The malacologicalsocietymalacological society of Japan

HM VENUS (Jap. Jour. Malac.) Vol 39,No. 3 (l9BO): 131-13S

M g Original Articles

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New Species of () from the Indian Ocean, the Philippines and Brasil

x Hans BERTscH and Anthony D'ATTiLIo (Department of Marine Invertebrates, Natural History Museum, Balboa Park, P.O. Box 1390, San Diego, California 92112, USA)

Through the eourtesy of various eonchologists and professional malacolegists, we have been given specimen of 3 new mollusk species from a variety of loca- tions. In this paper we describe these new taxa, all representatives of the family Muricidae RAFINEsQuE, 1815. This continues our investigations into the of tropical murieid gastropods Csee EMERsoN & D'ATTIuo, 1979; D'ATTILio 1980a & BERTscH, and 1980b), References for taxa authered prior to 1975 can be found in the literature cited of RADwiN & D'ATTIuo (1976) and hence are not repeated here.

Muricinae RAFINEs'QuE, 1815

Pterynetzas SwAINsoN, 1833

PteT:gnotus atbobrunneus n. sp. (Figs. 1 and 2) Description: Shell 41mm long, strongly fusiform; spire high; protoconch not preserved; teleoconch of 6 weakly convex whorls, the suture impressed. Aperture ovate but eompressed above at the anal sulcus; outer lip with crenula- tions that are the terminations ef the spiral sculpture; 6 weakly swollen dentieles inside the outer lip, a weak additional dentiele in the anal sulcus; inner lip mostly erect, expanded above into a moderate callus; canal broad, of moderate length, open, its lowest third terminally reeurved; terminal parts of two previous canals on the siphonal faseiole.

Varices 3, robust, low, each with a large mid-varical node-like costa; flange rising above varix cQmparatively low, extending from suture to anterior part

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132 VENUS: VoL 39, No.3(1980)

of canal. Fine spiral cords numerous, 14 or 15 major ones, with weaker ones between, Growth striae extremely fine, close set, undulating and sealy where

not, abraded. Shell bicolored externally, white with rust to bro"rn staining, the brownish

color strongest on the mid-varical eostae and low varical flanges, narrow whitish

diffuse band on shoulder below suture; aperture white.

Operculum unguiculate, with basal terminal nueleus. The paratype specimen was apparently more mature than the holotype, judging from its rnore thiekened outer lip. The parat･ype also has a rust color in the concave portions of the erenulations of the outer lip. Maten-ctt Examined: (1) Holotype, San Diego Natural History Museum,

Department of Marine Invertebrates, Type Series: SDNHM, T.S. 516. Shell

41mm long; collected 1978 off Kiltan Island, Laceadive Islands, Indian Ocean

by H. MAKAME, C3) One specimen, cited by VoKEs (1978: 397-398; pl, 6, fig. 3) as Pterynotus cf. pelLzecielu･s ; colleeted southwest Conducia Bay, northeast of Choeas, northern Mozambique, on rock in fine sand and Tha･lassodendron, 5m; by K. GRoscH (in eolleetion of the Natal Museum). Type Locaeity: Off Kiltan Island, Laccadive Islands, Indian Ocean (ap- proximately 11e 30' N; 73e E).

Figs. 1-2. PteTynot?ts atbobru･nne'us n. sp., Holotype specimen.

Photograph by Barbara MyERs,

Figs. 3-4. F'ava･rtia o'eanae n. sp., Holotype specimen. Photograph by David MuLuNER.

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BERTscH･D'ATTIuo: Three New Species of Muricidae 133

Etymology: The specific name combines albus (white) and brhzenneus

(brown) in referenee to the distinetive bieoloration. Disctession: D'ATTILIo & BERTscH (1980:169) have treated three closely

related groups of trivaricate muricine shells as subgenera of Pterynotus. Pterynotus (Pterynotus) a.ebobTunneus can immediately be separated from species

of the subgenus Pterochelus JoussEAuME, 1880, because the varical webbing does

not end at･ the shoulder in a spine with a well-developed median ehannel and

beeause the siphonal canal is elosed. A combination of characters distinguishes

this new speeies from the subgenus Ma.rchia JoussEAuME, 1880, and from other

species of Pterynottts sensu stricto: it has Telatively Iow varices, with the whorls

evenly increasing in size cn the body whorl, and the body whorl flange is not

markedly set off frem the ethers. These traits are shared with Pterynotus

a･latus (R6DING, 1798) and P. (R.) bibbeyi (RAD"rlN & D'ATTILIo, 1976), but

their white coloration and columellar dentieles respeetively differentiate these

species from P. albobrunneus. Another speeies with relatively low varices,

Pterynotus aparTii D'ATTmlo & BERTscH, 1980, has many differentiating features

from P. atbobrunneus, ineluding columellar denticles.

