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Morphology, Taxonomy, and Biology of Larval Scarabaeoidea
Digitized by the Internet Archive in 2011 with funding from University of Illinois Urbana-Champaign http://www.archive.org/details/morphologytaxono12haye ' / ILLINOIS BIOLOGICAL MONOGRAPHS Volume XII PUBLISHED BY THE UNIVERSITY OF ILLINOIS *, URBANA, ILLINOIS I EDITORIAL COMMITTEE John Theodore Buchholz Fred Wilbur Tanner Charles Zeleny, Chairman S70.S~ XLL '• / IL cop TABLE OF CONTENTS Nos. Pages 1. Morphological Studies of the Genus Cercospora. By Wilhelm Gerhard Solheim 1 2. Morphology, Taxonomy, and Biology of Larval Scarabaeoidea. By William Patrick Hayes 85 3. Sawflies of the Sub-family Dolerinae of America North of Mexico. By Herbert H. Ross 205 4. A Study of Fresh-water Plankton Communities. By Samuel Eddy 321 LIBRARY OF THE UNIVERSITY OF ILLINOIS ILLINOIS BIOLOGICAL MONOGRAPHS Vol. XII April, 1929 No. 2 Editorial Committee Stephen Alfred Forbes Fred Wilbur Tanner Henry Baldwin Ward Published by the University of Illinois under the auspices of the graduate school Distributed June 18. 1930 MORPHOLOGY, TAXONOMY, AND BIOLOGY OF LARVAL SCARABAEOIDEA WITH FIFTEEN PLATES BY WILLIAM PATRICK HAYES Associate Professor of Entomology in the University of Illinois Contribution No. 137 from the Entomological Laboratories of the University of Illinois . T U .V- TABLE OF CONTENTS 7 Introduction Q Economic importance Historical review 11 Taxonomic literature 12 Biological and ecological literature Materials and methods 1%i Acknowledgments Morphology ]* 1 ' The head and its appendages Antennae. 18 Clypeus and labrum ™ 22 EpipharynxEpipharyru Mandibles. Maxillae 37 Hypopharynx <w Labium 40 Thorax and abdomen 40 Segmentation « 41 Setation Radula 41 42 Legs £ Spiracles 43 Anal orifice 44 Organs of stridulation 47 Postembryonic development and biology of the Scarabaeidae Eggs f*' Oviposition preferences 48 Description and length of egg stage 48 Egg burster and hatching Larval development Molting 50 Postembryonic changes ^4 54 Food habits 58 Relative abundance. -
Quick Guide for the Identification Of
Quick Guide for the Identification of Maryland Scarabaeoidea Mallory Hagadorn Dr. Dana L. Price Department of Biological Sciences Salisbury University This document is a pictorial reference of Maryland Scarabaeoidea genera (and sometimes species) that was created to expedite the identification of Maryland Scarabs. Our current understanding of Maryland Scarabs comes from “An Annotated Checklist of the Scarabaeoidea (Coleoptera) of Maryland” (Staines 1984). Staines reported 266 species and subspecies using literature and review of several Maryland Museums. Dr. Price and her research students are currently conducting a bioinventory of Maryland Scarabs that will be used to create a “Taxonomic Guide to the Scarabaeoidea of Maryland”. This will include dichotomous keys to family and species based on historical reports and collections from all 23 counties in Maryland. This document should be cited as: Hagadorn, M.A. and D.L. Price. 2012. Quick Guide for the Identification of Maryland Scarabaeoidea. Salisbury University. Pp. 54. Questions regarding this document should be sent to: Dr. Dana L. Price - [email protected] **All pictures within are linked to their copyright holder. Table of Contents Families of Scarabaeoidea of Maryland……………………………………... 6 Geotrupidae……………………………………………………………………. 7 Subfamily Bolboceratinae……………………………………………… 7 Genus Bolbocerosoma………………………………………… 7 Genus Eucanthus………………………………………………. 7 Subfamily Geotrupinae………………………………………………… 8 Genus Geotrupes………………………………………………. 8 Genus Odonteus...……………………………………………… 9 Glaphyridae.............................................................................................. -
The Asian Mango Flower Beetle, Protaetia Fusca (Herbst), and Euphoria Sepulcralis (Fabricius) in Florida and the West Indies (Coleoptera: Scarabaeidae: Cetoniinae)
