Silva Gabreta vol. 22 p. 79–98 Vimperk, 2016

Vegetation succession of a post-harvested peat bog and survival of and butterflies (): a case study of the Soumarský Most bog (Bohemian Forest, South Bohemia, )

Josef Jaroš*, Karel Spitzer & Hana Zikmundová Institute of Entomology, Biology Centre CAS, Branišovská 31, CZ-37005 České Budějovice, * [email protected] Abstract Moths and butterflies (Lepidoptera) communities were investigated in the post-harvested valley peat bog Soumarský Most in the Bohemian Forest during vegetation season of 2011–2016. Total 371 of all peat bog ecological categories were recorded. The survival of moths and butterflies was interpreted under natural vegetation succession and revitalization. The Soumarský Most bog seems to be colonized by bog specialists (tyrphobionts and tyrphophiles) from surrounding pristine peatlands of the Vltavský Luh complex (including the Mrtvý Luh bog), which is the unique source of recolonization by relict taxa. Only small insect species proportion probably survived the heavy peat exploitation of the Soumarský Most bog in the past time.

Key words: moths, butterflies, peat bog, ecological succession, conservation, Šumava Mountains, Vltavský Luh reserve

INTRODUCTION Peat bogs of central Europe are relict unique ecosystems existing continuously since the early Holocene period at the same sites. The conservation priorities are the virgin and semi- virgin localities in South Bohemia, namely the peat bogs of the Bohemian Forest (Šumava Mountains) – cf. SCHREIBER (1924), BUFKOVÁ et al. (2005), SPITZER & DANKS (2006), SPITZER & BUFKOVÁ (2013). The extrazonal character of such peat bogs considering the existence of a large zone of similar peatland ecosystems in the North and Northeast Europe, gives the central European peat bogs a large biogeographical context in entomofauna and vegetation (for the list of Lepidoptera and bibliography see SPITZER & DANKS 2006, JAROŠ et al. 2014). The most important human-made impacts are change of hydrological conditions, extrac- tion of horticulture peat, and forest plantations. Even in the Bohemian Forest are some lo- calities disturbed by human activities. The most important conservation of pristine peatlands needs supplementary projects of restoration ecology including studies of insect bioindica- tors. The extremely useful seems to be the Soumarský Most bog, which is under a project of revitalization and restoration of hydrological conditions. Hydrology of the Soumarský Most bog was disturbed several times for peat exploitation and forest management (LANTA et al. 2004, HORN 2009, HORN & BASTL 2012). Data of insect bioindicators (namely Lepidoptera) associated with restoration of the Soumarský Most bog are badly needed in continuation of our previous research in virgin/semivirgin peat bogs of the Bohemian Forest.

79 MATERIAL AND METHODS Study site All revised Lepidoptera records from the Soumarský Most bog were noted (2011–2016) in two basic revitalized units of wet and dry parts of the investigated bog (cf. HORN 2009). The locality Soumarský Most bog is a large peat bog (90 ha) – 48°54'N 13°50'E, altitude 745 m a.s.l. The original climax formation had to be association Pino rotundatae-Sphagnetum Kästner et Flössner 1933, surrounded by association Vaccinio uliginosi-Pinetum Kleist 1949. But only small fragments survived after large scale peat exploitation (HORN 2009). The lo- cality is a part of the large wetland complex Vltavský Luh with several well preserved and mostly pristine valley peat bogs, e.g. the Mrtvý Luh bog (cf. BUFKOVÁ et al. 2005, SPITZER & BUFKOVÁ 2013, JAROŠ et al. 2014). First original description of vegetation was published by SCHREIBER (1924) with notes about partial bog exploitation. The impact of peat exploitation was extremely strong between 1958–1999 (LANTA et al. 2004, HORN 2009). The vegetation of the Soumarský Most bog before revitalization and restoration was described by HORN (2009) who recorded also plant colonization of a peat barren ground. From entomological point of view, the most important have been improvement of hydrological conditions and introduction of basic peat plants: spp., spp., and Carex rostrata (Fig. 1, cf. HORN 2009). Relatively rare records of spp. (namely V. uliginosum, Fig. 2) form food plants complex for some tyrphobiontic and tyrphophilous . Surveys The species of Lepidoptera were studied for a period 2011–2016. The main method of sam- pling moths was UV light trapping by modified Pennsylvanian light trap (8 W UV lamp). The light was placed directly on the ground. Diurnally active butterflies and moths were sampled by netting. Larvae were collected from food plants individually and reared in the laboratory. Nomenclature of Lepidoptera follows JAROŠ et al. (2014). The four basic ecological categories were used (see e.g. MIKKOLA & SPITZER 1983, SPITZER & DANKS 2006): (1) tyrphobiontic species – bog specialists (stenotopic cold-adapted species of bogs); (2) tyrphophilous species (characteristic for bog, but not confined to them, more abundant on bogs than in adjacent habitat); (3) tyrphoneutral species (resident in bogs, but also common in other habitats); (4) tyrphoxenous species (non resident vagrants, often mi- gratory species that cannot live permanently in bogs).

RESULTS AND DISCUSSION During our five-year study, 371 species and 2758 individuals of Lepidoptera were recorded: ten species of which are tyrphobionts, 27 species tyrphophilous, 281 species tyrphoneutral, and 53 tyrphoxenous species (Appendix 1). Our results of the Soumarský Most bog con- forms low number of tyrphobiontic species and high proportion of tyrphophiles and tyr- phoneutrals (cf. JAROŠ et al. 2014). Tyrphoxens are associated with early succession and/or temporary colonization by migrants from other habitat (Appendix 1 – e.g. Eurrhypara hor- tulata and Inachis io). The most abundant are tyrphoneutrals in all stages of the succession, wet and dry habitats (see Appendix 1). From conservation point of view, the most interesting groups are tyrphobionts, which are recorded from pristine bogs, some taxa are common in the nearest large peat bog Mrtvý Luh and adjacent localities. All the Mrtvý Luh populations of tyrphobionts are more di- verse and abundant compared to Soumarský Most bog populations with exception of Am- phipoea lucens associated with wet peaty grasslands (see ELSNER et al. 1981, SPITZER &

80 Fig. 1. The Soumarský Most bog (May 2011) – communities of natural peatland succession, dominant suc- cession of .

Fig. 2. The Soumarský Most bog (July 2016) – relatively rare communities of .