Although the new speeies is similar in coloration to Pterynotus miyokoae

KosuGE, 1979, its shorter flange length immediately distinguishes it.

VoKEs (1978: 397-398) referred a specimen of this species to Pterynotus (M.) peleucielus (REEvE, 1845), but the two species are not sympatrie and are

distinct. RADwlN & D'ATTmlo (1976: 60) considered P. petlucidus to range from

southeast Japan to east Africa, based on the same misidentified speeimen from

Mozambique that Vokes reported. Pterynotus peltuci･dus is apparently restricted

to the western Paeific from southeast Japan to the central Philippines. The shell

color of P. pellucidus is generally white, occasionally pink, not white and golden

brown like the new speeies; and the shell has a much more flaring flange on

the body whorl.

Murieopsinae RADwlN & D'ATTILIo, 1971

F'avartia･ JoussEAuME, 1880

]Plzvartia jeanae n. sp.

(Figs. 3-5)

Description: A small murieopsine 11 mm long, slenderly fusiform; teleoeoneh

ot 6 eonvex whoris; protoconch (Fig. 5) of 2i,･(.,. to 2% weakly convex whorls.

Spire very high, the suture weakly defined, Aperture ovate; outer lip with 5

or 6 large knobby denticles inside, somewhat set away from the weakly erenulate

]abrum; anal sulcus shallow and broad. Inner lip strongly erect with 4 large

knobby denticles on the lower half of eolumella, the lowest denticles on the labrum

and eolumella are paired above entranee to eanal. Canal open, broad above, becoming tube-like and sharply recurved distally.

Siphonal faseiole with earlier eanals forming a rostrate strueture on left side.

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134 VENUS: Vol. 39, No.3<1980)

Varices on body whorl 4, broad at base, sloping on their leading side; final two varices thiekest with broad interspaces, the first 2 similar to varices on

spire; vances on spire 8 per whorl, appearing as a series of costae with barely

any intervarical areas.

Spiral seulpture of 5 majer cords on body divided by single minor inter-

callary eords. Major posterior cords divided centrally by a groove; three addi-

tional major eords with minor cords on shoulder, a major and minor cord on canal.

Leading side of varices white, covered by strongly vaulted scales. Growth striae well defined, not eonspieuously seabrous, but forming thickened nodes on

the spiral sculpture.

Shell essentially orange, ranging from yellow orange to red orange ; aperture white and posterior leading side of main body whorl varices also white. Materia･l Examined: (1) Holotype, San Diego Natural History Museum, Department of Marine Invertebrates, Type Series: SDNHM, T.S, 796. Shell

11 mm long; collected in about 80-100m off Punta Engafio, Mactan Island, Cebu,

Philippine Islands, in tangle nets.

(2) Paratypes, SDNHM T,S. 797. 15 specimens collected with holotype. (3) Paratypes, Bishop Museum, Malacology Catalogue No. 207077; 2 speeimens eollected with holotype. Type Locality: Punta Engafio, Cebu Island, Bohol Straits, Philippine

Islands (approximately 10e20'N; 124eE).

Etymotogy: The patronym honors Mrs. Jeanne PisoR, of San Diego, Cali-

"ne" fornia, who brought these speeimens to our attention (the letters have been

omitted for the sake of euphony).

Discussion: In a previous work (D'ATTILIo & BERTscH, 1980) we studied

the morphological characters useful for the generie assignment of speeies in

FTava･rtia. On' the basis of oveTlapping traits Cradular and shell characters), we

concluded that 1;arurexiella CLENcH & FARFANTE, 1945, be restricted, as a subgenus

of Fava･rtia, to include only those speeies strongly resembling the type Favartia

(Murexiettct) hidalgoi (CRossE, 1869).

