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Papers in Entomology Museum, University of Nebraska State

9-16-2005 An electronic checklist of the New World chafers (Coleoptera: : ) Arthur V. Evans Smithsonian Institution, [email protected]

Andrew B. T. Smith University of Nebraska State Museum & Canadian Museum of Nature, [email protected]

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Evans, Arthur V. and Smith, Andrew B. T. , "An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae)" (2005). Papers in Entomology. Paper 2. http://digitalcommons.unl.edu/entomologypapers/2

This Article is brought to you for free and open access by the Museum, University of Nebraska State at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Papers in Entomology by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae)

Version 1 - released 16 September 2005

Arthur V. Evans and Andrew B. T. Smith

Please email us < [email protected] > if you find any errors or omissions in this checklist. We would greatly appreciate copies of new papers pertaining to New World Melolonthinae to help us keep this list as up-to-date as possible.

Suggested citation:

Evans, A. V. and A. B. T. Smith. 2005. An Electronic Checklist of the New World Chafers (Coleoptera: Scarabaeidae: Melolonthinae). Version 1. Electronically published, Ottawa, Canada. 344 pp.

AN ELECTRONIC CHECKLIST OF THE NEW WORLD CHAFERS (COLEOPTERA: SCARABAEIDAE: MELOLONTHINAE)

ARTHUR V. EVANS

Research Associate, Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC; Department of Recent Invertebrates, Virginia Museum of Natural History, Martinsville, VA; c/o1600 Nottoway Ave., Richmond, VA 23227, USA; [email protected].

and

ANDREW B. T. SMITH

Canadian Museum of Nature, P.O. Box 3443, Station D, Ottawa, ON, K1P 6P4, Canada; asmith@mus- nature.ca

INTRODUCTION

The following is a checklist of all Melolonthinae (Coleoptera: Scarabaeidae) found in the New World. It has been modified from Evans (2003) and Smith and Evans (2005) and has been updated to 1 September 2005. Included in this checklist are all of the available names given for New World Melolonthinae (both valid and invalid). Tribes are listed in traditional order (pseudo-phylogenetically) with genera, species, and subspecies listed alphabetically within. Under each valid generic name the subgenera and synonymies are listed as are type species and, in some cases, citations for keys, checklists, and bibliographies. Listed under each valid species are synonymies, distributional data by country, and citations for new combinations and spellings. A complete bibliography is included in the “References” section of all papers mentioned in the checklist. The purpose of this checklist is to present accurate and complete information for all the names of Melolonthinae in the New World. The herein is based on the current literature (even if we have unpublished data contradicting what has been published) and the nomenclature carefully follows the International Code of Zoological Nomenclature. It is our intent to update this checklist periodically as errors are corrected and new papers are published requiring changes. Please email us any errors, additions, or suggestions for this checklist. We would greatly appreciate copies of new papers pertaining to New World Melolonthinae to help us keep this list as up- to-date as possible.

ACKNOWLEDGMENTS

We would like to thank the following people for assistance in reviewing previous versions of the checklist, supplying us with pertinent literature, discussing various aspects of Melolonthinae classification, and/or helping with nomenclatural issues and determining publication dates: Charles L. Bellamy (California Department of Food and Agriculture, Sacramento, California, USA), Aleš Bezdĕk (Institute of Entomology, České Budějovice, Czech Republic), Tristao Branco (Porto, Portugal), Ev Britton (CSIRO, Canberra, Australia, now deceased), Phillip Harpootlian, (Simpsonville, South Carolina, USA), Mary Liz Jameson (University of Nebraska, Lincoln, Nebraska, USA), Paul Lago CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

(University of Mississippi, Oxford, Mississippi, USA), Scott McCleve (Douglas, Arizona, USA), Jose Mondaca (Santiago, Chile), Miguel Angel Morón (Instituto de Ecología, Xalapa, Veracruz, México), Federico Ocampo (University of Nebraska, Lincoln, Nebraska, USA), Matt Paulsen (University of Nebraska, Lincoln, Nebraska, USA), Brett Ratcliffe (University of Nebraska, Lincoln, Nebraska, USA), and Paul Robbins (Cornell University, Ithaca, New York, USA), Preparation of the checklist was supported, in part, by an NSF/BS&I grant (DEB-0342189) to A. B. T. Smith and F. C. Ocampo, an NSF/PEET grant (DEB9712447) to B. C. Ratcliffe and M. L. Jameson, the Natural History Museum of Los Angeles County, and our own resources.

SUBFAMILY MELOLONTHINAE Samouelle, 1819

Catalogues: Dejean, 1821, 1833, 1836; Haldeman and LeConte, 1853; LeConte, 1863a; Harold, 1869a; Crotch, 1873; Henshaw, 1885, 1887, 1889, 1895; Dalla Torre, 1912a, b, c 1913; Leng, 1920; Leng and Mutchler, 1914, 1917, 1927, 1933; Blackwelder, 1939, 1944; Blackwelder and Blackwelder, 1948; Blackwelder and Arnett, 1974; Bousquet, 1991; Poole and Gentili, 1996; Smith, 2001; Evans, 2003; Smith and Evans, 2005.

Tribe ONCERINI LeConte and Horn, 1861

Keys to genera: Saylor, 1938b: 101

Genus ONCERUS LeConte Oncerus LeConte, 1856: 283. Type species Oncerus floralis LeConte, by monotypy.

Oncerus floralis LeConte Oncerus floralis LeConte, 1856: 283 Distribution: United States

Genus NEFONCERUS Saylor Nefoncerus Saylor, 1938b: 102. Type species Oncerus convergens Horn, by monotypy.

Nefoncerus convergens (Horn) Oncerus convergens Horn, 1894: 394 Nefoncerus convergens (Horn); Saylor, 1938b: 102 Distribution: México

Tribe PODOLASIINI Howden, 1997

Keys to genera: Howden, 1997: 226

Genus PODOLASIA Harold Podolasia Harold, 1869b: 122 (for LeConte, 1856: 282). Type species Lasiopus ferruginea LeConte, by monotypy.

Synonym: Lasiopus LeConte, 1856: 282 (preoccupied). Type species Lasiopus ferruginea LeConte, by monotypy.

Keys to species: Saylor, 1948a: 340; Howden, 1954: 3; 1997: 234 2 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Podolasia emarginata Howden Podolasia emarginata Howden, 1954: 6 Distribution: México

Podolasia ferruginea (LeConte) Lasiopus ferrugineus LeConte, 1856: 283 Podolasia ferruginea (LeConte); Harold, 1869a: 1098 Distribution: United States

Podolasia involucris Howden Podolasia involucris Howden, 1998: 275 Distribution: México

Podolasia lavignei Howden Podolasia lavignei Howden, 1997: 238 Distribution: México

Podolasia longipenis Howden Podolasia longipenis Howden, 1997: 248 Distribution: México

Podolasia peninsularis Howden Podolasia peninsularis Howden, 1954: 9 Distribution: México

Synonym: Podolasia saylori Howden, 1954: 10

Podolasia pilosa Howden Podolasia pilosa Howden, 1954: 6 Distribution: México, United States

Podolasia rotundipenis Howden Podolasia rotundipenis Howden, 1997: 241 Distribution: México

Podolasia stillwellorum Howden Podolasia stillwellorum Howden, 1997: 239 Distribution: México

Podolasia varicolor Saylor Podolasia varicolor Saylor, 1948a: 339 Distribution: México

Genus PODOSTENA Howden Podostena Howden, 1997: 226. Type species: Podolasia bottimeri Howden, by original designation.

Key to species: Howden, 1997: 227 3 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Podostena bottimeri (Howden) Podolasia bottimeri Howden, 1958a: 291 Podostena bottimeri (Howden); Howden, 1997: 229 Distribution: United States

Podostena litoralis Howden Podostena litoralis Howden, 1997: 232 Distribution: México

Podostena rileyi Howden Podostena rileyi Howden, 1997: 230 Distribution: United States

Podostena sleeperi Howden Podostena sleeperi Howden, 1997: 231 Distribution: México

Tribe SERICINI Kirby, 1837

Keys to genera: Martínez, 1967; 331; Frey, 1973b: 315 note: see Machatschke (1959: 741) for an informative presentation on the relationships of the sericine and melolonthine scarabs.

Genus ASTAENA Erichson Astaena Erichson, 1847a: 101. Type species Astaena tridentata Erichson, by monotypy.

Synonym: Temnostoma Blanchard, 1850: 84. Type species Temnostoma sulcatipennis Blanchard, by monotypy.

Keys to species: Frey, 1973b: 325

Astaena abaca Saylor Astaena abaca Saylor, 1946d: 223 Distribution: Colombia

Astaena abcora Saylor Astaena abcora Saylor, 1946d: 224 Distribution: Colombia

Astaena acuticollis Frey Astaena acuticollis Frey, 1973b: 350 Distribution: Colombia

Astaena aequatorialis Kirsch Astaena aequatorialis Kirsch, 1885: 213 Distribution: Ecuador 4 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Astaena andicola Frey Astaena andicola Frey, 1973b: 354 Distribution: Peru

Astaena andina Frey Astaena andina Frey, 1973b: 345 Distribution: Ecuador

Astaena apolinarmaria Saylor Astaena apolinar-maria Saylor, 1946d: 225 Distribution: Colombia

Astaena argentina Moser Astaena argentina Moser, 1921b: 133 Distribution: Argentina

Astaena bahiana Moser Astaena bahiana Moser, 1918c: 326 Distribution: Brazil

Astaena baroni Moser Astaena baroni Moser, 1918c: 324 Distribution: Ecuador

Astaena biciliata Saylor Astaena biciliata Saylor 1946d: 219 Distribution: Peru

Astaena bogotana Saylor Astaena bogotana Saylor, 1946d: 228 Distribution: Colombia

Astaena boliviana Frey Astaena boliviana Frey, 1974b: 132 Distribution: Bolivia

Astaena boliviensis Moser Astaena boliviensis Moser, 1918c: 319 Distribution: Bolivia

Astaena brasiliana Moser Astaena brasiliana Moser, 1918c: 325 Distribution: Brazil

Astaena brasiliensis (Blanchard) Serica brasiliensis Blanchard, 1850: 82 Astaena brasiliensis (Blanchard); Frey, 1973b: 339 5 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Astaena bruchi Moser Astaena bruchi Moser, 1924a: 120 Distribution: Argentina

Astaena callosipygus Frey Astaena callosipygus Frey, 1973b: 332 Distribution: Bolivia

Astaena capillata Moser Astaena capillata Moser, 1918c: 311 Distribution: Bolivia

Astaena castanea Moser Astaena castanea Moser, 1918c: 313 Distribution: Colombia

Astaena catharinensis Frey Astaena catharinensis Frey, 1973b: 347 Distribution: Brazil

Astaena clypealis Moser Astaena clypealis Moser, 1921b: 135 Distribution: Ecuador

Astaena cochabamba Frey Astaena cochabamba Frey, 1973b: 333 Distribution: Bolivia

Astaena cognata Burmeister Astaena cognata Burmeister, 1855: 129 Distribution: Colombia

Astaena columbiana (Blanchard) Serica columbiana Blanchard, 1850: 82 Serica columbica [sic] (Blanchard); Brenske, 1902: 33 Astaena columbiana (Blanchard); Frey, 1973b: 354 Distribution: Colombia

Synonym: Astaena norrisi Burmeister, 1855: 128

Astaena columbiensis Moser Astaena columbiensis Moser, 1918c: 318 Distribution: Colombia

Astaena conformis Blanchard Astaena conformis Blanchard, 1850: 84 6 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Paraguay

Astaena cordobana Moser Astaena cordobana Moser, 1921b: 133 Distribution: Argentina, Bolivia

Astaena corumbana Moser Astaena corumbana Moser, 1921b: 135 Distribution: Brazil

Astaena crassitarsis Frey Astaena crassitarsis Frey, 1973b: 357 Distribution: Brazil

Astaena crinicollis Frey Astaena crinicollis Frey, 1973b: 346 Distribution: Colombia

Astaena cuyabana Moser Astaena cuyabana Moser, 1918c: 317 Astaena cujabana [sic] Moser; Frey, 1973b: 336 Distribution: Brazil, Paraguay

Synonym: Astaena rufobrunnea Moser, 1921b: 133

Astaena divergens Frey Astaena divergens Frey, 1973b: 357 Distribution: Brazil

Astaena elongata Burmeister Astaena elongata Burmeister, 1855: 129 Distribution: Brazil

Astaena excisicollis Frey Astaena excisicollis Frey, 1973b: 343 Distribution: Ecuador

Astaena excisipes Saylor Astaena excisipes Saylor, 1947b: 433 Distribution: Ecuador

Astaena explaniceps Saylor Astaena explaniceps Saylor, 1947b: 436 Distribution: Argentina

Astaena exquisita Frey Astaena exquisita Frey, 1973b: 363 Distribution: Peru 7 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Astaena fassli Moser Astaena fassli Moser, 1918c: 327 Distribution: Colombia

Astaena ferrugata Blanchard Serica ferrugata Blanchard, 1850: 82 Astaena ferrugata (Blanchard); Frey, 1973b: 364 Distribution: Bolivia

Astaena ferruginea Moser Astaena ferruginea Moser, 1918c: 312 Distribution: Colombia

Astaena forsteri Frey Astaena forsteri Frey, 1974e: 280 Distribution: Bolivia

Astaena foveicollis Kirsch Astaena foveicollis Kirsch, 1885: 212 Distribution: Colombia

Astaena fusagona Saylor Astaena fusagona Saylor, 1946d: 221 Distribution: Colombia

Astaena glabroclypealis Frey Astaena glabroclypealis Frey, 1974b: 134 Distribution: Argentina, Peru

Astaena guanabarae Frey Astaena guanabarae Frey, 1973b: 349 Distribution: Brazil

Astaena heterophylla Moser Astaena heterophylla Moser, 1921b: 137 Distribution: Bolivia

Astaena hiekei Frey Astaena hiekei Frey, 1973b: 331 Distribution: Costa Rica

Astaena hirsuta Frey Astaena hirsuta Frey, 1973b: 328 Distribution: Ecuador

Astaena hirtella Frey Astaena hirtella Frey, 1973b: 360 8 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Bolivia

Astaena incachaca Saylor Astaena incachaca Saylor, 1946d: 215 Distribution: Bolivia

Astaena insulana Moser Astaena insulana Moser, 1918c: 328 Distribution: Trinidad

Astaena insularis Moser Astaena insularis Moser, 1918c: 328 Distribution: Trinidad, Venezuela

Astaena iridescens Frey Astaena iridescens Frey, 1974b: 131 Distribution: Brasil

Astaena kuehnelti Frey Astaena kuehnelti Frey, 1973b: 361 Distribution: Venezuela

Astaena kuntzeni Moser Astaena kuntzeni Moser, 1921b:134 Distribution: Bolivia

Astaena leechi Frey Astaena leechi Frey, 1973b: 340 Distribution: Argentina

Astaena lojana Frey Astaena lojana Frey, 1973b: 334 Distribution: Ecuador

Astaena longicornis Frey Astaena longicornis Frey, 1975c: 259 Distribution: Venezuela

Astaena longula Moser Astaena longula Moser, 1921a: 53 Distribution: Bolivia

Astaena lurida Moser Astaena lurida Moser, 1918c: 310 Distribution: Bolivia

Astaena macilenta Bates Astaena macilenta Bates, 1887: 136 9 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Costa Rica, Honduras

Astaena maqueta Saylor Astaena maqueta Saylor, 1947b: 434 Distribution: Argentina

Astaena marginicollis Frey Astaena marginicollis Frey, 1973b: 325 Distribution: Argentina

Astaena marginithorax Frey Astaena marginithorax Frey, 1973b: 341 Distribution: Ecuador

Astaena micans Frey Astaena micans Frey, 1973b: 356 Distribution: Brazil

Astaena montivaga Frey Astaena montivaga Frey, 1973b: 358 Distribution: Bolivia

Astaena moseri Frey Astaena moseri Frey, 1973b: 354 Distribution: Peru

Astaena neglecta Frey Astaena neglecta Frey, 1976b: 376 Distribution: Brazil

Astaena negligens Frey Astaena negligens Frey, 1973b: 349 Distribution: Peru

Astaena nigroana Saylor Astaena nigroana Saylor, 1947b: 435 Distribution: Colombia

Astaena nitens Frey Astaena nitens Frey, 1976b: 375 Distribution: Brazil

Astaena nitidula Moser Astaena nitidula Moser, 1918c: 314 Distribution: Colombia

Astaena oblonga Moser Astaena oblonga Moser, 1918c: 323 10 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Colombia

Astaena obscurata Moser Astaena obscurata Moser, 1918c: 321 Distribution: Colombia

Astaena obscurifrons Moser Astaena obscurifrons Moser, 1921b: 135 Distribution: Bolivia

Astaena ohausi Moser Astaena ohausi Moser, 1921b: 136 Distribution: Brazil

Astaena opaca Frey Astaena opaca Frey, 1973b: 327 Distribution: Ecuador

Astaena opalicauda Bates Astaena opalicauda Bates, 1887: 136 Astaena opalicaudata [sic] Bates; Frey, 1973b: 331 Distribution: Nicaragua

Astaena opalipennis Frey Astaena opalipennis Frey, 1973b; 355 Distribution: Brazil

Astaena pauloensis Frey Astaena pauloensis Frey, 1973b: 348 Distribution: Brazil

Astaena pectoralis Moser Astaena pectoralis Moser, 1918c: 309 Distribution: Brazil

Astaena penai Frey Astaena penai Frey, 1973b: 345 Distribution: Peru

Astaena peruana Moser Astaena peruana Moser, 1918c: 315 Distribution: Peru

Astaena peruensis Frey Astaena peruensis Frey, 1973b: 354 Distribution: Peru

Astaena pilicollis Frey 11 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Astaena pilicollis Frey, 1973b: 342 Distribution: Colombia

Astaena pilosa Moser Astaena pilosa Moser, 1921b: 136 Distribution: Argentina

Astaena pilosella Kirsch Astaena pilosella Kirsch, 1885: 213 Distribution: Colombia

Astaena pinguis Burmeister Astaena pinguis Burmeister, 1855: 128 Distribution: Colombia

Astaena plaumanni Frey Astaena plaumanni Frey, 1973b: 331 Distribution: Brazil

Astaena postnodata Frey Astaena postnodata Frey, 1973b: 339 Distribution: Brazil

Astaena pottsi Saylor Astaena pottsi Saylor, 1946d: 217 Distribution: Peru

Astaena producta Bates Astaena producta Bates, 1891: 26 Distribution: Ecuador

Astaena pruinosa Moser Astaena pruinosa Moser, 1918c: 320 Distribution: Venezuela

Astaena pubescens Burmeister Astaena pubescens Burmeister, 1855: 126 Distribution: Brazil

Astaena pusilla Frey Astaena pusilla Frey, 1973b: 355 Distribution: Panama

Astaena pygidia Saylor Astaena pygidia Saylor, 1946c: 230 Distribution: Peru

Astaena pygidialis Kirsch 12 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Astaena pygidialis Kirsch, 1885: 214 Distribution: Colombia

Astaena robusta Buremister Astaena robusta Buremister, 1855: 127 Distribution: Brazil

Astaena rockefelleri Frey Astaena rockefelleri Frey, 1973b: 336 Distribution: Colombia

Astaena rosettae Frey Astaena rosettae Frey, 1973b: 334 Distribution: Brazil

Astaena rotundiceps Frey Astaena rotundiceps Frey, 1973b: 357 Distribution: Brazil

Astaena rufa Moser Astaena rufa Moser, 1921b: 134 Distribution: Brazil

Astaena rufescens Frey Astaena rufescens Frey, 1973b: 351 Distribution: Brazil

Astaena ruficollis Moser Astaena ruficollis Moser, 1926: 202 Distribution: Argentina

Astaena rugithorax Saylor Astaena rugithorax Saylor, 1946d: 222 Distribution: Colombia

Astaena salta Saylor Astaena salta Saylor, 1946d: 231 Distribution: Argentina

Astaena santaecrucis Frey Astaena santaecrucis Frey, 1973b: 327 Distribution: Brazil

Astaena saylori Frey Astaena saylori Frey, 1973b: 335 Distribution: Argentina

Astaena schereri Frey 13 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Astaena schereri Frey, 1973b: 363 Distribution: Bolivia

Astaena schneblei Frey Astaena schneblei Frey, 1973b: 340, 354 Distribution: Colombia

Astaena semiopaca Frey Astaena semiopaca Frey, 1973b: 361 Distribution: Ecuador

Astaena sericea Frey Astaena sericea Frey, 1973b: 352 Distribution: Argentina

Astaena setosa Frey Astaena setosa Frey, 1973b: 329 Distribution: Peru

Astaena simulatrix Frey Astaena simulatrix Frey, 1974b: 133 Distribution: Bolivia

Astaena sparsesetosa Frey Astaena sparsesetosa Frey, 1976b: 375 Distribution: Brazil

Astaena splendens Frey Astaena splendens Frey, 1973b: 351 Distribution: Brazil

Astaena sulcatipennis (Blanchard) Temnostoma sulcatipennis Blanchard, 1850: 84 Symmela sulcatipennis (Blanchard); Lacordaire, 1856: 208 Astaena sulcatipennis (Blanchard); Harold, 1869a: 1127 Distribution: Brazil

Synonym: Astaena fuscipennis Burmeister, 1855: 130

Astaena suturalis (Kirsch) Symmela suturalis Kirsch, 1865: 50 Astaena suturalis (Kirsch); Frey, 1973b: 346 Distribution: Colombia

Astaena tarsalis Moser Astaena tarsalis Moser, 1918c: 322 Distribution: Colombia

14 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Astaena tenella Burmeister Astaena tenella Burmeister, 1855: 131 Distribution: Brazil

Synonym: Astaena variolata Burmeister, 1855: 128

Astaena tenellula Moser Astaena tenellula Moser, 1918c: 316 Distribution: Brazil, Paraguay

Astaena tomentosa Frey Astaena tomentosa Frey, 1973b: 335 Distribution: Ecuador

Astaena tridentata Erichson Astaena tridentata Erichson, 1847a: 101 Distribution: Ecuador, Peru

Astaena truncaticeps Moser Astaena truncaticeps Moser, 1924a: 120 Distribution: Brazil

Astaena tucumana Frey Astaena tucumana Frey, 1974b: 134 Distribution: Argentina

Astaena tumidiceps Frey Astaena tumidiceps Frey, 1974b: 132 Distribution: Brazil

Astaena valida Burmeister Astaena valida Burmeister, 1855: 127 Distribution: Colombia

Astaena vicina Frey Astanea vicina Frey, 1973b: 343 Distribution: Ecuador

Astaena yungasa Saylor Astaena yungasa Saylor, 1946d: 217 Distribution: Bolivia

Astaena zischkai Frey Astanea zischkai Frey, 1973b: 359 Distribution: Bolivia

Astaena zyrota Saylor Astaena zyrota Saylor, 1946d: 232 15 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Guyana

Genus ATHLIA Erichson Athlia Erichson, 1835: 266. Type species Athlia rustica Erichson, by monotypy.