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Center for Systematic Entomology, Gainesville, Insecta Mundi Florida 9-1-2006 The Asian mango flower beetle, Protaetia fusca (Herbst), and Euphoria sepulcralis (Fabricius) in Florida and the West Indies (Coleoptera: Scarabaeidae: Cetoniinae) Robert E. Woodruff Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services, [email protected] Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi Part of the Entomology Commons Woodruff, Robert E., "The Asian mango flower beetle, Protaetia fusca (Herbst), and Euphoria sepulcralis (Fabricius) in Florida and the West Indies (Coleoptera: Scarabaeidae: Cetoniinae)" (2006). Insecta Mundi. 114. https://digitalcommons.unl.edu/insectamundi/114 This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 20, No. 3-4, September-December, 2006 227 The Asian mango flower beetle, Protaetia fusca (Herbst), and Euphoria sepulcralis (Fabricius) in Florida and the West Indies (Coleoptera: Scarabaeidae: Cetoniinae) Robert E. Woodruff Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P.O. Box 147100, Gainesville, FL 32614-7100 Abstract. The mango flower beetle, Protaetia fusca (Herbst), an Asiatic pest of several commercial crops, was first found in Florida in 1985. All subsequent Florida specimens are recorded here, as well as the first documented records from the Bahamas and Barbados. Illustrations are provided and comparisons are made to the similar native Euphoria sepulcralis (Fabricius), which is also recorded for the first time from the Bahamas. -
Insect Egg Size and Shape Evolve with Ecology but Not Developmental Rate Samuel H
ARTICLE https://doi.org/10.1038/s41586-019-1302-4 Insect egg size and shape evolve with ecology but not developmental rate Samuel H. Church1,4*, Seth Donoughe1,3,4, Bruno A. S. de Medeiros1 & Cassandra G. Extavour1,2* Over the course of evolution, organism size has diversified markedly. Changes in size are thought to have occurred because of developmental, morphological and/or ecological pressures. To perform phylogenetic tests of the potential effects of these pressures, here we generated a dataset of more than ten thousand descriptions of insect eggs, and combined these with genetic and life-history datasets. We show that, across eight orders of magnitude of variation in egg volume, the relationship between size and shape itself evolves, such that previously predicted global patterns of scaling do not adequately explain the diversity in egg shapes. We show that egg size is not correlated with developmental rate and that, for many insects, egg size is not correlated with adult body size. Instead, we find that the evolution of parasitoidism and aquatic oviposition help to explain the diversification in the size and shape of insect eggs. Our study suggests that where eggs are laid, rather than universal allometric constants, underlies the evolution of insect egg size and shape. Size is a fundamental factor in many biological processes. The size of an 526 families and every currently described extant hexapod order24 organism may affect interactions both with other organisms and with (Fig. 1a and Supplementary Fig. 1). We combined this dataset with the environment1,2, it scales with features of morphology and physi- backbone hexapod phylogenies25,26 that we enriched to include taxa ology3, and larger animals often have higher fitness4. -
SCARABS “Hvisdet Lugter, Vil De Komme”
SCARABS “Hvisdet Lugter, Vil De Komme” Occasional Issue Number 15 November, 2004 Scarabs is Back!!! WITHIN THIS ISSUE As If Life Wasnʼt Bad Enough....Now This! Bad News ........................... 1 Okay, so we haven’t published a and coming face to face with the Bug-Proof Clothes ............ 2 Scarabs newsletter since Occasional drivel and forced humor within Issue 14, dated October, 1998. these pages. By posting Scarabs on Glorious Art ...................... 2 We apologize, but we have been the web, distribution is much more Revised Classifification busy. Editor Rich has been busy efficient. Only those foolish enough of the Scarabaeoidea ........ 3 chasing Phobetus panamintensis to tempt fate would dare to click and planning construction of his on our file. The PDF file containing Scatalogical Ramblings . 11 basement bug room. His stint each issue can be printed at will, if Notable Publications ..... 