About 20 specimens of the new speeies were examined. Speeimens of 6

whorls range from 9 to 11mm. The reduet,ion from 8 vai'ices on the spire to

4 varices on the body, coupled with the slender form, separates this new speeies

frem any other Favartia known from the Indo-Paeific. By using several other

basic characteTistics, Favartia deanae ean be further distinguished from its

known Indo-Pacific congeners. Favartia deanae laeks the long spines of F.

batteata (SowERBy, 1841), F. a'udithae D'ATTmlo & BERTscH, 1980, and IF'.

peeepili D'ATTILIo & BERTscH, 1980. It does not have the flanges or highly

developed variees eharacteristic of F. eonfusa CBRAzlER, 1877), F. cyctosto7n.a

(SowERBy, 1841), jF'. fnna.fz`tiensis (HEDLEy, 1899), F. garrettii (PEAsE, 1868),

and Ii'. peasei (TRyoN, 1880), nor does it have the heavy

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BERTscH-D,ATTImo: ThreeNew Species ef Muricidae 135

/tt.

fx.i... X '. x. x･ "K' ' x

Fig. 5. Pavctrtia J'ectuae n. sp., close-up diagram of pivtoconeh. Drawing by A. D'ATTImo.

Figg. 6-7. Typhina riosi n. sp., Holotype specimen. Drawing by A. D'ATTmlo.

mina.tauros RADwlN & D'ATTiuo, 1976, and F. tetf'a･gona (BRoDERIp, 1833).

fi'avartia etatensis EMERSON & D'ATTILIo, 1979, and IiT. guomensis EMERSON &

D'ATTmlo, 1979, incised at the suture, 17a･va･rtia

g'eanae has denticles laeking in all the above species and in F'. brevicuea･ (SOWERBY, 1834), IiT. crossei (LiENARD, 1873), F'. kuroelai

NAKAMIGAwA& HABE, 1964, F. maioriae (MELvlLL&STANDEN, 1903), F. munda (REEvE, 1849), and F. sa･lmonea･ (MELvlLL &

STANDEN, 1899) D'ATTImo, 1979, has only one

tooth on the columella and has a proportionately broader and larger shell than F. 2'eanae. Favartia denticles on the columella but

nope on the outer apertural lip; whereas P. has 4 and 5-6 denticles respec-

tiveEy; it also has long spines. 'Typhinae COSSMANN, 1903

Typhzna･ JOuSSEAuME, 1880

7Iyphina riosi n. sp.

(Figs. 6 and 7)

DescTiption : Shell rnoderately small, up to 11,5mm long, slenderly fusi-

form; teleoconch of 4 whorls, angulate at shoulders; protoeoneh of 1% swollen

polished whorls ; suture impressed and clearly defined. Aperture ovate, the

peristome entire, strongly extruded; canal broad above and quickly narrowing

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136 VENUS: VoL 39, No.3(1980)

into a Iong tube barely bent back. SpiTal sculpture Iacking. Axial sculpture of 4 varices, with the four tubes closer to preceding varices. Functional tube long, projecting obliquely upward and weakly back from the shoulder angle, similar

to all preeeding tubes. A sloping shoulder area remains between the seulpture on sheulder angle the and the suture. Varieal margin simple, adherent, crossing shoulder the behind tube and terminating at suture. An additional pseudovarix between varical margin and tube, very elose to or eontiguous to true varix on lower part of body whorl, pseudovarix raised at the shoulder into a bread spine trough-like and after crossing shoulder beeoming appressed again and turning obliquely to join and overlap true varix. This pseudovarix with its broad extended trough is a charaeteristic sculpture feature between peristome and funetional tube. Siphonal faseiole with three previous canals, the earliest one

the least preminent. Siphon with weakly erect and eurved flange below aperture. Growth striae moderately developed. Partition (sensu GERTMAN, 1969: 145, text-fig. 1) absent. Shell semi-translucent, pale oehre brown. Material Examined: (1) Holotype, San Diego Natural History Museum, Marine Invertebrates, T.S. 794. Shell 11.5 mm Iong, collected off Tramandai, RS, southerm Brasil (30e04'S; 47e55'W), from 280m depth, sandy mud bottorn, by N, Oe. W. BEsNARD, 3 August 1972. Paratypes, (2) SDNHM T.S. 795. 13 specimens (total lengths of specimens: 5.5, 5.5, 6.5, 7, 7.5, 7.5, 7.5, 8, 8, 9, 9, 10.5, 11.5 mm) collected with holotype. Two specimens from this lot have been sent to eaeh of the following institutions: Museu Naeional, Rio de Janeiro, Brasil; Museu de Zoologia, Universidade de S. Paulo, Brasil; Museu de Ciencias Naturais, Fundaeao Zoobotanica, Porto Alegre,

RS, Brasil; and Museo Nacional de H. Natural, Montevideo, Uruguay.