Synonym: Rivera Germain, 1903a: 392. Type species Athlia plebeja Burmeister, by monotypy.

Subgenus: Dihymenonyx Gutiérrez, 1949:16 (as genus). Type species Dihymenonyx herrerai Gutiérrez, by original designation.

Keys to species: Saylor, 1946b: 21; Gutiérrez, 1949: 13; Martínez, 1967: 337; 1974: 352

Athlia bollei Martínez Athlia bollei Martínez, 1955: 418 Athlia (Athlia) bollei Martínez; Martínez, 1974: 345 Distribution: Argentina

Athlia brasilica Saylor Athlia brasilica Saylor, 1946b: 23 Athlia (Athlia) brasilica Saylor; Gutiérrez, 1949: 14 Distribution: Brazil

Athlia bruchi Moser Athlia bruchi Moser, 1924a: 121 Athlia (Athlia) bruchi Moser; Martínez, 1974: 345 Distribution: Argentina

Athlia freyi Martínez Athlia (Athlia) freyi Martínez, 1974: 347 Distribution: Brazil

Athlia giaii Martínez Athlia giaii Martínez, 1959a: 23 Athlia (Athlia) giaii Martínez; Martínez, 1974: 345 Distribution: Argentina

Athlia herrerai (Gutiérrez) Dihymenonyx herrerai Gutiérrez, 1949: 16 Athlia (Dihymenonyx) herrerai (Gutiérrez); Martínez, 1974: 345 Distribution: Chile

Athlia parvissima Saylor Athlia parvissima Saylor, 1946b: 24 Athlia (Athlia) parvissima Saylor; Martínez, 1974: 345 Distribution: Argentina

Athlia plebeja Burmeister Athlia plebeja Burmeister, 1855: 125 16 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Rivera plebeja (Burmeister); Germain, 1903a: 392 Athlia (Athlia) plebeja Burmeister; Martínez, 1974: 345 Distribution: Chile

Synonym: Athlia rivera Saylor, 1946a: 23

Athlia problematica Gutiérrez Athlia problematica Gutiérrez, 1950: 270 Athlia (Athlia) problematica Gutiérrez; Martínez, 1974: 345 Distribution: Chile

Athlia rotundata Gutiérrez Athlia rotundata Gutiérrez, 1949: 12 Athlia (Athlia) rotundata Gutiérrez; Martínez, 1974: 345 Distribution: Chile

Athlia rustica Erichson Athlia rustica Erichson, 1835: 267 Athlia (Athlia) rustica Erichson; Martínez, 1974: 345 Distribution: Chile

Genus MALADERA Mulsant and Rey Maladera Mulsant and Rey, 1871: 343. Type species Serica holosericea (Scopoli), by monotypy.

Synonym: Aserica Lewis, 1895: 394. Type species Autoserica secreta Brenske, by subsequent designation (Arrow, 1927: 69; 1933a: 71; see Pope, 1961: 550).

Synonym: Autoserica Brenske, 1897: 377. Type species Serica piceorufa Fairmaire, by subsequent designation (Arrow, 1927: 69; see Pope, 1961: 550).

Maladera castanea (Arrow) Autoserica castanea Arrow, 1913a: 398 Aserica castanea (Arrow); Arrow, 1927: 70 Maladera castanea (Arrow); Pope, 1961: 545 Distribution: United States [Japan]

Genus NIPPONOSERICA Nomura

Nipponoserica Nomura, 1973: 139. Type species Serica similis Lewis, by original designation.

Nipponoserica peregrina (Chapin) Serica peregrina Chapin, 1938: 68 Nipponoserica peregrina (Chapin); Nomura, 1976: 188 Distribution: United States [Japan]

Genus MIOTEMNA Lacordaire

17 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Miotemna Lacordaire, 1856: 210. Type species Octotemna singularis Blanchard, by monotypy (for Blanchard, 1850: 84).

Synonym: Octotemna Blanchard, 1850: 84 (preoccupied). Type species: Octotemna singularis Blanchard, by monotypy.

Miotemna singularis (Blanchard) Octotemna singularis Blanchard, 1850: 84 Miotemna singularis (Blanchard); Lacordarie, 1856: 210 Distribution: Bolivia

Genus RAYSYMMELA Saylor Raysymmela Saylor 1947a: 161. Type species Symmela bruchi Moser, by original designation.

Keys to species: Frey, 1973b: 32

Raysymmela bruchi (Moser) Symmela bruchi Moser, 1924a: 120 Raysymmela bruchi (Moser); Saylor, 1947a: 164 Distribution: Argentina

Raysymmela curtula (Erichson) Symmela curtula Erichson, 1835: 266 Raysymmela curtula (Erichson); Saylor, 1947a: 165 Distribution: Brazil

Raysymmela huanuca Saylor Raysymmela huanuca Saylor, 1947a: 162 Distribution: Peru

Raysymmela pallipes (Blanchard) Symmela pallipes Blanchard, 1850: 85 Raysymmela pallipes (Blanchard); Saylor, 1947a: 165 Distribution: Bolivia

Genus SERICA MacLeay Serica MacLeay, 1819: 146. Type species Melolontha brunnea Linné, by monotypy.

Keys to species: Blatchley, 1910: 957; Lago et al., 1979: 24 (North Dakota); Hardy and Andrews, 1987: 175 (Nevada sand dunes); Ratcliffe, 1991: 159,162 (Nebraska); Harpootlian, 2001: 66 (South Carolina).

Synonym: Camptorhina Kirby, 1837: 128. Type species Camptorhina atracapilla Kirby, by monotypy.

Synonym: Stilbolema Harris, 1827: 7. Type species Melolontha vespertina Gyllenhal, subsequent designation (Evans 2003: 35).

18 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Serica abdita Dawson Serica abdita Dawson, 1921: 166 Distribution: United States

Serica acicula Dawson Serica acicula Dawson, 1932: 533 Distribution: United States

Serica acontia Dawson Serica acontia Dawson, 1933: 438 Distribution: United States

Serica adversa Dawson Serica adversa Dawson, 1967: 161 Distribution: United States

Serica aemula Dawson Serica aemula Dawson, 1947: 228 Distribution: United States

Serica alabama Dawson Serica alabama Dawson, 1967: 162 Distribution: United States

Serica alleni Saylor Serica alleni Saylor, 1939c: 457 Distribution: United States

Synonym: Serica searli Saylor, 1939c: 459

Serica alternata alternata LeConte Serica alternata LeConte, 1856: 276 Maladera (Paramaladera) alternata (LeConte); Nikolaev, 1979: 191 Distribution: México, United States

Serica alternata exolita Dawson Serica alternata exolita Dawson, 1933: 437 Distribution: United States

Serica alternata patruela Dawson Serica alternata patruela Dawson, 1933: 437 Distribution: United States

Serica ammomenisco Hardy Serica ammomenisco Hardy, 1987: 173 Distribution: United States

Serica antediluviana Wickham 19 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Serica antediluviana Wickham, 1912: 23 Distribution: United States Remark: fossil species

Serica anthracina LeConte Serica anthracina LeConte, 1856: 276 Distribution: Canada, United States

Synonym: Serica crassata Walker, 1866: 323

Synonym: Serica frontalis, 1856: 276

Synonym: Serica valida Harold, 1869: 123 (for LeConte, 1856: 276)

Synonym: Serica robusta LeConte, 1856: 276 (not Blanchard, 1850: 79)

Serica apatela Dawson Serica apatela Dawson, 1922: 158 Distribution: United States

Serica arkansana Dawson Serica arkanasa Dawson, 1947: 223 Distribution: United States

Serica aspera Dawson Serica aspera Dawson, 1922: 161 Distribution: United States

Serica atracapilla (Kirby) Camptorhina atracapilla Kirby, 1837: 129 Serica atricapilla [sic] (Kirby); Dawson, 1932: 542 Serica atracapilla (Kirby); Dawson, 1967: 162 Distribution: Canada, United States

Synonym: Serica cucullata Dawson, 1919a: 34

Serica atratula atratula LeConte Serica atratula LeConte, 1856: 274 Distribution: United States

Serica atratula monita Dawson Serica atratula monita Dawson, 1947: 226 Distribution: United States

Serica aviceps Dawson Serica aviceps Dawson, 1967: 162 Distribution: United States

20 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Serica barri Dawson Serica barri Dawson, 1967: 163 Distribution: United States

Serica blatchleyi Dawson Serica trociformis blatchleyi Dawson, 1932: 545 Serica blatchleyi Dawson; Dawson, 1967: 163 Distribution: United States

Serica bruneri Dawson Serica bruneri Dawson, 1967: 164 Distribution: United States

Serica calignosa Dawson Serica calignosa Dawson, 1932: 532 Distribution: United States

Serica campestris Dawson Serica campestris Dawson, 1919a: 33 Distribution: United States

Serica carolina Dawson Serica carolina Dawson, 1920: 209 Distribution: United States

Serica catalina Dawson Serica catalina Dawson, 1947: 234 Distribution: United States

Serica chaetosoma Dawson Serica chaetosoma Dawson, 1932: 529 Distribution: United States

Serica chicoensis Saylor Serica chicoensis Saylor, 1939c: 458 Distribution: United States

Serica coalinga Dawson Serica coalinga Dawson, 1952: 65 Distribution: United States

Serica cockerelli Wickham Serica cockerelli Wickham, 1914: 262 Distribution: United States Remark: fossil species

Serica concinna Dawson Serica concinna Dawson, 1947: 232 21 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: United States

Serica contorta Dawson Serica contorta Dawson, 1947: 227 Distribution: United States

Serica craighead Saylor Serica craighead Saylor, 1939a: 55

Synonym: Serica craigheadi Dawson, 1952: 73 (unjustified emendation)

Serica cruzi Saylor Serica cruzi Saylor, 1939a: 55 Distribution: United States

Serica curvata LeConte Serica curvata LeConte, 1856: 276 Distribution: Canada, United States

Serica cuyamaca Saylor Serica cuyamaca Saylor, 1939c: 459 Distribution: United States

Serica delicata Dawson Serica delicata Dawson, 1922: 161 Distribution: United States

Serica deserticola Dawson Serica deserticola Dawson, 1952: 66 Distribution: United States

Serica diablo Dawson Serica diablo Dawson, 1967: 164 Distribution: United States

Serica egregia Dawson Serica egregia Dawson, 1921: 165 Distribution: United States

Serica elmontea Saylor Serica elmontea Saylor, 1939a: 56 Distribution: United States

Serica elongata Nonfried Serica elongata Nonfried, 1891: 261 Distribution: Honduras

Synonym: Serica uniformis Nonfried, 1891: 260 22 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Serica elongulata Horn Serica elongatula Horn, 1870: 77 Distribution: United States

Serica elusa Dawson Serica elusa Dawson, 1919a: 36 Distribution: Canada, United States

Serica ensenada Saylor Serica ensenada Saylor, 1948a: 346 Distribution: México

Serica evidens Blatchley Serica evidens Blatchley, 1919: 153 (for Blatchley, 1910: 957) Distribution: United States

Synonym: Serica carinata Blatchley, 1910: 957 (not Burmeister, 1855)

Serica falcata Dawson Serica falcata Dawson, 1933: 439 Distribution: United States

Serica falli Dawson Serica falli Dawson, 1932: 534 Distribution: United States

Serica fimbriata LeConte Serica fimbriata LeConte, 1856: 275 Distribution: México, United States

Synonym: Serica michelbacheri Saylor, 1948a: 345

Serica floridana Dawson Serica floridana Dawson, 1967: 165 Distribution: United States

Serica frosti Dawson Serica frosti Dawson, 1967: 166 Distribution: United States

Serica georgiana georgiana Leng Serica georgiana Leng, 1911: 214 Distribution: Canada, United States

Serica georgiana lecontei Dawson Serica lecontei Dawson, 1921: 160 Serica georgiana lecontei Dawson; Dawson, 1952: 70 23 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: United States

Serica heteracantha Dawson Serica heteracantha Dawson, 1967: 166 Distribution: United States

Serica howdeni Dawson Serica howdeni Dawson, 1967: 167 Distribution: United States

Serica humboldti Gordon Serica humboldti Gordon, 1975: 173 Distribution: United States

Serica imitans Chapin Serica imitans Chapin, 1931: 5 (for Uhler, 1855: 415) Distribution: United States

Synonym: Omaloplia trogiformis Uhler, 1855: 415 (not Burmeister, 1855: 179; see Dawson, 1932: 546)

Serica intermixta Blatchley Serica intermixta Blatchley, 1910: 957 Distribution: United States

Serica iricolor (Say) Melolontha iricolor Say, 1824: 246 Stilbolemma iricolor (Say); Harris, 1827: 7 Serica iricolor (Say); Burmeister, 1855: 178 Distribution: United States

Serica kanakoffi Pierce Serica kanakoffi Pierce 1946: 132 Distribution: United States Remark: fossil species

Serica laguna Saylor Serica laguna Saylor, 1935a: 1 Distribution: United States

Serica ligulata ligulata Dawson Serica ligulata Dawson, 1932: 535 Distribution: United States

Serica ligulata praetermissa Dawson Serica praetermissa Dawson, 1932: 536 Serica ligulata praetermissa Dawson; Dawson, 1952: 71 Distribution: United States 24 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Serica lodingi Dawson Serica lodingi Dawson, 1952: 66 Distribution: United States

Serica loxia Dawson Serica loxia Dawson, 1920: 208 Distribution: United States

Serica mckenziei Saylor Serica mckenziei Saylor, 1935a: 2 Distribution: United States

Serica mixta LeConte Serica mixta LeConte, 1856: 276 Distribution: United States

Serica mystaca Dawson Serica mystaca Dawson, 1922: 160 Distribution: United States

Serica ochrosoma Dawson Serica ochrosoma Dawson, 1919a: 38 Distribution: United States

Serica oliver Saylor Serica oliver Saylor, 1939a: 56 Distribution: United States

Synonym: Serica joaquinella Saylor, 1939a: 56

Synonym: Serica oliveri Dawson, 1952: 73 (unjustified emendation)

Serica opposita Dawson Serica opposita Dawson, 1921: 163 Distribution: United States

Serica panda Dawson Serica panda Dawson, 1952: 67 Distribution: United States

Serica parallela Casey Serica parallela Casey, 1885a: 176 Distribution: Canada, United States

Serica parvula Blanchard Serica parvula Blanchard, 1850: 82 Distribution: United States 25 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Serica pavonia Dawson Serica pavonia Dawson, 1932: 533 Distribution: United States

Serica peleca Dawson Serica peleca Dawson, 1952: 68 Distribution: United States

Serica perigonia eremicola Dawson Serica perigonia eremicola Dawson, 1967: 168 Distribution: México

Serica perigonia perigonia Dawson Serica perigonia Dawson, 1920: 210 Distribution: United States

Serica pilifera Horn Serica pilifera Horn, 1894: 397 Distribution: México

Serica porcula Casey Serica porcula Casey, 1885a: 177 Serica perculla [sic] Casey; Nonfried, 1892b: 257 Distribution: United States

Serica prava Dawson Serica prava Dawson, 1933: 440 Distribution: México, United States

Serica pruinipennis Saylor Series [sic] prunipennis [sic] Saylor, 1936: 4 (for Saylor, 1935a: 2) Serica pruinipennis Saylor; Saylor, 1940a: 30 (emendation) Distribution: United States

Synonym: Serica mendota Saylor, 1939c: 457

Synonym: Serica pruinosa Saylor, 1935a: 2

Serica psammobuna Hardy Serica psammobunus Hardy, 1987: 174 Distribution: United States

Serica pullata Dawson Serica pullata Dawson, 1967: 170 Distribution: United States

Serica pusilla Dawson 26 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Serica pusilla Dawson, 1922: 162 Distribution: United States

Synonym: Serica errans Blatchley, 1929: 35 Serica pusilla errans Blatchley; Dawson, 1932: 538

Serica repanda Dawson Serica repanda Dawson, 1933: 439 Distribution: United States

Serica rhypha Dawson Serica rhypha Dawson, 1952: 68 Distribution: United States

Serica rossi Saylor Serica rossi Saylor, 1948a: 345 Distribution: United States

Serica sandiegensis Saylor Serica sandiegensis Saylor, 1939c: 458 Distribution: United States

Serica satrapa Dawson Serica satrapa Dawson, 1947: 231 Distribution: United States

Serica scaphia Dawson Serica scaphia Dawson, 1952: 69 Distribution: United States

Serica sculptilis Dawson Serica sculptilis Dawson, 1922: 162 Distribution: United States

Serica senta Dawson Serica senta Dawson, 1933: 440 Distribution: United States

Serica serensia Saylor Serica serensia Saylor, 1948a: 347 Distribution: México

Serica sericea (Illiger) Melolontha sericea Illiger, 1802: 75 Stilbolemma sericea (Illiger); Harris, 1827: 7 Omaloplia sericea (Illiger); Dejean, 1833: 182 Serica sericea (Illiger); Burmeister, 1855: 176 Distribution: Canada, United States 27 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Serica sericeoides Dawson Serica sericeoides Dawson, 1967: 170 Distribution: United States

Serica serotina LeConte Serica serotina LeConte, 1856: 275 Camptorhina serotina (LeConte); LeConte, 1857: 40 Distribution: United States

Serica solita Dawson Serica solita Dawson, 1922: 163 Distribution: United States

Serica spicula Dawson Serica spicula Dawson, 1921: 162 Distribution: United States

Serica sponsa Dawson Serica sponsoa Dawson, 1919b: 223 Distribution: Canada, United States

Serica stygia Dawson Serica stygia Dawson, 1933: 439 Distribution: United States

Serica subnisa Dawson Serica subnisa Dawson, 1947: 233 Distribution: United States

Serica tantula Dawson Serica tantula Dawson, 1922: 162 Distribution: United States

Serica texana LeConte Serica texana LeConte, 1856: 274 Distribution: United States

Serica tristis LeConte Serica tristis LeConte, 1850: 226 Distribution: Canada, United States

Serica trociformis Burmeister Serica trociformis Burmeister, 1855: 179 Distribution: United States

Synonym: Melolontha aphodiina Billberg, 1820: 386 (nomen oblitum)

28 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Serica ventura dorsalis Dawson Serica ventura dorsalis Dawson, 1952: 69 Distribution: United States

Serica ventura personata Dawson Serica personata Dawson 1932: 531 Serica ventura personata Dawson; Dawson, 1952: 69 Distribution: United States

Serica ventura ventura Dawson Serica ventura Dawson, 1932: 530 Distribution: United States

Serica vespertina accola Dawson Serica vespertina accola Dawson, 1921: 164 Distribution: United States

Serica vespertina vespertina (Gyllenhal) Melolontha vespertina Gyllenhal, 1817: 180 Stilbolemma vespertina (Gyllenhal); Harris, 1827: 7 Serica vespertina (Gyllenhal); Dejean, 1833: 183 Omaloplia vespertina (Gyllenhal); Harris, 1841: 29 Distribution: United States

Serica watson Saylor Serica watson Saylor, 1939a: 58 Distribution: México, United States

Synonym: Serica watsoni Dawson, 1952: 73 (unjustified emendation)

Genus SYMMELA Erichson Symmela Erichson, 1835: 261. Type species Symmela instabilis Erichson, subsequent designation (Evans 2003: 47).

Subgenus: Sayloria Frey, 1973b: 324 (as subgenus). Type species Astaena bicoloripes Saylor, by monotypy.

Keys to species: Frey, 1973b: 321

Symmela angustula Moser Symmela angustula Moser, 1919b: 7 Symmela (Symmela) angustula Moser; Frey, 1973b: 323 Distribution: Brazil

Symmela bicoloripes (Saylor) Astaena bicoloripes Saylor, 1946d: 226 Symmela (Sayloria) bicoloripes (Saylor); Frey, 1973b: 324 Distribution: Peru 29 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Symmela boliviensis Moser Symmela boliviensis Moser, 1919b: 8 Symmela (Symmela) boliviensis Moser; Frey, 1973b: 322 Distribution: Bolivia

Symmela brasiliensis Moser Symmela brasiliensis Moser, 1919b: 8 Symmela (Symmela) brasiliensis Moser; Frey, 1973b: 323 Distribution: Brazil

Symmela clypeata Erichson Symmela clypeata Erichson, 1835: 264 Symmela (Symmela) clypeata Erichson; Frey, 1973b: 323 Distribution: Brazil

Symmela corumbana Moser Symmela corumbana Moser, 1921b: 137 Symmela (Symmela) corumbana Moser; Frey, 1973b: 323 Distribution: Brazil

Symmela costaricensis Moser Symmela costaricensis Moser, 1924a: 119 Symmela (Symmela) costaricensis Moser; Frey, 1973b: 322 Distribution: Costa Rica

Symmela elegans Erichson Symmela elegans Erichson, 1835: 263 Symmela (Symmela) elegans Erichson; Frey, 1973b: 322 Distribution: Brazil

Symmela instabilis flavimana Gory Symmela instabilis var. flavimana Gory, 1841: 98 Symmela instabilis var. flavimarogo [sic] Gory; Frey, 1973b: 321 Distribution: Brazil

Symmela instabilis instabilis Erichson Symmela instabilis Erichson, 1835: 262 Symmela (Symmela) instabilis Erichson; Frey, 1973b: 321 Distribution: Brazil

Symmela jatahyensis Frey Symmela (Symmela) jatahyensis Frey, 1973b: 323 Distribution: Brazil

Symmela longula Erichson Symmela longula Erichson, 1835: 265 Symmela (Symmela) longula Erichson, Frey, 1973b: 322 30 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Symmela mutabilis Erichson Symmela mutabilis Erichson, 1835: 263 Symmela (Symmela) mutabilis Erichson; Frey, 1973b: 323 Distribution: Brazil

Symmela nitida Erichson Symmela nitida Erichson, 1835: 264 Symmela (Symmela) nitida Erichson; Frey, 1973b: 323 Distribution: Brazil

Symmela nitidicollis Burmeister Symmela nitidicollis Burmeister, 1855: 132 Symmela (Symmela) nitidicollis Burmeister; Frey, 1973b: 324 Distribution: Brazil

Symmela opaca Erichson Symmela opaca Erichson, 1835: 264 Symmela (Symmela) opaca Erichson; Frey, 1973b: 324 Distribution: Brazil

Symmela seticollis Moser Symmela seticollis Moser, 1921b: 137 Symmela (Symmela) seticollis Moser; Frey, 1973b: 321 Distribution: Brazil

Symmela tarsalis Moser Symmela tarsalis Moser, 1919b: 9 Symmela (Symmela) tarsalis Moser; Frey, 1973b: 322 Distribution: Brazil

Symmela tenella Erichson Symmela tenella Erichson, 1835: 265 Distribution: Brazil

Symmela uniformis Blanchard Symmela uniformis Blanchard, 1850: 85 Distribution: Bolivia

Symmela varians Erichson Symmela varians Erichson, 1847a: 102 Symmela (Symmela) varians Erichson; Frey, 1973b: 321 Distribution: Peru

Tribe SERICOIDINI Burmeister, 1855

Keys to genera: Gutiérrez, 1952: 208 31 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Genus APTERODEMIDEA Gutiérrez Apterodemidea Gutiérrez, 1952: 209. Type species Apterodema paraguayensis Arrow, by monotypy.