15 at a pro wrestling (“El Mierdo”) desired. proved to be short-lived when A Collecting Tip for “The Next Big Thing” threw Rich Thanks go to Scarab Central at Bradycinetulus .................16 into the crowd, wrenching Rich’s University of Nebraska for hosting back. Editor Barney was tied up for Scarabs on their web page. Pin Labels, Macs and Mi- crosoft Word ................... 17 awhile in a house-building project, and is building up a huge supply of Another change is that Scarabs About Scarabs ..................17 his “special bait” while praying that is now in color. To celebrate our airport security does not decide to first color issue, we are including Insect Pins ....................... 17 open all containers. Editor Bill has artwork of arguably one of the most Plea for Phanaeus ...........18 been busy on the scarab speaking beautiful scarab beetles on the circuit. -
New Larval Descriptions for Two Species of <I>Euphoria</I> Burmeister
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Papers in Entomology Museum, University of Nebraska State July 2000 New larval descriptions for two species of Euphoria Burmeister (Coleoptera: Scarabaeidae: Cetoniinae: Cetoniini: Euphoriina) with a key to the known larvae and a review of the larval biology for species in the genus. Estefanía Micó Universidad de Alicante, Spain, [email protected] Andrew Smith University of Nebraska State Museum, [email protected] Miguel Angel Morón Instituto de Ecologia, Veracruz, Mexico, [email protected] Follow this and additional works at: https://digitalcommons.unl.edu/entomologypapers Part of the Entomology Commons Micó, Estefanía; Smith, Andrew; and Morón, Miguel Angel, "New larval descriptions for two species of Euphoria Burmeister (Coleoptera: Scarabaeidae: Cetoniinae: Cetoniini: Euphoriina) with a key to the known larvae and a review of the larval biology for species in the genus. " (2000). Papers in Entomology. 11. https://digitalcommons.unl.edu/entomologypapers/11 This Article is brought to you for free and open access by the Museum, University of Nebraska State at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Papers in Entomology by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. SYSTEMATICS New Larval Descriptions for Two Species of Euphoria Burmeister (Coleoptera: Scarabaeidae: Cetoniinae: Cetoniini: Euphoriina) with a Key to the Known Larvae and a Review of the Larval Biology 1 2 ESTEFANI´A MICO´ , ANDREW B. T. SMITH, AND MIGUEL ANGEL MORO´ N Systematics Research Collections, University of Nebraska State Museum,W436 Nebraska Hall, Lincoln, NE 68588Ð0514 Ann. Entomol. Soc. Am. 93(4): 795Ð801 (2000) ABSTRACT The larvae of Euphoria devulsa Horn and Euphoria lurida (F.) (Coleoptera: Scarabaeidae: Cetoniinae: Cetoniini: Euphoriina) are described for the Þrst time. -
~E of Melolonthinae
"Goklla7~ Institute of Politics & Economlc1 Bul!etjg 5~ June, 1949 ---~ .. e.... ~e of Melolonthinae With Keys to Tribes, Genera, and Species (Coleoptera: Scarabaeidae) By P. 0. Ritcher Kent~ ~ky Agriculr~ral Experiment Station University of Kentucky l-exington Fig. 1.-Third-stage larva of Phyllophaga hirticula (Knoch), left lateral view. Much enlarged. CONTENTS Page PagG Subfamily Melolonthinae 4 Tribe Macrodactylini . 15 Key to Tribes of the Tribe Pachydemini . ........ : l6 Subfamily Melolonthinae . 4 Tribe Plectrini . 17 Tribe Sericini . 5 Tribe Melolonthini . 18 Tribe Dippltaxini . 11 Bibliography . 26 Tribe Hopliini . 12 Plates, with lists of Tribe Dichelonycini . 13 symbols used ..... ...... i . · 30 1949] .,. LARVAF. OF 1\.fEI.OI.ONTHINAF. •' BIBLIOGRAPHY BOVING, A. G. 1936 Description of the larva of Plectris aliena Chapin and explanation of new terms applied to the epipharynx and raster. Proc. Ent. Soc. \Vash~ 38(8):169-185. 1942a Descriptions of the larvae of some \Vest Indian melolonthine beetles and a key to the known larvae of the tribe. Proc. U.S. Nat. 92(3146):167-176. 1942b Description of the third-stage larva of Ampllimallon majali.{ (Razoumow ski). Proc. Ent. Soc. Wash. 44(6):111-121. 1942c A classification of larvae and adults o£ the genus Ph)'llop~aga. Mem. Ent. Soc. Wash. 2:95 pages. 1945 Description of the larva and pupa or the scarab beetle Anclyonyclra mindanaona (Brenske). Jour. Wash. Acad. Sci. 35 (1):13-15. ------ 3nd CliArGHEAD, F. C. 1931 An illustrated synopsis of the principal larval forms of the order Cole optera. Ent. Amer. II (N. S.) (1·4):351 pages. BOYER, B. L. -
