Descriptions of Some Neotropical Euliini and Archipini (Lepidoptera, Tortricidae)
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DESCRIPTIONS OF SOME NEOTROPICAL EULIINI AND ARCHIPINI (LEPIDOPTERA, TORTRICIDAE) Razowski, J., 1990. Descriptions of some Neotropical Euliini and Archipini (Lepidoptera, Tortn- cidae). Misc. Zool., 14: 105-114. Descriptions of some Neotropical Euliini and Archipini (Lepidoptera, Tortricidae).- Four new ge- nera (Hyptiharpa n. gen., Hypenolobosa n. gen., Netechma n. gen., Icteralaria n. gen.) and seven new species (Hyptiharpa hypostas n. sp., Hypenolobosa glechoma n. sp.,Atepa colaptes n. sp., Ic- teralaria idiochroma n. sp., 1. ichnobursa n. sp., Clepsis naucinum n. sp. and C. misgurna n. sp.) are described. Key words: Euliini, Archipini, Tortncidae, Lepidoptera, Neotropical. (Rebut: 4 VI 90; Acceptació condicional: 11 11 91; Acc. definitiva: 21 V 91) Józef Razowski, Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17,31-016 Kraków, Polska (Poland). INTRODUCTION (USA). The holotypes and the majority of the paratypes are deposited in the collection of The present paper contains the descriptions that institution and some paratypes are in the of new taxa of Euliini and Archipini. The list- collection of the Institute of Systematics and ing of their genera and species is given in the Evolution of Animals, PAS, Kraków (Po- checklist of Neotropical Lepidoptera (Po- land). WELL et al., in press). Whilst to Euliini belong The number given in the descriptions for 59 described Neotropical genera and several the labial palpus indicates the proportion of further genera are expected (four new genera its total length to the diameter of the eye. are described below), only six genera of Ar- chipini are recorded of that region. Four of them (Argyrotaenia Stephens, 1852, Choris- RESULTS toneura Lederer, 1859, ArchipsHübner, 1822 and Clepsis Guenée, 1845) are widely distri- Tribe Euliini buted in the Holarctic and Oriental regions and only Durangarchips Powell, 1988 and The genera described in this paper belong to Idolatteria Walsingham, 1913 are exclusively two groups: one (Netechma, Icteralaria) is Neotropical in distribution. rather close to other Neotropical Euliini as mentioned in their descriptions, and the other (Hyptiharpa, Hypenolobosa) has a se- MATERIAL AND METHODS parate position within the tribe. They show some characters similar to those of Cochylini, The studied material was collected by the such as the shape of the valva and its strength, staff of the Division of Enthomology and Pa- or in Hyptiharpa the complete reduction of rasitology, University of California, Berkeley the gnathos. However, the juncture among Misc. Zool. 14,1990 the tegumen, valva and vinculum is different pica1 tortricine shape; genital segment almost from that in the cochyline moths. The shape uniform, in large part membranous, with of the aedeagus recalls that there are some dense groups of sceit scales on ventral and la- Chlidanotini in which the opening for the teral surfaces. Sterigma, a sclerotized median ductus ejaculatorius is ventral. These two ge- plate distinctly concave medially, with long nera are included in Euliini on the basis of the lateral ribs not connecting with apophyses an- forewing pattern which is close to that of the teriores; ostium in anterior part of segment; majority of the species of this tribe. Their dor- distal portioin of ductus bursae slightly broad- sal blotch is a part of the usual tortricine basal ening, not sclerotized; ductus seminalis blotch, the other remainder of which is at cos- postmedian; accessory bursa and signa ab- ta and wing base. Another, more posterior sent. dorsal blotch is probably derived of the me- Bio1ogy.- Moths collected in July at alti- dian pattern and is connected with the me- tude of 800'. dian fascia. The males have no scale pencil on Distribution.- Sinaloa and Chihuahua the foreleg, but that is often reduced in the re- (Mexico) . presentatives of the tribe. Comments.-The probable autapomorph- ies of this genus are the shape of the sacculus, the presence of the membranous, plicate and setose ventral portion or the disc of valva, the Hyptiharpa n. gen. form of the proximal part of aedeagus, the form of the subgenital segment of the female Type-species: Hyptiharpa hypostas n. sp. and the shape of the sterigma. The genitalia of the two sexes show some distinct reduc- Labial palpus 2. Venation: in forewing al1 tions, the most important of which is a com- veins separate, R5to termen below apex; trace plete atrophy of the gnathos. of chorda at 113 distance R, - R,; Cu, originat- ing opposite mid-length of R, - R,; in hind- wing M, - Cu, very close to one another at Hyptiharpa hypostas n. sp. discal cell. No sexual dimorphism. Male genitalia.