Apterodemidea paraguayensis (Arrow) Apterodema paraguayensis Arrow, 1903b: 255 Apterodemidea paraguayensis (Arrow); Gutiérrez, 1952: 210 Distribution: Paraguay

Genus SERICOIDES Guérin-Méneville Sericoides Guérin-Méneville, 1839: 301. Type species Melolontha glacialis Fabricius, subsequent designation (Evans 2003: 50).

Synonym: Apterodema Fairmaire, 1884: 490. Type species Apterodema acuticollis Fairmaire, by monotypy.

Synonym: Macrosoma Hope, 1837: 109 (preoccupied). Type species Melolontha glacialis Fabricius, by original designation.

Synonym: Maypa Blanchard, 1850: 115. Type species Maypa chlorosticta Blanchard, by monotypy.

Synonym: Paramaypa Brenske, 1906: 30. Type species Maypa orsornoana Brenske, by monotypy.

Subgenus: Accia Curtis, 1845: 453 (as genus). Type species Accia lucida Curtis, by monotypy.

Subgenus: Listronyx Guérin-Méneville, 1839: 302 (as genus). Type species Listronyx nigriceps Guérin-Méneville, a junior synonym of Sericoides testacea (Fabricius), by monotypy.

Subgenus: Paralistronyx Brenske, 1906: 30 (as genus). Type species Listronyx livida Germain, by monotypy.

Keys: Fairmaire and Germain, 1863: 724, 741

Sericoides acuticollis (Fairmaire) Apterodema acuticollis Fairmaire, 1884: 491 Sericoides (Listronyx) acuticollis (Fairmaire); Martínez, 1971: 119 Distribution: Argentina, Chile

Sericoides andina (Germain) Maypa andina Germain, 1863: 726 Sericoides andina (Germain); Dalla Torre, 1912b: 130 Distribution: Chile

Sericoides antarctica (Brenske) Listronyx antarcticus Brenske, 1900: 109 Sericoides antarctica (Brenske); Dalla Torre, 1912b: 130 Distribution: Argentina, Chile

32 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Sericoides argentinensis Martínez Sericoides (Listronyx) argentinensis Martínez, 1971: 115 Distribution: Argentina

Sericoides atra (Solier) Maypa atra Solier, 1851: 108 Sericoides atra (Solier); Dalla Torre, 1912b: 130 Distribution: Chile

Sericoides castanea Guérin-Méneville Sericoides castanea Guérin-Méneville, 1839: 302 Distribution: Argentina

Sericoides chilena (Germain) Maypa chilena Germain, 1863: 736 Sericoides chilena (Germain); Dalla Torre, 1912b: 130 Distribution: Chile

Sericoides chilensis Blanchard Sericoides chilensis Blanchard, 1850: 114 Sericoides chiliensis [sic] Blanchard; Dalla Torre, 1912b: 130 Distribution: Chile

Sericoides chlorosticta (Blanchard) Maypa chlorosticta Blanchard, 1850: 115 Sericoides chlorosticta (Blanchard); Dalla Torre, 1912b; 130 Sericoides (Paramaypa) chlorosticta (Blanchard); Gutiérrez, 1949: 18 Distribution: Chile

Synonym: Sericoides similis Philippi, 1861: 737

Sericoides comata (Germain) Maypa comata Germain, 1863: 739 Sericoides comata (Germain); Dalla Torre, 1912b: 130 Distribution: Chile

Sericoides convexa (Germain) Maypa convexa Germain, 1863: 729 Sericoides convexa (Germain); Dalla Torre, 1912b: 130 Distribution: Chile

Sericoides delicatula (Germain) Maypa delicatula Germain, 1863: 731 Sericoides delicatula (Germain); Dalla Torre, 1912b: 130 Sericoides (Paramaypa) delicatula (Germain); Gutiérrez, 1949: 18 Distribution: Chile

Sericoides dubia (Germain) 33 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Maypa dubia Germain, 1863: 738 Sericoides dubia (Germain); Dalla Torre, 1912b: 130 Distribution: Chile

Sericoides faminaei (Blanchard) Listronyx faminaei Blanchard, 1850: 114 Maypa faminaei (Blanchard); Harold, 1869a: 1138 Sericoides faminaei (Blanchard); Dalla Torre, 1912b: 130 Distribution: Argentina, Chile

Sericoides frigida (Germain) Listronyx frigida Germain, 1863: 743 Listronyx frigidus Germain; Brenske, 1906: 30 Sericoides frigida (Germain); Dalla Torre, 1912b: 131 Sericoides (Listronyx) frigida (Germain); Martínez, 1971: 115 Distribution: Chile

Sericoides germaini Dalla Torre Sericoides germaini Dalla Torre, 1912b: 131 (for Germain, 1863: 741) Distribution: Argentina, Chile

Synonym: Listronyx castanea Germain, 1863: 741 (not Guérin-Méneville, 1839: 301)

Sericoides glacialis (Fabricius) Melolontha glaciale Fabricius 1775: 35 Macrosoma glacialis (Fabricius): Hope, 1837: 109 Sericoides glacialis (Fabricius); Blanchard, 1850: 114 (emendation) Sericoides (Listronyx) glacialis (Fabricius); Gutiérrez, 1949: 19 Distribution: Argentina, Chile

Synonym: Sericoides atricapilla Curtis, 1845: 451

Synonym: Sericoides reichei Guérin-Méneville, 1839: 301

Sericoides hirsuta (Brenske) Listronyx hirsutus Brenske, 1900: 110 Sericoides hirsuta (Brenske); Dalla Torre, 1912b: 131 Distribution: Argentina, Chile

Sericoides intermedia (Philippi) Maypa intermedia Philippi, 1864: 452 Sericoides intermedia (Philippi); Dalla Torre, 1912b: 131 Distribution: Chile

Sericoides lineolata (Germain) Maypa lineolata Germain, 1863: 732 Sericoides lineolata (Germain); Dalla Torre, 1912b: 131 Sericoides (Paramaypa) lineolata (Germain); Gutiérrez, 1949: 18 34 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Chile

Sericoides livida (Germain) Listronyx livida Germain, 1863: 745 Listronyx lividus Harold, 1869a: 1138 Macrosoma (Paralistronyx) lividus (Germain); Brenske, 1906; 30 Sericoides livida (Germain); Dalla Torre, 1912b: 131 Sericoides (Paralistronyx) lividus (Germain); Martínez, 1959a: 25 Distribution: Chile

Synonym: Sericoides (Listronyx) aenigmaticus Martínez, 1959a: 27 (for Philippi 1864: 453)

Synonym: Listronyx obscura Philippi, 1864: 453 (not Germain, 1863: 728)

Synonym: Macrosoma (Paramaypa) osornoana Brenske, 1906: 30 Sericoides osornoana (Brenske); Dalla Torre, 1912b: 131

Sericoides lloydi Martínez Sericoides lloydi Martínez, 1965: 89 Distribution: Argentina

Sericoides longipes (Germain) Maypa longipes Germain, 1863: 727 Sericoides longipes (Germain); Dalla Torre, 1912b: 131 Distribution: Chile

Sericoides lucida (Curtis) Accia lucida Curtis, 1845: 454 Sericoides lucida (Curtis); Dalla Torre, 1912b: 131 Distribution: Argentina, Chile

Sericoides monticola (Germain) Maypa monticola Germain, 1863: 736 Sericoides monticola (Germain); Dalla Torre, 1912b: 131 Distribution: Chile

Sericoides multicolor Martínez Sericoides multicolor Martínez, 1956: 45 Distribution: Argentina, Chile

Sericoides nitida Philippi and Philippi Sericoides nitida Philippi and Philippi, 1864: 325 Distribution: Chile

Sericoides nossi Martínez Sericoides nossi Martínez 1972b: 50 Distribution: Argentina

35 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Sericoides obesa (Germain) Maypa obesa Germain, 1863: 728 Sericoides obesa (Germain); Dalla Torre, 1912b: 131 Distribution: Chile

Sericoides obscura (Germain) Maypa obscura Germain, 1863: 728 Sericoides obscura (Germain); Dalla Torre, 1912b: 131 Distribution: Argentina, Chile

Sericoides olivacea (Germain) Maypa olivacea Germain, 1863: 735 Sericoides olivacea (Germain); Dalla Torre, 1912b: 131 Distribution: Chile

Sericoides opacipennis (Germain) Maypa opacipennis Germain, 1863: 738 Sericoides opacipennis (Germain); Dalla Torre, 1912b: 131 Distribution: Argentina, Chile

Sericoides pallida (Germain) Listronyx pallida Germain, 1863: 744 Sericoides pallida (Germain); Dalla Torre, 1912b: 131 Distribution: Chile

Sericoides palpalis (Germain) Maypa palpalis Germain, 1863: 725 Sericoides palpalis (Germain); Dalla Torre, 1912b: 131 Distribution: Chile

Sericoides philippiana Dalla Torre Sericoides philippiana Dalla Torre, 1912b: 132 (for Philippi, 1864: 451) Distribution: Chile

Synonym: Sericoides andina Philippi, 1864: 451 (not Germain, 1863: 726)

Sericoides picea (Kolbe) Accia picea Kolbe, 1907: 111 Sericoides picea (Kolbe); Dalla Torre, 1912b: 132 Distribution: Chile

Sericoides piligera (Germain) Maypa piligera Germain, 1863: 731 Sericoides piligera (Germain); Dalla Torre, 1912b: 132 Distribution: Chile

Sericoides pubescens (Germain) Maypa pubescens Germain, 1863: 733 36 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Sericoides pubescens (Germain); Dalla Torre, 1912b: 132 Distribution: Chile

Sericoides punctata (Solier) Maypa punctata Solier, 1851: 107 Macrosoma punctatum (Solier); Harold, 1869a: 1137 Sericoides punctata (Solier); Dalla Torre, 1912b: 132 Distribution: Chile

Sericoides rechencqui Martínez Sericoides rechencqui Martínez 1972b: 45 Distribution: Argentina

Sericoides rufeola (Solier) Maypa rufeola Solier, 1851: 108 Macrosoma rufeola (Solier); Harold, 1869a: 1137 Sericoides rufeola (Solier); Dalla Torre, 1912b: 132 Distribution: Chile

Sericoides rufocastanea (Germain) Maypa rufo-castanea Germain, 1863: 737 Sericoides rufocastanea (Germain); Dalla Torre, 1912b: 132 Distribution: Chile

Sericoides rugosula (Germain) Maypa rugosula Germain, 1863: 733 Sericoides rugosula (Germain); Dalla Torre, 1912b: 132 Distribution: Chile

Synonym: Maypa cuprea Philippi, 1864: 450

Sericoides sinuaticollis (Germain) Maypa sinuaticollis Germain, 1863: 737 Sericoides sinuaticollis (Germain); Dalla Torre, 1912b: 132 Distribution: Chile

Sericoides striata (Fabricius) Melolontha striata Fabricius, 1775: 35 Macrosoma striata (Fabricius); Hope, 1837: 109 Sericoides striata (Fabricius); Dalla Torre, 1912b: 132 Distribution: Chile

Synonym: Sericoides striata Philippi, 1864: 450

Sericoides subcostata (Germain) Maypa subcostata Germain, 1863: 729 Sericoides subcostata (Germain); Dalla Torre, 1912b: 132 Distribution: Chile 37 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Maypa opaca Philippi and Philippi, 1864: 324

Sericoides sulcatopunctata (Germain) Listronyx sulcato-punctata Germain, 1863: 740 Sericoides sulcatopunctata (Germain); Dalla Torre, 1912b: 132 Distribution: Chile

Sericoides sylvatica (Germain) Maypa sylvatica Germain, 1863: 730 Sericoides silvatica [sic] (Germain); Dalla Torre, 1912b: 132 Distribution: Argentina, Chile

Sericoides testacea (Fabricius) Melolontha testacea Fabricius, 1775: 35 Macrosoma testacea (Fabricius); Hope, 1837: 109 Listronyx testacea (Fabricius); Blanchard, 1850: 114 Maypa testacea (Fabricius); Solier, 1851: 111 Sericoides testacea (Fabricius); Dalla Torre, 1912b: 132 Sericoides (Listronyx) testacea (Fabricius); Martínez, 1971: 115 Distribution: Argentina, Chile

Synonym: Melolontha fusca Gmelin, 1790: 1569

Synonym: Listronyx melanocephala Blanchard, 1846: plate 8, fig. 10

Synonym: Maypa nigriceps Guérin-Méneville, 1839: 302

Sericoides variegata (Germain) Listronyx variegata Germain, 1863: 742 (for Solier, 1851: 109) Sericoides variegata (Germain); Dalla Torre, 1912b: 133 Distribution: Chile

Synonym: Maypa chlorosticta Solier, 1851: 109 (not Blanchard, 1850: 115)

Sericoides variolosa (Solier) Maypa variolosa Solier, 1851: 109 Sericoides variolosa (Solier); Dalla Torre, 1912b: 133 Distribution: Chile

Synonym: Sericoides rugulosa Philippi 1861: 736

Sericoides vestita (Germain) Listronyx vestita Germain, 1863: 744 Sericoides vestita (Germain); Dalla Torre, 1912b: 133 Sericoides (Listronyx) vestita (Germain); Gutiérrez, 1949: 19 Distribution: Chile

38 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Sericoides viridis (Solier) Maypa viridis Solier, 1851: 107 Listronyx viridis (Solier); Redtenbacher, 1868: 66 Sericoides viridis (Solier); Dalla Torre, 1912b: 133 Distribution: Argentina, Chile

Genus ULATA Saylor Ulata Saylor, 1945c: 441. Type species Ulata argentina Saylor, by original designation.

Ulata argentina Saylor Ulata argentina Saylor, 1945c: 442 Distribution: Argentina

Tribe LIPARETRINI Burmeister, 1855

Genus APLODEMA Blanchard Aplodema Blanchard, 1850: 115. Type species Brachyphylla magellanica Blanchard, subsequent designation (Evans 2003: 58)..

Synonym: Haplodema Harold, 1869a: 1136 (unjustified emendation)

Aplodema angustata Blanchard Aplodema angustata Blanchard, 1850: 115 Distribution: Bolivia

Aplodema magellanica (Blanchard) Brachyphylla magellanica Blanchard, 1846: plate 8, fig. 9 Aplodema magellanica (Blanchard) 1850: 115 Distribution: Argentina, Chile

Aplodema rufescens (Saylor) Haplodema rufescens Saylor, 1938a: 73 Distribution: Guyana

Genus BLEPHAROTOMA Blanchard Blepharotoma Blanchard, 1850: 115. Type species Blepharotoma tarsalis Blanchard, by monotypy.

Keys to species: Frey, 1973b: 317

Blepharotoma argentina Frey Blepharotoma argentina Frey, 1973b: 319 Distribution: Argentina

Blepharotoma boccaine Frey Blepharotoma boccaine Frey, 1973b: 319 Distribution: Brazil

Blepharotoma calvicollis Frey 39 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Blepharotoma calvicollis Frey, 1973b: 321 Distribution: Argentina

Blepharotoma confusa Martínez Blepharotoma confusa Martínez, 1959a: 27 Distribution: Argentina

Blepharotoma martinezi Frey Blepharotoma martinezi Frey, 1973b: 319 Distribution: Argentina

Blepharotoma nitens Frey Blepharotoma nitens Frey, 1973b: 320 Distribution: Brazil

Blepharotoma nitidula Moser Blepharotoma nitidula Moser, 1918b: 117 Distribution: Brazil

Blepharotoma ohausiana (Saylor) Heteronyx ohausiana Saylor, 1938d: 135 Blepharotoma ohausiana (Saylor); Martínez, 1959a: 29 Distribution: Brazil

Blepharotoma petropolisana Moser Blepharotoma petropolisana Moser, 1924b: 158 Distribution: Brazil

Blepharotoma plaumanni Frey Blepharotoma plaumanni Frey, 1973b: 319 Distribution: Brazil

Blepharotoma tarsalis Blanchard Blepharotoma tarsalis Blanchard, 1850: 115 Distribution: Argentina, Bolivia

Genus HETERONYX Guérin-Méneville Heteronyx Guérin-Méneville, 1831b: pl. 3. Type species Heteronyx australis Guérin-Méneville, by monotypy. note: Britton (2000) removed the Neotropical species from Heteronyx, but did place them in a new or existing genus.

Heteronyx boliviana Moser Heteronyx bolivianus Moser, 1919b: 6 Heteronyx boliviana Moser; Blackwelder, 1944: 221 (justified emendation) Distribution: Bolivia

40 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Heteronyx corumbana Moser Heteronyx corumbanus Moser, 1921b: 138 Heteronyx corumbana Moser; Blackwelder, 1944: 221 (justified emendation) Distribution: Brazil

Heteronyx cuyabana Moser Heteronyx cuyabanus Moser, 1919b: 5 Heteronyx cuyabana Moser ; Blackwelder, 1944: 221 (justified emendation) Distribution: Brazil

Heteronyx heynei Moser Heteronyx heynei Moser, 1919b: 5 Distribution: Paraguay

Heteronyx schencklingi Moser Heteronyx schencklingi Moser, 1919b: 4 Distribution: Brazil

Genus ZABURINA Saylor Zaburina Saylor, 1945a: 79. Type species Zaburina colombiana Saylor, by original designation.

Zaburina colombiana Saylor Zaburina colombiana Saylor, 1945a: 80 Distribution: Colombia

Tribe Samouelle, 1819

Keys to genera: Howden, 1968: 546; Hardy, 1974: 4; Ratcliffe, 1991: 171

Genus AMPHIMALLON Berthold Amphimallon Berthold, 1827: 362. Type species Scarabaeus solstitialis Linné, by subsequent designation. (see Baraud, 1992: 458)

Amphimallon majalis (Razoumowsky) Melolontha majalis Razoumowsky, 1789: 249 Amphimallon majalis (Razoumowsky); Reitter, 1902: 248 Distribution: Canada, United States (western Europe)

Genus DINACOMA Casey Dinacoma Casey, 1889: 174. Type species Thyce marginata Casey, by monotypy.

Keys to species: Blaisdell, 1930: 177; Hardy, 1974: 25

Dinacoma caseyi Blaisdell Dinacoma caseyi Blaisdell, 1930: 174 Distribution: United States

Dinacoma marginata (Casey) 41 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Thyce marginata Casey, 1887: 39 Dinacoma marginata (Casey); Casey, 1889: 174 Distribution: United States

Genus FOSSOCARUS Howden Fossocarus Howden, 1961: 807. Type species Fossocarus creoleorum Howden, by original designation.

Fossocarus creoleorum Howden Fossocarus creoleorum Howden, 1961: 808 Distribution: United States

Genus GRONOCARUS Schaeffer Gronocarus Schaeffer, 1927: 213. Type species Gronocarus autumnalis Schaeffer, by monotypy.

Gronocarus autumnalis Schaeffer Gronocarus autumnalis Schaeffer, 1927: 213 Distribution: United States

Synonym: Gronocarus multispinosus Howden, 1961: 810

Gronocarus inornatus Skelley Gronocarus inornatus Skelley, 2005: 141 Distribution: United States

Genus HOWDENOCARUS Hardy Howdenocarus Hardy, 1978b: 47. Type species Howdenocarus mexicanus Hardy, by original designation.

Howdenocarus mexicanus Hardy Howdenocarus mexicanus Hardy, 1978b: 47 Distribution: México

Genus HYPOTHYCE Howden Hypothyce Howden, 1968: 542. Type species Hypothyce mixta Howden, by original designation.

Keys to species: Hardy, 1974: 8.

Hypothyce burnei Skelley Hypothyce burnei Skelley, 2005: 153 Distribution: United States

Hypothyce mixta Howden Hypothyce mixta Howden, 1968: 543 Distribution: United States

Hypothyce osburni (Cartwright) Thyce osburni Cartwright, 1967: 238 42 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Hypothyce osburni (Cartwright); Hardy, 1974: 9 Distribution: United States

Genus HYPOTRICHIA LeConte Hypotrichia LeConte, 1862: 137. Type species Hypotrichia spissipes LeConte, by monotypy.

Hypotrichia spissipes LeConte Hypotrichia spissipes LeConte, 1861: 137 Distribution: United States

Genus MIOLACHNOSTERNA Wickham Miolachnosterna Wickham, 1914: 458. Type species Miolachnosterna tristoides Wickham, by monotypy.

Miolachnosterna tristoides Wickham Miolachnosterna tristoides Wickham, 1914: 458 Distribution: United States Remark: fossil species

Genus MODIALIS Fairmaire and Germain Modialis Fairmaire and Germain, 1860: 267. Type species Modialis prasinella Fairmaire and Germain, by monotypy.

Synonym: Acanthosternum Philippi, 1861: 739. Type species Acanthosternum splendens Philippi, by monotypy.

Modialis prasinella Fairmaire and Germain Modialis prasinella Fairmaire and Germain, 1860: 267 Distribution: Chile

Synonym: Acanthosternum splendens Philippi, 1861: 739

Genus PARATHYCE Hardy Parathyce Hardy, 1974: 12. Type species Plectrodes carpenteri LeConte, by original designation.

Keys to species: Hardy, 1974: 12

Parathyce bidentata (Fall) Thyce bidentata Fall, 1932: 201 Parathyce bidentata (Fall); Hardy, 1974: 15 Distribution: United States

Parathyce carpenteri (LeConte) Plectrodes carpenteri LeConte, 1876: 516 Thyce carpenteri (LeConte); Casey, 1889: 170 Parathyce carpenteri (LeConte); Hardy, 1974: 17 Distribution: United States

43 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Thyce crinicollis Casey, 1914: 310

Synonym: Thyce pistoria Casey, 1895: 607

Parathyce fieldi (Fall) Thyce fieldi Fall, 1908: 160 Parathyce fieldi (Fall); Hardy, 1974: 13 Distribution: México, United States

Synonym: Thyce angusticollis Casey, 1914: 320

Synonym: Thyce simplicipes Casey, 1914: 319

Parathyce harfordi (Casey) Thyce harfordi Casey 1889: 173 Parathyce harfordi (Casey): Hardy, 1974: 21 Distribution: United States

Synonym: Thyce harfordi nanella Casey, 1914: 318

Synonym: Thyce longipalpis Casey, 1914: 316

Synonym: Thyce squamosa Casey, 1891: 20

Parathyce palpalis (Horn) Plectrodes palpalis Horn, 1880: 146 Thyce palpalis (Horn); Casey, 1889: 170 Parathyce palpalis (Horn); Hardy, 1974: 19 Distribution: United States

Synonym: Thyce angustula Casey, 1914: 315

Synonym: Thyce aurata Van Dyke, 1943: 105

Synonym: Thyce blaisdelli Casey, 1891: 19

Synonym: Thyce blaisdelli nitidula Casey, 1914: 313

Synonym: Thyce cinerea Van Dyke, 1943: 106

Synonym: Thyce clypeata Van Dyke, 1943: 107

Synonym: Thyce fossiger Casey, 1889: 172

Synonym: Thyce fossiger aperta Casey, 1914: 314

Synonym: Thyce fossiger brevitarsus Casey, 1914: 314

44 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Thyce fossiger ochreata Casey, 1914: 314

Synonym: Thyce pulverea Casey, 1889: 171

Synonym: Thyce rotundicauda Casey, 1914: 311

Synonym: Thyce vestita Casey, 1914: 312

Parathyce riversi (Casey) Thyce riversi Casey, 1895: 608 Parathyce riversi (Casey); Hardy, 1974: 16 Distribution: United States

Genus PHYLLOPHAGA Harris Phyllophaga Harris, 1827: 7. Type species Melolontha hirticula Knoch, by subsequent designation (Glasgow, 1916: 370).

