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The Head Morphology of Micromalthus Debilis (Coleoptera: Micromalthidae) – an Archostematan Beetle with an Un Usual Morphology and a Unique Life Cycle
76 (3): 475 – 486 11.12.2018 © Senckenberg Gesellschaft für Naturforschung, 2018. The head morphology of Micromalthus debilis (Coleoptera: Micromalthidae) – an archostematan beetle with an un usual morphology and a unique life cycle Margarita I. Yavorskaya 1, Thomas Hörnschemeyer 2 & Rolf G. Beutel*, 1 1 Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität Jena, 07743 Jena, Germany; Margarita Yavorskaya [margojavor @gmail.com]; Rolf Georg Beutel * [[email protected]] — 2 Senckenberg Gesellschaft für Naturforschung, Senckenberg an lage 25, 60325 Frankfurt a.M., Germany; Thomas Hörnschemeyer [[email protected]] — * Corresponding author Accepted 25.vii.2018. Published online at www.senckenberg.de/arthropod-systematics on 27.xi.2018. Editors in charge: Joseph McHugh & Klaus-Dieter Klass Abstract. Cephalic features of Micromalthus debilis were examined and described in detail for the first time. The head displays several seemingly plesiomorphic features compared to other extant species of Archostemata, especially representatives of Cupedidae and Ommat idae. Cephalic protuberances characteristic for species of these two families are missing and antennal grooves are also absent. The surface of the head capsule is largely smooth, without the characteristic tubercles found in stemgroup beetles and ommatid and cupedid species. Cuticular scales, probably ancestral for Archostemata and possibly for Coleoptera, are also completely absent. The arrangement of three mandibular teeth in a vertical row and an immobilized labrum are derived features shared with Ommatidae. The maxillary endite lobes are absent, as in the very small Crowsoniella relicta (Crowsoniellidae). Like in all other examined archostematan species, mandibular molae and prosthecae are missing. The simplified maxillae apparently play no role in the food uptake but rather function as accessory “ventral antennae”. -
The Evolution and Phylogeny of Beetles
Darwin, Beetles and Phylogenetics Rolf G. Beutel1 . Frank Friedrich1, 2 . Richard A. B. Leschen3 1) Entomology group, Institut für Spezielle Zoologie und Evolutionsbiologie mit Phyletischem Museum, FSU Jena, 07743 Jena; e-mail: [email protected]; 2) Biozentrum Grindel und Zoologisches Museum, Universität Hamburg, 20144 Hamburg; 3) New Zealand Arthropod Collection, Private Bag 92170, Auckland, NZ Whenever I hear of the capture of rare beetles, I feel like an old warhorse at the sound of a trumpet. Charles R. Darwin Abstract Here we review Charles Darwin’s relation to beetles and developments in coleopteran systematics in the last two centuries. Darwin was an enthusiastic beetle collector. He used beetles to illustrate different evolutionary phenomena in his major works, and astonishingly, an entire sub-chapter is dedicated to beetles in “The Descent of Man”. During his voyage on the Beagle, Darwin was impressed by the high diversity of beetles in the tropics and expressed, to his surprise, that the majority of species were small and inconspicuous. Despite his obvious interest in the group he did not get involved in beetle taxonomy and his theoretical work had little immediate impact on beetle classification. The development of taxonomy and classification in the late 19th and earlier 20th centuries was mainly characterised by the exploration of new character systems (e.g., larval features, wing venation). In the mid 20th century Hennig’s new methodology to group lineages by derived characters revolutionised systematics of Coleoptera and other organisms. As envisioned by Darwin and Ernst Haeckel, the new Hennigian approach enabled systematists to establish classifications truly reflecting evolution. -
