Coleoptera: Chrysomelidae): the Generic Classification and Type Specimens of the New World Species

Home , Donacia, Donaciinae, Neohaemonia, Plateumaris

AN ANNOTATED LIST OF NEARCTIC DONACIINAE (COLEOPTERA: CHRYSOMELIDAE): THE GENERIC CLASSIFICATION AND TYPE SPECIMENS OF THE NEW WORLD SPECIES.

BY INGOLF S. ASKEVOLD Department of Entomology, University of Manitoba, Winnipeg, Manitoba, Canada R3T 2N21

INTRODUCTION During systematic revision of Nearctic donaciines, the majority of type specimens were examined. Some lectotype and neotype designations, made where necessary in the context of revisions of genera (Plateumaris Thomson and Neohaemonia Sz6kessy) or solution of specific taxonomic problems, were detailed elsewhere (Askevold 1987a,b, 1988, 1990, 1991). Some taxonomic problems remain despite relatively recent treatment by Marx (1957) of Donacia F. (sensu lato). I indicated earlier that Donacia is repre- sented in North America by 10 species in Donacia (s. str.), and 21 described species in Donacia (Donaciomima) Medvedev, but details were not given then (Askevold 1990). The purpose of the present paper is to detail remaining lectotype designations among Nearctic members of the genus Donacia F., to list the Nearctic members of Donaciinae, and to detail relevant taxonomic references and information (i.e., synonymies, taxonomic changes since prior authors, and data about type specimens). Schaeffer (1919, 1925) indicated only if he based his species on a type and allotype, but not that there were also many paratypes; paratype information is included insofar as such specimens have been found. Species are listed within genera and subgenera alpha- betically rather than phylogenetically, because phylogenies have been reconstructed only for members of Plateumaris and Neo- haemonia (Askevold 1991, 1988).

1present address: Entomology-Biological Control, Division of Agricultural Sci- ences, Florida A & M University, Tallahassee, FL 32307-2001 Manuscript received June 7, 1991.

165 166 Psyche [Vol. 98

ANNOTATED CATALOGUE OF NEW WORLD DONACIINAE The following list includes all described extant New World Donaciinae, serving to summarize the many taxonomic changes that have resulted from revision of North American Donaciinae. Details of historical use of species names is given where substan- tial changes in name use occurred [i.e., changes since Schaeffer (1925), Marx (1957) and Jolivet (1970)]. The taxonomic history of some names is complicated, often crossing generic boundaries; some species were synonymized with more than one taxon at once, based on composition of the original type series. I have judged that what is important is current synonymy and whether or not that dif- fers from the most recent uses of names, and therefore have not pried deeply into tracing the varied uses of names since their original description. Data of individual labels borne by type specimens, where recorded, are stated verbatim and are separated by a slash ("/"). The museums and private collections in which type specimens of any kind are known to be preserved are identified by correspond- ing codens:

AMNH. Department of Entomology, American Museum of Natural History, Central Park West at 79th St., New York, NY 10024, U.S.A. APHIS. Animal and Plant Health Inspection Service, USDA, Room 308, 40 South Gay St., Baltimore, MD 21202, U.S.A. BMNH. Department of Entomology, British Museum of Natural History, Cromwell Road, London SW7 5BD, England. CARR. J.L. and A.F. Carr, 24 Dalrymple Green, N.W. Calgary, Alta. Canada, T3A 1Y2. CAS. Department of Entomology, California Academy of Sci- ences, Golden Gate Park, San Francisco, CA 94118, U.S.A. CDAS. California Department of Food and Agriculture, California State collection of Arthropods, Insect Taxonomy Laboratory, 1220 N Street, Sacramento, CA 95814, U.S.A. CLCH. C. Chantal, B.P. 2072, St. Nicolas-est, P.Q., Canada, G0S 3L0. CMP. Section of Entomology, Carnegie Museum of Natural His- tory, 4400 Forbes Ave., Pittsburgh, PA 15213, U.S.A. 1991 Askevold 167 CNC. Biosyntematics Research Centre, Agriculture Canada, Ottawa, Ont., Canada K1A 0C6. CUIC. Department of Entomology, Comstock Hall, Cornell Uni- versity, Ithaca, NY 14853, U.S.A. DEFW. Department of Entomology, Fisheries and Wildlife, Uni- versity of Minnesota, 219 Hodson Hall, 1980 Folwell Ave., St. Paul, MN 55108, U.S.A. EGRC. E.G. Riley, Department of Entomology, Texas A & M Uni- versity, College Station, TX 77843-2475, U.S.A. FMNH. Field Museum of Natural History, Roosevelt Road at Lake Shore Drive, Chicago, IL 60605, U.S.A. FSCA. Florida State Collection of Arthropods, Division of Plant Industry, Florida Department of Agriculture, 1911 S.W. 34th St., P.O. Box 1269, Gainesville, FL 32602, U.S.A. INHS. Department of Entomology, Illinois Natural History Survey, Natural Resources Bldg., Urbana, IL 61801, U.S.A. ISAC. I.S. Askevold, Entomology-Biological Control, Division of Agricultural Sciences, Florida A & M University, Tallahas- see, FL 32307, U.S.A. JBWM. J.B. Wallis Museum, Department of Entomology, Univer- sity of Manitoba, Winnipeg, Man. R3T 2N2, Canada. LEM. Collection Provancher, D6partement de biologie, Universit6 Laval, Qu6bec, P.Q., G1K 7P4, Canada. LSU. Department of Entomology, Louisiana State University, Baton Rouge, LA 70803, U.S.A. MCZ. Department of Entomology, Museum of Comparative Zool- ogy, Harvard University, Cambridge, MA 02138, U.S.A. MLU. Section Biowissenschaften, WB Zoologie, Martin-Luther- Universitit Halle-Wittenberg, Domplatz 4, 4010 Halle (Saale) Germany. NMDC. N.M. Downie, 505 Lingle Terrace, Lafayette, IN 47901, U.S.A. OSUC. Systematic Entomology Laboratory, Department of Ento- mology, Oregon State University, Corvallis, OR 97331, U.S.A. PUR. Department of Entomology, Entomology Hall, Purdue Uni- versity, West Lafayette, IN 47907, U.S.A. ROM. Department of Entomology, Royal Ontario Museum, 100 Queen's Park, Toronto, Ont. M5S 2C6, Canada. 168 Psyche [Vol. 98 SCSU. Department of Biological Sciences, St. Cloud State Univer- sity, St. Cloud, MN 56301, U.S.A. SMCL. S.M. Clark, Pest Identification Laboratory, West Virginia Department of Agriculture, Charleston, WV 25305, U.S.A. TAMU. Department of Entomology, Texas A & M University, College Station, TX 77843-2516, U.S.A. UAE. Department of Entomology, University of Alberta, Edmon- ton, Alta. T6G 2E3, Canada. UMHF. Universitetets Zoologiska Museum, Entomologiska Avdel- ningen, N. Jirnvigsgatan 13, SF-00100 Helsingfors 10, Fin- land. UMMA. Museum of Zoology, Insect Division, The University of Michigan, Ann Arbor, MI 48109, U.S.A. USNM. National Museum of Natural History, Smithsonian Institu- tion NHB-105, Washington, D.C. 20560, U.S.A. UVDZ. Department of Zoology, Marsh Life Science Bldg., Uni- versity of Vermont, Burlington, VT 05401, U.S.A. ZMHB. Zoologisches Museum, Museum ftir Naturkunde der Hum- bolt-Universitit zu Berlin, Invalidenstr. 43, 1040 Berlin, Ger- many.