Synonym: Abcrana Saylor, 1942a: 159. Type species Phyllophaga crinitissima Moore, by monotypy.

Synonym: Clemora Saylor, 1942a: 159. Type species Phytalus smithi Arrow, by original designation.

Synonym: Endrosa LeConte, 1856: 234. Type species Melolontha quercus Knoch, by subsequent designation (Saylor, 1942a: 162).

Synonym: Eugastra LeConte, 1856: 233. Type species Tostegoptera cribrosa LeConte, by subsequent designation (Bloeker, 1937: 84; see Saylor, 1942a: 162)

Synonym: Gynnis LeConte, 1856: 262. Type species Gynnis debilis LeConte, by monotypy.

Synonym: Lachnosterna Hope, 1837: 100. Type species Melolontha fervida Fabricius, by original designation

Synonym: Trichesthes Erichson, 1847b: 658. Type species Melolontha pilosicollis Knoch, a junior synonym of Melolontha tristis Fabricius, by monotypy.

Subgenus: Chirodines Bates, 1888: 169 (as genus). Type species Chirodines zunilensis Bates, by monotypy.

Subgenus: Chlaenobia Blanchard, 1850: 116 (as genus). Type species Chlaenobia ciliatipes Blanchard, by monotypy.

Subgenus: Cnemarachis Saylor, 1942a: 159 (as genus). Type species Lachnosterna vandinei Smyth, by original designation.

Subgenus: Listrochelus Blanchard, 1851: 141 (as genus). Type species Listrochelus castlenaui Blanchard, by monotypy.

45 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Subgenus: Phytalus Blanchard, 1851: 130 (as genus). Type species Melolontha pubereus Mannerheim, by subsequent designation (Saylor, 1942a: 162).

Subgenus: Tostegoptera Blanchard, 1851: 149 (as genus) . Type species Melolontha lanceolata Say, by monotypy.

Subgenus: Triodonyx Saylor, 1942a: 158 (as genus). Type species Triodonyx gigantissima Saylor, by original designation.

Keys to species: Horn, 1885: 119 (United States); Forbes, 1894: 139 (Illinois); Blatchley, 1910 (Indiana); Langston, 1927b: 8 (Mississippi); Luginbill, 1928: 55 (South Carolina); Chapin, 1932b: 179 (Cuba); 1935a: 2 (subgenus Chlaenobia); Travis, 1934: 315 (Iowa); Saylor, 1939b: 90 (sociatus species group); 1939c: 160 (subgenus Phytalus); 1940c: 63 (subgenus Listrochelus); Böving, 1942: 21 (larvae); Fattig, 1944: (Georgia); Reinhard, 1946: 70 (apterous species); 1950: 31 (Texas); Sanderson, 1951: 254 (Hispaniola); 1958: 159 (Arizona subgenera and species groups); Luginbill and Painter, 1953: 4 (United States); Rosander and Werner, 1970: 1136 (Arizona larvae); Frey, 1975a: 202 (South American Phyllophaga); Lago et al., 1979: 42 (North Dakota); King, 1983: 45 (Costa Rica); Morón 1986: 189 (Mexican subgenera and species groups); Woodruff and Beck, 1989: 78 (Florida); Ratcliffe, 1991: 186 (Nebraska); Morón and Deloya 1991: 216 (Durango); Warner and Morón, 1992: 323 (subgenus Triodonyx); Morón, 1991: 406 (subgenus Chirodines); 1993: 76 (Veracruz); 1994: 39 (Hidalgo); Morón, Aragón, Tapia-Rojas, Rojas-Garcia, 2000: 96 (Puebla); Aragón, Morón, Tapia- Rojas, Rojas-Garcia 2001: 155 (Puebla); Harpootlian, 2001: 75 (South Carolina).

Annotated lists: Morón, Hernandez-Rodriguez, and Ramiriez Campos, 1996: 37 (Nayarit); Cano and Morón, 1998: 9 (Guatemala); Morón, Vallejo and Restrepo-Giraldo 1998: 31 (Colombia); Solís and Morón, 1998: 21 (Costa Rica)

Bibliography: Pike et al., 1976: 2.

Phyllophaga abcea Saylor Phyllophaga (Listrochelus) abcea Saylor, 1940d: 112 Phyllophaga (Phyllophaga) abcea Saylor; Cano and Morón, 1998: 14 Distribution: Guatemala

Phyllophaga abdominalis (Moser) Lachnosterna abdominalis Moser, 1921c: 254 Phyllophaga abdominalis (Moser); Blackwelder, 1944: 223 Phyllophaga (Phyllophaga) abdominalis (Moser); Morón, 1986: 205 Distribution: México

Phyllophaga abudantuni Chalumeau and Gruner Phyllophaga (Cnemarachis) abudantuni Chalumeau and Gruner, 1976: 89 Distribution: Martinique

Phyllophaga acapulca Saylor Phyllophaga (Phyllophaga) acapulca Saylor, 1943d: 135 Distribution: México

46 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga aceitillar Woodruff Phyllophaga aceitillar Woodruff, 2005: 31 Distribution: Dominican Republic

Phyllophaga acinosa (Arrow) Lachnosterna acinosa Arrow, 1920: 192 Phyllophaga acinosa (Arrow): Blackwelder, 1944: 224 Distribution: Trinidad

Phyllophaga acunai Chapin Phyllophaga acuñai Chapin, 1937: 3 Cnemarachis acuñai (Chapin): Blackwelder, 1944: 223 Phyllophaga acunai (Chapin); Evans, 2003: 67 Distribution: Cuba

Phyllophaga adjuntas Saylor Phyllophaga (Phyllophaga) adjuntas Saylor, 1940e: 312 Cnemarachis adjuntas (Saylor); Blackwelder, 1944: 223 Phyllophaga adjuntas (Saylor); Evans, 2003: 67 Distribution: Puerto Rico

Phyllophaga aemula (Horn) Lachnosterna aemula Horn, 1887b: 271 Phyllophaga aemula (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga aenea (Moser) Lachnosterna aenea Moser, 1921c: 252 Phyllophaga aenea (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) aenea (Moser); Cano and Morón, 1998: 13 Distribution: Guatemala

Phyllophaga aeneotincta Chapin Phyllophaga aeneotincta Chapin, 1932: 201 Cnemarachis aeneotincta (Chapin); Blackwelder, 1944: 223 Phyllophaga aeneotincta (Chapin); Evans, 2003: 68 Distribution: Cuba

Phyllophaga aequalis (LeConte) Tostegoptera aequalis LeConte, 1853b: 440 Lachnosterna aequalis (LeConte); LeConte, 1856: 238 Phyllophaga aequalis (LeConte); Glasgow, 1916: 372 Distribution: United States

Synonym: Lachnosterna exorata Horn, 1887b: 278 Phyllophaga exorata (Horn): Evans, 2003: 68

Phyllophaga aequata (Bates) 47 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Chlaenobia aequata Bates, 1888: 168 Phyllophaga aequata (Bates); Blackwelder, 1944: 224 Phyllophaga (Chlaenobia) aequata (Bates); Morón, 1986: 209 Distribution: Costa Rica, México, Nicaragua

Phyllophaga aequatorialis (Moser) Lachnosterna aequatorialis Moser, 1921c: 257 Phyllophaga aequatorialis (Moser); Blackwelder, 1944: 224 Distribution: Ecuador

Phyllophaga aeruginosa (Burmeister) Ancylonycha aeruginosa Burmeister, 1855: 338 Phyllophaga aeruginosa (Burmeister); Chapin, 1932b: 204 Cnemarachis aeruginosa (Burmeister); Blackwelder, 1944: 223 Phyllophaga (Cnemarachis) aeruginosa (Burmeister); Woodruff, 1961: 12 Distribution: Cuba

Phyllophaga affabilis (Horn) Lachnosterna affabilis Horn, 1887b: 283 Phyllophaga affabilis (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga afflicta (Blanchard) Ancylonycha afflicta Blanchard, 1851: 134 Phyllophaga afflicta (Blanchard); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) afflicta (Blanchard); Frey, 1975a: 213 Distribution: Brazil

Phyllophaga aguadita Saylor Phyllophaga (Phyllophaga) aguadita Saylor, 1942b: 156 Distribution: Colombia

Phyllophaga ahlbrandti Garcia-Vidal Phyllophaga ahlbrandti Garcia-Vidal, 1988: 104 Distribution: Cuba

Phyllophaga alayoi Garcia-Vidal Phyllophaga (Cnemarachis) alayoi Garcia-Vidal, 1978: 3 Distribution: Cuba

Phyllophaga albina (Burmeister) Ancylonycha albina Burmeister, 1855: 328 Lachnosterna albina (Burmeister); LeConte, 1856: 258 Phyllophaga albina (Burmeister); Glasgow, 1916: 372 Distribution: United States

Phyllophaga alcoa Woodruff Phyllophaga alcoa Woodruff, 2005: 33 48 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Dominican Republic

Phyllophaga aliada Sanderson Phyllophaga aliada Sanderson, 1951: 279 Distriubution: Dominican Republic, Haiti

Phyllophaga almada Saylor Phyllophaga (Listrochelus) almada Saylor, 1941a: 25 Distribution: México.

Phyllophaga alquizara Chapin Phyllophaga alquizara Chapin, 1932b: 204 Cnemarachis alquizara (Chapin): Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga amazonica (Moser) Lachnosterna amazonica Moser, 1918a: 57 Phyllophaga amazonica (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) amazonica (Moser); Frey, 1975a: 219 Distribution: Brazil

Phyllophaga ambigenus (Bates) Phytalus ambigenus Bates, 1888: 176 Phyllophaga ambigenus (Bates): Blackwelder, 1944: 224 Phyllophaga (Phytalus) ambigenus (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga amplicornis Reinhard Phyllophaga tristis amplicornis Reinhard, 1939: 59 Phyllophaga amplicornis Reinhard; Riley and Wolfe, 2003: 21 Distribution: United States

Phyllophaga analis (Burmeister) Ancylonycha analis Burmeister, 1855: 340 Phyllophaga analis (Burmeister); Chapin, 1932b; 197 Cnemarachis analis (Burmeister): Blackwelder, 1944: 223 Distribution: Cuba

Synonym: Ancylonycha bifoveolata Jacquelin du Val, 1856: 133 Phyllophaga bifoveolata (Jacquelin du Val); Evans, 2003: 67

Synonym: Ancylonycha subsericans Jacquelin du Val, 1856: 132 Phyllophaga subsericans (Jacquelin du Val); Evans, 2003: 67

Phyllophaga androw Woodruff Phyllophaga androw Woodruff, 2005: 39 Distribution: Dominican Republic

49 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga angulicollis (Bates) Lachnosterna angulicollis Bates, 1888: 203 Phyllophaga angulicollis (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) angulicollis (Bates): Morón, 1986: 204 Distribution: México

Phyllophaga angusta (Blanchard) Ancylonycha angusta Blanchard, 1851: 138 Phyllophaga angusta (Blanchard); Chapin, 1932b: 205 Cnemarachis angusta (Blanchard); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga anodentata (Bates) Lachnosterna anodentata Bates, 1888: 208 Phyllophaga (Phyllophaga) anodentata (Bates); Morón, 1986: 250 Distribution: México

Phyllophaga anolaminata (Moser) Lachnosterna anolaminata Moser, 1921c: 254 Phyllophaga anolaminata (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) anolaminata (Moser); Cano and Morón, 1998: 15 Distribution: Guatemala, Honduras, México

Phyllophaga anomaloides (Bates) Lachnosterna anomaloides Bates, 1888: 192 Phyllophaga anomaloides (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) anomaloides (Bates); Morón, 1986: 202 Distribution: México

Phyllophaga antennalis (Moser) Lachnosterna antennalis Moser, 1921c: 255 Phyllophaga antennalis (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) antennalis (Moser); Morón, 1986: 205 Distribution: México

Phyllophaga antennata (Smith) Lachnosterna antennata Smith, 1889a: 99 Phyllophaga antennata (Smith); Glasgow, 1916: 374 Distribution: United States

Phyllophaga antiguae (Arrow) Lachnostera antiguae Arrow, 1920: 191 Phyllophaga antigua [sic] (Arrow); Blackwelder, 1944: 223 [error] Phyllophaga antiguae (Arrow); Chalumeau, 1980: 87 Distribution: Antigua

Phyllophaga anxia (LeConte) Lachnosterna anxia LeConte, 1850: 226 50 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga anxia (LeConte); Glasgow, 1916: 371 Distribution: Canada, United States

Synonym: Lachnosterna alpina Linell, 1896: 726 Phyllophaga alpina (Linell); Evans, 2003: 71

Synonym: Ancylonycha brevicollis Blanchard, 1851: 132 Phyllophaga brevicollis (Blanchard); Evans, 2003: 71

Synonym: Lachnosterna cephalica LeConte, 1856: 245 Phyllophaga cephalica (LeConte); Evans, 2003; 71

Synonym: Lachnosterna dubia Smith, 1888: 183 Phyllophaga dubia (Smith); Evans, 2003: 71

Synonym: Ancylonycha guadulpensis Blanchard, 1851: 132 Phyllophaga guadulpiensis [sic] (Blanchard); Paulian, 1947: 53

Synonym: Lachnosterna insperata Smith, 1889a: 93 Phyllophaga insperata (Smith); Evans, 2003: 72

Synonym: Ancylonycha puncticollis Blanchard, 1851: 133 Phyllophaga puncticollis (Blanchard); Evans, 2003: 72

Synonym: Ancylonycha uninotata Walker, 1866: 323 Phyllophaga uninotata (Walker); Evans, 2003: 72

Phyllophaga apicalis (Blanchard) Phytalus apicalis Blanchard, 1851: 131 Clemora apicalis (Blanchard); Saylor, 1942a: 160 Phyllophaga apicalis (Blanchard); Chalumeau, 1985: 31 Distribution: Puerto Rico, US Virgin Islands

Synonym: Phyllophaga insularis Smyth, 1917: 163

Phyllophaga apicata Reinhard Phyllophaga tristis apicata Reinhard, 1939: 58 Phyllophaga apicata Reinhard; Riley and Wolfe, 2003: 9 Distribution: United States

Phyllophaga apolinari Saylor Phyllophaga apolinari Saylor, 1940d: 110 Distribution: Colombia

Phyllophaga apolinaria Saylor Phyllophaga apolinaria Saylor, 1942b: 154 Distribution: Colombia

51 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga approxima Woodruff and Sanderson Phyllophaga approxima Woodruff and Sanderson, 2005: 42 Distribution: Dominican Republic

Phyllophaga arcta (Horn) Lachnosterna arcta Horn, 1887b: 271 Phyllophaga arcta (Horn); Glasgow, 1916: 373 Distribution: United States

Synonym: Phyllophaga opacita Reinhard, 1939: 56

Phyllophaga ardara Saylor Phyllophaga (Phyllophaga) ardara Saylor, 1943e: 268 Distribution: México

Phyllophaga arenicola Howden Phyllophaga (Listrochelus) arenicola Howden, 1960: 457 Distribution: México, United States

Phyllophaga arizona Saylor Phyllophaga (Listrochelus) arizona Saylor, 1940c: 112 Distribution: United States

Phyllophaga arkansana (Schaeffer) Lachnosterna arkansana Schaeffer, 1906: 257 Phyllophaga arkansana (Schaeffer); Glasgow, 1916: 374

Synonym: Lachnosterna lenta Fall, 1908: 162 Phyllophaga lenta (Fall); Evans, 2003: 73

Phyllophaga arribans Saylor Phyllophaga (Phyllophaga) arribans Saylor, 1943e: 262 Distribution: México

Phyllophaga arrowi (Saylor) Lachnosterna (Phytalus) arrowi Saylor, 1935f: 206 Phyllophaga arrowi (Saylor); Blackwelder, 1944: 224 Phyllophaga (Phytalus) arrowi (Saylor); Frey, 1975c: 225 Distribution: Brazil

Phyllophaga assmani Saylor Phyllophaga assmani Saylor, 1936d: 294 Phyllophaga (Phyllophaga) assmani Saylor; Solís and Morón, 1998: 26 Distribution: Costa Rica

Phyllophaga atra (Moser) Lachnosterna atra Moser, 1918b: 166 Phyllophaga atra (Moser); Blackwelder, 1944: 224 52 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phyllophaga) atra (Moser); Morón, 1986, 204 Distribution: México

Phyllophaga atrata (Moser) Lachnosterna atrata Moser, 1918a: 34 Phyllophaga atrata (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) atrata (Moser); Morón, 1986, 205 Distribution: México

Phyllophaga atratoides Morón Phyllophaga (Phyllophaga) atratoides Morón, 2003a: 247 Distribution: México

Phyllophaga audanti Wolcott Phyllophaga audanti Wolcott, 1928: 27 Distribution: Dominican Republic, Haiti

Phyllophaga aurea Luginbill and Painter Phyllophaga aurea Luginbill and Painter, 1941: 716 Distribution: United States

Phyllophaga austera (Erichson) Ancyclonycha austera Erichson, 1847a: 101 Lachnosterna (Ancylonycha) austera (Erichson); Kirsch, 1874a: 342 Lachnosterna austera (Erichson); Harold, 1869a: 1166 Phyllophaga austera (Erichson): Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) austera (Erichson); Frey, 1975a: 208 Distribution: Peru

Phyllophaga babicora Morón Phyllophaga (Listrochelus) babicora Morón, 2004: 86 Distribution: México

Phyllophaga babis Saylor Phyllophaga (Phyllophaga) babis Saylor, 1943e: 269 Distribution: México

Phyllophaga bahiana Saylor Lachnosterna (Phytalus) bahiana Saylor, 1935f: 207 Phyllophaga (Phytalus) bahiana (Saylor); Frey, 1975a: 276 Distribution: Brazil

Synonym: Phyllophaga (Phytalus) glabratus Frey, 1964c: 694

Phyllophaga balia (Say) Melolontha balia Say, 1825: 194 Lachnosterna balia (Say); LeConte, 1856: 255 Phyllophaga balia (Say); Glasgow, 1916: 371 53 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Canada, United States

Synonym: Ancylonycha comata Burmeister, 1855: 337 Phyllophaga comata (Burmeister); Evans, 2003: 74

Phyllophaga baneta Saylor Phyllophaga (Phyllophaga) baneta Saylor, 1943d: 133 Distribution: Guatemala, Honduras

Phyllophaga baoruco Woodruff Phyllophaga baoruco Woodruff, 2005: 45 Distribution: Dominican Republic

Phyllophaga baracoana Chapin Phyllophaga baracoana Chapin, 1932b: 188 Cnemarachis baracoana (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga baraguensis Chapin Phyllophaga baraguensis Chapin, 1932b: 185 Cnemarachis baraguensis (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga barda (Horn) Lachnosterna barda Horn, 1887b: 248 Phyllophaga barda (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga baroni (Bates) Lachnosterna baroni Bates, 1889: 401 Phyllophaga baroni (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) baroni (Bates); Morón, 1986: 194 Distribution: México

Phyllophaga barrerana Aragón and Morón Phyllophaga (Listrochelus) barrerana Aragón and Morón, 2003: 563 Distribution: México

Phyllophaga barrosa Sanderson Phyllophaga barrosa Sanderson, 1951: 280 Distribution: Haiti

Phyllophaga batillifer (Bates) Phytalus batillifer Bates, 1888: 180 Phyllophaga batillifer (Bates); Blackwelder, 1944: 224 Phyllophaga (Phytalus) batillifer (Bates); Morón, 1986: 200 Distribution: México

54 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga beameri Sanderson Phyllophaga (Phyllophaga) beameri Sanderson, 1958: 171 Phyllophaga (Phytalus) beameri Sanderson; Morón, 1986: 200 Distribution: México

Phyllophaga beckeri (Moser) Lachnosterna beckeri Moser, 1921c: 248 Phyllophaga beckeri (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) beckeri (Moser); Sanderson, 1958: 166 Distribution: México, United States

Synonym: Phyllophaga inopia Sanderson, 1942: 51

Phyllophaga bellamyi Warner and Morón Phyllophaga (Triodonyx) bellamyi Warner and Morón, 1992: 331 Distribution: United States

Phyllophaga benexonana Morón Phyllophaga (Phytalus) benexonana Morón, 2000a: 59 Distribution: México

Phyllophaga bicavifrons Chapin Phyllophaga bicavifrons Chapin, 1932b: 187 Phyllophaga (Cnemarachis) bicavifrons Chapin; Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga bicolor (Moser) Lachnosterna bicolor Moser, 1921c: 258 Phyllophaga bicolor (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) bicolor (Moser); Frey, 1975a: 212 Distribution: Ecuador

Phyllophaga bilobatata Saylor Phyllophaga (Phytalus) bilobatata Saylor, 1939c: 165 (for Horn, 1885: 120) Phyllophaga (Phyllophaga) bilobatata Saylor; Sanderson, 1958: 166 Distribution: México, United States

Synonym: Phytalus cephalicus Horn, 1885: 120 (not LeConte, 1856: 245)

Phyllophaga bilyi Chalumeau Phyllophaga bilyi Chalumeau, 1990: 338 Distribution: Cuba

Phyllophaga bimaculata Garcia-Vidal Phyllophaga (Cnemarachis) bimaculata Garcia-Vidal, 1984: 313 Distribution: Cuba

Phyllophaga bimammifrons Saylor 55 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phyllophaga) bimammifrons Saylor, 1940e: 305 Cnemarachis bimammifrons (Saylor); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga bipartita (Horn) Lachnosterna bipartita Horn, 1887b: 242 Phyllophaga bipartita (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga blackwelderi Saylor Phyllophaga blackwelderi Saylor, 1940e: 309 Cnemarachis blackwelderi (Saylor); Blackwelder, 1944: 223 Distribution: St. Lucia

Phyllophaga blanchardi (Arrow) Lachnosterna blanchardi Arrow, 1933b: 149 (for Blanchard, 1851: 133) Phyllophaga (Phyllophaga) blanchardi (Arrow); Sanderson, 1958: 166 Distribution: México

