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W' D FFO;Mpt of Muricidae (Gastropoda)

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W' D FFO;Mpt of Muricidae (Gastropoda) The malacologicalsocietymalacological society of Japan HM VENUS (Jap. Jour. Malac.) Vol 39,No. 3 (l9BO): 131-13S M g Original Articles -r )/ Ft?･7aV ltf )/ ptU]ftas 7" i7 t2 ivilk 7 . =i; w' d FFO;Mpt iJ=, .li . .I- - . 7z 77V == -S" yti 1) t '-' ('V ･ t' i =L ii' e thWwoehaee) New Species of Muricidae (Gastropoda) from the Indian Ocean, the Philippines and Brasil x Hans BERTscH and Anthony D'ATTiLIo (Department of Marine Invertebrates, Natural History Museum, Balboa Park, P.O. Box 1390, San Diego, California 92112, USA) Through the eourtesy of various eonchologists and professional malacolegists, we have been given specimen of 3 new mollusk species from a variety of loca- tions. In this paper we describe these new taxa, all representatives of the family Muricidae RAFINEsQuE, 1815. This continues our investigations into the taxonomy of tropical murieid gastropods Csee EMERsoN & D'ATTIuo, 1979; D'ATTILio 1980a & BERTscH, and 1980b), References for taxa authered prior to 1975 can be found in the literature cited of RADwiN & D'ATTIuo (1976) and hence are not repeated here. Muricinae RAFINEs'QuE, 1815 Pterynetzas SwAINsoN, 1833 PteT:gnotus atbobrunneus n. sp. (Figs. 1 and 2) Description: Shell 41mm long, strongly fusiform; spire high; protoconch not preserved; teleoconch of 6 weakly convex whorls, the suture impressed. Aperture ovate but eompressed above at the anal sulcus; outer lip with crenula- tions that are the terminations ef the spiral sculpture; 6 weakly swollen dentieles inside the outer lip, a weak additional dentiele in the anal sulcus; inner lip mostly erect, expanded above into a moderate callus; canal broad, of moderate length, open, its lowest third terminally reeurved; terminal parts of two previous canals on the siphonal faseiole. Varices 3, robust, low, each with a large mid-varical node-like costa; flange rising above varix cQmparatively low, extending from suture to anterior part NII-Electronic Library Service The malacologicalsocietymalacological society of Japan 132 VENUS: VoL 39, No.3(1980) of canal. Fine spiral cords numerous, 14 or 15 major ones, with weaker ones between, Growth striae extremely fine, close set, undulating and sealy where not, abraded. Shell bicolored externally, white with rust to bro"rn staining, the brownish color strongest on the mid-varical eostae and low varical flanges, narrow whitish diffuse band on shoulder below suture; aperture white. Operculum unguiculate, with basal terminal nueleus. The paratype specimen was apparently more mature than the holotype, judging from its rnore thiekened outer lip. The parat･ype also has a rust color in the concave portions of the erenulations of the outer lip. Maten-ctt Examined: (1) Holotype, San Diego Natural History Museum, Department of Marine Invertebrates, Type Series: SDNHM, T.S. 516. Shell 41mm long; collected 1978 off Kiltan Island, Laceadive Islands, Indian Ocean <approximately 11" 30' N; 73D E), brought up in shrimp nets from 100 m. (2) Paratype, SDNHM T.S. 793. Shell 45mm long; beach-worn speeimen eollected at Zanzibar, Tanzania (approximately 6e S; 39e E), November 1961, by H. MAKAME, C3) One specimen, cited by VoKEs (1978: 397-398; pl, 6, fig. 3) as Pterynotus cf. pelLzecielu･s ; colleeted southwest Conducia Bay, northeast of Choeas, northern Mozambique, on rock in fine sand and Tha･lassodendron, 5m; by K. GRoscH (in eolleetion of the Natal Museum). Type Locaeity: Off Kiltan Island, Laccadive Islands, Indian Ocean (ap- proximately 11e 30' N; 73e E). Figs. 1-2. PteTynot?ts atbobru･nne'us n. sp., Holotype specimen. Photograph by Barbara MyERs, Figs. 3-4. F'ava･rtia o'eanae n. sp., Holotype specimen. Photograph by David MuLuNER. NII-Electronic Library Service The malacologicalsocietymalacological society of Japan BERTscH･D'ATTIuo: Three New Species of Muricidae 133 Etymology: The specific name combines albus (white) and brhzenneus (brown) in referenee to the distinetive bieoloration. Disctession: D'ATTILIo & BERTscH (1980:169) have treated three closely related groups of trivaricate muricine shells as subgenera of Pterynotus. Pterynotus (Pterynotus) a.ebobTunneus can immediately be separated from species of the subgenus Pterochelus JoussEAuME, 1880, because the varical webbing does not end at･ the shoulder in a spine with a well-developed median ehannel and beeause the siphonal canal is elosed. A combination of characters distinguishes this new speeies from the subgenus Ma.rchia JoussEAuME, 1880, and from other species of Pterynottts sensu stricto: it has Telatively Iow varices, with the whorls evenly increasing in size cn the body whorl, and the body whorl flange is not markedly set off frem the ethers. These traits are shared with Pterynotus a･latus (R6DING, 1798) and P. (R.) bibbeyi (RAD"rlN & D'ATTILIo, 1976), but their white coloration and columellar dentieles respeetively differentiate these species from P. albobrunneus. Another speeies with relatively low varices, Pterynotus aparTii D'ATTmlo & BERTscH, 1980, has many differentiating features