PLATEUMARINI ASKEVOLD, 1990 Plateumaris Thomson (1859:154). Revision of genus by Askevold (1991). Rather than making numerous, repetitive statements about taxonomic details for this genus, users are referred to Askevold (1991) for details about type specimens. Unless other- wise stated, lectotype and neotype designations were made by Askevold (1991); a detailed account of the historical use of names is given there also, and not repeated here. P. aurifera (LeConte, 1851:313). Lectotype ( MCZ #4247. Vali- dated, and removed from synonymy with P. germari by Askevold (1991). P. balli Askevold (1991). Holotype c, CNC #19497, and 23 paratypes, ISAC, USNM, AMNH, CNC, APHIS, UAE, UVDZ, CAS and SMCL. P. diversa (Schaeffer, 1925:143). Holotype'9, USNM #42397 and allotype c, USNM. P. dubia (Schaeffer, 1925:152). Holotype c), USNM #4240-C, allotype 9 USNM #42400 and 4 paratypes, USNM. 1991] Askevold 169

Donacia idola Hatch (1938:110). Holotype c, USNM type #103893, allotype 90SUC and 45 paratypes, OSUC. Syn- onymy with P. dubia by Askevold (1991). P. flavipes (Kirby, 1837:223). Holotype 9, BMNH. This name was used incorrectly by previous authors; the taxon previously known as P. flavipes is now called P. shoemakeri (Schaef- fer). The taxon formerly called P. wallisi is now correctly called P. flavipes. Donacia wallisi (Schaeffer, 1925:147). Holotype ( USNM #42399, allotype 9 USNM #42399, and 13 paratypes, USNM. Synonymy with P. flavipes by Askevold (1991). P. frosti (Schaeffer, 1925). Plateumaris frosti was formerly placed as a subspecies of P. emarginata (now P. nitida). Donacia emarginata var. frosti Schaeffer (1925:136). Holotype c, USNM #42395, allotype 9, USNM #42395, 28 paratypes, USNM, and 3 paratypes, MCZ. P. fulvipes (Lacordaire, 1845:192). Holotype c, BMNH. P. germari (Mannerheim, 1843:306). Lectotype 9, MCZ #4250, and 1 syntype, UMHF. Donacia flavipennis Mannerheim (1843"306). Lectotype c, MCZ #4251, and 1 syntype, UMHF. Specimens from western Canada possessing flavous elytra and/or pronotum were given varietal status by Schaeffer (1925); Askevold (1991) gave no taxonomic status to specimens showing such variation. Synonymy with P. germari by Askevold (1991). Donacia dives LeConte (1851:314). Holotype 9, MCZ #4245. Donacia serricauda Schaeffer (1919:318). Holotype 9, USNM #42396. P. metallica (Ahrens, 1810:33). Neotype cy, CNC #19493. Donaciafemoralis Kirby (1837:223). Lectotype 9, BMNH. Donacia cataractae Newman (1838:391). Lectotype c, BMNH. Donacia parva Lacordaire (1845:190), Lectotype c, BMNH. Donacia indica Melsheimer (1847:158) (not Clark, 1866:1). Lectotype 9, MCZ #33363. Donacia nana Melsheimer (1847:160). Neotype c, MCZ #32953. Donacia gentilis LeConte (1851:314). Lectotype (y, MCZ #4249, and two paralectotypes MCZ #4249. 170 Psyche [Vol. 98

P. neomexicana (Schaeffer, 1925:154). Holotype or, USNM #42401-1, allotype 9 USNM #42401, three paratypes, USNM, and two paratypes, MCZ #27227 [Fall colin.]. Donacia longicollis Schaeffer (1925:156). Holotype cr, USNM #4248-2, allotype USNM #42402 and 22 paratypes, USNM. Synonymized with P. neomexicana by Askevold (1991). Donacia vermiculata Schaeffer (1925:158). Holotype 9, USNM. Synonymized with P. neomexicana given by Askevold (1991). P. nitida (Germar, 1811:31). Lectotype 9 MLU. This name was used incorrectly by Schaeffer (1925); the species misidenti- fled by Schaeffer as P. nitida is now called P. schaefferi. The taxon to which the name P. nitida applies was called P. emarginata by Schaeffer (1925) and other authors. Donacia emarginata Kirby (1837:224). Lectotype or, BMNH. Synonymized with P. nitida by Askevold (1991). Donacia binodosa LeConte (1851:316) (cf. Suffrian, 1872). Unjustified emendation or nomen nudum. Synonymy with P. nitida by Askevold (1991). Donacia juncina Couper (1864:88). Lectotype 9, LEM, designated by Becker (1974). Synonymized with P. nitida by Askevold (1991). Donacia emarginata var. pacifica Schaeffer (1925:135). Holo- type c, USNM #42394, allotype 9, USNM #42394, and 9 paratypes, USNM. Synonymized with P. nitida by Askevold (1991). P. notmani (Schaeffer, 1925:132). Holotype cr, USNM #42393, allotype 9 #42393, and five paratypes, USNM, 1 paratype MCZ #27226 [Fall Colln.]. P. pusilla (Say, 1827:293). Neotype c, MCZ type #32950. Donacia rugifrons Newman (1838:391). Types of this name have not been found, but should be in BMNH. Three specimens (of P. rufa, however, not P. pusilla) were found in the MCZ that are labelled Trenton Falls, New York, collected by Doubleday. Similar labels are borne by specimens of two other Newman species D. cincticornis and D. cataractae and I consider the possibility that these might be specimens of Newman's D. rugifrons. 1991] Askevold 171

Donacia pyritosa LeConte 1857(1860):66. Lectotype or, MCZ type #4246. This name was formerly used as a variety of P. pusilla, but synonymized by Askevold (1991). P. robusta (Schaeffer, 1919:318). This name was used as a variety of P. pusilla until elevated to species status by Askevold (1991). Donacia pusilla var. robusta Schaeffer (1919:318). Holotype c, USNM #42398, and 3 paratypes USNM #42398. P. rufa (Say, 1827:283). Neotype c, MCZ #32947. This species was formerly called P. sulcicollis Lac. The name D. rufa has been used incorrectly by authors (Marx 1957:250, Schaeffer 1925:117), and was transferred from Donacia to Plateumaris by Askevold (1991) [cf. also D. (Donaciomima) tuberculata Lacord., below]. Donacia affinis Kirby (1837:224) (not Kunze, 1818). Lectotype 9, BMNH. Transferred to synonymy with P. rufa from synonymy with P. sulcicollis by Askevold (1991). Donacia sulcicollis Lacordaire (1845:186). Lectotype cr, BMNH. Synonymized with P. rufa by Askevold (1991). Donacia chalcea Lacordaire (1845:188). Holotype 9, MLU. Transferred to synonymy with P. rufa from P. flavipes, by Askevold (1991). Donacia kirbyi Lacordaire (1845:202). Replacement name for D. affinis Kirby; transferred to synonymy with P. rufa from P. sulcicollis by Askevold (1991). Donacia jucunda LeConte (1851:315). Holotype c, MCZ #4248. Transferred to synonymy of P. rufa from P. sulcicollis by Askevold (1991). P. schaefferi Askevold (1991). Holotype c, USNM, and 7 paratypes, USNM, CUIC and UMMA. Donacia nitida, sensu Schaeffer (1925:130) (see P. nitida, above). P. shoemakeri (Schaeffer, 1925:129). This species was formerly incorrectly known as P. flavipes (see P. flavipes, above), val- idated by Askevold (1991). Donacia flavipes var. shoemakeri Schaeffer (1925"129). Holo- type cr, USNM #4239-1 and 5 paratypes USNM #42391. Donaciaflavipes var. lodingi Schaeffer (1925"129). Holotype USNM #4239-2 and 1 paratype, USNM #42392. Southern 172 Psyche [Vol. 98 specimens, from Alabama, were given varietal status by Schaeffer (1925), but Askevold (1991) did not give this geographic variation taxonomic status.