Synonym: Ancylonycha cribricollis Blanchard, 1851: 133 (not Redtenbacher, 1842)

Phyllophaga blanda Sanderson Phyllophaga (Phyllophaga) blanda Sanderson, 1958: 169 Distribution: México, United States

Phyllophaga bolacoides (Bates) Phytalus bolacoides Bates, 1888: 182 Phyllophaga bolacoides (Bates); Blackwelder, 1944; 224 Phyllophaga (Phytalus) bolacoides (Bates); Morón, 1986: 213 Distribution: México

Phyllophaga boliviensis (Blanchard) Phytalus boliviensis Blanchard, 1851: 131 Phyllophaga boliviensis (Blanchard); Blackwelder, 1944: 224 Phyllophaga (Phytalus) boliviensis (Blanchard); Frey, 1975a: 225 Distribution: Bolivia, Paraguay

Synonym: Phyllophaga schultzei Saylor, 1935h: 496

Phyllophaga bonfils Woodruff and Sanderson Phyllophaga bonfils Woodruff and Sanderson, 2005: 52 Distribution: Haiti

Phyllophaga boruca Morón Phyllophaga (Phyllophaga) boruca Morón, 2003a: 223 Distribution: Costa Rica

Phyllophaga bottimeri Reinhard 56 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Listrochelus) bottimeri Reinhard, 1950: 29 Distribution: United States

Phyllophaga bowditchi Saylor Phyllophaga (Phyllophaga) bowditchi Saylor, 1938e: 189 Distribution: Belize

Phyllophaga brama Saylor Phyllophaga (Phyllophaga) brama Saylor, 1943e: 267 Distribution: México

Phyllophaga brevicornis Garcia-Vidal Phyllophaga brevicornis Garcia-Vidal, 1988: 101 Distribution: Cuba

Phyllophaga brevidens (Bates) Lachnosterna brevidens Bates, 1888: 210 Phyllophaga brevidens (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) brevidens (Bates); Morón, 1986: 204 Distribution: México

Phyllophaga brevipilosa (Moser) Lachnosterna brevipilosa Moser, 1918a: 46 Phyllophaga brevipilosa (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) brevipilosa (Moser); Frey, 1975a: 206 Distribution: Venezuela

Phyllophaga brevisetosa (Moser) Lachnosterna brevisetosa Moser, 1918a: 38 Phyllophaga brevisetosa (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) brevisetosa (Moser); Frey, 1975a: 211 Distribution: Colombia

Phyllophaga brownella Saylor Phyllophaga (Phyllophaga) brownella Saylor, 1942a: 160 Distribution: Panama

Phyllophaga bruneri Chapin Phyllophaga bruneri Chapin, 1932b: 203 Cnemarachis bruneri (Chapin); Blackwelder, 1944: 223 Phyllophaga (Cnemarachis) bruneri Chapin; Woodruff, 1961: 12 Distribution: Cuba, United States

Phyllophaga bucephala (Bates) Phytalus bucephalus Bates, 1888: 182 Phyllophaga bucephala (Bates); Blackwelder, 1944: 224 Phyllophaga (Phytalus) bucephala (Bates); Morón, 1986: 200 Distribution: Guatemala, México 57 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga bueta Saylor Phyllophaga (Listrochelus) bueta Saylor, 1940d: 115 Distribution: México

Phyllophaga caanchaki Morón Phyllophaga caanchaki Morón, 1998: 43 Phyllophaga (Phyllophaga) caanchaki Morón; Cano and Morón, 1998: 16 Distribution: Guatemala, México

Phyllophaga cahitana Morón Phyllophaga (Phyllophaga) cahitana Morón, 2001a: 169 Distribution: México

Phyllophaga calceata (LeConte) Lachnosterna calceata LeConte, 1856: 250 Phyllophaga calceata (LeConte); Glasgow, 1916: 372 Distribution: United States

Phyllophaga calculiventris Saylor Phyllophaga calculiventris Saylor, 1935g: 34 Phyllophaga (Phyllophaga) calculiventris Saylor; Solís and Morón, 1998: 25 Distribution: Costa Rica, México, Panama

Phyllophaga calderasa (Saylor) Lachnosterna (Phytalus) calderasa Saylor, 1937c: 31 Phyllophaga (Phytalus) calderasa Saylor; Cano and Morón, 1998: 10 Distribution: Guatemala, México

Phyllophaga callosiventris (Moser) Lachnosterna callosiventris Moser, 1921c: 249 Phyllophaga callosiventris (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) callosiventris (Moser); Morón, 1986: 203 Distribution: México

Phyllophaga cambeforti Cartwright and Chalumeau Phyllophaga cambeforti Cartwright and Chalumeau, 1977: 92 Phyllophaga (Cnemarachis) cambeforti Cartwright and Chalumeau, 1978: 21 Distribution: Dominica

Phyllophaga candelaria Morón Phyllophaga (Phyllophaga) candelaria Morón, 2001b: 778 Distribution: México

Phyllophaga caneyensis Garcia-Vidal Phyllophaga caneyensis Garcia-Vidal, 1984: 315 Distribution: Cuba

58 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga canipolea Saylor Phyllophaga (Phyllophaga) canipolea Saylor, 1948a: 349 Distribution: México

Phyllophaga canoa Sanderson Phyllophaga (Cnemarachis) canoa Sanderson, 1951: 279 Distribution: Dominican Republic

Phyllophaga canoana Morón Phyllophaga (Phyllophaga) canoana Morón, 2003a: 289 Distribution: Guatemala

Phyllophaga capillata (Blanchard) Ancylonycha capillata Blanchard, 1851: 135 Phyllophaga capillata (Blanchard); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) capillata (Blanchard); Frey, 1975a; 206 Distribution: Bolivia, Brazil

Phyllophaga caraga Saylor Phyllophaga (Phyllophaga) caraga Saylor, 1943d: 134 Distribution: Costa Rica, Nicaragua, Panama

Phyllophaga cardini Chapin Phyllophaga cardini Chapin, 1932b: 197 Cnemarachis cardini (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga carminator (Horn) Listrochelus carminator Horn, 1894: 398 Phyllophaga (Listrochelus) carminator (Horn); Saylor, 1940c: 75 Distribution: México

Phyllophaga carnegie Woodruff Phyllophaga carnegie Woodruff, 2005: 57 Distribution: Dominican Republic

Phyllophaga cartaba Sanderson Phyllophaga (Cnemarachis) cartaba Sanderson, 1951: 274 Distribution: Dominican Republic

Phyllophaga cartiginesa Morón Phyllophaga (Phyllophaga) cartiginesa Morón, 2001b: 795 Distribution: Costa Rica

Phyllophaga castaniella (Bates) Lachnosterna castaniella Bates, 1888: 186 Phyllophaga (Phyllophaga) castaniella (Bates): Blackwelder. 1994: 224 Phyllophaga (Phyllophaga) castaniella (Bates): Morón, 1986: 194 59 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Phyllophaga castineirasi Garcia-Vidal Phyllophaga (Cnemarachis) castineirasi Garcia-Vidal, 1978: 5 Distribution: Cuba

Phyllophaga catemacoana Morón Phyllophaga (Phyllophaga) catemacoana Morón, 2003a: 293 Distribution: México

Phyllophaga cavata cavata (Bates) Listrochelus cavatus Bates, 1888: 170 Phyllophaga (Listrochelus) cavata (Bates); Saylor, 1940c: 103 Distribution: México, United States

Synonym: Listrochelus cavatus var. durangoensis Bates, 1888: 170 Phyllophaga cavata durangoensis (Bates); Evans, 2003: 81

Phyllophaga cavata michoacana Saylor Phyllophaga (Listrochelus) cavatus michoacana Saylor, 1943a: 24 Distribution: México

Phyllophaga caviceps (Moser) Lachnosterna caviceps Moser, 1918a: 53 Phyllophaga caviceps (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) caviceps (Moser); Frey, 1975a: 213 Distribution: Colombia

Phyllophaga cayennensis (Moser) Lachnosterna cayennensis Moser, 1918a: 54 Phyllophaga cayennensis (Moser); Blackwelder, 1944: 224 Distribution: French Guiana

Phyllophaga caymanensis Sanderson Phyllophaga caymanensis Sanderson, 1939b: 274 Cnemarachis caymanensis (Sanderson); Blackwelder, 1944: 224 Distribution: Grand Cayman

Phyllophaga cazierana Saylor Phyllophaga cazierana Saylor, 1938f: 346 (for Saylor, 1936b: 63) Phyllophaga (Phyllophaga) cazierana Saylor; Morón, 1986: 225 Distribution: México

Synonym: Phyllophaga cazieri Saylor, 1936b: 63 (not Bloeker, 1936: 57)

Phyllophaga centralis (Nonfried) Lachnosterna centralis Nonfried, 1894: 119 Phyllophaga centralis (Nonfried); Blackwelder, 1944: 224 60 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Honduras

Phyllophaga certanca Saylor Phyllophaga (Phyllophaga) certanca Saylor, 1943a: 25 Distribution: México

Phyllophaga chada Saylor Phyllophaga (Phyllophaga) chada Saylor, 1948b: 355 Distribution: Brazil

Phyllophaga chalumeaui Garcia-Vidal Phyllophaga (Cnemarachis) chalameaui Garcia-Vidal, 1984: 317 Distribution: Cuba

Phyllophaga chamacayoca Morón Phyllophaga (Phyllophaga) chamacayoca Morón, 1992a: 126 Distribution: México

Phyllophaga chamula Morón Phyllophaga (Phyllophaga) chamula Morón, 1999: 384 Distriubution: México

Phyllophaga changuena Morón Phyllophaga (Phyllophaga) changuena Morón, 2003a: 290 Distribution: Costa Rica, Nicaragua, Panama

Phyllophaga chapini Saylor Phyllophaga (Listrochelus) chapini Saylor, 1940c: 84 Distribution: United States

Phyllophaga chiapas Saylor Phyllophaga (Phyllophaga) chiapas Saylor, 1943e: 276 Distribution: México

Phyllophaga chiapensis (Chapin) Chlaenobia aequata chiapensis Chapin, 1935a: 18 Phyllophaga aequata chiapensis (Chapin); Blackwelder, 1944: 224 Phyllophaga (Chlaenobia) chiapensis (Chapin); Morón, 2003b: 5 Distribution: Costa Rica, El Salvador, Guatemala, Honduras, México

Phyllophaga chimoxtila Morón Phyllophaga (Phyllophaga) chimoxtila Morón, 2003a: 257 Distribution: México

Phyllophaga chiblacana Morón Phyllophaga (Phyllophaga) chiblacana Morón, 2003a: 295 Distribution: Guatemala

61 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga chinanteca Morón and Nogueira Phyllophaga (Phyllophaga) chinanteca Morón and Nogueira, 1997: 258 Distribution: México

Phyllophaga chippewa Saylor Phyllophaga chippewa Saylor, 1939d: 455 Distribution: United States

Phyllophaga chiriquina (Bates) Lachnosterna chiriquina Bates, 1888: 196 Phyllophaga (Phyllophaga) chiriquina (Bates); Saylor, 1943d: 137 Distribution: Colombia, Costa Rica, México, Nicaragua, Panama

Phyllophaga chlaenobiana Saylor Phyllophaga (Phytalus) chlaenobiana Saylor, 1936b: 62 Distribution: México

Phyllophaga cholana Morón Phyllophaga (Phyllophaga) cholana Morón, 2003a: 287 Distribution: México

Phyllophaga chorotega Morón Phyllophaga (Phyllophaga) chorotega Morón, 2001b: 785 Distribution: Costa Rica

Phyllophaga chortiana Morón Phyllophaga (Phyllophaga) chortiana Morón, 2003a: 253 Distribution: Guatemala

Phyllophaga ciliatipes (Blanchard) Chlaenobia ciliatipes Blanchard, 1850: 116 Phyllophaga ciliatipes (Blanchard); Blackwelder, 1944: 224 Phyllophaga (Chlaenobia) ciliatipes (Blanchard); Morón, 1986: 206 Distribution: Guatemala, México

Synonym: Chlaenobia aegrota Bates, 1888: 167 Phyllophaga aegrota (Bates); Blackwelder, 1944: 224 Phyllophaga (Chlaenobia) aegrota (Bates); Morón, 1986; 206

Phyllophaga cinnamomea (Blanchard) Ancylonycha cinnamomea Blanchard, 1851: 134 Lachnosterna cinnamomea (Blanchard); Bates, 1888: 197 Phyllophaga (Phyllophaga) cinnamomea (Blanchard); Saylor, 1943d: 138 Distribution: Guatemala, México

Phyllophaga ciudadensis (Bates) Phytalus ciudadensis Bates, 1888: 183 Phyllophaga ciudadensis (Bates); Blackwelder, 1944: 224 62 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phytalus) ciudadensis (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga clemens (Horn) Lachnosterna clemens Horn, 1887b: 227 Phyllophaga clemens (Horn), Glasgow, 1916: 373 Distribution: United States

Synonym: Phyllophaga howei Sanderson , 1937a: 17

Phyllophaga clypeata (Horn) Lachnosterna clypeata Horn, 1887a: 145 (for LeConte, 1856: 258) Phyllophaga clypeata (Horn); Glasgow, 1916: 373 Distribution: United States

Synonym: Lachnosterna integra LeConte, 1856: 258 (not Say, 1835: 180) Phyllophaga integra (LeConte); Evans, 2003: 83

Phyllophaga cneda Saylor Phyllophaga (Phyllophaga) cneda Saylor, 1940e: 310 Distribution: Montserrat, St. Kitts-Nevis

Phyllophaga cochisa Saylor Phyllophaga (Listrochelus) cochisa Saylor, 1940d: 105 Distribution: México, United States

Phyllophaga colima Saylor Phyllophaga (Phyllophaga) colima Saylor, 1943e: 273 Distribution: México

Phyllophaga colimana (Moser) Lachnosterna colimana Moser, 1921c: 256 Phyllophaga colimana (Moser); Blackwelder, 1944: 224 Distribution: México

Phyllophaga collaris (Moser) Lachnosterna collaris Moser, 1921c: 247 Phyllophaga collaris (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) collaris (Moser); Morón, 1986: 202 Distribution: México

Synonym: Phyllophaga (Phyllophaga) gestis Saylor, 1943e: 267

Phyllophaga columbiana (Blanchard) Ancylonycha columbiana Blanchard, 1851: 134 Phyllophaga columbiana (Blanchard); Blackwelder, 1944: 224 Distribution: Colombia

63 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga comaltepecana Morón Phyllophaga (Phyllophaga) comaltepecana Morón, 2003a: 277 Distribution: México

Phyllophaga cometes (Bates) Phytalus cometes Bates, 1888: 177 Phyllophaga cometes (Bates); Blackwelder, 1944: 224 Phyllophaga (Phytalus) cometes (Bates); Morón, 1986: 200 Distribution: Costa Rica, Guatemala, México, Nicaragua, Panama

Phyllophaga complexipennis Garcia-Vidal Phyllophaga complexipennis Garcia-Vidal, 1984: 317 Distribution: Cuba

Phyllophaga compostela Robinson Phyllophaga compostela Robinson, 1948: 34 Distribution: México

Phyllophaga conformis (Blanchard) Ancylonycha conformis Blanchard, 1851: 135 Phyllophaga conformis (Blanchard); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) conformis (Blanchard); Frey, 1975a: 207 Distribution: Argentina

Phyllophaga congrua (LeConte) Lachnosterna congrua LeConte, 1856: 243 Phyllophaga congrua (LeConte); Glasgow, 1916: 372 Distribution: United States

Phyllophaga contaminata Fall Phyllophaga contaminata Fall, 1932: 197 Phyllophaga (Phyllophaga) contaminata Fall; Saylor, 1948a: 349 Distribution: México

Synonym: Phyllophaga extranea Fall, 1932: 198

Phyllophaga coronadis Saylor Phyllophaga (Phyllophaga) coronadis Saylor, 1941e: 538 Distribution: Costa Rica

Phyllophaga corrosa (LeConte) Lachnosterna corrosa LeConte, 1856: 249 Phyllophaga corrosa (LeConte); Glasgow, 1916: 372 Distribution: United States

Synonym: Lachnosterna affinis LeConte, 1856: 252 Phyllophaga affinis (LeConte); Evans, 2003: 85

64 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga costaricensis (Moser) Lachnosterna costaricensis Moser, 1918a: 25 Phyllophaga costaricensis (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) costaricensis (Moser); Solís and Morón, 1998: 23 Distribution: Costa Rica

Phyllophaga costura Sanderson Phyllophaga costura Sanderson, 1951: 264 Distribution: Dominican Republic

Phyllophaga crassa (Burmeister) Phytalus crassus Burmeister, 1855: 347 Phyllophaga crassa (Burmeister); Evans, 2003: 85 Distribution: Brazil

Phyllophaga crassissima (Blanchard) Ancylonycha crassissima Blanchard, 1851: 133 Lachnosterna crassissima (Blanchard); Horn, 1887b: 238 Lachnosterna crasissima [sic] (Blanchard); Smith, 1888: 499 Phyllophaga crassissima (Blanchard); Glasgow, 1916: 372 Distribution: United States

Synonym: Lachnosterna generosa Horn, 1887b: 222

Synonym: Lachnnosterna obesa LeConte, 1856: 251

Synonym: Lachnnosterna robusta LeConte, 1856: 257

Phyllophaga crena Saylor Phyllophaga (Phyllophaga) crena Saylor, 1941e: 538 Distribution: Costa Rica

Phyllophaga crenaticollis (Blanchard) Ancylonycha crenaticollis Blanchard, 1851: 137 Ancylonycha crenatocollis [sic] Blanchard; Chevrolat, 1865: 23 [error] Phyllophaga crenaticollis (Blanchard); Chapin, 1932b: 183 Cnemarachis crenaticollis (Blanchard); Saylor, 1942a: plate 17 Distribution: Cuba

Phyllophaga crenonycha Saylor Phyllophaga (Phyllophaga) crenonycha Saylor, 1943c: 25 Distribution: México

Phyllophaga crenulata (Froelich) Melolontha crenulata Froelich, 1792: 94 Ancylonycha crenulata (Froelich); Blanchard, 1851: 133 Lachnosterna crenulata (Froelich); LeConte, 1856: 258 Phyllophaga crenulata (Froelich); Glasgow, 1916: 370 65 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Canada, United States

Synonym: Melolontha georgicana Gyllenhal, 1817: 77 Phyllophaga georgicana (Gyllenhal); Harris, 1841: 29

Phyllophaga cribrosa (LeConte) Tostegoptera cribrosa LeConte, 1853a: 231 Eugastra cribrosa (LeConte); LeConte, 1856: 234 Lachnosterna cribrosa (LeConte); Horn, 1887b: 217 Phyllophaga cribrosa (LeConte); Glasgow, 1916: 372 Phyllophaga (Phyllophaga) cribrosa (LeConte); Sanderson, 1958: 164 Phyllophaga (Eugastra) cribrosa (LeConte); Saylor, 1942a: 162 Distribution: México, United States

Synonym: Tostegoptera ventricosa LeConte, 1853b: 440 Eugastra ventricosa (LeConte); LeConte, 1856: 234 Phyllophaga cribrosa ventricosa (LeConte): Bloeker, 1937: 85

Phyllophaga crinalis (Bates) Lachnosterna crinalis Bates, 1888: 199 Phyllophaga crinalis (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) crinalis (Bates); Morón, 1986: 203 Distribution: México

Phyllophaga crinipennis (Bates) Lachnosterna crinipennis Bates, 1889: 402 Phyllophaga crinipennis (Bates): Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) crinipennis (Bates): Morón, 1986: 40 Distribution: México

Phyllophaga crinita (Burmeister) Trichestes crinita Burmeister, 1855: 359 Phyllophaga crinita (Burmeister); Glasgow, 1916: 372 Phyllophaga (Phyllophaga) crinita (Bates): Morón, 1986: 204 Distribution: México

Synonym: Lachnosterna glabripennis LeConte, 1856: 260

Synonym: Listrochelus longiclavus Fall, 1922: 173

Phyllophaga crinitissima More Phyllophaga crinitissima More, 1924: 105 Cnemarachis (Abrcrana) crinitissima (More); Saylor, 1942a: 159 Cnemarachis crinitissima (More); Blackwelder, 1944: 223 Abcrana crinitissima (More); Sanderson, 1950: 92 Distribution: Puerto Rico

Phyllophaga cristagalli (Arrow) 66 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Listrochelus crista-galli Arrow, 1933b: 146 Phyllophaga (Listrochelus) cristagalli (Arrow); Morón, 1986: 201 Distribution: México

Phyllophaga cubana Chapin Phyllophaga cubana Chapin, 1932b: 194 Cnemarachis cubana (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga cuicateca Morón and Aragón Phyllophaga (Listrochelus) cuicateca Morón and Aragón, 1997: 40 Distribution: México

Phyllophaga culminata (Bates) Listrochelus culminatus Bates, 1888: 172 Phyllophaga culminata (Bates); Blackwelder, 1944: 224 Phyllophaga (Listrochelus) culminata (Bates), Morón, 1986: 202 Distribution: México

Phyllophaga cupuliformis Langston Phyllophaga micans var. cupuliformis Langston, 1924: 450 Phyllophaga cupuliformis Langston; Woodruff, 1973: 28 Distribution: United States

Phyllophaga curialis Reinhard Phyllophaga curialis Reinhard, 1939: 54 Distribution: United States

Phyllophaga cushmani Saylor Phyllophaga (Listrochelus) cushmani Saylor, 1940d: 109 Distribution: México, United States

Phyllophaga cuyabana (Moser) Lachnosterna cuyabana Moser, 1918a: 52 Phyllophaga cuyabana (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) cuyabana (Moser); Frey, 1975a: 207 Distribution: Brazil

Phyllophaga cylindrica (Burmeister) Trichestes cylindrica Burmeister, 1855: 361 Lachnosterna cylindrica (Burmeister); Bates, 1888: 214 Cnemarachis cylindrica (Burmeister); Blackwelder, 1944: 223 Phyllophaga cylindrica (Burmeister); Evans, 2003: 88 Distribution: Cuba, United States

Synonym: Phyllophaga emusa Saylor, 1937d: 321 (unjustified emendation for Burmeister, 1855: 361)

Phyllophaga darlingtoni Sanderson 67 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga darlingtoni Sanderson, 1939: 14 Cnemarachis darlingtoni Sanderson; Blackwelder, 1944: 223 Distribution: Jamaica

Phyllophaga dasypoda (Bates) Lachnosterna dasypoda Bates, 1888: 193 Phyllophaga dasypoda (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) dasypoda (Bates); Morón, 1986: 202 Distribution: Costa Rica, El Salvador, Guatemala, México

Phyllophaga davidsoni Woodruff Phyllophaga davidsoni Woodruff, 2005: 60 Distribution: Dominican Republic

Phyllophaga davisi Langston Phyllophaga davisi Langston, 1927a: 221 Distribution: United States