from P. atbobrunneus, ineluding columellar denticles. Although the new speeies is similar in coloration to Pterynotus miyokoae KosuGE, 1979, its shorter flange length immediately distinguishes it. VoKEs (1978: 397-398) referred a specimen of this species to Pterynotus (M.) peleucielus (REEvE, 1845), but the two species are not sympatrie and are distinct. RADwlN & D'ATTmlo (1976: 60) considered P. petlucidus to range from southeast Japan to east Africa, based on the same misidentified speeimen from Mozambique that Vokes reported. Pterynotus peltuci･dus is apparently restricted to the western Paeific from southeast Japan to the central Philippines. The shell color of P. pellucidus is generally white, occasionally pink, not white and golden brown like the new speeies; and the shell has a much more flaring flange on the body whorl. Murieopsinae RADwlN & D'ATTILIo, 1971 F'avartia･ JoussEAuME, 1880 ]Plzvartia jeanae n. sp. (Figs. 3-5) Description: A small murieopsine 11 mm long, slenderly fusiform; teleoeoneh ot 6 eonvex whoris; protoconch (Fig. 5) of 2i,･(.,. to 2% weakly convex whorls. Spire very high, the suture weakly defined, Aperture ovate; outer lip with 5 or 6 large knobby denticles inside, somewhat set away from the weakly erenulate ]abrum; anal sulcus shallow and broad. Inner lip strongly erect with 4 large knobby denticles on the lower half of eolumella, the lowest denticles on the labrum and eolumella are paired above entranee to eanal. Canal open, broad above, becoming tube-like and sharply recurved distally. Siphonal faseiole with earlier eanals forming a rostrate strueture on left side. NII-Electronic Library Service The malacologicalsocietymalacological society of Japan 134 VENUS: Vol. 39, No.3<1980) Varices on whorl 4, body broad at base, sloping on their leading side; final two varices thiekest with broad interspaces, the first 2 similar to varices on spire; vances on spire 8 per whorl, appearing as a series of costae with barely any intervarical areas. Spiral seulpture of 5 majer cords on body divided by single minor inter- callary eords. Major posterior cords divided centrally by a groove; three addi- tional major eords with minor cords on shoulder, a major and minor cord on canal. Leading side of varices white, covered by strongly vaulted scales. Growth striae well defined, not eonspieuously seabrous, but forming thickened nodes on the spiral sculpture. Shell essentially orange, ranging from yellow orange to red orange ; aperture white and posterior leading side of main body whorl varices also white. Materia･l Examined: Holotype, (1) San Diego Natural History Museum, Department of Marine Invertebrates, Type Series: SDNHM, T.S, 796. Shell 11 mm long; collected in about 80-100m off Punta Engafio, Mactan Island, Cebu, Philippine Islands, in tangle nets. (2) Paratypes, SDNHM T,S. 797. 15 specimens collected with holotype. (3) Paratypes, Bishop Museum, Malacology Catalogue No. 207077; 2 speeimens eollected with holotype. Type Locality: Punta Engafio, Cebu Island, Bohol Straits, Philippine Islands (approximately 10e20'N; 124eE). Etymotogy: The honors of patronym Mrs. Jeanne PisoR, San Diego, Cali- "ne" fornia, who brought these speeimens to our attention (the letters have been omitted for the sake of euphony). Discussion: In a previous work (D'ATTILIo & BERTscH, 1980) we studied the morphological characters useful for the generie assignment of speeies in FTava･rtia. On' the basis of oveTlapping traits Cradular and shell characters), we concluded that 1;arurexiella CLENcH & FARFANTE, 1945, be restricted, as a subgenus of Fava･rtia, to include only those speeies strongly resembling the type Favartia (Murexiettct) hidalgoi (CRossE, 1869). About 20 specimens of the new speeies were examined. Speeimens of 6 whorls range from 9 to 11mm. The reduet,ion from 8 vai'ices on the spire to 4 varices on the body, coupled with the slender form, separates this new speeies frem any other Favartia known from the Indo-Paeific. By using several other basic characteTistics, Favartia deanae ean be further distinguished from its known Indo-Pacific congeners. Favartia deanae laeks the long spines of F. batteata (SowERBy, 1841), F. a'udithae D'ATTmlo & BERTscH, 1980, and IF'. peeepili D'ATTILIo & BERTscH, 1980. It does not have the flanges or highly developed variees eharacteristic of F. eonfusa CBRAzlER, 1877), F. cyctosto7n.a (SowERBy, 1841), jF'. fnna.fz`tiensis (HEDLEy, 1899), F. garrettii (PEAsE, 1868), and Ii'. peasei (TRyoN, 1880), nor does it have the heavy <thick) varices of F. NII-Electronic Library Service The malacologicalsocietymalacological society ofJapanof Japan BERTscH-D,ATTImo: ThreeNew Species ef Muricidae 135 /tt. fx.i... X '. x. x･ "K' ' x Fig. 5. Pavctrtia J'ectuae n. sp., close-up diagram of pivtoconeh. Drawing by A. D'ATTImo. Figg. 6-7. Typhina riosi n. sp., Holotype specimen. Drawing by A. D'ATTmlo. mina.tauros RADwlN & D'ATTiuo, 1976, and F. tetf'a･gona (BRoDERIp, 1833). fi'avartia etatensis EMERSON & D'ATTILIo, 1979, and IiT. guomensis EMERSON & D'ATTmlo, 1979, incised at the suture, 17a･va･rtia g'eanae has denticles laeking in all the above species and in F'.
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