Poecilocera Schaeffer (1919:316). Elevated to genus status by Askevold (1990) P. harrisii (LeConte, 1851:316). Lectotype 9 MCZ #4244, designated by Askevold (1990:635); the abdomen is now missing but was present when I first examined it. LeConte stated he had at least two specimens, and 9, although only a single specimen remains in the LeConte collection. P. harrisii was placed in Sominella by Goecke (1931), but Marx (1957) reverted to place- ment in Donacia (Poecilocera), evidently unaware of Goecke's treatment.

DONACIINI KIRBY, 1837 Donaciella Reitter (1920:38). Elevated to genus by Askevold (1990), this name has been treated by European authors as a subgenus of Donacia, which has not been recognized in North America despite direct comparisons by Schaeffer (1925) with species occurring in the Palaearctic Region. D. pubicollis (Suffrian, 1872:21). Transferred to Donaciella by Askevold (1990). Suffrian did not indicate the number of specimens before him, but given a range of size measure- ments, had at least two. A series of 12 specimens are preserved in MLU, associated with a Suffrian label. As determined previously (see P. chalcea and P. nitida, above) these specimens are also likely original Suffrian material, and are here designated. LECTOTYPE , following a Suf- frian label ["pubicollis m. Illinois", on large, green, black- bordered paper]: "25519 [red handwriting on small white label] / LECTOTYPE r Donacia pubicollis Suffrian 1872 design. I.S. Askevold 1991". The remaining 11 specimens are here designated paralectotypes. These bear the same red, handwritten number and my paralectotype label: 25516 (r), 25517 (c), 25520 (cy), 25521 (c), 25522 (cr), 25523 (9), 25524 (9), 25525 (9), 25526 (9), 33256 (9) and 25518 (sex undetermined, abdomen missing). 1991] Askevold 173 Donacia pubicollis Crotch (1873:21) (not Suffrian, 1872:21). Synonymy with Donacia pubicollis Suffrian by Marx (1957). No specimens of Crotch's species were found at MCZ indicated as type, probably because it was not recognized that Crotch had also described the species, until Marx (1957). I presume the type must be lost, but it is hardly necessary to designate a neotype for such a well-known species.

Donacia (Donaciomima) Medvedev (1973:876). The subgenus Donaciomima was originally based on an eastern Palaearctic species, but was disregarded in subsequent treatments. All Nearctic species of Donacia were formerly placed in Donacia s. str., but Askevold (1990) revalidated Donacia (Donaciomima), and defined it to include the majority of species hitherto placed in the nominate subgenus. Most Nearctic species (23) of Dona- cia are included, of which two remain undescribed. D. assimilis Lacordaire (1845:153). Holotype c, BMNH. Lacor- daire had before him a single specimen of undetermined sex: "TYPE [red] / TYPE [white] / E. Coll. Lafert6 / Donacia assimilis type Lacord. / Donacia assimilis Dej. h. in Ameri. bor. J. Leconte". Donacia glabrata Schaeffer (1919:313) (not Solsky, 1872). Based on single c, USNM holotype: "Type / Clementon, VIII. 6. N.J. / Brooklyn Museum Colln. 1929 / TYPE USNM 42387 / D. glabrata type Schffr. / Donacia assimilis Lac." Synonymy with D. assimilis by Schaeffer (1925). D. biimpressa Melsheimer (1847:159). Lectotype cr, MCZ #33365. Melsheimer had before him at least two specimens; the single remaining specimen is here designated as lectotype" "Melsh. biimpres / Type [red] / biimpressa [large Melsheimer label, handwritten] / LECTOTYPE Donacia biimpressa Melsheimer design. I.S. Askevold 1990 [red] / MCZ. Lecto- type 33365 [red]". Donacia aurichalcea Melsheimer (1847:159). Evidently based on a single specimen. Holotype or, MCZ #33364: "Melsh. aurichalcea / Type [red] / dissected and remounted IS Askevold 1983 / aurichalcea [large handwritten Melsheimer label] / Donacia biimpressa Melsheimer 1847 det. I.S. 174 Psyche [Vol. 98 Askevold 1983 /M.C.Z. Lectotype 33364 [red]". Synonymy with D. biimpressa by Schaeffer (1925). Donacia torosa LeConte (1851:313). LeConte did not specify number of specimens, but three specimens found in the LeConte collection are here designated as types. Lectotype c, MCZ: "pink disc / LECTOTYPE [blue trimmed disc] / TYPE 4243 [red] / D. torosa Lec. / J. LECONTE COLLECTION / LECTOTYPE cy Donacia torosa LeConte 1851 designated IS Askevold 1983 [red]". Paralectotype cy: "pink disc / Type 4243 [red] / torosa 2 / J. LECONTE COLLECTION / PAR- ALECTOTYPE cy Donacia torosa LeConte [red]"; paralectotype [sex undetermined, abdomen missing] "pink disc / torosa 3 / J. LECONTE COLLECTION / PARALECTO- TYPE Donacia torosa LeConte [red]". Synonymy with D. biimpressa by Schaeffer (1925). D. caerulea Olivier (1795:10). Neotype cy, CNC #19492, designated by Askevold (1987b:347). Discussed by Askevold (1987b), this species was formerly known as D. aequalis; D. caerulea was used incorrectly by Schaeffer (1925) and Marx (1957), which they applied to certain specimens of D. proxima. Donacia aequalis Say (1824:248). Neotype c, MCZ #32948, designated by Askevold (1987b:347); placed in synonymy with D. caerulea by Askevold (1987b). Donacia confusa Lacordaire (1845; 109) (not LeConte, 1851). Lectotype cy, BMNH, designated by Askevold (1987b:347); placed in synonymy with D. caerulea by Askevold (1987b). Donacia confusa LeConte (1851:313) (not Lacordaire, 1845). Lectotype cy, MCZ #32949, designated by Askevold (1987b:347); placed in synonymy with D. caerulea by Askevold (1987b). D. cazieri Marx (1957:213). Holotype cy, CNC, allotype , CNC, four paratopotypes, CNC, and one paratopotype, AMNH, Goose Bay, Labrador (various dates and collectors, see Marx 1957). D. confluenta Say (1827:293). Neotype cy, MCZ #32956, designated by Askevold (1987a:634); removed from synonymy with D. subtilis Kunze by Askevold (1987a). 1991] Askevold 175