Fauna Europaea: Coleoptera 2 (Excl. Series Elateriformia, Scarabaeiformia, Staphyliniformia and Superfamily Curculionoidea)
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Digital.CSIC Biodiversity Data Journal 3: e4750 doi: 10.3897/BDJ.3.e4750 Data Paper Fauna Europaea: Coleoptera 2 (excl. series Elateriformia, Scarabaeiformia, Staphyliniformia and superfamily Curculionoidea) Paolo Audisio‡, Miguel-Angel Alonso Zarazaga§, Adam Slipinski|, Anders Nilsson¶#, Josef Jelínek , Augusto Vigna Taglianti‡, Federica Turco ¤, Carlos Otero«, Claudio Canepari», David Kral ˄, Gianfranco Liberti˅, Gianfranco Sama¦, Gianluca Nardi ˀ, Ivan Löblˁ, Jan Horak ₵, Jiri Kolibacℓ, Jirí Háva ₰, Maciej Sapiejewski†,₱, Manfred Jäch ₳, Marco Alberto Bologna₴, Maurizio Biondi ₣, Nikolai B. Nikitsky₮, Paolo Mazzoldi₦, Petr Zahradnik ₭, Piotr Wegrzynowicz₱, Robert Constantin₲, Roland Gerstmeier‽, Rustem Zhantiev₮, Simone Fattorini₩, Wioletta Tomaszewska₱, Wolfgang H. Rücker₸, Xavier Vazquez- Albalate‡‡, Fabio Cassola §§, Fernando Angelini||, Colin Johnson ¶¶, Wolfgang Schawaller##, Renato Regalin¤¤, Cosimo Baviera««, Saverio Rocchi »», Fabio Cianferoni»»,˄˄, Ron Beenen ˅˅, Michael Schmitt ¦¦, David Sassi ˀˀ, Horst Kippenbergˁˁ, Marcello Franco Zampetti₩, Marco Trizzino ₵₵, Stefano Chiari‡, Giuseppe Maria Carpanetoℓℓ, Simone Sabatelli‡, Yde de Jong ₰₰,₱₱ ‡ Sapienza Rome University, Department of Biology and Biotechnologies 'C. Darwin', Rome, Italy § Museo Nacional de Ciencias Naturales, Madrid, Spain | CSIRO Entomology, Canberra, Australia ¶ Umea University, Umea, Sweden # National Museum Prague, Prague, Czech Republic ¤ Queensland Museum, Brisbane, -
Taxonomy of the Reticulate Beetles of the Subfamily Cupedinae (Coleoptera: Archostemata), with a Review of the Historical Development A.G
Invertebrate Zoology, 2016, 13(2): 61–190 © INVERTEBRATE ZOOLOGY, 2016 Taxonomy of the reticulate beetles of the subfamily Cupedinae (Coleoptera: Archostemata), with a review of the historical development A.G. Kirejtshuk1,2, A. Nel2, P.A. Kirejtshuk3 1 Zoological Institute of the Russian Academy of Sciences, Universitetskaya emb. 1, St. Petersburg, 199034, Russia. E-mail: [email protected], [email protected] 2 Institut de Systématique, Évolution, Biodiversité, ISYEB - UMR 7205 – CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France. E-mail: [email protected] 3 6th Linia, 49-34, St. Petersburg, 199004, Russia. E-mail: [email protected] ABSTRACT:The paper is mainly devoted to the studies of the fossil Cupedidae from the Paleocene of Menat (Puy-de-Dôme, France) and to a preliminary generic analysis of the subfamily Cupedinae. All modern and fossil taxa of genus rank which could belong to Cupedinae are revised. The previous division of this subfamily into tribes (including Priacmini and Mesocupidini) is regarded as not advisable. An annotated list of the fossil species of the subfamily Cupedinae is compiled and revised. New keys to subfamilies of Cupedidae and to genera and subgenera of Cupedinae, and also keys for species of Cainomerga subgen.n. and Cupes found in deposits from Menat are given. As a result, several new genera and subgenera are proposed, i.e. the fossil Apriacma gen.n., from the early Cretaceous (the type species: Priacma tuberculosa Tan, Ren et Shin, 2006), Cre- tomerga gen.n., from the early Cretaceous (the type species: Priacmopsis subtilis Tan et Ren, 2006), Cupopsis gen.n. -
Mckenna2009chap34.Pdf