81 Table 1. Tyrphobiontic Lepidoptera of the revitalized Soumarský Most bog (SM) and adjacent pristine Mrtvý Luh bog (ML) and Velká Niva bog (VN) Species SM ML VN haworthana (Stephens, 1834) + + + kilmunella Stainton, 1849 0 + + uliginosella Glitz, 1872 0 + + Athrips pruinosellus (Lienig & Zeller, 1846) 0 + + Chionodes nebulosellus (Heinemann, 1870) Gelechiidae 0 + 0 Chionodes viduellus (Fabricius, 1794) Gelechiidae 0 + 0 fraterculana Krogerus 1946 0 0 + gimmerthaliana (Lienig & Zeller, 1846) Tortricidae 0 + + Olethreutes turfosanus (Herrich-Schäffer, 1851) Tortricidae 0 + 0 luedersiana (Sorhagen, 1885) Tortricidae 0 + + alienellus (Germar & Kaulfuss, 1817) 0 + 0 Pediasia truncatella (Zetterstedt, 1839) Crambidae + + 0 Colias palaeno (Linnaeus, 1761) Pieridae + + + Boloria aquilonaris (Stichel, 1908) Nymphalidae 0 + 0 Proclossiana eunomia (Esper, 1799) Nymphalidae 0 + + Vacciniina optilete (Knoch, 1781) Lycaenidae + + + melanaria (Linnaeus, 1758) Geometridae + + + Carsia sororiata (Hübner, 1813) Geometridae 0 + + menyanthidis (Esper, 1789) + + + lucens (Freyer, 1845) Noctuidae + + + haworthii (Curtis, 1829) Noctuidae + + + subrosea (Stephens, 1829) Noctuidae + + + cordigera (Thunberg, 1788) Noctuidae 0 + 0 Lithophane lamda (Fabricius, 1787) Noctuidae + + + Xestia alpicola (Zetterstedt, 1839) Noctuidae 0 + 0 Total number of tyrphobiontic species 10 24 17

DANKS 2006, JAROŠ et al. 2014, and Table 1). Glyphipteryx haworthana (Figs. 3,4) and Ce- laena haworthii are species with high probability associated with early succession of Erio- phorum spp. and expanding Sphagnum cover after revitalization. Several other tyrphobionts are characteristic for drier habitats of relict Vaccinium uliginosum, which is the food plant of Colias palaeno. Already ELSNER et al. (1981) recorded several peatland Lepidoptera from the Soumarský Most bog including the tyrphobiontic moth Crambus alienellus, which disap- peared after heavy exploitation and disturbance of water system of the bog. Some forest species (F in Appendix 1) are also associated with Vaccinium formations. The tyrphophiles are often common (Appendix 1), some are important for conservation and rare outside peat- lands. The only exception is Crambus uliginosellus discovered in the Soumarský Most bog, which is a rare species in Czechia and has only one locality in the Bohemian Forest (Ap- pendix 1).

82 Fig. 3. Eriophorum vaginatum and freshly emerged male of , the Soumarský Most bog, 10 May 2012.

Fig. 4. Glyphipterix haworthana is common in all peatlands near Soumarský Most bog (e.g. the Velká Niva bog, 16 May 2013) and strongly associated with the Sphagnum and Eriophorum spp. complex.

83 CONCLUSIONS Revitalization of the Soumarský Most bog (see HORN 2009) seems to be important from lepidopterological point of view. Most of the species colonize the locality (wet parts espe- cially) from near virgin peatlands (Mrtvý Luh bog, Malá Niva bog and some peaty frag- ments, “stepping stones” for some biota). If the revitalization of the Soumarský Most bog is successful, the territory of the large reserve Vltavský Luh will be completed. The whole Vltavský Luh reserve is one of the most important parts of the core zone of the Šumava National Park with unique Lepidoptera. The preservation of peatland hydrological condi- tions and elimination of closed forest vegetation are basic implementations for con- servation of relict insects, not only butterflies and moths (SPITZER & DANKS 2006).

Acknowledgements. The Biology Centre CAS, Institute of Entomology provided facilities for our research (RVO: 60077344).

REFERENCES

BUFKOVÁ I., PRACH K. & BASTL M., 2005: Relationships between vegetation and environment within the montane floodplain of the Upper Vltava River (Šumava National Park, Czech Republic). Silva Gabreta, 2 (suppl.): 5– 76. ELSNER G., KRAMPL F., NOVÁK I. & SPITZER K., 1981: K poznání mikrolepidopter (Lepidoptera) šumavských rašelinišť [Microlepidoptera of the Šumava Mountains peat bogs]. Sborník Jihočeského Muzea v Českých Budějovicích, Přírodní Vědy, 21: 73–88 (in Czech). HORN P., 2009: Ekologie rašelinišť na Šumavě [Mire ecology in the Šumava Mountains]. Ms., Ph.D. thesis, Fac- ulty of Science, University of South Bohemia, České Budějovice, 98 pp. HORN P. & BASTL M., 2012: Restoration of the mined peatbog Soumarský Most. In: Ecological Restoration in the Czech Republic, JONGEPIEROVÁ I., PEŠOUT P., JONGEPIER J.W. & PRACH K. (eds) Nature Conservation Agency of the Czech Republic, Prague: 82–84. JAROŠ J., SPITZER K. & ZIKMUNDOVÁ H., 2014: Variability of Lepidoptera communities (moths and butterflies) along an altitudinal gradient of peat bogs from the Třeboň Basin up to the Bohemian Forest (South Bohemia, Central Europe). Silva Gabreta, 20: 55–95. LANTA V., DOLEŽAL J. & ŠAMATA J., 2004: Vegetation patterns in a cut-away peatland in relation to abiotic and biotic factors: a case study from the Šumava Mts., Czech Republic. Suo, 55: 33–43. MIKKOLA K. & SPITZER K., 1983: Lepidoptera associated with peatlands in central and northern Europe: a synthe- sis. Nota Lepidopterologica, 6: 216–229. SCHREIBER H., 1924: Moore des Böhmerwaldes und des deutschen Südböhmen. IV. Sebastianberg, 119 pp. SPITZER K. & BUFKOVÁ I., 2013: Peatlands of Šumava. Administration of the Šumava National Park and Protected Landscape Area, Vimperk, 212 pp. SPITZER K. & DANKS H.V., 2006: Insect biodiversity of boreal peat bogs. Annual Review of Entomology, 51: 137– 161. Received: 18 August 2016 Accepted:26 September 2016