D. cuprea Kirby (1837:225) (not Melsheimer, 1847). Lectotype BMNH, designated, and removed from synonymy with D. quadricollis Say, by Askevold (1987a:633). Donacia curticollis Knab (1905:122) (not Haupt, 1956). Removed from synonymy of D. quadricollis Say by Askevold (1987a). Lectotype 9, and three paratypes, INHS; designation and details given by Frison (1927:160). D. dissimilis Schaeffer (1925:115). Holotype 9, USNM: "Florida ex Liebeck / Charles Schaeffer Collection / HS Barber Bequest 1950 / dissimilis Holotype C. Schaeffer coll. 1935 Barber / Donacia dissimilis Schffr.". Allotype c, PUR: "Dunedin [Florida] 4.16.24 W.S.B. coll./2724 / c Allotype / Donacia dissimilis cr allotype Schaeffer". D. distincta LeConte (1851:313). LeConte did not state the number or sex specimens before him; two specimens were found in the LeConte collection. Lectotype cr and paralectotype MCZ #4242, here designated: "cr [pale blue disc] / LECTO- TYPE [blue trimmed disc] / Type 4242 [red] / D. distincta Lec. aequalis Kirby / LECTOTYPE cy Donacia distincta LeConte 1851 designated 1983 IS Askevold [red]" [specimen dissected, with genitalia placed in microvial pinned beneath specimen]; paralectotype 9, "9 [pale blue disc] / Type 4242 [red] / distincta 2 / J. LECONTE COLLECTION / PAR- ALECTOTYPE 9 Donacia distincta LeConte [red]". Donacia distincta occidentalis Mead (1938:115)..Holotype CAS #4691, allotype 9, CAS #4692, and paratype, MCZ [ex Fall Coll.], Luthers Pass, Cal. vi-15-1930 AT McClay. Syn- onymy with D. distincta by Marx (1957), to which I agree. D. fulgens LeConte (1851:312). Lectotype cr, MCZ #4240, designated by Askevold (1987a:634). At the time of this designation, another syntype specimen was not sent to me; I here add a 9 paralectotype from the LeConte collection: "pale blue disc, two cuts / Type #4240 [red] / D. fulgens Lec. / J. LECONTE COLLECTION / PARALECTOTYPE 9 Donacia fulgens LeConte [red]". Removed from synonymy of D. subtilis Kunze by Brivio and Balsbaugh (1984). D. hirticollis Kirby (1837:226). Holotype c, BMNH. Kirby stated he had before him a single specimen, which bears the following labels: "TYPE [red trimmed disc] / N. Amer. 5954a 176 Psyche [Vol. 98 [white disc] / Donacia hirticollis Kirby N. Ameri. 5954 Rev. Wm. Kirby [type, on underside]". The specimen lacks the right fore leg and left hind leg, and right elytron smashed by pin. Donacia rudicollis Lacordaire, 1845:108. LECTOTYPE 9, BMNH, by present designation. Lacordaire stated he had before him two, supposedly female, specimens. The single specimen found is here designated: "LECTOTYPE [blue trimmed disc] /TYPE [red trimmed disc] /TYPE [white] /E. Coll. Lafert6 / [green, square paper, 4 mm] / 67.56 / Donacia rudicollis Type Lac. / Donacia rudicollis. Reiche Amer. bor./ LECTOTYPE 9 Donacia rudicollis Lacordaire 1845 designated 10.vii.1984 I.S. Askevold [red] / Donacia hirticollis Kirby det. I.S. Askevold 1984". I have not determined the original source of synonymy. D. liebecki Schaeffer (1919:314). Holotype c, USNM #42390: "Type cr / Wyandanch L Isld. / Brooklyn Museum Colin. 1929 / liebecki type USNM 42390 [red] / Donacia liebecki Schffr. [red bordered]". Allotype 9: "Type 9 Allotype / Wyandanch L Isld./liebecki Schffr. Allotype USNM 42390 [red] / Brooklyn Museum Colin. 1929". Five paratypes MCZ #27228 (two ex Fall Colin.), Malaga 1.8 N.J. One specimen labelled as a paratype (USNM) may be a paratype, though it does not bear labels indicating it is part of the original syntype series of 11 specimens: "Lakehurst NJ / Ed. A. Bischoff Collection / Brooklyn Museum Colin. 1929 / liebecki USNM paratype ? 42390 [red]". Another specimen labelled as a paratype (USNM) cannot be a paratype because it is from Wading River, Long Island, a locality not mentioned by Schaeffer. Donacia pallipes Lacordaire (1845:149) (not Kunze, 1818). Holotype 9, BMNH. Lacordaire had before him a single specimen which he supposed to be male. It is female, however. "TYPE [red trimmed disc] / TYPE [white] / 67.56 / D. pallipes Lac. cr type / Donacia type 32. pallipes Lac. 119 39 Philadelp Am. Sept. Boston D. Green [green, folded] / Dona- cia liebecki Schaeffer det. IS Askevold 1985." D. pallipes was listed in early catalogues as a synonym of D. aequalis Say; Schaeffer (1925) realized that D. pallipes Lac. was a 1991] Askevold 177 senior synonym of D. liebecki, but preoccupied, thus D. liebecki remains valid. Synonymy with D. liebecki by Schaeffer (1925). D. limonia Schaeffer (1925). This name was described as a variety of D. biimpressa Melsh.; Marx (1957) reduced it to synonymy with D. biimpressa. I here remove it from synonymy, and recognize D. limonia as a valid species on the basis of distinctive structure of male genitalia; further treatment of this species as a member of the D. distincta-Group is planned. NEW STATUS D. biimpressa var. limonia Schaeffer (1925:100). Holotype c, USNM: "Indian Pass N.Y. VII. 11. / Essex Co. Quirsfeld / Brooklyn Museum Coll. 1929 / limonia Schffr. TYPE USNM 42388 / Donacia biimpressa limonia Schffr.". Allotype 9, USNM, Van Cortlandt Park, N.Y. (additional details not recorded), and five paratypes, USNM (label details not recorded). D. rnagnifica LeConte (1851:310). LeConte did not state the number of specimens before him, but stated he had both male and female specimens. Four specimens in the Leconte collection bear appropriate label data, and are here designated, lectotype c, MCZ #4237, one cr and two 99 paralectotypes, MCZ #4237: "LECTOTYPE [blue trimmed disc] / cr [pale blue disc] / Type 4237 [red] / Donacia magnifica Lec./LECTO- TYPE c Donacia magnifica LeConte 1851 designated I.S. Askevold 1984 [red]"; the three paralectotypes are labelled "magnifica 2", "magnifica 3" and "magnifica 4", each with the labels "Type 4237" [red] and "PARALECTOTYPE Donacia magnifica LeConte [red]". In earlier catalogues, D. magnifica was listed under D. cincticornis. D. megacornis Blatchley (1910:1103). Lectotype cr, PUR. Blatch- ley did not state the number of specimens before him, but designated a lectotype c later (Blatchley 1930): "Lake Co., Ind. W.S.B. 5-28-1903 / Donacia megacornis, sp. nov. [handwritten]". Donacia megatocera [sic] Weise (1913:118). Unnecessary replacement name. Clavareau (1913:23) altered the spelling of megatocera to megalocera, probably supposing that it was a printer's error. Weise proposed the name D. megalocera to 178 Psyche [Vol. 98 replace Blatchley's D. megacornis, on the basis that megacornis is a name formed from two languages; the name megacornis is valid, and Schaeffer (1925) restored use of the name D. megacornis, and synonymized D. megatocera. D. porosicollis Lacordaire (1845:150). Holotype 9, BMNH. Lacor- daire described this species from a single, apparently female specimen (which it is): "TYPE [red trimmed disc] / TYPE [white] /Donacia Chvt longipennis parraceicollis Lac. type America bor. Boston D. Greene 627 [green] / 67.56 / E. Coll. Chevt. / 21. [folded paper] / D. porosicollis Lac. type". The specimen is a little crushed and is missing the right middle leg. D. pubescens LeConte (1868:55). Described from a unique specimen, which LeConte stated was from the Smoky Hill River. Holotype 9, MCZ #4234: "Type [red] / D. pubescens Lec.". D. rugosa LeConte (1878:415). Described from a unique specimen, from Enterprise, Florida. Holotype 9, MCZ #4241: "Enterprise May 20 Fla. / 402 / Type 4241 [red] / D. rugosa Lec./J. LECONTE COLLECTION". D. subtilis Kunze (1818:12) (replacement name for D. aenea Ahrens). Donacia aenea Ahrens (1810:21) (not Hoppe 1795:44; not Gis- tel 1857:524). Neotype cy, CMP; designation and discussion by Askevold (1987a:633). Donacia quadricollis Say (1827:282). Neotype cy, MCZ #32951, designated by Askevold (1987a:633). The name D. quadricollis was formerly used by authors (e.g. Schaeffer 1925, Wilcox 1954, Marx 1957), applied to the species now known as D. cuprea Kirby (see above) but was synonymized with D. subtilis by Askevold 1987a). Donacia aerea Lacordaire (1845:148) (not Schrank, 1798). Lec- totype cr, BMNH, designated by Askevold (1987a). I have not determined the original source of synonymy. Donacia subtilis magistrigata Mead (1938:113). Holotype CAS #4689, allotype 9, CAS #4690, two paratypes, CAS, and one paratype, MCZ #27230 (ex Fall Colin.), Clarksburg, Cal, June-4-1931, A.T. McClay. Synonymized with D. fulgens by Marx (1957) but transferred to synonymy with D. subtilis by Askevold (1987a). 1991 Askevold 179