(3) 2 specimens, also eollected with holotype; in collection of Emily H.

VoKEs, Tulane University.

(4) 4 specimens; collected off Albradao, RS, southern Brasil (3.3022'S;

51"05'W), from 128m depth, sandy mud bottom, by N. Oc. W. BESNARD, 19

January 1972.

Type Locaeity: Off Tramandai, Brasil (30D04'S; 47055'W), sandy mud bottom.

Etymoeogy: The patronym honors our friend and colleague, Dr. Eliezer

de Carvalho RIos, who has made many contributions to knowledge of Brasilian mollusks.

Discussion: Typhiua is widely-distributed chronologieally and spatially.

It has been reported from the Eocene to the Recent (KEEN, 1944: 53-55;

GERTMAN, 1969: 148-150). Modern species are known from the western ?acific

(espeeially Japan, Australia, and New Zealand) and the Atlantic (VELLA, 1961:

365; KEEN & CAMpBELL, 1964: 46-48; and RADwlN & D'ATTILIo, 1976: 206-210).

As a representative of the Typhina, this new species has a compara-

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・ ， BERTSCH D ATTILIo ： Three New Species of Muricidae 137

tively simple ， smooth ， unornamented surface 且acking ， spiral elements and the

， usual presence of frilly characters on the varix or see D ATTII．10 pseudovarix （ ， 1975 for ， a discussion of typhine shell ornamentation ）． The only notable feature besides its noncomplex form is the spine −1ike fold at the shoulder of the pseudo −

varix ． 1・engths of the tubes and canal are variable characters entirely dependent

， on fortuitous conditions of the individua1s living condition ．

Between 七he varical margin and the anal tube is a small secondary spine ， appressed basally but pinching distany away from the shoulder as a very short

open trough ． By contrast to Typhilnα γゼos 琶， this secondary fold is prominent on

the early whorls in Typhina betcheri （BRODERIP ，1833 ）， T ． bivaricα tes （VERco ，

1909 T ． imperialis KEEN 恥 − ）， （ ＆ CAMPBE ， 1964 ）， and T ．pa uperis （MEsTAYER ，

1916 ． T α riosi has smooth varices in contrast to the frilly ornamentation ） ！lphin ，

and crenulations on outer margin mont orti the lip in T ． ∫ （A ．ADAMs ， 1863 ）， T ． ・ nitens ． ・ ＄ ． r α mos α ＆ （HINDS ，1843 ）， T phitipp inen is （WATSON ， 1883 ）， T （HABE

and α tesi KosuGE ， 1971 ）， T ．野 （CRossE ＆ FIscHER ， 1865 ），

cleryi 廿 Typhis （Rugotyphis ） （Pe t de la SAussAYE， 1840 ） from southern

， Brasil was assigned to Typhin α by RADwlN ＆ D ATTILIo （1976 ： 207 ）， Its earlier

（ ： more placement in RugoteJphis VELLA ， 1961 ，by GERTMAN 1969 15 ， 154 ） is

appropriate among other characters a ． ん because， ， large partition is present Typ is

〜．cl−eryi a robust species much （1 ） is distinguishedby its broader proportions ， the possession of a strong flange or partition connecting the apertural varix and the

spire ， and the strong ， spirally Huted varices retained on the body whorl ．

Acknowledgement ： We thank the following people who gave specimens to the San

： 乢 Llps of Diego Natural History Museum Mr ．John PH ． Little River ， Califomia ； Mrs ． ， and and 且 ald エ of Rose D ATTIHo Mr ． Mrs 。DQ P soR San Diego ， California ； ，Dr ．Eliezer

de C ． Rlos of the Museum Oceanografico de Furg BrasiL Dr ． Emily H ． VoKEs Tulane ， ， 」 ・ us speclmens 上 collection ． are to s ， University ， kindly loaned from he We grateful MI