Beetles (Coleoptera) Duane D. McKenna* and Brian D. Farrell and Polyphaga (~315,000 species; checkered beetles, Department of Organismic and Evolutionary Biology, 26 Oxford click beetles, A reP ies, ladybird beetles, leaf beetles, long- Street, Harvard University, Cambridge, MA 02138, USA horn beetles, metallic wood-boring beetles, rove beetles, *To whom correspondence should be addressed scarabs, soldier beetles, weevils, and others) (2, 3). 7 e ([email protected]) most recent higher-level classiA cation for living beetles recognizes 16 superfamilies and 168 families (4, 5). Abstract Members of the Suborder Adephaga are largely preda- tors, Archostemata feed on decaying wood (larvae) and Beetles are placed in the insect Order Coleoptera (~350,000 pollen (adults), and Myxophaga are aquatic or semi- described species). Recent molecular phylogenetic stud- aquatic and feed on green and/or blue-green algae ( 6). ies defi ne two major groups: (i) the Suborders Myxophaga Polyphaga exhibit a diversity of habits, but most spe- and Archostemata, and (ii) the Suborders Adephaga and cies feed on plants or dead and decaying plant parts Polyphaga. The time of divergence of these groups has (1–3). 7 e earliest known fossil Archostemata are from been estimated with molecular clocks as ~285–266 million the late Permian (7), and the earliest unequivocal fossil years ago (Ma), with the Adephaga–Polyphaga split at ~277– Adephaga and Polyphaga are from the early Triassic (1). 266 Ma. A majority of the more than 160 beetle families Myxophaga are not known from the fossil record, but are estimated to have originated in the Jurassic (200–146 extinct possible relatives are known from the Permian Ma). -
Inizio Anno 2018 MM
ATTI DELLA ACCADEMIA NAZIONALE ITALIANA DI ENTOMOLOGIA RENDICONTI Anno LXVI 2018 TIPOGRAFIA COPPINI - FIRENZE ATTI DELLA ACCADEMIA NAZIONALE ITALIANA DI ENTOMOLOGIA RENDICONTI Anno LXVI 2018 TIPOGRAFIA COPPINI - FIRENZE ISSN 0065-0757 Direttore Responsabile: Prof. Romano Dallai Presidente Accademia Nazionale Italiana di Entomologia Coordinatore della Redazione: Dr. Roberto Nannelli La responsabilità dei lavori pubblicati è esclusivamente degli autori Registrazione al Tribunale di Firenze n. 5422 del 24 maggio 2005 INDICE Rendiconti Consiglio di Presidenza . Pag. 5 Elenco degli Accademici . »6 Verbali delle adunanze del 23-24 febbraio 2018 . »9 Verbali delle adunanze del 8-9 giugno 2018 . »19 Verbali delle adunanze del 16-17 novembre 2018 . »27 Commemorazione ROBERTO NANNELLI – Breve ricordo di Fausta Pegazzano . »51 Letture GIUSEPPE GARGIULO – Interazione ospite-parassitoide: analisi dei fattori di virulenza nel modello Drosophila . »57 EMILIO BALLETTO, FRANCESCA BARBERO, SIMONA BONELLI,LUCA PIETRO CASACCI – La mirmecofilia nelle farfalle diurne europee . »63 ROBERTO A. PANTALEONI, MAURO RUSTICI – Il nemico condiviso e l’entomologia applicata: alcune chiavi di lettura . »69 Tavola Rotonda su: DISCESE TERMICHE E TEMPERATURE BASSE E ULTRABASSE PER LA CREAZIONE DI COLLEZIONI VIVENTI DI INTERESSE AGROFORESTALE PIO FEDERICO ROVERSI, FRANCESCO PAOLI, SILVIA LANDI – Embriologia e crioconservazione di insetti . »77 MAURIZIO LAMBARDI – La crioconservazione di specie arboree: dal laboratorio alla criobanca . »81 SAURO SIMONI, ENRICO DE LILLO – Note su conservazione a basse temperature di ceppi di acari utilizza- bili per il controllo biologico . »89 GIAN PAOLO BARZANTI, VALERIA FRANCARDI – Tecniche per la crioconservazione di microrganismi ento- mopatogeni . »95 SILVIA LANDI, GIULIA TORRINI, EUSTACHIO TARASCO – Temperature ultrabasse e protocolli cryo per la costituzione di collezioni viventi di nematodi entomopatogeni . -
From the Middle Mesozoic of China Illuminating the Phylogeny of Ommatidae Jingjing Tan1,2, Yongjie Wang2, Dong Ren2* and Xingke Yang1*