84 Vegetation type Vegetation W W / F W / HG W / HG W / F W / F W / HG W / F W / F HG / W W W W W W W W W W W W W W W W W W 2 1 1 1 5 2 6 1 1 2 3 3 2 1 2 2 1 2 1 4 26 18 15 81 53 115 Total 0 0 9 0 0 4 1 5 1 0 1 0 1 6 1 0 1 0 0 9 1 1 3 11 23 13 2016 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 6 0 0 0 2014 0 0 3 1 0 0 1 0 1 0 1 1 1 0 1 0 0 0 1 1 0 0 0 0 65 24 2013 1 0 0 0 0 0 0 0 0 0 1 2 7 0 2 1 2 0 0 1 0 0 1 10 52 10 2012 0 1 4 0 1 1 1 0 0 0 0 1 0 4 1 0 0 3 0 0 0 4 1 0 0 39 2011 Tortricidae Tortricidae Tortricidae Family Glyphipterigidae Crambidae Pieridae Geometridae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Psychidae Elachistidae Gelechiidae Gelechiidae Crambidae Crambidae Saturnidae Satyridae Lycaenidae Tortricidae Tortricidae Tortricidae Tortricidae (Denis & 1775) Schiffermüller, Zeller, 1850 Zeller, (Humphries 1845 & Westwood, (Nylander, 1848) (Nylander, (Herrich-Schäffer, 1854) (Herrich-Schäffer, (Geyer, 1832) (Geyer, (Müller, 1764) (Müller, (Zetterstedt, 1839) (Clerck, 1759) (Fabricius, 1787) (Knoch, 1781) (Hübner, 1799) (Hübner, (Linnaeus, 1758) (Treitschke, 1830) Acronicta Acronicta menyanthidis 1789) (Esper, Biselachista albidella Neofaculta infernella myrtillana Ancylis unguicella (Frölich, 1828) Bactra lancealana margaritella Coenonympha tullia Pediasia truncatella Colias palaeno (Linnaeus, 1761) optilete Vacciniina Arichanna melanaria (Linnaeus, 1758) Amphipoea lucens 1845) (Freyer, Celaena haworthii (Curtis, 1829) (Stephens, 1829) Lithophane lamda Tyrphophiles Sterrhopterix 1851) standfussi (Wocke, Pleurota bicostella Neofaculta ericetella Olethreutes micanus (Denis & 1775) Schiffermüller, Olethreutes palustranus (Lienig & 1846) Zeller, Rhopobota myrtillana Crambus uliginosellus Saturnia pavonia (Linnaeus, 1758) Species Tyrphobionts Glyphipterix haworthana (Stephens, 1834) Ap pendix 1. List of Lepidoptera of the Soumarský Most bog. W F – Wetland, – Forest, HG – Heathland-Grassland, tyrphoxenous species are not classified, their habitat association is not permanent (e.g. many migratory species).

85 Vegetation type Vegetation HG / W W / HG W / F HG / W W / F W / HG F / W W / HG / F W / F W W W F W F HG F F F F F F F W F HG W 3 1 1 1 5 1 1 3 1 1 4 1 3 1 1 1 3 1 1 1 1 1 2 1 26 29 10 Total 3 0 0 1 3 0 1 1 0 1 1 1 0 3 1 0 0 0 0 0 0 0 0 2 0 11 16 2016 0 0 0 0 0 1 6 0 0 0 1 0 0 0 0 5 0 0 0 0 0 1 0 0 0 0 0 2014 0 0 0 0 0 0 5 0 0 2 0 0 0 0 3 1 0 0 1 2 1 0 0 0 0 0 1 2013 0 0 1 1 0 0 2 1 0 0 0 0 3 0 0 1 0 0 0 1 0 0 1 1 1 0 0 2012 0 0 0 0 1 5 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 10 2011 Lycaenidae Yponomeutidae Family Geometridae Geometridae Geometridae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Psychidae Gracillariidae Gracillariidae Gracillariidae Gracillariidae Bucculatricidae Oecophoridae Oecophoridae Oecophoridae Oecophoridae Elachistidae Elachistidae Yponomeutidae Yponomeutidae (Hübner, 1817) (Hübner, Zeller, 1847 Zeller, (Toll, 1956) (Toll, (Hübner, 1813) (Hübner, (Hübner, 1813) (Hübner, (Linnaeus, 1758) (Hübner, 1813) (Hübner, (Denis & 1775) Schiffermüller, Zeller, 1839 Zeller, (Hübner, 1803) (Hübner, (Fabricius, 1781) (Hering, 1928) (Linnaeus, 1758) (Stainton, 1849) Stainton, 1854 (Stainton, 1854) (Esper, 1788) (Esper, (Linnaeus, 1761) (Linnaeus, 1758) (Linnaeus, 1758) (Linnaeus, 1761) Callophrys rubi (Linnaeus, 1758) Ematurga Ematurga atomaria Eulithis testata Anarta myrtilli Eurois occultus Lycophotia porphyrea (Denis & 1775) Schiffermüller, Protolampra sobrina (Duponchel, 1843) Parornix betulae stipella Elachista canapennella Semiothisa brunneata (Thunberg, 1784) Hypenodes humidalis 1850 Doubleday, Hyppa rectilinea Lithomoia solidaginis Tyrphoneutrals Nematopogon metaxellus Epichnopterix plumella Caloptilia betulicola Caloptilia stigmatella Phyllonorycter ulmifoliellus Bucculatrix demaryella (Duponchel, 1840) conjugella Argyresthia goedartella Argyresthia Ocnerostoma piniariellum angelicella Agonopterix ciliella Pseudatemelia josephinae Elachista alpinella Species Appendix 1. Continued.