Barbara MYERs and ML DaVid K 。 MuLLINER for the photographs and to Dr ． Reid

MoRAN San Diego Natural History Museum for editorial comments on the manuseript ． ， ，

要 約

フ タ ロ バ シ ・ PterPtnotus αlbobrunneus n ．sp ． イ ョ ウ ガ イ （新 種 新 称 ） − （Figs ．1 2）

殻 は 紡 錘 形 で 螺 塔 高 く 6 階 。縫 合 深 い 。殻 口 卵 形 ，外 唇 は 螺 状 彫 刻 の 末 端 に 対 応 し て 刻 ま れ ， 内面

it3 に 6 つ の 小 突 起が あ る 。前溝 は 広 く中 庸 の 長 さ ，下 方 は 反 る 。糊 帯 状 に 旧 い 2 水 管 が 乗 る 。縦 張 ゆ 々 つ 〜 肋 は 低 い 翼 状 で 3 ， 堅 牢 で ， 各 の 間 に 縦 張 肋 問 肋 を も 。 螺 状 腺 は 主 な も の は 14 15 本 で あ る か

細 い も の が 多 数 あ る 。成 長 腺 は 極 め て 細 い 。殻 表 は 白 と 羯 色 の 2 色 染 め 。 模 式 標 本 ： 殻 高 43mm 。

ン ン バ ル モ ン ビ ー 模式 産 地 ： イ ド洋 ラ ッ カ デ ィ ブ 諸 島，分 布 ： ザ ジ ， ザ ク 。

バ シ い で 比 較 ：他 の ョ ウ ガ イ 類 と は 比較 的 低 い 縦 張 肋 と そ れ ら が 各 螺 層 と ず れ が 少 な こ と 及 び 色 彩 ’ ヒ ロ ビ レ ・ミ シ ガ 区 別 出来 る 。 P ．Pelt” cida REEVE ョ ウ イ は 分 布域 を 異 に す る 。

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1980 138 VENUS ： Vol ．39 ， No ．3 （ ）

・ Favartia 佃 砌 α ε n ． sp ． オ レ ン ジ ヨ ウ ラ ク ガ イ （新種 新 称 ） − （Figs．3 5）

で 6 は む 口 壁 に は 〜 個 の 小 突 起 が あ る 。内 殻 は 小 形 堅 牢 ， 紡 錘形 階 。 胎 殻 弱 く膨 ら 。殻 外 唇 内 5 6

で 唇 ド半 分 に 4 個 の 瘤状小 歯 が あ り ，前 溝 は 管状 強 く反 る 。 旧 い 水 管 溝 が 左 側 に 在 る 。縦張 肋 は 体 層

の 2 が の に は 1 ご と に 月力 あ る 体 層 の 螺 状 彫 刻 は 5 本 の 4 本 ， 最 後 肋 は 強 く間隔 あ く。 E 方 螺 層 階 8 。

主螺 肋 と 1 本 宛 の 問 肋 で ，白 い 縦 張 肋 は 鱗状 。成 長 脉 は 明 瞭 で 螺 肋 上 で 肥 厚 す る 。殻 の 地 色 オ レ ン ジ

： 11 mm ： セ ブ ブ ン タ ・エ ン ガ ニ 色 。 模 式 標 本 殻 高 ， 模式産 地 島 ， ョ 。

二 の に で の イ ン ド の 同 比 較 全 形 高 く．ヒ方 螺 層 8 縦 肋 が あ り， そ れ が 体 層 で 4 と な る 点 他 太 平 洋 産 属

他 種 と 区 別 出来 る 。 ・ TPtPhina riosi n ． sp ， ブ ラ ジ ル エ ソ ト ツ ヨ ウ ラ ク （新 種 新 称 ） （Figs．6 ・7）

で い 口 は は く 状 僅 か に 曲 殻 は 小 形 で ， 紡 錘 型 4 階 。 胎 殻 は 1 平 滑 。縫 合 深 。殻 卵 円形 ， 前 溝 長 管

に か に に る 。螺 状 彫 刻 は な く，各 層 に 4 本 の ．縦 張 肋 と 4 本管 が あ る 。管 は 長 く．E 斜 方 伸 び 僅 後方 反 る 。