Phyllophaga debilis (LeConte) Gynnis debilis LeConte, 1856: 262 Lachnosterna debilis (LeConte); Horn, 1887a: 145 Phyllophaga debilis (LeConte); Glasgow, 1916: 373 Distribution: United States

Synonym: Phyllophaga austricola Fall, 1929b: 216

Phyllophaga delata (Horn) Lachnosterna delata Horn, 1887b: 267 Phyllophaga delata (Horn); Glasgow, 1916: 373. Distribution: United States

Phyllophaga delplanquei Chalumeau and Gruner Phyllophaga (Cnemarachis) delplanquei Chalumeau and Gruner, 1976: 88 Distribution: Martinique

Phyllophaga densata (Moser) Lachnosterna densata Moser, 1918a: 36 Phyllophaga densata (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) densata (Moser); Cano and Morón, 1998: 15 Distribution: Costa Rica, Guatemala

Phyllophaga densepunctata (Moser) Lachnosterna densepunctata Moser, 1918a: 50 Phyllophaga densepunctata (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) densepunctata (Moser); Morón, Vallejo and Restrepo-Giraldo, 2001:32 Distribution: Colombia

Phyllophaga densicollis (LeConte) 68 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Lachnosterna densicollis LeConte, 1863b: 77 Phyllophaga (Listrochelus) densicollis (LeConte); Saylor, 1940d: 74 Distribution: México

Phyllophaga dentex (Bates) Lachnosterna dentex Bates, 1888: 192 Phyllophaga (Lachnosterna) dentex (Bates); Saylor, 1935i: 66 Phyllophaga (Phyllophaga) dentex (Bates); Sanderson, 1958: 164 Distribution: México, United States

Phyllophaga denticulata (Blanchard) Ancylonycha denticulata Blanchard, 1851: 137 Lachnosterna denticulata (Blanchard); Dalla Torre, 1912c: 187 Cnemarachis denticulata (Blanchard); Blackwelder, 1944: 223 Phyllophaga denticulata (Blanchard); Evans, 2003: 90 Distribution: Martinique

Synonym: Phyllophaga (Lachnosterna) citri Smyth, 1917: 65

Synonym: Lachnosterna insulicola Moser, 1918a: 61 Cnemarachis insulicola (Moser); Blackwelder, 1944: 223

Phyllophaga dieteriana Deloya and Morón Phyllophaga (Phytalus) dieteriana Deloya and Morón, 1999: 110 Distribution: México

Phyllophaga diffinis (Blanchard) Ancyclonycha diffinis Blanchard, 1851: 138 Lachnosterna diffinis (Blanchard); Blatchley, 1910: 968 Phyllophaga diffinis (Blanchard); Glasgow, 1916: 372 Distribution: United States

Synonym: Trichestes comans Burmeister, 1855: 358 Lachnosterna comans (Burmeister); Horn, 1887b: 261

Synonym: Lachnosterna rufiola LeConte, 1856: 256

Synonym: Lachnosterna sororia LeConte, 1856: 246

Phyllophaga diminuta Evans Phyllophaga (Phytalus) diminuta Evans, 2003: 91 (for Saylor, 1943a: 22) Distribution: México

Synonym: Phyllophaga (Phytalus) minutissima Saylor, 1943a: 22 (not Wolcott, 1928: 28)

Phyllophaga disca Saylor Phyllophaga (Phyllophaga) disca Saylor, 1943e: 274 Distribution: México 69 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga discalis (Chapin) Phyllophaga discalis Chapin, 1935c: 70 Cnemarachis discalis (Chapin): Blackwelder, 1944: 223 Distribution: Puerto Rico

Phyllophaga dispar (Burmeister) Trichestes dispar Burmeister, 1855: 361 Lachnosterna dispar (Burmeister); LeConte, 1856: 240 Phyllophaga dispar (Burmeister); Glasgow, 1916: 372 Distribution: United States

Synonym: Lachnosterna boops Horn, 1887b: 284

Phyllophaga disparilis (Horn) Listrochelus disparilis Horn, 1878: 141 Phyllophaga (Listrochelus) disparilis, Saylor, 1940d: 81 Distribution: México, United States

Phyllophaga disrupta Cockerell Phyllophaga disrupta Cockerell, 1927: 587 Distribution: United States Remark: fossil species

Phyllophaga dissimilis (Chevrolat) Ancylonycha dissimilis Chevrolat, 1865: 26 Phyllophaga dissimilis (Chevrolat); Chapin, 1932b: 201 Cnemarachis dissimilis (Chevrolat); Saylor, 1942a: plate 17 Distribution: Cuba

Phyllophaga divertens (Bates) Lachnosterna divertens Bates, 1888: 206 Phyllophaga (Phyllophaga) divertens (Bates); Butler and Werner, 1961: 18 Distribution: México, United States

Synonym: Lachnosterna setipennis Bates 1888: 206 Phyllophaga setipennis (Bates): Blackwelder, 1944: 226

Phyllophaga dominicensis Cartwright and Chalumeau Phyllophaga dominicensis Cartwright and Chalumeau, 1977: 102 Phyllophaga (Cnemarachis) dominicensis Cartwright and Chalumeau, 1978: 19 Distribution: Dominica

Phyllophaga drakii (Kirby) Rhizotrogus drakii Kirby, 1837: 133 Phyllophaga drakii (Kirby); Glasgow, 1916: 371 Distribution: Canada, United States

70 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Lachnosterna consimilis LeConte, 1850: 226

Synonym: Lachnosterna grandis Smith, 1888: 181

Phyllophaga dsaimana Morón Phyllophaga (Phyllophaga) dsaimana Morón, 2003a: 281 Distribution: México

Phyllophaga dubitata Garcia-Vidal Phyllophaga (Cnemarachis) dubitatus Garcia-Vidal, 1978: 8 Distribution: Cuba

Phyllophaga duenas Saylor Phyllophaga (Phyllophaga) duenas Saylor, 1941d: 385 Distribution: Guatemala

Phyllophaga dugesiana Morón, Rivera, and López Phyllophaga (Phytalus) dugesiana Morón, Rivera, and López, 2002: 494 Distribution: México

Phyllophaga dulcis (Bates) Lachnosterna dulcis Bates, 1888: 189 Phyllophaga dulcis (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) dulcis (Bates); Morón, 1986: 203 Distribution: México

Phyllophaga duncani (Barrett) Listrochelus duncani Barrett, 1933: 129 Phyllophaga (Listrochelus) duncani (Barrett); Saylor, 1940d: 111 Distribution: United States

Phyllophaga durango Saylor Phyllophaga (Listrochelus) durango Saylor, 1940e: 114 Distribution: México

Phyllophaga durangoana (Moser) Lachnosterna durangoanus Moser, 1921c: 261 Phyllophaga durangoana (Moser): Blackwelder, 1944: 224 Distribution: México

Phyllophaga durangosa Saylor Phyllophaga (Phyllophaga) durangosa Saylor, 1943e: 275 Distribution: México

Phyllophaga ecostata (Horn) Lachnosterna ecostata Horn, 1887b: 284 Triodonyx ecostata (Horn); Saylor, 1942a: 158 Phyllophaga ecostata (Horn); Glasgow, 1916: 373 71 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: United States

Phyllophaga ecuadorae (Blackwelder) Lachnosterna ecuadorae Blackwelder, 1944: 224 (for Moser, 1921c: 258) Phyllophaga ecuadorae (Blackwelder); Evans, 2003: 93 Distribution: Ecuador

Synonym: Lachnosterna densicollis Moser, 1921c: 258 (not LeConte, 1863b: 77) Phyllophaga (Phyllophaga) densicollis (Moser); Frey, 1975a: 203

Phyllophaga eladio Woodruff Phyllophaga eladio Woodruff, 2005: 65 Distribution: Dominican Republic

Phyllophaga elegans (Nonfried) Lachnosterna elegans Nonfried, 1891: 264 Phyllophaga elegans (Nonfried): Blackwelder, 1944: 224 Distribution: Honduras

Phyllophaga elenans Saylor Phyllophaga (Phyllophaga) elenans Saylor, 1938e: 190 Distribution: Costa Rica, El Salvador, Guatemala, Honduras

Phyllophaga eligia Sanderson Phyllophaga (Listrochelus) eligia Sanderson, 1958: 161 Distribution: México, United States

Phyllophaga elizoria Saylor Phyllophaga elizoria Saylor, 1937d: 321 (for Schaeffer, 1906: 257) Distribution: United States

Synonym: Lachnosterna pygidialis Schaeffer, 1906: 257 (not Brenske, 1892) Phyllophaga pygidialis (Schaeffer); Glasgow, 1916: 374

Phyllophaga elongata (Linell) Lachnosterna elongata Linell, 1896: 725 Phyllophaga elongata (Linell); Glasgow, 1916: 374 Distribution: United States

Synonym: Phyllophaga murrea Sanderson, 1950: 90

Phyllophaga emburyi Saylor Phyllophaga emburyi Saylor, 1938f: 344 Phyllophaga (Phytalus) emburyi Saylor; Morón, 1986: 218 Distribution: México

Phyllophaga eniba Saylor Phyllophaga (Phyllophaga) eniba Saylor, 1943e: 277 72 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Phyllophaga enkerliniana Deloya and Morón Phyllophaga (Phyllophaga) enkerliniana Deloya and Morón, 1999: 114 Distribution: México

Phyllophaga ephilida ephilida (Say) Melolontha ephilida Say, 1825: 196 Trichestes ephilidia [sic] (Say); Burmeister, 1855; 359 [error] Lachnosterna ephilida (Say); LeConte, 1856: 241 Phyllophaga ephilida (Say); Glasgow, 1916: 371 Phyllophaga (Phyllophaga) ephilida (Say); Morón, 1986: 204 Distribution: Canada, México, United States

Synonym: Lachnosterna burmeisteri LeConte, 1856: 242

Synonym: Ancylonycha laticeps Blanchard, 1851: 136 Lachnosterna laticeps (Blanchard); Bates, 1888: 213 Phyllophaga laticeps (Blanchard): Blackwelder, 1944; 225

Phyllophaga ephilida virilis Reinhard Phyllophaga ephilida virilis Reinhard, 1939: 51 Distribution: United States

Phyllophaga epigaea (Wickham) Lachnosterna epigaea Wickham, 1903: 71 Phyllophaga epigoea (Wickham); Glasgow, 1916: 374 [error] Phyllophaga epigaea (Wickham); Sanderson, 1939: 11 Phyllophaga (Phyllophaga) epigaea (Wickham); Sanderson, 1958: 164 Phyllophaga (Eugastra) epigea [sic](Wickham); Morón, 1986: 206 [error] Distribution: United States

Phyllophaga epulara Sanderson Phyllophaga (Phyllophaga) epulara Sanderson, 1958: 169 Phyllophaga (Phytalus) epulara Sanderson; Morón, 1986: 200 Distribution: México

Phyllophaga erota Saylor Phyllophaga (Phyllophaga) erota Saylor, 1942a: 157 Phyllophaga (Phyllophaga) erosa [sic] Saylor; Frey, 1975a: 218 [error] Distribution: Ecuador

Phyllophaga errans (LeConte) Lachnosterna errans LeConte, 1860: 283 Phyllophaga errans (LeConte); Glasgow, 1916: 373 Distribution: United States

Phyllophaga espina Sanderson 73 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga espina Sanderson, 1951: 264 Distribution: Dominican Republic

Phyllophaga esquinada Sanderson Phyllophaga esquinada Sanderson, 1951: 268 Distribution: Haiti

Phyllophaga estacea Saylor Phyllophaga (Listrochelus) estacea Saylor, 1943c: 31 (for Moser, 1921c: 261) Distribution: México

Synonym: Phyllophaga ponerosus Blackwelder, 1944: 226 (for Moser, 1921c: 261)

Synonym: Lachnostera testaceipennis Moser, 1921c: 261 (not Blanchard, 1851: 134)

Phyllophaga etabatesiana Morón Phyllophaga (Chlaenobia) etabatesiana Morón, 1992b: 28 Distriubtion: México

Phyllophaga euryaspis (Bates) Lachnosterna euryaspis Bates, 1888: 204 Phyllophaga (Phyllophaga) euryaspis (Bates); Morón, 1986: 205 Distribution: México

Phyllophaga exorata (Horn) Lachnosterna exorata Horn, 1887b: 278 Phyllophaga exorata (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga expansa Chapin Phyllophaga expansa Chapin, 1932b: 182 Cnemarachis expansa (Chapin): Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga explanicollis Chapin Phyllophaga explanicollis Chapin, 1935b: 69 Cnemarachis explanicollis (Chapin): Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga extincta (Wickham) Lachnosterna extincta Wickham, 1916: 9 Phyllophaga extincta (Wickham); Wickham, 1920: 358 Distribution: United States

Phyllophaga falsa (LeConte) Listrochelus falsus LeConte, 1856: 264 Phyllophaga (Listrochelus) falsa (LeConte); Saylor, 1940d: 97 Distribution: United States 74 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Listrochelus langeri Chapin, 1934b: 93

Phyllophaga falta Sanderson Phyllophaga falta Sanderson, 1950: 92 Distribution: United States

Phyllophaga farcta (LeConte) Lachnosterna farcta LeConte, 1856: 238 Phyllophaga farcta (LeConte); Glasgow, 1916: 372 Phyllophaga (?Eugastra) farcta (LeConte); Morón, 1986: 206 Distribution: United States

Phyllophaga favosa Saylor Phyllophaga (Phyllophaga) favosa Saylor, 1941e: 539 Distribution: Costa Rica

Phyllophaga ferrugata (Moser) Lachnosterna ferrugata Moser, 1918a: 42 Phyllophaga ferrugata (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) ferrugata (Moser); Frey, 1975a: 211 Distribution: Bolivia

Phyllophaga ferruginea (Moser) Lachnosterna ferruginea Moser, 1918a: 41 Phyllophaga ferruginea (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) ferruginea (Moser); Frey, 1975a: 203 Distribution: Venezuela

Phyllophaga ferupilis Saylor Phyllophaga (Phyllophaga) ferupilis Saylor, 1941d: 386 Distribution: Guatemala

Phyllophaga fervida (Fabricius) Melolontha fervida Fabricius, 1775: 32 Ancylonycha fervida (Fabricius); Blanchard, 1851: 133 Phyllophaga fervida (Fabricius); Glasgow, 1916: 370 Cnemarachis fervida (Fabricius): Blackwelder, 1944: 223 Distribution: Canada, Hispaniola, Jamaica, United States

Synonym: Lachnosterna arcuata Smith, 1889: 183

Synonym: Melolontha quercina Knoch, 1801: 74 Lachnosterna quercina (Knoch); Hope, 1837: 100 Ancyclonycha quercina (Knoch): Burmeister, 1855: 319

Phyllophaga fesina Saylor Phyllophaga (Phyllophaga) fesina Saylor, 1948b: 356 75 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Argentina

Phyllophaga fimbriata Chapin Phyllophaga fimbriata Chapin, 1932b: 182 Cnemarachis fimbriata (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga fimbripes (LeConte) Listrochelus fimbripes LeConte, 1856: 264 Phyllophaga (Listrochelus) fimbripes (LeConte); Saylor, 1940d: 95 Distribution: United States

Phyllophaga fissilabris (Bates) Phytalus fissilabris Bates, 1888: 184 Phyllophaga fissilabris (Bates); Blackwelder, 1944: 224 Phyllophaga (Phytalus) fissilabris (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga flavipennis (Horn) Listrochelus flavipennis Horn, 1885: 123 Phyllophaga (Listrochelus) flavipennis (Horn); Saylor, 1940d: 84 Distribution: United States

Phyllophaga flavipilosa (Moser) Lachnosterna flavipilosa Moser, 1921c: 252 Phyllophaga flavipilosa (Moser); Blackwelder, 1944: 224 Distribution: México

Phyllophaga flavisetis (Bates) Diplotaxis flavisetis Bates, 1888: 165 Phyllophaga flavisetis (Bates); Vaurie, 1958: 280 Distribution: México

Phyllophaga floridana Robinson Phyllophaga floridana Robinson, 1938: 110 Distribution: United States

Phyllophaga foralita Saylor Phyllophaga (Listrochelus) foralita Saylor, 1938e: 187 Distribution: México

Phyllophaga forbesi Glasgow Phyllophaga forbsei Glasgow, 1916: 378 Distribution: United States

Phyllophaga forcipata (Burmeister) Phytalus forcipatus Burmeister, 1855: 346 Phyllophaga forcipata (Burmeister); Blackwelder, 1944: 224 76 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phytalus) forcipata (Burmeister); Frey, 1975a: 222 Distribution: Argentina, Brazil

Synonym: Phytalus simplex Burmeister, 1855: 347 Phyllophaga simplex (Burmeister); Blackwelder, 1944: 226

Phyllophaga forsteri (Burmeister) Ancylonycha forsteri Burmeister, 1855: 325 Phyllophaga forsteri (Burmeister); Glasgow, 1916: 372 Distribution: United States

Synonym: Lachnosterna lugubris LeConte, 1856: 248 Phyllophaga lugubris (LeConte); Evans, 2003: 99

Synonym: Lachnosterna lutescens LeConte, 1856: 249 Phyllophaga lutescens (LeConte); Evansk 2003: 99

Synonym: Lachnosterna nova Smith, 1889a: 95 Phyllophaga nova (Smith); Evans, 2003: 99

Synonym: Lachnosterna politula Horn, 1887b: 248 Phyllophaga politula (Horn); Evans, 2003: 99

Synonym: Lachnosterna semicribrata LeConte, 1856: 247 Phyllophaga semicribrata (LeConte); Evans, 2003: 99

Phyllophaga fossoria Sanderson Phyllophaga fossoria Sanderson, 1951: 258 Distribution: Dominican Republic

Phyllophaga foveicollis (Bates) Lachnosterna foveicollis Bates, 1888: 190 Phyllophaga foveicollis (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) foveicollis (Bates); Morón, 1986: 259 Distribution: México

Phyllophaga foxii Davis Phyllophaga foxii Davis, 1920: 334 Distribution: United States

Phyllophaga fragilipennis (Blanchard) Ancylonycha fragilipennis Blanchard, 1851: 135 Phyllophaga fragilipennis (Blanchard); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) fragilipennis (Blanchard); Frey, 1975a: 206 Distribution: Colombia

Phyllophaga franciscana Garcia-Vidal Phyllophaga franciscana Garcia-Vidal, 1984: 319 77 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Cuba

Phyllophaga fraterna fraterna Harris Phyllophaga fraterna Harris, 1841: 29 Ancyclonycha fraterna (Harris); Burmeister, 1855: 322 Lachnosterna fraterna (Harris); LeConte, 1856: 249 Phyllophaga fraterna (Harris); Glasgow, 1916: 371 Distribution: Canada, United States

Synonym: Ancylonycha cognata Burmeister, 1855: 323 Lachnosterna cognata (Burmeister); LeConte, 1856: 248

Phyllophaga fraterna mississippiensis Davis Phyllophaga fraterna var. mississippiensis Davis, 1920: 330 Distribution: United States

Phyllophaga fraternaria Cano and Morón Phyllophaga (Chlaenobia) fraternaria Cano and Morón, 2002: 353 Distribution: Guatemala

Phyllophaga freeborni (Saylor) Lachnosterna freeborni Saylor, 1937c: 32 Phyllophaga (Phyllophaga) freeborni (Saylor); Morón, 1986: 205 Distribution: México

Phyllophaga fucata (Horn) Lachnosterna fucata Horn, 1887b: 278 Phyllophaga fucata (Horn); Glasgow, 1916: 373 Phyllophaga (Phyllophaga) fucata (Horn); Sanderson, 1958: 172 Distribution: México, United States

Synonym: Phyllophaga linsleyi Saylor, 1936a: 4

Phyllophaga fulvipennis (Blanchard) Ancylonycha fulvipennis Blanchard, 1851: 135 Phyllophaga fulvipennis (Blanchard); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) fulvipennis (Blanchard); Frey, 1975a: 206 Distribution: Brazil

Phyllophaga fulviventris (Moser) Lachnosterna fulviventris Moser, 1918a: 27 Phyllophaga fulviventris (Moser); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) fulviventris (Moser); Morón, 1986: 202 Distribution: México

Phyllophaga fusca (Froelich) Melolontha fusca Froelich, 1792: 99 (see note below) Ancylonycha fusca (Froelich); Blanchard, 1851: 133 78 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Lachnosterna fusca (Froelich); Horn, 1887b: 245 Phyllophaga fusca (Froelich); Glasgow, 1916: 371 Phyllophaga (Phyllophaga) fusca (Froelich); Riley and Wolfe, 2003: 22 Distribution: Canada, United States

Synonym: Melolontha fervens Gyllenhal, 1817: 74 Ancylonycha fervens (Gyllenhal); Blanchard, 1851: 133

Remark: the senior homonym Melolontha fusca Gmelin, 1788: 1569 is a junior synonym of Sericoides testacea (Fabricius) 1775: 35.