Bumble Flower Beetle
Published by Utah State University Extension and Utah Plant Pest Diagnostic Laboratory ENT-052-11 December 2011 Bumble Flower Beetle Taun Beddes, Cache County Horticulture Agent, and Ryan S. Davis, Arthropod Diagnostician DO YOU KNOW? • Bumble flower beetles are common throughout the growing season on flowers, oozing sap, and other sweet, overripe, or fermenting matter. • Bumble flower beetles seldom warrant the use of chemicals for control. • Control methods include removing organic material from near affected plants, and hand removal of the adult beetles from plants. INTRODUCTION The bumble flower beetle (BFB) is a common member of Fig. 2. Adult bumble flower beetle (Euphoria inda)2. the Scarabaeidae (scarab) beetle family. Its common name originated because adult BFB’s often fly close to the ground and emit a loud buzzing sound similar to that of a bumble bee. Larvae (grubs) and adults are often GENERAL BIOLOGY found in or near fermenting and/or decomposing organic Bumble Flower Beetle matter. Because BFB’s feed on and further break down organic matter they are beneficial in many situations. Scientific Name:Euphoria inda (Scarabaeidae) Economic damage caused by BFB in Utah is rare and Range: Widely distributed in the United States from they seldom require control, though larvae (Fig. 1) may Connecticut to Florida and westward to Oregon and be mistaken for other damaging pests such as Japanese Arizona. beetle and various lawn feeding white grubs. Hosts: Identification Adult:Adult BFB’s (Fig. 2) are 12-16 mm (7/16 - 5/8 inch) in length and 8-10 mm (5/16 - 3/8 inch) wide. -
Species List for Garey Park-Inverts
Species List for Garey Park-Inverts Category Order Family Scientific Name Common Name Abundance Category Order Family Scientific Name Common Name Abundance Arachnid Araneae Agelenidae Funnel Weaver Common Arachnid Araneae Thomisidae Misumena vatia Goldenrod Crab Spider Common Arachnid Araneae Araneidae Araneus miniatus Black-Spotted Orbweaver Rare Arachnid Araneae Thomisidae Misumessus oblongus American Green Crab Spider Common Arachnid Araneae Araneidae Argiope aurantia Yellow Garden Spider Common Arachnid Araneae Uloboridae Uloborus glomosus Featherlegged Orbweaver Uncommon Arachnid Araneae Araneidae Argiope trifasciata Banded Garden Spider Uncommon Arachnid Endeostigmata Eriophyidae Aceria theospyri Persimmon Leaf Blister Gall Rare Arachnid Araneae Araneidae Gasteracantha cancriformis Spinybacked Orbweaver Common Arachnid Endeostigmata Eriophyidae Aculops rhois Poison Ivy Leaf Mite Common Arachnid Araneae Araneidae Gea heptagon Heptagonal Orbweaver Rare Arachnid Ixodida Ixodidae Amblyomma americanum Lone Star Tick Rare Arachnid Araneae Araneidae Larinioides cornutus Furrow Orbweaver Common Arachnid Ixodida Ixodidae Dermacentor variabilis American Dog Tick Common Arachnid Araneae Araneidae Mangora gibberosa Lined Orbweaver Uncommon Arachnid Opiliones Sclerosomatidae Leiobunum vittatum Eastern Harvestman Uncommon Arachnid Araneae Araneidae Mangora placida Tuft-legged Orbweaver Uncommon Arachnid Trombidiformes Anystidae Whirligig Mite Rare Arachnid Araneae Araneidae Mecynogea lemniscata Basilica Orbweaver Rare Arachnid Eumesosoma roeweri -
A Field Study of the Nesting Ecology of the Thatching Ant, Formica Obscuripes Forel, at High Altitude in Colorado
Great Basin Naturalist Volume 56 Number 4 Article 6 11-21-1996 A field study of the nesting ecology of the thatching ant, Formica obscuripes Forel, at high altitude in Colorado John R. Conway University of Scranton, Scranton, Pennsylvania Follow this and additional works at: https://scholarsarchive.byu.edu/gbn Recommended Citation Conway, John R. (1996) "A field study of the nesting ecology of the thatching ant, Formica obscuripes Forel, at high altitude in Colorado," Great Basin Naturalist: Vol. 56 : No. 4 , Article 6. Available at: https://scholarsarchive.byu.edu/gbn/vol56/iss4/6 This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. Great Basin Naturalist 56(4), © 1996, pp. 326-332 A FIELD STUDY OF THE NESTING ECOLOGY OF THE THATCHING ANT, FORMICA OBSCURIPES FOREL, AT HIGH ALTITUDE IN COLORADO John R. Conwayl