were either Swamp Bloodwood – E. ptychocarpa F. Muell. sampled by pulling down sheets of loose bark from a Long-fruited Bloodwood – E. polycarpa F. Muell. number of bark-shedding eucalypts (Red River Gum, Coolibah – E. microtheca F. Muell. Ghost Gum, White Gum) into an umbrella, or by Pyre- Terminalia – Terminalia platyphylla F. Muell. thrum fogging the rough-barked trunks of a variety of Satinash – Syzygium sp. eucalypt and non-eucalypt trees (Darwin Stringybark, Acacia – Acacia spp. Darwin Whoolybutt, Smooth-stemmed Bloodwood, Grevillea – Grevillea sp. Swamp Bloodwood, Long-fruited Bloodwood, Cooli- Casuarina – Casuarina sp. 232 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de Abbreviations of collections and of states New records AMS Australian Museum, Sydney The new records are arranged in the sequence of CBM Working Collection of M. Baehr in Zoologische the species in the respective revisions (Baehr 1992, Staatssammlung, München 1997). NTM Museum and Art Gallery of the Northern Ter- ritory, Darwin QDPIB Queensland Department of Primary Industries, Genus Sphallomorpha Westwood Brisbane QMB Queensland Museum, Brisbane UQIC University of Queensland Insect Collection, Sphallomorpha Westwood, 1837: 414; Baehr 1992: 15. Brisbane With about 150 described species this is the larg- WAM Western Australian Museum (sampled by De- est genus of the subfamily Pseudomorphinae
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