86 Vegetation type Vegetation F F W F W W F F W F W F F F F HG F F F F W F HG F F F F W 2 1 3 8 2 2 1 2 2 1 4 1 1 1 1 2 6 1 1 7 1 8 3 11 17 13 24 21 Total 0 0 0 0 0 0 2 0 0 0 0 0 0 0 0 0 1 0 0 0 1 1 0 4 1 2 9 0 2016 0 0 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 1 1 1 0 0 0 0 2 4 0 2014 1 0 0 3 0 1 0 0 0 0 0 1 1 0 0 1 0 2 5 0 0 2 0 1 1 2 11 17 2013 1 1 1 0 4 1 3 1 1 0 2 2 0 2 0 1 0 0 1 2 3 0 1 1 0 3 5 1 2012 0 0 0 4 1 0 0 1 1 0 0 1 1 0 0 0 0 0 1 1 0 0 0 0 0 2 0 16 2011 Family Gelechiidae Gelechiidae Coleophoridae Coleophoridae Coleophoridae Coleophoridae Coleophoridae Coleophoridae Coleophoridae Coleophoridae Coleophoridae Gelechiidae Gelechiidae Gelechiidae Gelechiidae Gelechiidae Gelechiidae Gelechiidae Gelechiidae Gelechiidae Gelechiidae Gelechiidae Gelechiidae Cossidae Tortricidae Tortricidae Tortricidae Nylander, 1848 Nylander, Herrich-Schäffer, 1855 Herrich-Schäffer, (Rebel, 1937) Wood, 1892 Wood, Zeller, 1839 Zeller, (Linnaeus, 1758) Zeller, 1839 Zeller, Wood, 1892 Wood, (Hübner, 1813) (Hübner, (Linnaeus, 1761 (Zeller, 1839) (Zeller, Zeller, 1849 Zeller, (Clerck, 1759) (Zeller, 1839) (Zeller, (Hübner, 1796) (Hübner, Heinemann, 1876 (Hübner, 1793) (Hübner, (Zeller, 1839) (Zeller, (Linnaeus, 1758) (Thunberg, 1794) (Stainton, 1854) (Clerck, 1759) (Linnaeus, 1758) Chionodes electellus Chionodes luctuellus Coleophora alticolella Coleophora betulella Coleophora caespititiella Coleophora glaucicolella Coleophora milvipennis Coleophora serratella Coleophora striatipennella Coleophora sylvaticella Coleophora taeniipennella binotella Acanthophila latipennella Anacampsis populella affinis (Haworth, 1828) Bryotropha galbanella Bryotropha senectella Bryotropha similis Exoteleia dodecella rufescens (Haworth, 1828) Hypatima rhomboidella Sophronia semicostella proximellus Teleiodes Cossus cossus notana (Donovan, 1806) Adoxophyes orana (Fischer von Röslerstamm, 1834) Aethes 1854) cnicana (Westwood, Species Appendix 1. Continued.

87 Vegetation type Vegetation W W / F HG / F W / HG / F W / HG / F W / F / HG W / F / HG W / F / HG W / HG W / F F F F W F F HG F F F F F F F F F F F F 1 1 1 2 1 3 5 1 1 1 1 1 3 1 3 1 8 1 3 2 1 4 8 4 1 17 13 30 Total 0 0 0 0 1 2 3 1 0 0 0 0 0 1 3 1 0 0 4 0 0 1 0 1 4 8 3 1 2016 0 0 0 0 0 0 1 0 0 2 2 0 0 0 0 0 0 0 0 0 0 0 0 0 1 7 1 0 2014 1 0 0 2 0 1 1 0 1 0 1 0 0 0 0 0 3 0 0 0 2 0 0 2 1 6 0 0 2013 0 1 1 0 0 0 0 0 0 7 1 1 0 0 0 0 0 2 1 0 0 1 0 2 8 0 0 15 2012 0 0 0 0 0 0 0 0 0 0 3 0 0 0 0 0 0 1 2 0 1 1 0 1 0 1 0 0 2011 Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Family Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae Tortricidae (Treitschke, 1835) (Haworth, 1811) (Haworth, 1811) (Fabricius, 1794) (Linnaeus, 1758) (Clerck, 1759) (Clerck, 1759) (Fabricius, 1775) (Denis & 1775) Schiffermüller, Ancylis achatana (Denis & 1775) Schiffermüller, Ancylis laetana Celypha striana (Denis & 1775) Schiffermüller, Cnephasia stephensiana 1849) (Doubleday, Cochylis dubitana 1799) (Hübner, Epiblema hepaticanum Epinotia brunnichana (Linnaeus, 1767) Olethreutes rivulanus (Scopoli, 1763) Orthotaenia undulana (Denis & 1775) Schiffermüller, Pandemis cerasana 1786) (Hübner, Ancylis geminana (Donovan, 1806) Ancylis uncella Aphelia unitana 1799) (Hübner, 1825 Hübner, Archips podanus (Scopoli, 1763) Cnephasia asseclana (Denis & 1775) Schiffermüller, Epinotia demarniana (Fischer von Röslerstamm, 1840) Epinotia nanana (Treitschke, 1835) Epinotia nisella Epinotia solandriana (Linnaeus, 1758) Epinotia tedella Epinotia tetraquetrana Epinotia trigonella Eulia ministrana (Linnaeus, 1758) Olethreutes dissolutanus (Stange, 1886) Olethreutes lacunanus (Denis & 1775) Schiffermüller, Pandemis corylana Species Appendix 1. Continued.