Donacia tuberculata Lacordaire (1845:155). Lectotype c and paralectotype 9, BMNH. Two specimens were found that appeared to be authentic Lacordaire specimens, which are here designated: "LECTOTYPE [blue trimmed disc] / D. tuberculata Lac / 67.56 / LECTOTYPE cr Donacia tuberculata Lacordaire 1845 designated 10.vii.1984 I.S. Askevold [red]". A female is labelled: "PARALECTOTYPE 9 [pale blue-trimmed disc] / TYPE [red trimmed disc] / TYPE [white] / E. Coll. Lafert6 / Donacia tuberculata mihi h. in Ameri. bor. D. Leconte [green, folded] / 67.56 / PARALEC- TOTYPE 9 Donacia tuberculata Lacordaire 1845 designated 10.vii.1984 I.S. Askevold [red]." Donacia tuberculata was synonymized with D. rufa Say by Schaeffer (1925), an arrangement followed by Marx (1957). Leng (1891) had used D. rufa in the sense used above, as a species of Plateumaris to which it was transferred by Askevold (1991). With trans- ferral of D. rufa to Plateumaris, D. tuberculata becomes the senior, available name for the taxon previously recognized as D. rufa, sensu Schaeffer (1925) and Marx (1957). NEW STATUS Donacia rutila Melsheimer (1847:160). Lectotype 9, MCZ #32952, by present designation: "LECTOTYPE [blue trimmed disc] / Melsh. rutila / M.C.Z. Lectotype 32952 [red] / tuberculata [large handwritten Melsh. label] / LECTOTYPE 9 Donacia rutila Melsheimer, design. I.S. Askevold 1990 [red]". Earlier authors (e.g. Leng 1891) placedD, rutila under D. tuberculata. However, this name was transferred to synonymy of D. rufa. Because D. tuberculata is revalidated, D. rutila becomes its synonym. NEW SYNONYMY D. tuberculifrons Schaeffer (1919:315). Holotype c, USNM: "Toronto VII.7 Can. / Brooklyn Museum Coll. 1929 / TYPE USNM 42389 / D. tuberculifrons type Schffr.". Allotype 9 and one paratype, USNM (label details not recorded). Four paratypes, MCZ #27229 (1 ex Fall Colln.): "Toronto, Can. VII.I". This species was treated as a member of the D. subtilis- Group by Askevold (1987a). D. vicina Lacordaire (1845:154) (not Haupt, 1956). Holotype 9, BMNH. Lacordaire described the species from a single 180 Psyche [Vol. 98 specimen received from Reiche, incorrectly stating it to be male: "HOLOTYPE [red trimmed disc] / TYPE [white] / E. Coll. Lafert6 / Donacia vicina Dej. h. in Amer. bor. D. Leconte [green]".