- Pedunculi rather short, Alar expanse 9-10 mm. Head ochreous, labial top of tegumen slightly elongate; uncus well palpus more orange basally; thorax brownish sclerotized, without terminal brush; gnathos ochreous, with collar and distal part of tcgula absent; socius with base situated beyond mid- paler, ochreous. Costa of forewing convex, dle of its length, rounded dorsally, fusing with less so in male than in female; apex rather lateral folds of tegumen; vinculum fully deve- rounded; termen convex, slightly oblique. loped. Costa of valva well sclerotized except Ground colour pale ochreous cream suffused terminal portion; sacculus large, well sclerotiz- and sprinkled with brown especially along ed, with free, directed backwards terminal discal cell and posteriorly to it. Pattern portion; ventral portion of disc of valva mem- brown, pale edged, consisting of basal blotch branous, weakly plicate, connected to juxta, represented by triangular part situated at dor- provided with some small setae. Median part sum and small remainder at costa and base, of transtilla strongly sclerotized, long, lateral another triangular blotch in middle of dorsun parts small; juxta simple, concave posterior- with proximal edge convex and distal edge ly. Membrane of anellus with numerous mi- slightly concave connected to mid-part of crotrichia. Aedeagus simple, membranous proximal edge of slender median fascia, and ventrally; coecum penis rounded; caulis bilo- subapical fascia reaching mid-termen. Proxi- be distally; opening for ductus ejaculatorius mal edge of the latter abruptly concavc be- ventro-proximal; cornuti absent. yond costa, distal edge slightly convex. Dor- Fcmale genitalia.-Ovipositor short, of ty- sal strigulae and some dots at costa concolor- ous with pattern. Fringes concolorous with Biology.- No data except for date of co- ground colour, in female tinged orange. llection of holotype. Hindwing brown with rounded apex; fringes Distribution.- Chihuahua (Mexico). paler. Male genitalia (figs. 1,2).- Uncus strong- ly sclerotized, slender, bent, slightly tapering Hypenolobosa glechoma n. sp. subterminally, not setose; socius in dorsal part ovate, slender in ventral portion. Valva Alar expanse 2 mm; labial palpus brownish broad to beyond middle, then tapering termi- cream, remaining parts of head darker. Fore- nally; sacculus strongly sclerotized, setose, wing slender, with costa slightly convex and broad except for thin, strongly curved free termen somewhat oblique. Ground colour termination provided with a minute subter- cream with indistinct brown admixture; pat- mina1 thorn; ventro-caudal portion of valva tern brownish, darker on edges; a few costal well sclerotized. Median part of transtilla lar- spots concolorous with pattern; basal blotch ge, tapering terminally. Aedeagus small, ta- atrophied in basal and costal areas of wing, pering distally, with sharp apex. remaining part larger, triangular, with obli- Female genitalia (fig. 20).- Papilla analis que distal edge; median fascia reaching tor- rather small; apophyses posteriores slightly nus, with distal edge concave at cubital arm of longer than apophyses anteriores; anterior discal cell; proximal edge of dorsal blotch edge of ventral part of genital segment strong- straight; subapical fascia reaching tornus. Re- ly sclerotized laterally; media1 part of sterig- mainder of fringes concolorous with ground ma broad extending posteriorly and some- colour. Hindwing brown-grey; fringes proba- what so anteriorly, provided with a median bly cream (damaged), with basal line brown- pit deepened distally; small, spined membran- ish grey. ous fold beyond sterigma. Corpus bursae Male genitalia (figs. 3-3.- Uncus strongly elongate. curved, uniformly broad throughout, distinct- Ho1otype.- Male, "27 mi E Villa Union, ly flattened laterally; socius slender, hairy; la- 8007,Sin. aloa, Mex. ico, VII-26-64, J. Powell, teral lobe of tegumen extending medially, Black & White lights"; genitalia slide 6034. fused with arm of gnathos; median part of gnat- Paratype.- Female, "21 mi E Villa Union, hos plate-shaped; vinculum well developed, 300', Sin. aloa, Mex. ico, VII-25-64, J.A. sclerotized along proximal edge, with minute Chemsak &J. Powell, Black & White lights"; median prominence at its middle. Valva genitalia slide 11824. broad with terminal portion slender, tapering apicad; disc sparsely hairy; sacculus broad, with ventral edge convex, provided with post- Hypenolobosa n. gen. basal and subterminal thorns; termination of sacculus slender, sharp, minutely spined; dis- Type-species: Hypenolobosa glechoma n. sp. tinct thorn on ventral margin of left sacculus (asymmetry), medially; large, flat, sclerotiz- Wing venation and pattern as in Hyptiharpa. ed lobe with somewhat plicate surface and Male genita1ia.- Similar to those in that minutely dentate outher edge dorsally to sac- genus (cf notes below