88 Vegetation type Vegetation W / HG W W / HG W / F HG / W W / HG W / HG F / W F / W F F F F F F F F HG W HG HG HG HG F W F F F F 1 1 1 1 1 2 1 8 6 6 2 1 2 1 2 2 5 6 1 1 11 24 28 38 21 46 19 15 Total 1 1 1 0 0 1 1 2 1 0 1 0 0 1 0 8 0 6 1 1 7 1 6 1 0 12 15 18 2016 0 0 0 0 0 0 0 0 2 0 1 0 0 0 0 0 1 1 0 2 0 1 0 0 0 3 1 0 2014 0 0 1 0 0 0 0 0 2 5 4 0 0 1 0 0 1 0 0 0 2 6 2 0 4 1 0 12 2013 0 0 9 0 1 0 0 0 1 0 0 1 1 0 0 0 8 8 1 2 3 1 9 2 0 5 1 10 2012 0 0 2 0 0 1 1 0 1 0 1 0 0 1 0 2 3 9 1 1 1 0 0 1 2 0 11 13 2011 Tortricidae Tortricidae Tortricidae Family Pyralidae Pyralidae Pyralidae Pyralidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Crambidae Tortricidae Tortricidae (Linnaeus, 1758) (Hulst, 1887) (Herrich-Schäffer, 1848) (Herrich-Schäffer, Heinemann, 1863 (Stainton, 1849) (Zincken, 1817) (La Harpe, 1855) (Denis & 1775) Schiffermüller, (Goeze, 1778) (Scopoli, 1763) (Denis & 1775) Schiffermüller, (Denis & 1775) Schiffermüller, (Linnaeus, 1758) (Curtis, 1827) Pandemis dumetana (Treitschke, 1835) Opsibotys fuscalis (Denis & 1775) Schiffermüler, Retinia resinella Rhyacionia pinivorana (Lienig & 1846) Zeller, Agriphila tristella Crambus pascuellus (Linnaeus, 1758) Crambus perlellus Eudonia truncicolella Chrysoteuchia culmella Rhopobota naevana 1817) (Hübner, Syndemis musculana 1799) (Hübner, Apomyelois bistriatella abietella Dioryctria simplicella Metriostola betulae Pyla fusca (Haworth, 1811) Agriphila straminella (Denis & 1775) Schiffermüller, Cataclysta lemnata (Linnaeus, 1758) Catoptria falsella Catoptria osthelderi (de Lattin, 1950) Catoptria permutatella Crambus lathoniellus Dipleurina lacustrata 1804) (Panzer, Donacaula mucronellus (Denis & 1775) Schiffermüller, Eudonia pallida Scoparia ambigualis (Treitschke, 1829) Scoparia ancipitella Scoparia basistrigalis Knaggs, 1866 Species Appendix 1. Continued.

89 Vegetation type Vegetation W W / HG W / F HG / W W / HG W / HG W / F F F F F F W F HG W HG HG F F F F F F F F F F F 3 3 2 7 3 2 4 1 1 6 1 1 8 1 9 1 1 1 1 2 3 7 3 31 13 59 18 10 Total 0 2 0 3 3 2 0 1 1 1 2 0 0 0 0 1 0 0 5 0 1 2 0 2 2 2 13 47 2016 0 0 0 1 0 0 0 0 0 3 1 0 0 0 0 0 0 0 0 5 0 0 0 0 2 3 0 0 2014 0 0 0 0 0 0 0 0 0 0 1 1 8 1 8 1 1 0 2 1 0 0 0 1 0 1 0 15 2013 3 0 2 2 0 0 1 0 0 1 5 0 0 0 0 0 0 0 0 5 6 0 0 0 0 4 3 0 2012 0 1 0 1 0 0 3 0 0 1 8 0 0 0 0 0 0 0 0 0 6 1 0 0 0 1 1 1 2011 Family Lasiocampidae Lasiocampidae Endromidae Sphingidae Sphingidae Sphingidae Saturniidae Hesperiidae Pieridae Pieridae Pieridae Nymphalidae Nymphalidae Nymphalidae Nymphalidae Satyridae Satyridae Drepanidae Drepanidae Drepanidae Drepanidae Drepanidae Geometridae Geometridae Geometridae Geometridae Geometridae Lycaenidae (Hufnagel, 1766) (Linnaeus, 1758) (Denis & 1775) Schiffermüller, (Linnaeus, 1758) (Linnaeus, 1758) (Rottemburg, 1775) Dendrolimus Dendrolimus pini (Linnaeus, 1758) Endromis Endromis versicolora (Linnaeus, 1758) Sphinx pinastri Linnaeus, 1758 Anthocharis cardamines (Linnaeus, 1758) Brenthis ino (Rottemburg, 1775) Nymphalis antiopa (Linnaeus, 1758) Aphantopus hyperantus (Linnaeus, 1758) Macrothylacia Macrothylacia rubi (Linnaeus, 1758) Laothoe populi (Linnaeus, 1758) Smerinthus ocellatus Aglia tau (Linnaeus, 1758) Carterocephalus palaemon (Pallas, 1771) Gonepteryx rhamni (Linnaeus, 1758) Pieris napi (Linnaeus, 1758) Boloria euphrosyne (Linnaeus, 1758) Melitaea athalia Lasiommata maera (Linnaeus, 1758) Celastrina argiolus (Linnaeus, 1758) Drepana falcataria Falcaria lacertinaria Habrosyne pyritoides Ochropacha duplaris (Linnaeus, 1761) Thyatira batis (Linnaeus, 1758) Aethalura punctulata Alcis bastelbergeri (Hirschke, 1908) Alcis repandata (Linnaeus, 1758) Biston betularius (Linnaeus, 1758) Bupalus piniarius (Linnaeus, 1758) Species Appendix 1. Continued.