Phyllophaga fuscipennis (Moser) Lachnosterna fuscipennis Moser, 1918a: 64 Phyllophaga (Cnemarachis) fuscipennis (Moser); Chalumeau, 1983: 113 Distribution: Guadeloupe

Synonym: Phyllophaga vitraci Paulian, 1947: 54

Phyllophaga futilis (LeConte) Lachnosterna futilis LeConte, 1850: 226 Phyllophaga futilis (LeConte); Glasgow, 1916: 371 Distribution: Canada, United States

Synonym: Lachnosterna decidua LeConte, 1856: 246 Phyllophaga decidua (LeConte); Evans, 2003: 102

Synonym: Ancylonycha gibbosa Burmeister, 1855: 324 Lachnosterna gibbosa (Burmeister); Horn, 1887b: 230

Synonym: Lachnosterna serricornis LeConte, 1856: 247

Phyllophaga gaigei Sanderson Phyllophaga gaigei Sanderson, 1948: 4 Distribution: United States

Phyllophaga galeanae Saylor Phyllophaga galeanae Saylor, 1943c: 28 Phyllophaga (Phyllophaga) galeanae Saylor: Morón, 1986: 205 Distribution: México

Phyllophaga garrota Sanderson Phyllophaga (Cnemarachis) garrota Sanderson, 1951: 262 Distribution: Domincan Republic

Phyllophaga gaumeri (Bates) Lachnosterna gaumeri Bates, 1889: 404 Phyllophaga gaumeri (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) gaumeri (Bates); Morón, 1986: 205 79 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Phyllophaga gentryi Saylor Phyllophaga terminalis gentryi Saylor, 1936b: 64 Phyllophaga gentryi Saylor; Blackwelder, 1944: 224 Distribution: México

Phyllophaga georgiana (Horn) Phytalus georgianus Horn, 1885: 122 Phyllophaga (Phytalus) georgiana (Horn); Saylor, 1939c: 163 Phyllophaga (Phyllophaga) georgiana (Horn); Sanderson, 1958: 166 Distribution: United States

Phyllophaga ghentata (Saylor) Lachnosterna ghentata Saylor, 1937c: 33 Phyllophaga ghentata (Saylor); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) ghentata (Saylor); Morón, 1986: 205 Distribution: México

Phyllophaga gigantea (Bates) Lachnosterna gigantea Bates, 1888: 198 Phyllophaga gigantea (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) gigantea (Bates); Solís and Morón, 1998: 23 Distribution: Costa Rica, Guatemala, México, Nicaragua, Panama

Phyllophaga gigantissima Saylor Phyllophaga gigantissima Saylor, 1935g: 33 Triodonyx gigantissima Saylor; Saylor, 1942a: 158 Phyllophaga (Triodonyx) gigantissima Saylor; Morón, 1986: 259 Distribution: México

Phyllophaga ginigra Saylor Phyllophaga (Phyllophaga) ginigra Saylor, 1940d: 112 Distribution: México

Phyllophaga glaberrima (Blanchard) Ancylonycha glaberrima Blanchard, 1850: 136 Lachnosterna glaberrima (Blanchard); LeConte, 1856: 242 Phyllophaga glaberrima (Blanchard); Glasgow, 1916: 372 Distribution: United States

Synonym: Lachnosterna parva Linell, 1896: 726 (not Brenske, 1892) Phyllophaga parva (Linell); Glasgow, 1916: 374

Synonym: Phyllophaga pagilis Saylor, 1937d: 321 (for Linell, 1896: 726)

Phyllophaga glabricula (LeConte) Lachnosterna glabricula LeConte, 1856: 260 80 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga glabricula (LeConte); Glasgow, 1916: 373 Distribution: México, United States

Phyllophaga godmani (Bates) Lachnosterna godmani Bates, 1889: 403 Phyllophaga godmani (Bates); Blackwelder, 1944: 224 Phyllophaga (Phyllophaga) godmani (Bates); Morón and Nogueira, 1997: 260 Distribution: México

Phyllophaga gracilis angulata Glasgow Phyllophaga gracilis angulata Glasgow, 1925: 294 Distribution: United States

Phyllophaga gracilis gracilis (Burmeister) Trichestes gracilis Burmeister, 1855: 361 Phyllophaga gracilis (Burmeister); Glasgow, 1916: 372 Distribution: Canada, United States

Synonym: Lachnosterna inana LeConte, 1856: 242

Synonym: Endrosa volvula LeConte, 1856: 235 United States Phyllophaga gracillima (Dalla Torre) Lachnosterna gracillima Dalla Torre, 1912c: 191 (for Nonfried, 1891: 265) Phyllophaga gracillima (Dalla Torre); Blackwelder, 1944: 224 Distribution: Honduras

Synonym: Lachnosterna gracilis Nonfried, 1891: 265 (not Burmeister, 1855: 361)

Phyllophaga grancha Saylor Phyllophaga (Phyllophaga) grancha Saylor, 1948b: 354 Distribution: Argentina

Phyllophaga granti Saylor Phyllophaga (Listrochelus) granti Saylor, 1940d: 85 Distributon: México, United States

Phyllophaga gravidula (Moser) Lachnosterna gravidula Moser, 1921c: 247 Phyllophaga gravidula (Moser); Sanderson, 1942: 54 Phyllophaga (Phyllophaga) gravidula (Moser); Sanderson, 1958: 166 Distributon: México, United States

Synonym: Phyllophaga perita Sanderson , 1937a: 14

Phyllophaga grossepunctata (Moser) Lachnosterna grossepunctata Moser, 1918a: 33 Phyllophaga grossepunctata (Moser); Blackwelder, 1944: 224 81 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Costa Rica

Phyllophaga guanacasteca Morón and Solís Phyllophaga (Phyllophaga) guanacasteca Morón and Solís, 2000: 908 Distriubtion: Costa Rica

Phyllophaga guanicana Smyth Phyllophaga guanicana Smyth, 1917: 152 Cnemarachis guanicana (Smyth): Blackwelder, 1944: 223 Distribution: Peurto Rico

Phyllophaga guapilana Saylor Phyllophaga (Phytalus) guapilana Saylor, 1935h: 497 Distribution: Costa Rica

Phyllophaga guapiles Saylor Phyllophaga (Phyllophaga) guapiles Saylor, 1941e: 536 Distribution: Costa Rica

Phyllophaga guapilesea Saylor Phyllophaga (Phyllophaga) guapilesea Saylor, 1941e: 540 Distribution: Costa Rica

Phyllophaga guapiloides Morón and Solís Phyllophaga (Phyllophaga) guapiloides Morón and Solís, 2001: 26 Distribution: Costa Rica

Phyllophaga guatemala Saylor Phyllophaga (Listrochelus) guatemala Saylor, 1940e: 115 Phyllophaga (Phytalus) guatemala Saylor; Cano and Morón, 1998: 11 Description: Guatemala, México

Phyllophaga guayabillosa Saylor Phyllophaga guayabillos Saylor, 1936d: 293 Phyllophaga guayabillosa Saylor; Saylor, 1938f: 347 (emendation) Phyllophaga (Phyllophaga) guayabillosa Saylor; Solís and Morón, 1998: 25 Distribution: Costa Rica

Phyllophaga guerrocana Saylor Phyllophaga guerrocans Saylor, 1938f: 345 Phyllophaga (Phyllophaga) guerrocana Saylor; Morón, 1986: 205 Distribution: México

Phyllophaga haagi Saylor Phyllophaga (Phyllophaga) haagi Saylor, 1943a: 26 Distribution: México

Phyllophaga halffteriana Morón 82 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Chlaenobia) halffteriana Morón, 1992b: 16 Distribution: Costa Rica, Honduras, México

Phyllophaga hamata (Horn) Lachnosterna hamata Horn, 1887b: 220 Phyllophaga hamata (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga hardyi Garcia-Vidal Phyllophaga (Cnemarachis) hardyi Garcia-Vidal, 1984: 319 Distribution: Cuba

Phyllophaga hemilissa (Bates) Lachnosterna hemilissa Bates, 1888: 201 Phyllophaga hemilissa (Bates); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) hemilissa (Bates); Solís and Morón, 1998: 24 Distribution: Costa Rica, Panama

Phyllophaga hernandezi Garcia-Vidal Phyllophaga (Cnemarachis) hernandezi Garcia-Vidal, 1984: 321 Distribution: Cuba

Phyllophaga heteroclita (Burmeister) Phytalus heteroclitus Burmeister, 1855: 348 Phyllophaga heteroclita (Burmeister); Blackwelder, 1944: 225 Phyllophaga (Phytalus) heteroclita (Burmeister); Frey, 1975a: 223 Distribution: Brazil

Phyllophaga heteronycha heteronycha (Bates) Lachnosterna heteronycha Bates, 1888: 186 Phyllophaga heteronycha (Bates); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) heteronycha (Bates); Morón, 1998: 24 Distribution: México

Phyllophaga heteronycha pilidorsis Saylor Phyllophaga heteronycha pilidorsis Saylor, 1938f: 344 Phyllophaga pilidorsis Saylor; Blackwelder, 1944: 225 Distribution: México

Phyllophaga hidalgoana Saylor Phyllophaga (Phyllophaga) hidalgoana Saylor, 1943e: 270 Distribution: México

Phyllophaga hintonella Saylor Phyllophaga (Phyllophaga) hintonella Saylor, 1941a: 27 Distribution: México

Phyllophaga hintoni Saylor 83 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phytalus) hintoni Saylor, 1935h: 498 Distribution: México

Phyllophaga hirsuta (Knoch) Melolontha hirsuta Knoch, 1801: 78 Lachnosterna hirsuta (Knoch); LeConte, 1856: 254 Phyllophaga hirsuta (Knoch); Glasgow, 1916: 371 Distribution: Canada, United States

Phyllophaga hirticollis (Moser) Lachnosterna hirticollis Moser, 1921c: 249 Phyllophaga hirticollis (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) hirticollis (Moser); Morón , 1986: 203 Distribution: México

Phyllophaga hirticula comosa Davis Phyllophaga hirticula var. comosa Davis, 1920: 337 Distribution: United States

Phyllophaga hirticula hirticula (Knoch) Melolontha hirticula Knoch, 1801: 79 Ancylonycha hirticula (Knoch); Burmeister, 1855: 327 Lachnosterna hirticula (Knoch); Hope, 1837: 100 Phyllophaga hirticula (Knoch); Glasgow, 1916: 371 Distribution: Canada, United States

Phyllophaga hirtifrons (Moser) Phytalus hirtifrons Moser, 1921c: 259 Phyllophaga hirtifrons (Moser); Blackwelder, 1944; 225 Phyllophaga (Phyllophaga) hirtifrons (Moser); Morón , 1986: 200 Distribution: México

Phyllophaga hirtiventris (Horn) Lachnosterna hirtiventris Horn 1887b: 231 Phyllophaga hirtiventris (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga hoegei (Bates) Phytalus h gei Bates, 1888: 180 Phyllophaga hoegei (Bates); Blackwelder, 1944: 225 Phyllophaga (Phytalus) hoegei (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga hoegella Saylor Phyllophaga hoegella Saylor, 1943c: 26 Phyllophaga (Phyllophaga) hoagella Saylor; Morón, 1986: 203 Distribution: México

84 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga hoffmanita Morón Phyllophaga (Phyllophaga) hoffmanita Morón, 1996b: 332 Distribution: México

Phyllophaga hogardi (Blanchard) Ancylonycha hogardi Blanchard, 1851: 137 Phyllophaga hogardi (Blanchard); Wolcott, 1928: 26 Distribution: Dominican Republic, Haiti

Phyllophaga hondura Saylor Phyllophaga (Phyllophaga) hondura Saylor, 1943d: 132 Distribution: Belize, Costa Rica, United States

Phyllophaga hondurasana (Moser) Lachnosterna hondurasana Moser, 1921c: 255 Phyllophaga hondurasana (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) hondurasana (Moser); Solís and Morón, 1998: 25 Distribution: Costa Rica, Guatemala, Honduras

Phyllophaga hoogstraali Saylor Phyllophaga (Phytalus) hoogstraali Saylor, 1943a: 21 Distribution: México

Phyllophaga hornii (Smith) Lachnosterna hornii Smith, 1889a: 95 Phyllophaga hornii (Smith); Glasgow, 1916: 373 Phyllophaga (Phyllophaga) hornii (Smith); Distribution: United States

Phyllophaga howdeniana Morón Phyllophaga (Chlaenobia) howdeniana Morón, 1992b: 22 Distribution: México

Phyllophaga huachuca Saylor Phyllophaga (Listrochelus) huachuca Saylor, 1940d: 82 Distribution: México, United States

Phyllophaga hubbelli Cartwright Phyllophaga hubbelli Cartwright, 1946: 10 Distribution: United States

Phyllophaga humboldtiana Morón Phyllophaga (Phyllophaga) humboldtiana Morón, 2003a: 239 Distribution: México

Phyllophaga idonea Sanderson Phyllophaga idonea Sanderson, 1948: 1 Distribution: United States 85 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga ignava (Horn) Lachnosterna ignava Horn, 1887b: 280 Phyllophaga ignava (Horn); Glasgow, 1916: 373 Phyllophaga (Phyllophaga) ignava (Horn); Sanderson, 1958: 172 Distribution: México, United States

Phyllophaga ilhuicaminai Morón Phyllophaga (Phyllophaga) ilhuicaminai Morón, 1998: 40 Distribution: México

Phyllophaga ilicis (Knoch) Melolontha ilicis Knoch, 1801: 75 Ancyclonycha ilicis (Knoch); Blanchard, 1851: 133 Lachnosterna ilicis (Knoch); LeConte, 1856: 253 Phyllophaga ilicis (Knoch); Glasgow, 1916: 371 Distribution: Canada, United States

Synonym: Lachnosterna ciliata LeConte, 1856: 253

Synonym: Ancylonycha fimbriata Burmeister,1855: 326

Synonym: Phyllophaga jonesi Sanderson, 1939a: 5

Synonym: Melolontha porcina Hentz, 1830: 256

Phyllophaga imitans Frey Phyllophaga (Phyllophaga) imitans Frey, 1975a: 217 Distribution: Brazil

Phyllophaga imitatrix Chapin Phyllophaga imitatrix Chapin, 1932b: 199 Cnemarachis imitatrix (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga impar Davis Phyllophaga impar Davis, 1920: 335 Distribution: United States

Phyllophaga implicita (Horn) Lachnosterna implicita Horn, 1887b: 262 Phyllophaga implicita (Horn); Glasgow, 1916: 373

Synonym: Phyllophaga linelli Saylor, 1937d: 321

Synonym: Lachnosterna minor Linell, 1896: 728 (not Brenske, 1994)

Phyllophaga impressipyga Frey 86 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phyllophaga) impressipyga Frey, 1975: 213 Distribution: Colombia, Venezuela

Phyllophaga imprima Sanderson Phyllophaga (Cnemarchis) imprima Sanderson, 1951: 276 Distribution: Dominican Republic

Phyllophaga incuria Sanderson Phyllophaga incuria Sanderson, 1942: 53 (for Sanderson, 1937a: 16)

Synonym: Phyllophaga texana Sanderson, 1937a: 16 (not LeConte, 1856: 263)

Phyllophaga independentista Garcia-Vidal Phyllophaga (Cnemarachis) independentista Garcia-Vidal, 1984: 322 Distribution: Cuba

Phyllophaga inepta (Horn) Lachnosterna inepta Horn, 1887b: 282 Phyllophaga inepta (Horn); Glasgow, 1916: 373 Phyllophaga (Phyllophaga) inepta (Horn); Riley and Wolfe, 2003: 22 Distribution: United States

Phyllophaga infidelis (Horn) Lachnosterna infidelis Horn, 1887b: 253 Phyllophaga infidelis (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga inflativentris (Moser) Lachnosterna inflativentris Moser, 1918a: 22 Phyllophaga inflativentris (Moser); Blackwelder, 1944: 225 Distribution: México

Phyllophaga inflexa Barret Phyllophaga (Phyllophaga) inflexa Barret, 1935: 49 Distribution: United States

Phyllophaga instabilis Blackwelder Phyllophaga instabilis Blackwelder, 1944: 225 Phyllophaga (Chlaenobia) instabilis Blackwelder; Morón, 1986: 208 Distribution: México

Synonym: Chlaenobia arrowi Chapin, 1935a:9 (not Saylor, 1935f: 206)

Phyllophaga insulaepinora Garcia-Vidal Phyllophaga (Cnemarachis) insulaepinorum Garcia-Vidal, 1978: 6 Distribution: Cuba

Phyllophaga insulana (Moser) 87 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phytalus insulanus Moser, 1918a: 71 Phyllophaga insulanus (Moser); Blackwelder, 1944: 225 Phyllophaga (Phytalus) insulanus (Moser); Frey, 1975a: 220 Distribution: Trinidad

Synonym: Lachnosterna dilemma Arrow, 1920: 192

Phyllophaga integra (Say) Melolontha integra Say, 1835: 180 Lachnosterna integra (Say); Bates, 1888: 208 Phyllophaga integra (Say); Blackwelder, 1944: 225 Distribution: México, United States

Phyllophaga integriceps (Moser) Lachnosterna integriceps Moser, 1918a: 56 Phyllophaga integriceps (Moser); Blackwelder, 1944: 225 Distribution: México

Phyllophaga inversa (Horn) Lachnosterna inversa Horn, 1887b: 241 Phyllophaga inversa (Horn); Glasgow, 1916: 373 Distribution: Canada, United States

Phyllophaga invisa Riley and Wolfe Phyllophaga (Phyllophaga) invisa Riley and Wolfe, 1997: 426 Distribution: United States

Phyllophaga irazuana Saylor Phyllophaga irazuana Saylor, 1936d: 294 Phyllophaga (Phyllophaga) irazuana Saylor; Solís and Morón, 1998: 25 Distribution: Costa Rica

Phyllophaga itsmica Morón Phyllophaga (Phytalus) itsmica Morón, 2000b: 52 Distribution: México

Phyllophaga iviei Chalumeau Phyllophaga iviei Chalumeau, 1985: 32 Distribution: Virgin Islands

Phyllophaga izabalana Morón Phyllophaga (Phyllophaga) izabalana Morón, 2003a: 225 Distribution: Guatemala

Phyllophaga izucarana Morón and Aragón Phyllophaga (Phyllophaga) izucarana Morón and Aragón, 1997: 43 Distribution: México

88 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga jaliscensis Morón, Rivera, and López Phyllophaga (Phytalus) jaliscensis Morón, Rivera, and López, 2002: 497 Distribution: México

Phyllophaga jamaicana (Moser) Lachnosterna jamaicana Moser, 1918a: 63 Cnemarachis jamaicana (Moser); Blackwelder, 1944: 223 Phyllophaga jamaicana (Moser); Chalumeau, 1985: 22 Distribution: Jamaica

Synonym: Lachnosterna jamaicensis Arrow, 1920: 190 Cnemarachis jamaicensis (Arrow): Blackwelder, 1944: 223

Phyllophaga janzeniana Morón and Solís Phyllophaga (Phyllophaga) janzeniana Morón and Solís, 2000: 904 Distriubtion: Costa Rica

Phyllophaga jaragua Woodruff Phyllophaga jaragua Woodruff, 2005: 84 Distribution: Dominican Republic

Phyllophaga jaronua Chapin Phyllophaga jaronua Chapin, 1932b: 194 Cnemarachis jaronua (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga javepacuana Morón Phyllophaga (Phyllophaga) javepacuana Morón, 2003a: 241 Distribution: Guatemala, México

Phyllophaga jimenezi Woodruff and Sanderson Phyllophaga jimenezi Woodruff and Sanderson, 2005: 87 Distribution: Dominican Republic

Phyllophaga jovelana Morón and Cano Phyllophaga (Phyllophaga) jovelana Morón and Cano, 2000: 235 Distribution: México

Phyllophaga joyana Morón, Rivera, and López, Phyllophaga (Phyllophaga) joyana Morón, Rivera, and López, 2002: 492 Distribution: México

Phyllophaga jumberea Saylor Phyllophaga (Phyllophaga) jumberea Saylor, 1942a: 158 Distribution: Peru

Phyllophaga juquilana Morón Phyllophaga (Phytalus) juquilana Morón, 2000b: 41 89 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distriubtion: México

Phyllophaga juvenilis (Fall) 1932: 200 Listrochelus juvenilis Fall, 1932: 200 Phyllophaga (Listrochelus) juvenilis (Fall); Sanderson, 1958: 160 Distribution: México, United States

Synonym: Phyllophaga (Listrochelus) seri Morón, 2002: 459

Phyllophaga karlsioei (Linell) Lachnosterna karlsioei Linell, 1897: 400 Phyllophaga karlsioei (Linell); Glasgow, 1916: 374 Distribution: United States

Phyllophaga kenscoffi Wolcott Phyllophaga kenscoffi Wolcott, 1928: 24 Cnemarachis kenscoffi (Wolcott); Blackwelder, 194: 223 Phyllophaga kenscoffi Wolcott; Sanderson, 1951: 269 Distribution: Dominican Republic, Haiti

Phyllophaga kentuckiana Ritcher Phyllophaga kentuckiana Ritcher, 1937: 285 Distribution: United States

Phyllophaga knausii (Schaeffer) Listrochelus knausii Schaeffer, 1907: 67 Phyllophaga knausii (Schaeffer); Blackwelder and Arnett, 1974: 58 Distribution: United States

Phyllophaga knochii (Schoenherr and Gyllenhal) Melolontha knochii Schoenherr and Gyllenhal, 1817: 75 Ancylonycha knochii (Schoenherr and Gyllenhal); Burmeister, 1855: 325 Lachnosterna knochii (Schoenherr and Gyllenhal); LeConte, 1856: 252 Phyllophaga knochii (Schoenherr and Gyllenhal); Glasgow, 1916: 371 Distribution: United States

Phyllophaga koehleriana Saylor Phyllophaga (Listrochelus) koehleriana Saylor, 1940d: 99 Distribution: United States

Phyllophaga kohlmanniana Morón and Solís Phyllophaga (Phyllophaga) kohlmanniana Morón and Solís, 2000: 901 Distriubtion: Costa Rica

Phyllophaga kulzeri Frey Phyllophaga (Phyllophaga) kulzeri Frey, 1975a: 203 Distriubution: Venezuela

90 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga kuntzeni (Moser) Phytalus kuntzeni Moser, 1921c: 260 Phyllophaga kuntzeni (Moser); Blackwelder, 1944: 225 Distribution: Bolivia

Phyllophaga laboriosa Garcia-Vidal Phyllophaga laboriosa Garcia-Vidal, 1988: 102 Distribution: Cuba

Phyllophaga labrata Chapin Phyllophaga labrata Chapin, 1932b: 199 Cnemarachis labrata (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga lacroixi Paulian Phyllophaga (Cnemarachis) lacroixi Paulian, 1947: 54 Distribution: Martinique

Phyllophaga laeviscutata (Moser) Lachnosterna laeviscutata Moser, 1918a: 35 Phyllophaga laeviscutata (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) laeviscutata (Moser); Solís and Morón, 1998: 24 Distribution: Costa Rica, Panama

Phyllophaga lalanza Saylor Phyllophaga lalanza Saylor, 1941b: 67 Triodonyx lalanza (Saylor); Saylor, 1942a: 158 Phyllophaga (Triodonyx) lalanza Saylor; Morón, 1986: 259 Distribution: México

Phyllophaga laminata (Moser) Phytalus laminatus Moser, 1921c: 259 Phyllophaga laminata (Moser); Blackwelder, 1944: 225 Phyllophaga (Phytalus) laminata (Moser); Frey, 1975a; 223 Distribution: Brazil

Phyllophaga lanata (Blanchard) Ancylonycha lanata Blanchard, 1851: 135 Phyllophaga lanata (Blanchard): Blackwelder, 1944; 224 Phyllophaga (Phyllophaga) lanata (Blanchard): Frey, 1975a: 202 Distribution: Venezuela

Phyllophaga lanceolata (Say) Melolontha lanceolata Say, 1824: 242 Tostegoptera lanceolata (Say); Blanchard, 1851: 149 Ancylonycha lanceolata (Say); Lacordarie, 1856: 285 Lachnosterna lanceolata (Say); LeConte, 1856: 237 Phyllophaga lanceolata (Say); Glasgow, 1916; 371 91 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Tostegoptera) lanceolata (Say); Bloeker, 1936: 53 Distribution: Canada, United States

Synonym: Phyllophaga (Tostegoptera) grisiana Bloeker, 1936: 56

Synonym: Phyllophaga (Tostegoptera) lanceolata arizonae Bloeker, 1936: 55

Phyllophaga lanepta (Saylor) Lachnosterna lanepta Saylor, 1937c: 31 Phyllophaga lanepta (Saylor); Blackwelder, 1944: 225 Distribution: Guatemala

Phyllophaga laportaei (Blanchard) Listrochelus laportaei Blanchard, 1851: 141 Phyllophaga laportaei (Blanchard); Blackwelder, 1944: 225 Phyllophaga (Listrochelus) laportaei (Blanchard); Morón, 1986: 201 Distribution: México