Donacia (Donacia) Fabricius, 1775. All Nearctic species of Donacia were formerly placed in Donacia s. str. (e.g. Leng 1891, Jacoby and Clavareau 1904, Clavareau 1913, Jolivet 1970, Borowiec 1984), but Askevold (1990) restricted the nominate subgenus to include 10 Nearctic species. D. cincticornis Newman (1838"39i). Lectotype 9, BMNH, by present designation. Newman did not state the number of specimens before him. The single specimen remaining is therefore here designated: "LECTOTYPE [blue trimmed disc] / TYPE H.T. [red trimmed disc] / Ent. Club. 44-12 / R. Foster Tren- ton Falls, New York / Donacia cincticornis type Newm. / LECTOTYPE 9 Donacia cincticornis Newman 1838 designated 11.vii. 1984 I.S. Askevold 1984". Donacia lucida Lacordaire (1845:106). Lectotype, BMNH, by present designation. The species was described from at least two specimens. The single remaining specimen under this name is selected as lectotype, which bears the following labels: "LECTOTYPE [blue trim disc] / TYPE [red trim disc] / Donacia Lucida Lac Type / TYPE / E. Coll. Chevt. / D. lucida cr Lac. type / Donacia type 626 lucida (Chvt) Lac. mon 6 Am. bor. Boston D. Greene [green label] / 67.56 / LECTO- TYPE [abdomen missing, probably male] Donacia lucida Lacordaire 1845 designated 10.VII.1984 I.S. Askevold [red] / Donacia cincticornis Newman det. I.S. Askevold 1984". Syn- onymy with D. cincticornis was apparently first by Leng (1891). Donacia rufipennis Lacordaire (1845:110). Lectotype cy, BMNH, by present designation. Lacordaire described this species from four specimens. A male is here selected as lectotype, bearing the following labels: LECTOTYPE [blue trimmed disc, added by me] ! TYPE [red] ! Donacia rufipennis Type Lac. [white] ! TYPE [white] ! Donacia type picipen- nis Chevt rufipennis Lac. mon. Ameri. bor. Boston D. Greene [green] / 12 / 67.56 / LECTOTYPE Donacia rufipennis 19911 Askevold 181 Lacordaire 1845 designated 10.VII. 1984 I.S. Askevold [red] / Donacia cincticornis Newman det. I.S. Askevold 1984". Synonymy with D. lucida by Crotch (1873), therefore placed as synonym of D. cincticornis by later authors. Donacia pulchella LeConte (1851:312). LeConte apparently described a single specimen, stating at least that he had no males. The single specimen found in the LeConte collection is therefore assumed to be the holotype, which LeConte incorrectly sexed. Holotype 9, MCZ #4235: "pink disc / Type 4235 [red] / D. pulchella LeC.". Synonymy with D. lucida by Crotch (1873), therefore placed as synonym of D. cincticornis by later authors. Donacia antillarum Suffrian (1864:282). Suffrian described this from at least two specimens, because he gave a range of size. A single specimen here designated as lectotype was found in MLU, associated with the Suffrian label: "Antillarum m. Cuba [large, green, black-bordered label]". LECTOTYPE (sex undetermined, abdomen missing): "25004. [red handwriting on small label] / LECTOTYPE Donacia antillarum Suffrian 1864 design. I.S. Askevold 1991". Treated as a subspecies of D. cincticornis by Schaeffer (1925); synonymy with D. cincticornis by Marx (1957). Donacia cincticornis var. tryphera Schaeffer (1925:81). Holo- type cr, USNM: "Centerton VII N.J. / Brooklyn Museum Coll. 1929 / tryphera TYPE USNM 42385". Allotype 9, and 11 paratypes, USNM (label data not recorded). Marx (1957) reduced Schaeffer's subspecies to synonymy; I agree with this synonymy. Donacia cincticornis var. tenuis Schaeffer (1925:82). Holotype c, USNM" "Watershield/Swansea SO 11 Aug. 11 Fredk. Knab. / Type c / tenuis TYPE USNM 42386 / D. cincticornis tenuis Schffr.". Allotype 9, and 16 paratypes, USNM (label data not recorded). Two paratypes, MCZ #27232 (ex Fall Colin.): "Spencer L., N.Y. viii.26.17, H. Dietrich". Marx (1957) reduced Schaeffer's subspecies to synonymy; I agree with this synonymy. D. edentata Schaeffer (1919:313). Holotype or, USNM: "Crum Lk. N.Y. / U.S.N.M. Type No. 42383 / D. edentata type Schffr.". Allotype 9 and five paratypes, USNM (label data not 182 Psyche [Vol. 98 recorded). One paratype CAS, and eight paratypes, MCZ, "Centerton N.J., 5.31". D. hypoleuca Lacordaire, 1845:101. Lectotype c, BMNH, by present designation. Lacordaire lescribed this species from two, supposedly 9, specimens; however, there is only a single cr specimen under hypoleuca which bears an original Lacor- daire label: "Lectotype cr [blue trim disc added by me] / E. Coll. Lafert6 / 67.56 / LECTOTYPE c Donacia hypoleuca Lacordaire 1845 designated 11.VII.1984 I.S. Askevold [red, added by me]". Also in the BMNH is another specimen (c) which bears a type label; it bears the labels "TYPE [white] / This specimen cannot possibly be the type of hypoleuca Lac. The labels have evidently been misplaced. C.G. / Donacia hypoleuca. Van Wintheim h. in Amer. bor. Neu Orleans D. v. Wintheim [green Lacordaire label]". This latter label is evidently genuine, according to information given by Lacor- daire, mentioning van Wintheim specifically (1845:101); however, the specimen does not agree with the description at all, and is in fact a specimen of D. palmata Olivier. Whether the specimen is a syntype or not is irrelevant, for the other specimen is here designated lectotype in the interest of con- serving the traditional use of D. hypoleuca. D. militaris Lacordaire (1845"115). Lectotype c, BMNH, by present designation. Lacordaire did not state the number of specimens before him, only that he had no female specimens; acr specimen was found: "LECTOTYPE [blue trimmed disc] / TYPE [red trimmed disc] / Donacia militaris type Lac. / TYPE [white] / E. Coll. Chevt. / 67.56 / D. militaris Lac. [??] type / Donacia type 630 bispinosa (Chvt) 9 militaris Lac mon. America bor. Boston D. Greene [green, folded] LEC- TOTYPE Donacia militaris Lacordaire 1845 designated 10.vii.1984 I.S. Askevold [red]". Formerly misidentified as a synonym of D. palmata, but removed from synonymy by Schaeffer (1925). Donacia floridae Leng (1891:166). No type specimen specifically identified was found in USNM (R.E. White, pers. comm.); however, a specimen, USNM type #558 was found among specimens in "regular collection", which is assumed 1991] Askevold 183 to be a paratype. Synonymized by Schaeffer (1925), which is doubtless correct. D. palmata Olivier (1795:8). Type(s) almost certainly destroyed; no neotype is designated because Olivier provided a figure of the protarsus of male specimens; the species is unique among Donaciinae in this structure. Donacia claudicans Germar (1821:173). Germar evidently described this from a single male specimen, specifically dis- cussing the unusual and unmistakable protarsal structure. According to Suffrian (1872), he had the Germar collection in his possession (see extended discussion of P. nitida by Askevold, 1991). Thus, Germar's type, if existing, should be in the Suffrian collection. Three specimens (cc) are associated with the large, green, black-bordered Suffrian label in MLU: "palmata Oliv. Lec. claudicans Grm. Illinois". Neither of these fits especially the specific colours that Germar described, and so it is quite likely that none of these is the Germar specimen. However, it is not necessary to designate a neotype because the species is so distinctive and Germar described the distinguishing character. I have not traced the source of original synonymization. Donacia palmata angustipes Marx (1957"237). Holotype c and allotype 9 #8452, and 19 paratypes, CAS, "Devils Riv. near Del Rio, Texas, vi-16-32, J.O. Martin"; 2 paratypes, CAS, "Devils Riv. near Del Rio, Texas, v-25-32, J.O. Martin". These Texas specimens differ little from other, typical D. palmata from other regions, differing essentially only in that males have a less broadly palmate basal protarsomere. NEW SYNONYMY D. parvidens Schaeffer (1919:311). Holotype c, USNM: "Lahaway Ocean Co. V. N.J. / Brooklyn Museum Coll. 1929 / U.S.N.M. Holotype 42384 / D. parvidens type Schffr.". Four paratypes, USNM (label data not recorded) and one paratype, MCZ, "Lahaway, Ocean Co., N.Y." D. piscatrix Lacordaire (1845:113). Lectotype c, BMNH, by present designation. Lacordaire did not state the number of specimens before him, only that he had no male specimens. One 9 specimen was found: "LECTOTYPE [blue trimmed disc] / TYPE [red] / Donacia piscatrix Lacord. / E. Coll. Lafert6 / D. 184 Psyche [Vol. 98 piscatrix Lac./Donacia Am. bor. [green] / 67.56 / LECTO- TYPE 9 Donacia piscatrix Lacordaire 1845 designated 10.vii. 1984 I.S. Askevold [red]". Donacia carolina Lacordaire (1845:114). Lectotype or, BMNH. Lacordaire stated he had both a male and female before him. The single, apparently authentic, Lacordaire specimen found among specimens of D. piscatrix, is here designated" "LEC- TOTYPE [blue trimmed disc] / TYPE [red] / Donacia carolina Lacord. / TYPE [white] / 67.56 / Donacia carolina mihi. h. in Amer. bor. D. Latreille [green] / Donacia piscatrix det. I.S. Askevold '84". I did not trace the original source of synonymization. Donacia cuprea Melsheimer (1847:158) (not Kirby, 1837). Lec- totype or, MCZ, by present designation. Described from at least two specimens, for Melsheimer gave a size range and indicated a "var. a". The single remaining specimen is here designated: "cy [discoloured square] / Melsh. cuprea / Type [red] / cupria [sic] [large handwritten Melsheimer label] / LECTOTYPE c Donacia cuprea Melsheimer design. I.S. Askevold 1990 [red] / M.C.Z. Lectotype 33366 [red]". Syn- onymy with D. piscatrix apparently first by Schaeffer (1925). Donacia alutacea LeConte (1851:311). LeConte had at least two specimens before him. Two specimens found in the LeConte collection are here designated: lectotype cr, MCZ #4239, "LECTOTYPE [blue trimmed disc] / Type [partly red] / c [pink disc] / Type 4239 [red] / D. alutacea Lec. cuprea//Mels. / LECTOTYPE c Donacia alutacea LeConte 1851 designated I.S. Askevold 1984 [red]"; and paralectotype c, "9 [pink disc] / Type 4239 [red] alutacea 2 PARALECTO- TYPE cr Donacia alutacea LeConte [red]". Synonymy with D. piscatrix by Crotch (19873). Donacia congener LeConte (1851:310). LeConte did not state number of specimens before him, only that he had no males. Two specimens in the LeConte collection bear orange discs, so it appears he had at least two specimens, and I here designate them. Lectotype 9, MCZ #4238: "LECTOTYPE [blue trimmed disc] / 9 [orange disc] / Type 4238 [red] / D. congener Lec. / LECTOTYPE 9 Donacia congener LeConte 1851 designated I.S. Askevold 1984"; and one 9 paralectotype, MCZ #4238 "9 1991 Askevold 185 [orange disc] / Type 4238 [red] / congener 2 / PARALECTO- TYPE 9 Donacia congener LeConte [red]". Synonymy with D. piscatrix by Crotch (1873), treated as subspecies by Schaeffer (1925), resynonymized by Marx (1957). D. proxima Kirby (1837:225). Lectotype 9, BMNH, designated by Askevold (1987b:347). Donacia episcopalis Lacordaire (1845:105). Lectotype c, BMNH, designated by Askevold (1987b:348). Formerly placed in synonymy with D. proxima, transferred to synonymy with D. caerulea by Schaeffer (1925), who used this name used incorrectly; D. episcopalis was transferred back to junior synonymy with D. proxima by Askevold (1987b). Donacia californica LeConte (1861:357). Holotype 9, MCZ #4236 (cf. Askevold 1987b:348). Given subspecies status by Marx (1957), but this classification rejected by Askevold (1987b). Donacia caerulea, sensu Schaeffer (1925) and Marx (1957). See Askevold (1987b) for discussion. D. rufescens Lacordaire (1845:112). Lectotype c, BMNH, by present designation. Lacordaire had before him two specimens, supposedly female. However, both are male, of which one is here designated: "LECTOTYPE [blue trimmed disc] TYPE [red trimmed disc] TYPE [white] Donacia type 629 calcarata Chvt 8 rufescens Lac. mon. America bor. Boston D. Greene [green, folded] / 15. / 67.56 / LECTOTYPE cr Donacia rufescens Lacor- daire 1845 designated 10.vii.1984 I.S. Askevold [red]". D. texana Crotch (1873:22). Lectotype 9, MCZ #5032, by present designation. Crotch did not state the number of specimens before him, but three specimens already indicated as types were found in the MCZ. Lectotype 9, MCZ #5032, "Tex. Type 5032 [red] / D. texana Cr. / LECTOTYPE 9 Donacia texana Crotch design. IS Askevold 1990 [red]"; paralectotype 9, MCZ #5032, "Tex./ Type 5032 [red] / texana 2 / PARALECTOTYPE Donacia texana Crotch [red]" and paralectotype or, MCZ #5032, "Tex. Type 5032 [red] texana 3 PARALECTOTYPE Donacia texana Crotch [red]". Donacia texana var. minor Schaeffer (1925:72). Holotype c, USNM: "Centerton V. 28. N.J. / Donacia texana minor Schffr. c / U.S.N.M. Type No. 42382". Allotype 9, and 6 paratypes, USNM (label data not recorded). Marx (1957) did not recognize 186 Psyche [Vol. 98