Tan et al. BMC Evolutionary Biology 2012, 12:113 http://www.biomedcentral.com/1471-2148/12/113 RESEARCH ARTICLE Open Access New fossil species of ommatids (Coleoptera: Archostemata) from the Middle Mesozoic of China illuminating the phylogeny of Ommatidae Jingjing Tan1,2, Yongjie Wang2, Dong Ren2* and Xingke Yang1* Abstract Background: Ommatidae is arguably the “most ancestral” extant beetle family. Recent species of this group are only found in South America and Australia, but the fossil record reveals a much broader geographical distribution in the Mesozoic. Up to now, thirteen fossil genera with more than 100 species of ommatids have been described. However, the systematic relationships of the extant and extinct Ommatidae have remained obscure. Three constraint topologies were designed based on Kirejtshuk’s hypothesis, enforced the monophyly of Tetraphalerus + Odontomma, Pareuryomma + Notocupes and both respectively. Results: In this study, four new species, Pareuryomma ancistrodonta sp. nov., Pareuryomma cardiobasis sp. nov., Omma delicata sp. nov., and Tetraphalerus decorosus sp. nov., are described. Based on well-preserved fossil specimens and previously published data the phylogenetic relationships of extant and extinct lineages of Ommatidae were analyzed for the first time cladistically. Based on the results we propose a new classification with six tribes of Ommatidae: Pronotocupedini, Notocupedini, Lithocupedini, Brochocoleini, Ommatini and Tetraphalerini. These taxa replace the traditional four subfamilies. Conclusion: There is good support for the monophyly of the ingroup. Notocupedini, as defined by Ponomarenko, are paraphyletic. Notocupoides + Eurydictyon are the sister group of the remaining fossil and extant ommatids. Together they form the clade Pronotocupedini. Notocupedini and Lithocupedini are the next two branches. -
The Species-Level Phylogeny of Archostematan Beetles—Where Do Micromalthus Debilis and Crowsoniella Relicta Belong?
Systematic Entomology (2009), 34, 533–558 The species-level phylogeny of archostematan beetles—where do Micromalthus debilis and Crowsoniella relicta belong? THOMAS HORNSCHEMEYER¨ Department of Morphology and Systematics, Johann-Friedrich-Blumenbach-Institute for Zoology and Anthropology, Georg-August-University Gottingen,¨ Gottingen,¨ Germany Abstract. A species-level phylogenetic analysis comprising 37 of the 45 known extant species of archostematan beetles and a total of 110 morphological characters from adults and larvae is presented. For the first time, characters of the male genitalia are included in a phylogenetic analysis of Archostemata. The dataset is analysed with parsimony as well as with Bayesian algorithms. Analyses with differently arranged datasets, with larval characters included or excluded, and including or excluding Micromalthus debilis and Crowsoniella relicta are conducted. The resulting hypothesis of the species-level phylogeny of Archostemata confirms Cupedidae and Ommatidae as monophyletic taxa. Within Cupedidae, the South American Paracupes and the North American Priacma together are the sister group to all remaining Cupedidae. Among the latter, the identification of a clade comprising Rhipsideigma, Cupes capitatus and Tenomerga leucophaea renders Tenomerga polyphyletic. Ascioplaga scalena comb. nov. is transferred from Adinolepis. Characters of the male genitalia support phylogenetic affinities of Micromalthus debilis with Ommatidae and of Crowsoniella relicta with Cupedidae. Introduction and 36 species; and the monotypic families Crowsoniellidae, Micromalthidae and possibly Jurodidae (Lawrence & Newton, Archostemata is one of four suborders of beetles (Coleoptera) 1995; Kirejtshuk, 1999; Hornschemeyer,¨ 2005). (e.g. Crowson, 1960; Lawrence & Newton, 1995), and consti- Crowsoniella relicta and Micromalthus debilis are remark- tutes the smallest and the least investigated of these suborders. -