90 Vegetation type Vegetation W W / F / HG F / W W / F F / W W / F W / HG / F F HG F F F F F F F F F F HG F F F F F F F F F 3 5 1 3 5 1 2 1 3 4 2 1 6 2 1 2 1 5 1 6 2 4 1 4 1 11 11 12 Total 0 1 0 2 1 1 0 1 0 2 2 1 4 2 0 0 1 0 8 1 1 0 4 4 1 3 0 3 2016 2 2 0 0 0 0 0 0 0 1 0 0 1 0 0 1 0 1 0 0 1 0 0 0 0 0 0 4 2014 0 1 0 0 0 0 2 0 1 1 0 0 0 0 0 0 0 1 0 0 1 1 0 0 0 0 0 0 2013 0 1 1 0 1 0 0 0 0 0 0 0 1 0 1 0 0 0 2 0 2 1 6 0 0 0 1 1 2012 1 0 0 1 3 0 0 0 2 0 0 0 0 0 0 1 0 3 1 0 1 0 2 0 0 1 0 3 2011 Family Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae (Linnaeus, 1758) (Hufnagel, 1767) (Denis & 1775) Schiffermüller, (Werneburg, 1859) (Werneburg, (Linnaeus, 1758) (Scopoli, 1763) Boisduval, 1840 (Denis & 1775) Schiffermüller, (Scopoli, 1763) (Linnaeus, 1758) (Linnaeus, 1861) (Linnaeus, 1758) (Müller, 1764) (Müller, (Denis & 1775) Schiffermüller, (Haworth, 1809) (Denis & 1775) Schiffermüller, Cabera exanthemata Cabera pusaria (Linnaeus, 1758) Campaea margaritata (Linnaeus, 1767) miata Epirrhoe alternata Eupithecia tantillaria Geometra papilionaria Camptogramma bilineatum Chloroclysta citrata (Hufnagel, 1767) Chloroclysta truncata (Hufnagel, 1767) Crocallis elinguaria Cyclophora albipunctata Deileptenia ribeata (Clerck, 1759) Ecliptopera silaceata Ectropis crepuscularia (Denis & 1775) Schiffermüller, Ennomos autumnarius caesiata Epione repandaria (Hufnagel, 1767) Eulithis populata (Linnaeus, 1758) Eupithecia vulgata Hydrelia sylvata Hydriomena furcata (Thunberg, 1784) Hydriomena impluviata Hylaea fasciaria (Linnaeus, 1758) Hypomecis punctinalis Hypomecis roboraria (Denis & 1775) Schiffermüller, Idaea aversata (Linnaeus, 1758) Species Appendix 1. Continued.

91 Vegetation type Vegetation F / W W / F / HG W / F W / HG W / F W / HG F / W F F F W F W F F HG F F F HG F F F HG F F F F 5 1 6 9 3 1 4 3 2 2 1 1 1 2 1 8 3 3 2 1 1 9 1 7 1 1 20 16 Total 3 0 3 1 0 0 0 2 0 0 1 0 1 0 0 2 0 2 1 1 1 0 0 7 0 0 0 0 2016 0 0 0 5 0 0 0 1 0 0 0 0 0 0 0 0 1 2 0 0 0 0 0 0 0 2 1 0 2014 2 0 2 0 3 0 0 0 0 0 0 0 0 0 1 0 0 2 0 0 0 0 0 0 1 0 0 0 2013 0 1 1 3 0 1 3 0 2 1 0 0 1 0 0 0 1 2 0 0 0 1 0 0 2 0 1 14 2012 0 0 0 0 0 0 1 0 0 1 2 0 0 0 0 0 1 0 2 1 1 0 2 0 3 0 0 19 2011 Family Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae (Retzius, 1783) (Linnaeus, 1758) (Linnaeus, 1758) (Hufnagel, 1767) (Linnaeus, 1758) (Clerck, 1759) (Stephens, 1831) (Borkhausen, 1794) (Hufnagel, 1767) (Hufnagel, 1767) (Linnaeus, 1758) (Clerck, 1759) (Borkhausen, 1794) (Hübner, 1787) (Hübner, (Hufnagel, 1767) (Scopoli, 1763) Idaea biselata Lycia Lycia hirtaria (Clerck, 1759) Plagodis pulveraria (Linnaeus, 1758) Scopula immutata (Linnaeus, 1758) Scopula nigropunctata Scotopteryx chenopodiata Semiothisa liturata Lobophora halterata Lomaspilis marginata Odezia atrata (Linnaeus, 1758) Odontopera bidentata Orthonama vittata Peribatodes secundarius (Denis & 1775) Schiffermüller, Perizoma affinitatum Perizoma albulatum (Denis & 1775) Schiffermüller, Perizoma alchemillatum Perizoma didymatum (Linnaeus, 1758) Pterapherapteryx sexalata Scopula ternata (Schrank, 1802) Selenia tetralunaria Semiothisa clathrata Semiothisa signaria 1809) (Hübner, Siona lineata Thera firmata 1822) (Hübner, Thera obeliscata Thera variata (Denis & 1775) Schiffermüller, cambrica Curtis, Venusia 1839 Xanthorhoe biriviata Species Appendix 1. Continued.

92 Vegetation type Vegetation W W / F / HG F / W / HG F / W HG / F HG / W / F F / W / HG W / HG F / HG W / HG HG / W / F HG HG F F F F F F F F F F F F F HG HG HG 1 1 3 2 1 1 2 2 5 1 1 2 2 1 1 5 6 18 15 14 25 56 10 57 38 33 19 394 Total 0 0 0 0 0 0 0 9 1 1 1 0 0 1 1 5 5 1 1 5 11 15 12 28 10 32 18 183 2016 0 0 0 0 0 0 0 0 5 0 0 0 0 2 0 1 0 6 9 2 5 0 0 1 2 0 70 12 2014 0 0 0 2 0 1 1 1 1 0 0 0 1 0 0 0 0 3 3 3 6 1 0 0 0 0 14 62 2013 0 1 0 0 1 1 0 1 0 1 4 0 0 0 1 0 0 9 8 0 8 0 0 0 1 1 30 14 2012 1 0 3 0 0 1 0 0 0 0 0 0 0 0 0 1 0 2 0 7 2 0 0 0 1 0 14 49 2011 Family Geometridae Geometridae Geometridae Geometridae Notodontidae Notodontidae Notodontidae Notodontidae Notodontidae Notodontidae Notodontidae Notodontidae Notodontidae Notodontidae Arctiidae Arctiidae Arctiidae Arctiidae Arctiidae Arctiidae Arctiidae Arctiidae Arctiidae Arctiidae Arctiidae Lymantriidae Lymantriidae Lymantriidae (Clerck, 1759) (Linnaeus, 1758) (Linnaeus, 1767) (Linnaeus, 1758) (Linnaeus, 1758) Xanthorhoe Xanthorhoe designata (Hufnagel, 1767) Xanthorhoe Xanthorhoe montanata (Denis & 1775) Schiffermüller, Xanthorhoe quadrifasciata Eligmodonta ziczac (Linnaeus, 1758) Arctia caja (Linnaeus, 1758) Cybosia mesomella (Linnaeus, 1758) Diacrisia sannio (Linnaeus, 1758) Eilema complanum (Linnaeus, 1758) Eilema 1787) depressum (Esper, Spilosoma lubricipeda Xanthorhoe Xanthorhoe spadicearia (Denis & 1775) Schiffermüller, Clostera curtula (Linnaeus, 1758) Clostera pigra (Hufnagel, 1766) Notodonta dromedarius Phalera bucephala Pheosia gnoma (Fabricius, 1776) Pheosia tremula (Clerck, 1759) Pterostoma palpinum (Clerck, 1759) Ptilodon capucina (Linnaeus, 1758) Stauropus fagi (Linnaeus, 1758) Calliteara pudibunda (Linnaeus, 1758) Lymantria monacha (Linnaeus, 1758) Orgyia antiqua (Linnaeus, 1758) Eilema lurideolum (Zincken, 1817) Eilema lutarellum (Linnaeus, 1758) Eilema (Hufnagel, sororculum 1766) Lithosia quadra (Linnaeus, 1758) Phragmatobia fuliginosa Species Appendix 1. Continued.