D. minor as a subspecies and therefore synonymized D. minor, to which I agree.

HAEMONIINI CHEN, 1941 Neohaemonia Sz6kessy (1941:148). This genus was formerly consid- ered congeneric with the Palaearctic genus, Macroplea Samouelle. Revision of genus and establishment of genus status by Askevold (1988). N. flagellata Askevold (1988:390). Holotype or, CNC, and 72 paratypes of various locality data deposited in CARR, CAS, CNC, CUIC, EGRC, ISAC, JBWM, ROM, UAE, UMMA, USNM. N. flohri (Jacoby, 1884:126). Holotype 9, MCZ #8468. Details given by Askevold (1988). N. melsheimeri (Lacordaire, 1845:211). Holotype cr, BMNH. Details given by Askevold (1988). N. minnesotensis Askevold (1988:388). Holotype cr, CNC #19495, and 38 paratypes with various locality data deposited in AMNH, CAS, CMP, CNC, DEFW, EGRC, FMNH, FSCA, ISAC, JBWM, LSUC, MCZ, NMDC, SCSU, UMMA, USNM. N. nigricornis (Kirby, 1837:222). Holotype 9, BMNH. Details given by Askevold (1988). Haemonia americana Gu6rin-M6neville (1844:259). Holotype 9, BMNH. Details given by Askevold (1988).

ACKNOWLEDGEMENTS I owe the many curators of museums cited above profuse thanks for their inestimable help with my studies about Donaciinae. Without their assistance, this and other publications on this group would suffer significant incompleteness. Mostly I examined all types myself, but Richard E. White is owed particular thanks for providing details of Schaeffer and Leng types that I was neglectful in recording data about personally, or did not personally examine. Research over a number of years, and costs of publication, were supported by NSERC grant A0428 held by R.E. Roughley, University of Manitoba, and in part also by an NSERC PGS Scholarship, an Ernst Mayr Grant in aid of research at MCZ, and funds provided by CAS to 1991 Askevold 187 research at that institution. Some costs were supported by a grant from CSRS, USDA to Florida A. & M. University.

SUMMARY The Nearctic species of Donaciinae are catalogued according to the most recent classification of its members at tribal, generic and subgeneric levels. Annotations are provided where significant depar- tures have been made from their preceding classification. Donacia tuberculata Lacordaire is removed from synonymy with D. rufa Say (now placed in Plateumaris). Donacia limonia Schaeffer is removed from synonymy with D. biimpressa Melsheimer. Donacia angustipes Marx is reduced to synonymy of D. palmata Olivier; D. rutila Melsheimer is synonymized with D. tuberculata Lacordaire. Details of label data, number, kind and location of preservation of type specimens (if existing) of all described taxa are given. Lectotypes (and paralectotypes, if any) are designated for the following nominal taxa: D. alutacea LeConte, D. biimpressa Melsheimer, D. carolina Lacordaire, D. cincticornis Newman, D. congener LeConte, D. cuprea Melsheimer, D. distincta LeConte, D. hypoleuca Lacordaire, D. lucida Lacordaire, D. magnifica LeConte, D. militaris Lacordaire, D. piscatrix Lacordaire, D. rudicollis Lacordaire, D. rufescens Lacordaire, D. rufipennis Lacordaire, D rutila Melsheimer, D. texana Crotch, D. torosa LeConte and D. tuberculata Lacordaire. Types of D. palmata Olivier and D. claudicans Germar are believed lost, but are distinctive taxa whose identity is not in doubt.

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1990. Reconstructed phylogeny and reclassification of the genera of Donaciinae (Coleoptera: Chrysomelidae). Quaestiones Entomologicae 26: 601-664. 1991. Classification, reconstructed phylogeny and geographic history of the New World members of PIateumaris Thomson, 1859 (Coleoptera: Chrysomeli- dae). Memoirs of the Canadian Entomological Society 157: 1-175. BECKER, E.C. 1974. Designation of lectotypes for 13 species of Coleoptera described by Couper in 1864 and 1865. Annales de la Soci6t6 entomologique de Qu6bec 19: 62-73. BLATCHLEY, W.S. 1910. An illustrated descriptive catalogue of the Coleoptera or Beetles (exclusive of the Rhynchophora) known to occur in Indiana. Nature Publ. Co., Indi- anapolis. 1386 pp. 1930. Blatchleyana. A list of published writings of W.S. Blatchley, A.B., A.M., L.L.D. of Indianapolis, Indiana and Dunedin, Florida. Together with a chronology of his life: the fixation of types of new genera and species described by him, etc., etc. Nature Publ. Co., Indianapolis. 77 pp. BOROWlEC, L. 1984. Zoogeographical study on Donaciinae of the world (Coleoptera, Chrysomelidae). Polski Pismo Entomologiczne. 53:433-518. BRIVIO, C. AND E.U. BALSBAUGH, JR. 1984. New genitalic characters for distinguishing Donacia subtilis Kunze and D. fulgens LeConte (Coleoptera: Chrysomelidae). Coleopts. Bull. 38: 305-311. CHEN, S.H. 1941. Notes on donaciine beetles. Sinensia 12: 1-17. CLARK, H. 1866. A catalogue of Phytophaga (Coleoptera, Pseudotetramera). Appendix. Williams & Morgate, London and Edinburgh. + 88 pp. CLAVAREAU, H. 1913. Sagrinae, Donaciinae and Criocerinae, In Junk, W. and S. Schenkling, Edi- tors. Coleopterorum Catalogus 51: 1-103. COUPER, W. 1864. Paper V. List of Coleoptera and Diptera taken at Quebec, and other parts of Lower Canada. Transactions of the Library and Historical Society of Que- bec. New Series, Part 2. Session of 1863-1864: 75-93. CROTCH, G.R. 1873. Materials for the study of the Phytophaga of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 25: 19-83. FABRICIUS, J.C. 1775. Systerna Entornologicae, sistens insectorurn classes, ordines, genera, species, adjectis, synonymis, locis, descriptionibus, observationibus. Korte, Flensburgi et Lipsiae. 8. [30] / 832 pp [not seen]. FRISON, T.H. 1937. A list of the insect types in the collections of the Illinois State Natural His- tory Survey and the University of Illinois. Bulletin of the Illinois Natural History Survey 16: 137-309. GERMAR, E.F. 1811. II. Nachtrige zu Ahrens Monographie der Rohrkifer. Neue Schriften der Naturforschenden Gesellschaft zu Halle I: 26-38. 1991] Askevold 189