Evolution of the Insects
CY501-PIND[733-756].qxd 2/17/05 2:10 AM Page 733 Quark07 Quark07:BOOKS:CY501-Grimaldi: INDEX 12S rDNA, 32, 228, 269 Aenetus, 557 91; general, 57; inclusions, 57; menageries 16S rDNA, 32, 60, 237, 249, 269 Aenigmatiinae, 536 in, 56; Mexican, 55; parasitism in, 57; 18S rDNA, 32, 60, 61, 158, 228, 274, 275, 285, Aenne, 489 preservation in, 58; resinite, 55; sub-fossil 304, 307, 335, 360, 366, 369, 395, 399, 402, Aeolothripidae, 284, 285, 286 resin, 57; symbioses in, 303; taphonomy, 468, 475 Aeshnoidea, 187 57 28S rDNA, 32, 158, 278, 402, 468, 475, 522, 526 African rock crawlers (see Ambermantis wozniaki, 259 Mantophasmatodea) Amblycera, 274, 278 A Afroclinocera, 630 Amblyoponini, 446, 490 aardvark, 638 Agaonidae, 573, 616: fossil, 423 Amblypygida, 99, 104, 105: in amber, 104 abdomen: function, 131; structure, 131–136 Agaoninae, 423 Amborella trichopoda, 613, 620 Abies, 410 Agassiz, Alexander, 26 Ameghinoia, 450, 632 Abrocomophagidae, 274 Agathiphaga, 560 Ameletopsidae, 628 Acacia, 283 Agathiphagidae, 561, 562, 567, 630 American Museum of Natural History, 26, 87, acalyptrate Diptera: ecological diversity, 540; Agathis, 76 91 taxonomy, 540 Agelaia, 439 Amesiginae, 630 Acanthocnemidae, 391 ages, using fossils, 37–39; using DNA, 38–40 ametaboly, 331 Acari, 99, 105–107: diversity, 101, fossils, 53, Ageniellini, 435 amino acids: racemization, 61 105–107; in-Cretaceous amber, 105, 106 Aglaspidida, 99 ammonites, 63, 642 Aceraceae, 413 Aglia, 582 Amorphoscelidae, 254, 257 Acerentomoidea, 113 Agrias, 600 Amphientomidae, 270 Acherontia atropos, 585 -
202 • Stato Della Biodiversità in Italia
202 • STATO DELLA BIODIVERSITÀ IN ITALIA VEGETAZIONE E HABITAT PRIORITARI In base alla seconda concezione, la floristico-ecologica, [Edoardo Biondi] detta fitosociologica o sociologico vegetale, vengono in- dagati gli aspetti associativi delle piante, con l’individua- La vegetazione è la copertura vegetale della terra, il ri- zione di comunità vegetali, le associazioni appunto, che sultato della distribuzione e della combinazione delle pian- sono alla base di un sistema gerarchico di classificazione. te nei diversi luoghi determinata dai fattori ecologici, bio- Secondo il fondatore della fitosociologia, BRAUN-BLAN- tici e abiotici e dall’azione antropica. La vegetazione co- QUET (1915), «l’associazione è un aggruppamento vege- stituisce l’aspetto più rilevante della fitocenosi, in quan- tale più o meno stabile e in equilibrio con l’ambiente, ca- to definita dall’insieme delle piante che popolano il bio- ratterizzato da una composizione floristica determinata, topo, nel quale le singole specie trovano il necessario “spa- nel quale alcuni elementi esclusivi o quasi (specie carat- zio” vitale, la propria nicchia ecologica. La competizione teristiche) rivelano con la loro presenza un’ecologia par- tra specie è quindi alla base della costituzione della vege- ticolare e autonoma». tazione come la qualità e la quantità delle risorse dispo- Nella fitosociologia attuale vengono riconosciuti tre nibili presenti nel sito. principali livelli di analisi: La scienza della vegetazione studia le comunità vege- - quello della Fitosociologia classica, floristica ed ecolo- tali analizzandole principalmente per quanto concerne: gica, detta anche sigmatista o meglio braun-blanquet- - la composizione floristica e la struttura, tista, mediante il quale si definiscono le associazioni, i - le condizioni ecologiche che ne consentono la soprav- livelli gerarchici ad esse collegati (sintaxa) e la loro eco- vivenza e lo sviluppo, logia (sinecologia); - le modalità con le quali partecipano alla costruzione - quello della Sinfitosociologia o Fitosociologia seriale, del paesaggio vegetale. -