縦 張 肋 縁 は 管 の 後 方 を 通 り縫 合 で 終 る 。管 と縦 張 肋 の 間 に 擬 肋 が あ り肩 で 練状 に 立 ち 斜 行 し縦 張 肋 と

重 な る 。 こ の 擬 肋 と 棘 状 突 起 は 殻 口 と後 溝 の 間 で 顕 著 で あ る 。前管 繃 帯 に は 旧 い 水 管 が 3 こ 甫 る 。成

長 脉 は 中 庸 n 殻 は 半 透 明 ，淡 黄 褐 色 。 模 式 標 本 ： 殻 高 11．5 mm ，模 式産 地 ： 南 ブ ラ ジ ル ， ツ ラ マ ン

ダ ： ア ル ブ ラ ダ オ イ ， 280m ．分 布 1司 128m 。 ’ 比 較 ： 本 新 種 は 殻 表 ド滑 で 螺 状 彫 刻 は な く ，縦 張 肋 や 擬 肋 に フ リ ル が な い 点 と擬 肋 の 肩 が 棘 状 と な ． っ て い る 点 が 特 徴 で あ る 。

1． iterature Cited

’ ” ・’ and elaboration est 11vus D ATTILIo A ．1975 ．The Typhine shell ： tlleme development ， 。P ， ， − 6 （9 ）： 53 60 ，51figs ．

一 一 ． and BERTscH ， H ．1980a ．Comments on species of Homaeocantha （Gastropoda ： ． − ？ 2 ： text − Muricidae ）． Pest lvus 12 （ ） 14 20 ， 15 figs．（7 February 1980 ）

一 一 ・一一 ・ and 1980b ． Four species of P 亡e7 Ψ not u ．s and Favarti α （ ： Gastropoda ： ’ Muricidae ） from the Philippine Islands． Tr α nb ．8 α π Diego Soc ． N α ‘．超 紘 19 （12 ）： − − 1．69 179 ， 4 text figs．

， EMERsoN ， W ． K ．， and D ATTIL 正o ，A ．1979 ． Six new living species of muricacean

− 一figs gas 七ropods ．N α 臨 ‘粥 93 （1 ）： 1 10 ， 21 tex七 ．

GERTMAN ， L ．1969 ． Cenozoic Typhinae （Mollusca ： Gastr （，poda ） 〈）f the western Atlantic

region Tulane − − ． Stud ．（leol．P α ‘eo ．7 （3／4 ）： 143 191，8 pls ．， 3 text figs．

’’ KEEN ，A ．M ．1944 ．Catalogue and rev ｛sion of the gastropod subfamily Typllinae ．cJo tl，7 ． − Paleo 1 ： 50 72 20 text −figs ，18 （ ） ， ．

一 一 and CAMPB 肌 L G ． B ．1964 ． Ten new speeies ： ， of Typhinae （Gastropoda ・ − − 1 ： 46 57 11 −figs Muricidae ）．VeVigeor7 （ ） ， pls ．8 ， 3text ．

KosuGE ， S ．1979 ． Descriptions of two new species of the family Muricidae （Gas 七ropoda ： ・． − Mollusca ）． B ul 1．1π s む Malae ． Tok ？yr♪ 1 （1）： 1 2 ； 1 pl． （30 May 1979 ）．

， RADwlN G ． E ． and D ATTILIo A ．1976 ． ： 襴 加 磁 θ θ ， ， Murex shells o ／ the wortcl an 覗 d g 漉 d ・ 亡o 亡ノ〜召 M 「麗 麗 o ゼd α θ． ． text − ． ． 285pp ， 32 plsり 192 figs Stanfol d Univ Press ， Califomia．

V 肌 LA ， P 。1961 ． Australasian Typhinae （Gastropoda ） “ 厂 ith notes on the subl amily ． 1 − − ひ 3 ： 362 391 46 47 6text −figs Palaeunt loyy 4 （ ） ， pls ． ， ．

Vo 麗 s E ． H ．1978 Muricidae ： ， ． （Mollusca （｝astropoda ）from the eastern coast of Africa 。 ’ ＞α ‘α 乙 tcs 2 ： 375 −418 Ann ．1 M ．．23 （ ） ， 8 pls ．

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