93 Vegetation type Vegetation HG / W / F F / W HG / F HG / F F / W W / F HG / F W / F W / HG HG / W / F W HG HG HG HG HG W HG HG F F F W HG W F F W 2 8 3 1 2 1 1 1 2 4 1 1 1 3 3 1 1 3 4 1 2 9 3 50 14 24 10 10 Total 0 4 8 0 1 1 0 0 7 1 1 4 0 0 1 2 2 1 0 0 1 0 1 3 1 21 23 10 2016 2 0 3 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 10 2014 0 9 0 0 0 0 0 0 1 1 0 0 0 0 1 0 0 0 0 0 0 1 0 0 0 0 1 0 2013 0 6 2 1 0 0 1 0 0 0 2 0 1 0 0 1 0 1 1 0 0 1 2 0 0 1 5 2 2012 0 4 2 2 1 0 0 1 0 0 1 0 0 0 0 0 0 0 0 0 1 1 1 0 1 0 0 0 2011 Family Arctiidae Arctiidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae (Linnaeus, 1758) (Fabricius, 1776) (Linnaeus, 1758) (Hübner, 1808) (Hübner, (Denis & 1775) Schiffermüller, (Hufnagel, 1766) (Linnaeus, 1758) (Linnaeus, 1758) (Linnaeus, 1767) Spilosoma luteum (Hufnagel, 1766) Thumatha senex 1808) (Hübner, Agrotis exclamationis Amphipyra tragopoginis (Clerck, 1759) Anaplectoides prasinus (Denis & 1775) Schiffermüller, crenata (Hufnagel, 1766) Autographa bractea (Denis & 1775) Schiffermüller, Blepharita satura fraxini mendica (Fabricius, 1775) Acronicta Acronicta auricoma (Denis & 1775) Schiffermüller, Acronicta megacephala (Denis & 1775) Schiffermüller, Amphipyra pyramidea Apamea lateritia Apamea monoglypha (Hufnagel, 1766) Apamea 1809) remissa (Hübner, Apamea rubrirena (Treitschke, 1825) Apamea scolopacina 1788) (Esper, Autographa pulchrina (Haworth, 1809) Brachylomia viminalis Catocala nupta Celaena leucostigma Cerapteryx graminis (Linnaeus, 1758) Chortodes pygmina (Haworth, 1809) Colocasia coryli Cosmia trapezina (Linnaeus, 1758) Deltote uncula (Clerck, 1759) Diarsia brunnea (Denis & 1775) Schiffermüller, Species Appendix 1. Continued.

94 Vegetation type Vegetation W W / HG W / HG F / HG F / W W / HG / F HG / W F F F HG HG F HG HG HG HG HG HG W HG HG HG HG F W HG F F 1 1 5 1 2 5 3 2 4 2 1 1 1 1 1 1 1 1 5 3 1 5 1 16 24 14 27 180 Total 1 1 1 0 1 2 1 2 0 1 1 0 0 0 0 0 0 0 0 1 1 1 0 0 4 0 12 42 2016 0 0 1 0 0 3 2 0 3 2 0 0 1 0 0 0 1 0 0 1 0 0 0 0 0 0 0 36 2014 0 0 2 1 0 0 0 0 1 0 0 0 0 0 9 0 0 1 0 0 0 4 4 0 0 0 0 23 2013 0 0 0 0 1 0 0 0 0 1 1 0 0 0 9 0 0 8 0 0 0 7 0 3 1 1 0 54 2012 0 0 1 0 0 0 0 0 0 1 0 0 0 1 6 1 0 5 1 0 0 0 0 0 0 1 25 15 2011 Family Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae (Goeze, 1781) (Fabricius, 1794) (Knoch, 1782) (Linnaeus, 1761) (Esper, 1788) (Esper, (Denis & 1775) Schiffermüller, (Denis & 1775) Schiffermüller, (Clerck, 1759) (Linnaeus, 1758) (Vieweg, 1790) (Vieweg, Diarsia rubi Enargia Enargia paleacea Herminia tarsicrinalis Lygephila pastinum (Treitschke, 1826) Plusia putnami (Grote, 1873) Polymixis gemmea (Treitschke, 1825) pygarga (Hufnagel, Protodeltote 1766) imbecillus Herminia tarsipennalis Treitschke, 1835 Hoplodrina blanda (Denis & 1775) Schiffermüller, Hoplodrina octogenaria Hypena proboscidalis (Linnaeus, 1758) 1809) proxima (Hübner, Laspeyria flexula Melanchra pisi (Linnaeus, 1758) Mythimna conigera (Denis & 1775) Schiffermüller, Mythimna impura 1808 (Hübner, Mythimna pallens (Linnaeus, 1758) Mythimna pudorina (Denis & 1775) Schiffermüller, Naenia typica Noctua pronuba (Linnaeus, 1758) Ochropleura plecta Oligia latruncula Orthosia gothica (Linnaeus, 1758) Parastichtis suspecta 1718) (Hübner, Polia bombycina (Hufnagel, 1766) Polia hepatica Pseudoips prasinanus (Linnaus, 1758) Species Appendix 1. Continued.