Li, Y-D., Liu, Z-H., Jarzembowski, E. A., Yin, Z-W., Huang, D-Y., & Cai, C-Y
Li, Y-D., Liu, Z-H., Jarzembowski, E. A., Yin, Z-W., Huang, D-Y., & Cai, C-Y. (2019). Early evolution of Cupedidae revealed by a mid- Cretaceous reticulated beetle from Myanmar (Coleoptera: Archostemata). Systematic Entomology, 44(4), 777-786. https://doi.org/10.1111/syen.12355 Peer reviewed version Link to published version (if available): 10.1111/syen.12355 Link to publication record in Explore Bristol Research PDF-document This is the author accepted manuscript (AAM). The final published version (version of record) is available online via American Association for the Advancement of Science at https://science.sciencemag.org/content/365/6459/eaaw6974 . Please refer to any applicable terms of use of the publisher. University of Bristol - Explore Bristol Research General rights This document is made available in accordance with publisher policies. Please cite only the published version using the reference above. Full terms of use are available: http://www.bristol.ac.uk/red/research-policy/pure/user-guides/ebr-terms/ 1 Early evolution of Cupedidae revealed by a mid-Cretaceous reticulated beetle from Myanmar 2 (Coleoptera: Archostemata) 3 4 YAN-DA LI1,2, ZHEN-HUA LIU3,4, EDMUND A. JARZEMBOWSKI5,6, ZI-WEI YIN7, DI-YING HUANG5 5 and CHEN-YANG CAI8,2* 6 7 1School of Life Sciences, Peking University, Beijing, China, 2School of Earth Sciences, University of Bristol, 8 Bristol, UK, 3State Key Laboratory of Biocontrol, Key Laboratory of Biodiversity Dynamics and 9 Conservation of Guangdong Higher Education Institute, College of Ecology and Evolution, -
Coleoptera: Archostemata: Cupedidae)
73 (2): 239 – 258 20.8.2015 © Senckenberg Gesellschaft für Naturforschung, 2015. Morphology of the first instar larva of Tenomerga mucida (Chevrolat, 1829) (Coleoptera: Archostemata: Cupedidae) Margarita I. Yavorskaya 1, 2, Kazuki Kojima 3, Ryuichiro Machida 3 & Rolf G. Beutel *, 1 1 Institut für Spezielle Zoologie und Evolutionsbiologie, FSU Jena, 07743 Jena, Germany; Rolf G. Beutel * [[email protected]] — 2 Department of Entomology, Biological Faculty, Lomonosov Moscow State University, Leninskie gory 1 – 12, Moscow, Russia — 3 Sugadaira Montane Research Center, University of Tsukuba, Sugadaira Kogen, Ueda, Nagano 386-2204, Japan — * Corresponding author Accepted 05.vi.2015. Published online at www.senckenberg.de/arthropod-systematics on 07.viii.2015. Abstract External and internal structures of 1st instar larvae of Tenomerga mucida are described and illustrated in detail. The larvae display previous ly identified archostematan autapomorphies such as the posterodorsal and posteroventral cephalic emarginations, the median endocarina, a distinctly reduced epicranial suture, the lateral cardinal sclerite, the loss of M. craniocardinalis, a sclerotized prominent ligula, a distinctly reduced tentorium, the transverse muscle of the posterior head capsule, tergal and sternal ampullae, and the strongly muscularized loop of the hind gut. Unusual plesiomorphies are the presence of a well-developed M. frontolabralis (M8) and a tentoriomandibular muscle. The presence of two extrinsic premental muscles and a subdivided postlabium are also plesiomorphic features. A derived character state described for the first time in an archostematan larva is the presence of glands in the anterior and posterior abdomen. The postcephalic musculature is similar to the condition found in Micromalthus and Rhipsideigma. The neck musculature is strongly developed, and also the dorsal and ventral longitudinal muscles, probably in correlation with wood-boring habits.