AB~TRAGe-A field study of the thatching ant, Formica ohscuripes Forel, at 2560 m elevation in Colorado provided information on mound density, composition, dimensions, and temperatures; worker longevity; and mite parasitization. Density was 115 moundsftla. Mounds had 1-52 entrances and PeromysCU8 fecal pellets in the thatch. Mounds conserved heat and exhibited thermal stratification. Excavations of 4 nests revealed depths of 0.3 m to almost 1 m, novel myrme cophiles, and 0-198 wingless queens per nest. Marking experiments demonstrated that some workers overwinter and live more than a yea!: Key words: Formica obscuripes, thatching ant, Colorado, ant mouruls, myrmecophiles. -
Behavior and Food Habits of the Red-Tailed Hawk
THE CONDOR VOLUME 48 SEPTEMBER–OCTOBER, 1946 NUMBER 5 BEHAVIOR AND FOOD HABITS OF THE RED-TAILED HAWK By HENRY S. FITCH, FREEMAN SWENSON and DANIEL F. TILLOTSON Intensive study of certain phases of the habits of the Red-tailed Hawk (Buteo jamaicensis) was made by the writers at the San Joaquin Experimental Range in the foothills of Madera County, California, during 1939, 1940, and the early part of 1941. This wide-ranging, common, and important raptor is well known through such detailed studies as those of McAtee (1935), Bent (1937), and Sumner (1929). The present study was intended to reveal economic importance of the Red-tailed Hawk as reflected by its feeding habits in one particular region. The rather stereotyped behavior observed by us in such habits as choice of nest sites, choice of prey, territorial perches, and size of territory, should afford significant contrast with observations of this species under other environmental conditions elsewhere. Individual adaptability enables these hawks to take advantage of varying environmental conditions. Because of their ability to cover great distances, there must be continual interchange of individuals between popu lations of the narrow foothill belt where our study was made and nearby regions such as the San Joaquin Valley, the Coast Ranges, the timber belt of the Sierra Nevada and the arid Great Basin. Banding records show that individuals may travel hundreds of miles (Bond 1939: 56; Lincoln 1939: 360). Observations on general behavior were made incidentally as opportunities arose, but it was the primary purpose of our study to determine the effect of predation of the Red-tailed Hawk on rodent populations by gathering feeding records and data regarding the population density of the hawks. -
Coastal Sage Scrub at University of California, Los Angeles
BIOLOGICAL ASSESSMENT: COASTAL SAGE SCRUB AT UNIVERSITY OF CALIFORNIA, LOS ANGELES Prepared by: Geography 123: Bioresource Management UCLA Department of Geography, Winter 1996 Dr. Rudi Mattoni Robert Hill Alberto Angulo Karl Hillway Josh Burnam Amanda Post John Chalekian Kris Pun Jean Chen Julien Scholnick Nathan Cortez David Sway Eric Duvernay Alyssa Varvel Christine Farris Greg Wilson Danny Fry Crystal Yancey Edited by: Travis Longcore with Dr. Rudi Mattoni, Invertebrates Jesus Maldonado, Mammals Dr. Fritz Hertel, Birds Jan Scow, Plants December 1, 1997 TABLE OF CONTENTS CHAPTER 1: INTRODUCTION ..........................................................................................................................1 CHAPTER 2: PHYSICAL DESCRIPTION ........................................................................................................2 GEOLOGICAL FRAMEWORK.....................................................................................................................................2 LANDFORMS AND SOILS ..........................................................................................................................................2 The West Terrace ...............................................................................................................................................3 Soil Tests.............................................................................................................................................................4 SLOPE, EROSION, AND RUNOFF ..............................................................................................................................4