95 Vegetation type Vegetation W W / HG W / F / HG F / W W / HG W / HG / F W / HG HG / W F F F HG F HG 2 2 8 1 1 1 2 1 4 2 1 1 1 4 1 1 7 3 1 1 3 1 3 1 1 27 23 Total 1 6 1 0 0 0 1 7 0 1 0 0 1 0 1 0 0 0 1 3 0 0 1 1 0 0 0 2016 1 2 1 0 0 0 0 4 0 0 0 0 0 0 0 0 0 0 2 0 0 0 0 0 0 0 0 2014 0 7 0 0 0 0 0 1 0 0 2 0 0 0 0 2 1 0 0 0 0 0 0 0 0 0 1 2013 0 0 7 0 1 0 5 1 0 1 1 0 1 0 1 0 0 1 0 1 0 1 0 0 1 0 10 2012 0 2 0 1 1 0 0 6 1 0 1 1 0 0 0 1 0 1 3 0 0 1 1 0 3 0 0 2011 - Tineidae Roeslerstammii Yponomeutidae Yponomeutidae Family Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Adelidae Gracillariidae dae Plutellidae Oecophoridae Oecophoridae Oecophoridae Oecophoridae Coleophoridae Coleophoridae Batrachedridae (Zetterstedt, 1839) (Fabricius, 1787) (Linnaeus, 1758) (Stephens, 1835) (Linnaeus, 1758) Staudinger, 1871 Staudinger, (Standfuss, 1851) (Hübner, 1817) (Hübner, (Linnaeus, 1758) (Hummel, 1823) (Poda, 1761) (Linnaeus, 1758) (Linnaeus, 1758) (Hufnagel, 1766) (Esper, 1788) (Esper, (Scopoli, 1763) (Hübner, 1813) (Hübner, (Boisduval, 1840) Rivula sericealis (Scopoli, 1763) Tineola bisselliella Tineola Eucalybites aurogutellus Roeslerstammia erxlebella trifasciata Argyresthia evonymellus Yponomeuta Plutella xylostella Agonopterix (Haworth, liturosa 1811) Agonopterix petasitis minutella 1854 olerella Zeller, Coleophora laricella Coleophora sternipennella Batrachedra praeangusta (Haworth, 1828) Rusina ferruginea 1785) (Esper, Scoliopteryx libatrix Tholera decimalis atriplicis Trachea Xestia c-nigrum (Linnaeus, 1758) Xestia triangulum (Hufnagel, 1766) Tyrphoxens croesella Xanthia icteritia Xanthia togata Xestia baja (Denis & 1775) Schiffermüller, Xestia collina Xestia sexstrigata (Haworth, 1809) Xylena vetusta Species Appendix 1. Continued.

96 Vegetation type Vegetation 1 1 5 1 2 1 1 1 1 1 1 5 1 1 3 9 1 4 1 8 1 1 1 1 1 1 1 1 Total 0 0 1 0 0 0 1 0 0 0 0 1 0 1 1 0 0 3 0 0 1 1 1 1 1 0 0 0 2016 1 0 0 0 0 0 0 0 0 1 1 0 0 0 0 2 0 0 0 0 0 0 0 0 0 1 0 0 2014 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 1 8 0 0 0 0 0 0 1 0 2013 0 1 0 1 2 1 0 1 0 0 0 3 1 0 2 2 1 0 0 0 0 0 0 0 0 0 0 0 2012 0 0 0 0 0 0 0 0 1 0 0 1 0 0 0 5 0 0 0 0 0 0 0 0 0 0 0 1 2011 Tortricidae Tortricidae Lycaenidae Lycaenidae Family Momphidae Momphidae Gelechiidae Gelechiidae Gelechiidae Pyralidae Crambidae Crambidae Crambidae Sphingidae Nymphalidae Nymphalidae Nymphalidae Nymphalidae Nymphalidae Drepanidae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Geometridae Tortricidae (Duponchel, 1843) (Linnaeus, 1758) (Hübner, 1809) (Hübner, (Fabricius, 1775) (Hübner, 1813) (Hübner, (Sircom, 1850) (Denis & 1775) Schiffermüller, (Zeller, 1839) (Zeller, (Stephens, 1831) (Denis & 1775) Schiffermüller, (Denis & 1775) Schiffermüller, (Haworth, 1809) (Linnaeus, 1758) (Linnaeus, 1758) (Linnaeus, 1758) Mompha epilobiella Mompha raschkiella atrella Ilseopsis acuminatella Ancylis badiana (Denis & 1775) Schiffermüller, weirana (Douglas, 1850) Oncocera semirubella (Scopoli, 1763) Eurrhypara hortulata (Linnaeus, 1758) Phlyctaenia perlucidalis Pleuroptya ruralis (Scopoli, 1763) Sphinx ligustri Linnaeus, 1758 Aglais urticae Araschnia levana (Linnaeus, 1758) Inachis io (Linnaeus, 1758) Issoria lathonia atalanta Vanessa Cyaniris semiargus (Rotemburg, 1775) Polyommatus amandus 1792) (Schneider, cultraria (Fabricius, Watsonalla 1775) praeformata 1826) (Hübner, Eupithecia valerianata Eustroma reticulatum Chloroclystis v-ata Perizoma taeniatum Plemyria rubiginata (Denis & 1775) Schiffermüller, Rhinoprora rectangulata Species Appendix 1. Continued.

97 Vegetation type Vegetation 1 2 1 1 1 1 1 1 1 1 1 Total 0 1 1 0 1 1 1 0 1 1 0 2016 0 0 0 0 0 0 0 0 0 0 0 2014 0 1 0 0 0 0 0 0 0 0 1 2013 1 0 0 0 0 0 0 0 0 0 0 2012 0 0 0 1 0 0 0 1 0 0 0 2011 Lymantriidae Family Notodontidae Arctiidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae (Fabricius, 1775) (Thunberg, (Thunberg, 1784) (Clerck, 1759) Harpyia milhauseri Arctornis Arctornis l-nigrum 1764) (Müller, Nudaria mundana (Linnaeus, 1761) 1830) (Freyer, Callistege mi Dypterygia scabriuscula (Linnaeus, 1758) Charanyca trigrammica (Hufnagel, 1766) Ipimorpha retusa (Linnaeus, 1761) Meganola strigula (Denis & 1775) Schiffermüller, Nonagria typhae emortualis Trisateles (Denis & 1775) Schiffermüller, Species

98