1821. In Wiedemann, C.R.W. and E.F. Germar. Neue exotische Kifer beschrieben yon Wiedemann und Germar. Magazin der Entomologie 4: 107-183. GISTEL, J.N.F.X. 1857. Achthundert und zwanzig neue oder unbeschriebene wirbellose Thiere. Vacuna 2: 513-606. [Also published separately, Straubing, Schorner, 1857. $: 94 pp, under the title "Vacuna oder die Geheimnisse BUS der organischen und leblosen Welt." The former is usually cited.] GOECKE, H. 1931. Die Gattung Sominella Jocobson (Col. Chrys. Donaciini) (1. Beitrag zur Kenntnis der Donaciinen). Entomologische Bliitter ftir Biologie und Sys- tematik der Kiifer. 27: 152-161, plate. GUIRIN-MINIVILLE, F.E. 1944. Iconographie du Rgne Animal de G. Cuvier, repr6sentation d'aprs nature de l'une des espces les plus remarquables et souvent non encore figur6es, de chaque genre d'animaux. Tome III. Paris, J.B. Baillire, 1829-1844. 8. 576 pp, 104 plates. HATCH, M.H. 1938. A new species of Donacia from Washington. Pan-Pacific Entomologist 14: 110-112. 1971. The Beetles of the Pacific Northwest Part V. Rhipiceroidea, Sternoxi, Phy- tophaga, Rhynchophora, and Lamellicornia. University of Washington Publications in Biology Vol. 16. xiv + 662 pp. HAUPT, H. 1956. Beitrag zur Kenntnis der eozinen Arthropodenfauna des Geiseltales. Nova Acta Leopoldina (N.F.) 18:90 pp. HOPPE, D.H. 1795. Enumeration Insectorum elytratorum circa Erlangam indigenarum secun- dum systema Fabricianum observationibus iconibusque illustrata. Erlan- gae, Palm 1795. iv + 70 pp + pl. JACOBY, M. 1884. Beschreibung neuer Phytophagen. Stettiner entomologische Zeitung 45: 126-128. JACOBY, M. AND H. CLAVAREAU 1904. Coleoptera Phytophaga Fam. Donacidae. In P. Wytsman, Genera Insecto- rum 21: 1-14, plate. JOLIVET, P. 1970. Donaciinae. In Junk, W. Editor. Coleoptm. Cat. Suppl. Pars 51. Fasc. 2.71 pp. W.O. Steel, Gravenhage. KIRBY, W. 1837. Part the fourth and last. The Insects In Richardson, J. Fauna Boreali-Amer- icana; or the Zoology of the Northern Parts of British America: containing descriptions of the objects of natural history collected on the late northern land expedition under command of Cpt. Sir John Franklin, R.N. Norwich. xxxix + 325 pp, + 8 plates. KNAB, F. 1905. A new species of Donacia. Proceedings of the Entomological Society of Washington 7: 122-123. 190 Psyche [Vol. 98

KUNZE, G. 1818. I. Beitrige zur Monographie der Rohrkifer. Neue Schriften der naturforschenden Gesellschaft zu Halle. 2: 1-56. LACORDAIRE, TH. 1845. Monographie des col6optres subpentam6res de la famille des phy- tophages. Tome 1. M6moires de la Soci6t6 Royale des Sciences de Liege. Tome 3 (pt. 1). xiii + 740 pp. LECONTE, J.L. 1851. Synopsis of the species of Donacia (Fabr.) inhabiting the United States. Proceedings of the California Academy of Sciences 5: 310-316. 1857 (1860). No. 1. Report upon the insects collected on the Survey, pp 1-72 In, Report of Explorations and Surveys, to ascertain the most practicable and economical route for a railroad from the Mississippi River to the Pacific Ocean, made under the direction of .the Secretary of War in 1853-5. Vol- ume XII Book II. Washington, T.H. Ford 1860. 1861. New species of Coleoptera inhabiting the Pacific district of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 1861: 338-359. 1868. New Coleoptera collected on the Survey for the extension of the Union Pacific Railway, E.D. from Kansas to Fort Craig, New Mexico. Transac- tions of the American Entomological Society 2: 49-64. 1878. Additional descriptions of new species. Pp. 373-434 In, J.L. LeConte and E.A Schwarz, Editors. Coleoptera of Florida and Michigan. Proceedings of the American Philosophical Society 1878: 353-669. LENG, C.W. 1891. Revision of the Donaciae of Boreal America. Transactions of the American Entomological Society 18: 159-176. MANNERHEIM, C.G. 1843. Beitrag zur Kaefer-Fauna der Aleutischen Inseln, der Insel Sitkha und Neu- Californiens. Bulletin de la Soci6t6 Imp6riale des Naturalistes de Moscou 16: 175-314. MARX, E.J.F 1957. A Review of the subgenus Donacia in the Western Hemisphere (Coleoptera: Donaciidae). Bulletin of the American Museum of Natural History 112: 191-278. MEAD, A.R. 1938. New subspecies and notes on Donacia with key to the species of the Pacific States. The Pan-Pacific Entomologist 14:113-120. MEDVEDEV, L.N. 1973. New Leaf Beetles from Palaearctic. Entomological Review of Washington 52: 562-568. [Translated from: Ent. Obozr. 52: 876-885.] MELSHEIMER, F.E. 1847. Descriptions of new species of Coleoptera of the United States. Proceed- ings of the California Academy of Sciences 3: 158-181. NEWMAN, E. 1838. Entomological Notes. The Entomological Magazine V (1838): 532 pp. plates 17-18. 1991] Askevold 191

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INDEX TO SPECIES, GENUS AND FAMILY GROUP NAMES aenea, 178 flagellata, 186 pallipes, 176 aerea, 178 flavipennis, 169 palmata, 182, 183 aequalis, 174, 176 flavipes, 169, 171 parva, 169 affinis, 171 flohri, 186 parvidens, 183 alutacea, 184 floridae, 182 piscatrix, 183, 184 americana, 186 frosti, 169 Plateumarini, 168 angustipes, 183 fulgens, 175, 178 Plateumaris, 168 antillarum, 181 fulvipes, 169 Poecilocera, 172 assimilis, 173 gentilis, 169 porosicollis, 178 aurichalcea, 173 germari, 169 proxima, 174, 185 aurifera, 168 glab rata, 173 pubescens, 178 balli, 168 harrisii, 172 pubicollis, 172 biimpressa, 173, 177 hirticollis, 175 pulchella, 181 binodosa, 170 hypoleuca, 182 pusilla, 170, 171 caerulea, 174, 185 idola, 169 pyritosa, 171 californica, 185 indica, 169 quadricollis, 175, carolina, 184 juncina, 170 178 cataractae, 169, 170 jucunda, 171 robusta, 171 cazieri, 174 kirbyi, 171 rudicollis, 176 chalcea, 171 !iebecki, 176 rufa, 170, 171,179 cincticornis, 170, limonia, 177 rufescens, 185 177, 180, 181 lodingi, 171 rufipennis, 180 claudicans, 183 longicollis, 170 rugifrons, 170 confluenta, 174 lucida, 180, 181 rugosa, 178 confusa, 174 Macroplea, 186 rutila, 179 congener, 184 magistrigata, 178 serricauda, 169 cuprea, 175, 184 magnifica, 177 schaefferi, 170, 171 curticollis, 175 megacornis, 177 shoemakeri, 169, 171 dissimilis, 175 megalocera, 177 Sominella, 172 distincta, 175 megatocera, 177 subtilis, 174, 175, diversa, 168 melsheimeri, 186 178 dives, 169 metallica, 169 sulcicollis, 171 Donacia (Donacia), militaris, 182 tenuis, 181 180 minnesotensis, 186 texana, 185 Donacia (Dona- minor, 185, 186 torosa, 174 ciomima), 173 nana, 169 tryphera, 181 Donaciella, 172 Neohaemonia, 186 tuberculata, 171,179 Donaciini, 172 neomexicana, 170 tuberculifrons, 179 dubia, 168, 169 nigricornis, 186 vermiculata, 170 edentata, 181 nitida, 170, 171 vicina, 179 emarginata, 169, 170 notmani, 170 wallisi, 169 episcopalis, 185 occidentalis, 175 femoralis, 169 pacifica, 170 International Journal of Peptides

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