Phyllophaga larimar Woodruff Phyllophaga larimar Woodruff, 2005: 90 Distribution: Dominican Republic, Haiti

Phyllophaga latefissa (Moser) Phytalus latefissus Moser, 1918a: 66 Cnemarachis latefissus (Moser); Blackwelder, 1944: 223 Phyllophaga (Phytalus) latefissa (Moser); Frey, 1975a: 220 Distribution: Brazil, St. Croix

Synonym: Phytalus furcatus Moser, 1918a: 67 Phyllophaga furcata (Moser); Blackwelder, 1944: 224

Phyllophaga latens (Arrow) Lachnosterna latens Arrow, 1900: 176 Cnemarachis latens (Moser); Blackwelder, 1944: 223 Phyllophaga latens (Moser); Chalumeau, 1990: 336 Distribution: St. Vincent

Phyllophaga latidens (Schaeffer) Lachnosterna latidens Schaeffer, 1906: 258 Phyllophaga latidens (Schaeffer), Glasgow, 1916: 374 Phyllophaga (Phyllophaga) latidens (Schaeffer); Sanderson, 1958: 166 Distribution: United States

Phyllophaga latifrons (LeConte) Lachnosterna latifrons LeConte,1856: 241 Phyllophaga latifrons (LeConte); Glasgow, 1916: 372 Distribution: United States

92 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga latipes (Bates) Chlaenobia latipes Bates, 1888: 167 Phyllophaga latipes (Bates); Blackwelder, 1944: 225 Phyllophaga (Chlaenobia) latipes (Bates); Morón, 1986: 209 Distribution: El Salvador, Guatemala, México

Synonym: Chlaenobia bicallosa Bates, 1888: 168

Phyllophaga latiungula Wolcott Phyllophaga latiungula Wolcott, 1928: 28 Distribution: Haiti

Phyllophaga lebasii (Blanchard) Ancylonycha lebasii Blanchard, 1850: 134 Phyllophaga lebasi [sic] (Blanchard); Blackwelder, 1944: 225 Distribution: Colombia

Phyllophaga lempira Morón and Robbins Phyllophaga (Chlaenobia) lempira Morón and Robbins, 2005: 332 Distribution: Honduras

Phyllophaga lenis (Horn) Lachnosterna lenis Horn, 1887b: 287 Phyllophaga lenis (Horn); Glasgow, 1916: 373 Phyllophaga (Phyllophaga) lenis (Horn); Sanderson, 1958: 165 Distribution: Costa Rica, Guatemala, México, United States

Phyllophaga leonilae Morón Phyllophaga (Phyllophaga) leonilae Morón, 1996a: 72 Distribution: México

Phyllophaga leonina (Bates) Lachnosterna leonina Bates, 1888: 188 Phyllophaga leonina (Bates); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) leonina (Bates); Morón, 1986: 203 Distribution: México

Phyllophaga leporina (Erichson) Lachnosterna leporina Erichson, 1848: 563 Phyllophaga leporina (Erichson); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) leporina (Erichson); Frey, 1975a: 219 Distribution: Guyana

Phyllophaga leprieuri (Blanchard) Ancylonycha leprieuri Blanchard, 1851: 134 Phyllophaga leprieuri (Blanchard); Blackwelder, 1944: 225 Phyllophaga leprieuri (Blanchard): Frey, 1975a: 212 Distribution: Venezuela 93 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga leptospica Sanderson Phyllophaga leptospica Sanderson, 1951: 262 Distribution: Dominican Republic, Haiti

Phyllophaga lineata (Bates) Phytalus lineatus Bates, 1888: 183 Phyllophaga lineatus (Bates); Blackwelder, 1944: 225 Phyllophaga (Phytalus) lineata (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga lineatoides Morón Phyllophaga (Phytalus) lineatoides Morón, 2000: 46 Distribution: México

Phyllophaga linharensis Frey Phyllophaga (Phyllophaga) linharensis Frey, 1975a: 218 Distribution: Brazil

Phyllophaga lissopyge (Bates) Lachnosterna lissopyge Bates, 1888: 201 Phyllophaga lissopyge (Bates); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) lissopyge (Moser); Solís and Morón, 1998: 24 Distribution: Costa Rica, Nicaragua, Panama

Phyllophaga lobata (Fall) Lachnosterna lobata Fall, 1908: 163 Phyllophaga lobata (Fall); Glasgow, 1916: 374 Phyllophaga (Phyllophaga) lobata (Fall); Sanderson, 1958: 172 Distribution: México, United States

Phyllophaga lodingi Sanderson Phyllophaga lodingi Sanderson, 1939a: 4 Distribution: United States

Phyllophaga longicornis (Blanchard) Ancylonycha longicornis Blanchard, 1851: 134 Lachnosterna longicornis (Blanchard); Horn, 1887b: 280 Phyllophaga longicornis (Blanchard); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) longicornis (Blanchard); Frey, 1975a: 205 Distribution: Brazil, Colombia

Phyllophaga longifoliata (Moser) Lachnosterna longifoliata Moser, 1921c: 253 Phyllophaga longifoliata (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) longifoliata (Moser); Morón, 1986: 205 Distribution: México

94 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga longispina (Smith) Lachnosterna longispina Smith, 1889a: 97 Phyllophaga longispina (Smith); Glasgow, 1916: 374 Distribution: Canada, United States

Phyllophaga longitarsa (Say) Melolontha longitarsa Say, 1824: 241 Trichestes longitarsis (Say); Burmeister, 1855: 359 Lachnosterna longitarsis (Say); LeConte, 1856: 240 Phyllophaga longitarsa (Say); Glasgow, 1916: 371 Distribution: United States

Synonym: Lachnosterna frontalis LeConte, 1856: 239 Phyllophaga frontalis (LeConte); Glasgow, 1916: 372

Phyllophaga lorencita Morón and Solís Phyllophaga (Phyllophaga) lorencita Morón and Solís, 2001: 17 Distribution: Costa Rica

Phyllophaga lota Luginbill Phyllophaga lota Luginbill, 1928: 87 Distribution: United States

Phyllophaga luctuosa (Horn) Lachnosterna luctuosa Horn, 1887b: 254 Phyllophaga luctuosa (Horn); Glasgow, 1916: 373

Synonym: Lachnosterna rugosioides Linell, 1896: 728 Phyllophaga rugosioides (Linell); Glasgow, 1916: 374

Phyllophaga luginbilli Saylor Phyllophaga (Listrochelus) luginbilli Saylor, 1941c: 145 Distribution: United States

Phyllophaga lulaana Morón Phyllophaga (Phytalus) lulaana Morón, 2000: 43 Distribution: México

Phyllophaga luridipennis (Moser) Phytalus luridipennis Moser, 1918a: 68 Phyllophaga luridipennis (Moser); Blackwelder, 1944: 225 Phyllophaga (Phytalus) luridipennis (Moser); Frey, 1975a: 224 Distribution: Colombia

Synonym: Phyllophaga (Phytalus) penai Frey, 1964c: 695

Phyllophaga macasana (Moser) Lachnosterna macasana Moser, 1921c: 257 95 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga macasana (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) macasana (Moser); Frey, 1975a; 211 Distribution: Ecuador

Phyllophaga macgregori Morón Phyllophaga (Listrochelus) macgregori Morón, 2004: 90 Distribution: México

Phyllophaga macmurryi Saylor Phyllophaga (Listrochelus) macmurryi Saylor, 1940d: 110 Distribution: United States

Phyllophaga macrocera (Bates) Phytalus macrocerus Bates, 1888: 179 Phyllophaga macrocerus [sic] (Bates): Blackwelder, 1944: 225 Phyllophaga (Phytalus) macrocera (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga macrophylla (Bates) Lachnosterna macrophylla Bates, 1888: 193 Phyllophaga macrophylla (Bates): Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) macrophylla (Bates); Morón, 1986: 202 Distribution: México

Phyllophaga maculicollis (LeConte) Lachnosterna maculicollis LeConte, 1863b: 76 Phyllophaga maculicollis (LeConte); Glasgow, 1916; 373 Distribution: México

Synonym: Lachnosterna nitidula LeConte, 1863b: 77 Phyllophaga nitidula (LeConte); Glasgow, 1916; 373

Phyllophaga maestrensis Garcia-Vidal Phyllophaga (Cnemarachis) maestrensis Garcia-Vidal, 1984: 323 Distribution: Cuba

Phyllophaga magnicornis (Moser) Lachnosterna magnicornis Moser, 1921c: 253 Phyllophaga magnicornis (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) magnicornis (Moser); Morón, 1986: 205 Distribution: México

Phyllophaga mali Wolcott Phyllophaga mali Wolcott, 1928: 25 Cnemarachis mali (Wolcott); Blackwelder, 1944: 223 Distribution: Haiti

Phyllophaga manantleca Morón, Rivera, and López 96 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phyllophaga) manantleca Morón, Rivera, and López, 2002: 489 Distribution: México

Phyllophaga manaosana (Moser) Lachnosterna manaosana Moser, 1924b: 157 Phyllophaga manaosana (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) manaosana (Moser); Frey, 1975a: 219 Distribution: Brazil

Phyllophaga manchesterea Saylor Phyllophaga (Phyllophaga) manchesterea Saylor, 1940e: 313 Cnemarachis manchesterea (Saylor); Blackwelder, 1944: 223 Distribution: Jamaica

Phyllophaga mandevillea Saylor Phyllophaga (Phyllophaga) mandevillea Saylor, 1940e: 312 Cnemarachis mandevillea Saylor; Blackwelder, 1944: 223 Distribution: Jamaica

Phyllophaga marcano Woodruff Phyllophaga marcano Woodruff, 2005: 97 Distribution: Dominican Republic

Phyllophaga marcapatana (Moser) Phytalus marcapatanus Moser, 1918a: 69 Phyllophaga marcapatana (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) marcapatana (Moser); Frey, 1975a: 206 Distribution: Peru

Phyllophaga marginalis insolita Cartwright Phyllophaga marginalis var. insolita Cartwright, 1944: 33 Distribution: United States

Phyllophaga marginalis marginalis (LeConte) Lachnosterna marginalis LeConte, 1856: 250 Phyllophaga marginalis (LeConte); Glasgow, 1916: 372 Distribution: Canada, United States United States Phyllophaga mariaegalante Chalumeau and Gruner Phyllophaga (Cnemarachis) mariaegalante Chalumeau and Gruner, 1976: 90 Distribution: Guadeloupe

Phyllophaga mariana Fall Phyllophaga mariana Fall, 1929a: 111 Distribution: United States

Phyllophaga marilucasana Cano and Morón Phyllophaga (Chlaenobia) marilucasana Cano and Morón, 2002: 358 97 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Guatemala

Phyllophaga martiana Saylor Phyllophaga (Phyllophaga) martiana Saylor, 1943c: 30 Distribution: México

Phyllophaga marina Garcia-Vidal Phyllophaga marinus Garcia-Vidal, 1987: 303 Distribution: Cuba

Phyllophaga martinezi Frey Phyllophaga (Phyllophaga) martinezi Frey, 1975a: 207 Distribution: Colombia

Phyllophaga martinezpalaciosi Morón Phyllophaga (Phyllophaga) martinezpalaciosi Morón, 1989: 74 Distribution: México

Phyllophaga matacapana Morón Phyllophaga (Phyllophaga) matacapana Morón, 2003a: 266 Distribution: México

Phyllophaga maxima (Bates) Listrochelus maximus Bates, 1888: 172 Phyllophaga maxima (Bates): Blackwelder, 1944: 225 Phyllophaga (Listrochelus) maxima (Bates); Morón, 1986: 202 Distribution: México

Phyllophaga meadei Saylor Phyllophaga (Listrochelus) meadei Saylor, 1940d: 105 Distribution: México

Phyllophaga mella Sanderson Phyllophaga (Cnemarachis) mella Sanderson, 1951: 278 Distribution: Dominican Republic

Phyllophaga menetriesii (Blanchard) Ancylonycha menetriesii Blanchard, 1851: 135 Phyllophaga setifera var. mentriesi (Blanchard); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) mentriesi (Blanchard); Frey, 1975a: 202 Distribution: Colombia, Costa Rica, Guatemala, México, Panama

Phyllophaga mentalis Saylor Phyllophaga (Phyllophaga) mentalis Saylor, 1941d: 387 Distribution: Guatemala

Phyllophaga mesophylla Morón and Rivera Phyllophaga (Phytalus) mesophylla Morón and Rivera, 1992: 80 98 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Phyllophaga mexicana (Blanchard) Ancylonycha mexicana Blanchard, 1851: 138 Lachnosterna mexicana (Blanchard); Bates, 1888: 212 Phyllophaga mexicana (Blanchard); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) mexicana (Blanchard); Morón, 1986: 205 Distribution: México

Phyllophaga micans (Knoch) Melolontha micans Knoch, 1801: 77 Ancylonycha micans (Knoch); Blanchard, 1851: 138 Lachnosterna micans (Knoch); LeConte, 1856: 247 Phyllophaga micans (Knoch); Glasgow, 1916: 371 Distribution: United States

Phyllophaga michelbacheri Saylor Phyllophaga (Listrochelus) michelbacheri Saylor, 1940d: 76 Distribution: México

Phyllophaga microcera (Arrow) Lachnosterna microcerus Arrow, 1933b: 149 (for Bates, 1888: 189) Phyllophaga microcerus (Arrow); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) microcerus (Arrow); Morón, 1986: 203 Distribution: México

Synonym: Lachnosterna cribricollis Bates, 1888: 189 (not Blanchard, 1851: 133)

Phyllophaga microchaeta (Moser) Lachnosteran microchaeta Moser, 1924b: 154 Phyllophaga microchaeta (Moser); Blackwelder, 1944: 225 Distribution: México

Phyllophaga microdon (Bates) Lachnosterna microdon Bates, 1888: 211 Phyllophaga microdon (Bates); Fall, 1929a: 113 Phyllophaga (Phyllophaga) microdon (Bates); Morón, 1986: 204 Distribution: México

Phyllophaga microphylla (Moser) Lachnosterna microphylla Moser, 1918a: 60 Cnemarachis microphylla (Moser); Blackwelder, 1944: 223 Phyllophaga microphylla (Moser); Chalumeau, 1985: 26, 30 Distribution: St. Croix

Phyllophaga micros (Bates) Listrochelus micros Bates, 1888: 170 Phyllophaga (Listrochelus) micros (Bates); Saylor, 1940d: 104 99 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México, United States

Phyllophaga microsoma Chapin Phyllophaga microsoma Chapin, 1932b: 202 Cnemarachis microsoma (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga migratoria Garcia-Vidal Phyllophaga migratoria Garcia-Vidal, 1984: 324 Distribution: Cuba

Phyllophaga mimicana Saylor Phyllophaga (Listrochelus) mimicana Saylor, 1938e: 186 Distribution: México, United States

Phyllophaga minutissima Wolcott Phyllophaga minutissima Wolcott, 1928: 28 Cnemarachis minutissima Wolcott; Blackwelder, 1944: 223 Distribution: Haiti

Phyllophaga miraflora Saylor Phyllophaga (Listrochelus) miraflora Saylor, 1940d: 72 Distribution: México

Phyllophaga misteca (Bates) Lachnosteran misteca Bates, 1888: 209 Phyllophaga misteca (Bates); Blackwelder, 1944; 225 Phyllophaga (Phyllophaga) misteca (Bates); Morón, 1986: 204 Distribution: México

Phyllophaga mitlana Saylor Phyllophaga (Phyllophaga) mitlana Saylor, 1943e: 278 Distribution: México

Phyllophaga moei Saylor Phyllophaga moei Saylor, 1938f: 343 Phyllophaga (Phyllophaga) moei Saylor; Morón, 1986: 203 Distribution: México

Phyllophaga molopia (Bates) Lachnosterna molopia Bates, 1888: 205 Phyllophaga molopia (Bates); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) molopia (Bates); Morón, 1986: 205 Distribution: México

Phyllophaga monana (Moser) Lachnosterna monana Moser, 1921b: 181 Cnemarachis monana (Moser); Blackwelder, 1944: 223 100 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga monana (Moser); Chalumeau, 1985: 24 Distribution: Puerto Rico [Mona Island]

Phyllophaga monrosi Frey Phyllophaga (Phyllophaga) monrosi Frey, 1975a: 219 Distribution: Argentina

Phyllophaga monstrosa (Saylor) Listrochelus monstrosus Saylor, 1935g: 36 Phyllophaga monstrosa (Saylor); Blackwelder, 1944: 225 Phyllophaga (Listrochelus) monstrosa (Saylor); Morón, 1986: 266 Distribution: México

Phyllophaga monteverdosa Morón Phyllophaga (Phyllophaga) monteverdosa Morón, 2001b: 789 Distribution: Costa Rica

Phyllophaga montserratensis (Arrow) Lachnosterna montserratensis Arrow, 1920: 191 Cnemarachis montserratensis (Arrow); Blackwelder, 1944: 223 Phyllophaga montserratensis (Arrow); Chalumeau, 1985: 30 Distribution: Montserrat

Phyllophaga morganella Saylor Phyllophaga (Phyllophaga) morganella Saylor, 1942a: 158 Distribution: Costa Rica, Panama

Phyllophaga morgani Saylor Phyllophaga morgani Saylor, 1938f: 340 Distribution: Panama

Phyllophaga moseri Frey Phyllophaga (Phytalus) moseri Frey, 1965b: 149 Distribution: Bolivia

Phyllophaga mucorea (LeConte) Listrochelus mucoreus LeConte, 1856: 263 Phyllophaga mucorea (LeConte); Blackwelder, 1944: 225 Phyllophaga (Listrochelus) mucorea (LeConte); Saylor, 1940c: 86 Distribution: México, United States

Synonym: Listrochelus obtusus LeConte, 1856: 264 Phyllophaga obtusus (LeConte); Evans, 2003: 127

Synonym: Listrochelus texanus LeConte, 1856: 263 Phyllophaga texanus (LeConte); Evans, 2003: 127

Phyllophaga multipora (Bates) 101 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Lachnosterna multipora Bates, 1888: 211 Phyllophaga (Phyllophaga) multipora (Bates); Sanderson, 1958: 165 Distribution: Guatemala, México

Phyllophaga murina (Burmeister) Ancylonycha murina Burmeister, 1855: 334 Phyllophaga murina (Burmeister); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) murina (Burmeister); Frey, 1975a: 203 Distribution: Brazil

Phyllophaga nahui Morón and Solís Phyllophaga (Chirodines) nahui Morón and Solís, 1994: 182 Distribution: Costa Rica

Phyllophaga naranjina Morón and Solís Phyllophaga (Phyllophaga) naranjina Morón and Solís, 2001: 24 Distribution: Costa Rica

Phyllophaga nasalis Chapin Phyllophaga nasalis Chapin, 1935b: 72 Cnemarachis nasalis (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga necaxa Saylor Phyllophaga (Phyllophaga) necaxa Saylor, 1943e: 279 Distribution: México

Phyllophaga neglecta (Blanchard) Ancylonycha neglecta Blanchard, 1851: 137 Phyllophaga neglecta (Blanchard); Wolcott, 1928: 27 Cnemarachis neglecta (Blanchard); Blackwelder 1944: 223 Distribution: Dominican Republic, Haiti

Phyllophaga neomexicana Saylor Phyllophaga (Listrochelus) neomexicana Saylor, 1940d: 101 Distribution: United States

Phyllophaga nepida Saylor Phyllophaga (Phyllophaga) nepida Saylor, 1941d: 385 Distribution: Guatemala

Phyllophaga nevermannea Saylor Phyllophaga (Phyllophaga) nevermannea Saylor, 1941e: 534 Distribution: Costa Rica

Phyllophaga nevermanni Saylor Phyllophaga nevermanni Saylor, 1935h: 501 Phyllophaga (Phyllophaga) nevermanni Saylor; Solís and Morón, 1998: 25 102 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Costa Rica

Phyllophaga nigerrima (Bates) Lachnosterna nigerrima Bates, 1888: 207 Phyllophaga nigerrima (Bates): Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) nigerrima (Bates): Morón, 1986: 205 Distribution: México

Phyllophaga nigrita (Moser) Lachnosterna nigrita Moser, 1918a: 37 Phyllophaga nigrita (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) nigrita (Moser); Solís and Morón, 1998: 24 Distribution: Costa Rica, Nicaragua, Panama

Phyllophaga niquirana Morón Phyllophaga (Listrochelus) niquirana Morón, 1990: 59 Distribution: Costa Rica, Nicaragua

Phyllophaga nisuens (Saylor) Lachnosterna nisuens Saylor, 1937c: 32 Phyllophaga nisuens (Saylor); Blackwelder, 1944:225 Phyllophaga (Phyllophaga) nisuens (Saylor); Morón, 2001a: 182 Distribution: México

Phyllophaga nitida (LeConte) Lachnosterna nitida LeConte, 1856: 256 Phyllophaga nitida (LeConte); Glasgow, 1916: 372 Distribution: Canada, United States

Synonym: Lachnosterna innominata Smith, 1889a: 98

Synonym: Lachnosterna limula Horn, 1887b: 264

Phyllophaga nitidicauda (Arrow) Lachnosterna nitidicauda Arrow, 1913c: 464 Phyllophaga nitidicauda (Arrow); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) nitidicauda (Arrow); Frey, 1975a: 217 Distribution: Brazil

Phyllophaga nitidicollis (Blanchard) Ancylonycha nitidicollis Blanchard, 1851: 136 Phyllophaga nitidicollis (Blanchard); Evans, 2003: 136 Distribution: Brazil

Phyllophaga nitididorsis Frey Phyllophaga (Phytalus) nitididorsis Frey, 1975a: 226 Distribution: Brazil

103 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga nogales Saylor Phyllophaga (Listrochelus) falsa nogales Saylor, 1940d: 99 Phyllophaga (Listrochelus) nogales Saylor; Sanderson, 1958: 161 Distribution: United States

Phyllophaga nogueirai Warner and Morón Phyllophaga (Triodonyx) nogueirai Warner and Morón, 1992: 329 Distribution: México

Phyllophaga nogueirana Morón Phyllophaga (Listrochelus) nogueirana Morón, 2002: 455 Distribution: México

Phyllophaga nosa Blackwelder Phyllophaga nosa Blackwelder, 1944: 225 (for Bates, 1888: 182) Phyllophaga (Phytalus) nosa Blackwelder; Morón, 1986: 219 Distribution: México, United States

Synonym: Phytalus aegrotus Bates, 1888: 182 (not Bates, 1888: 167) Phyllophaga (Phytalus) aegrota (Bates); Sanderson, 1942: 54

Phyllophaga nubipennis (Bates) Phytalus nubipennis Bates, 1888: 176 Phyllophaga nubipennis (Bates); Blackwelder, 1944: 225 Phyllophaga (Phytalus) nubipennis (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga nuda (Moser) Lachnosterna nuda Moser, 1918a: 28 Phyllophaga nuda (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) nuda (Moser); Frey, 1975a: 212 Distribution: Guyana

Phyllophaga nudipennis Frey Phyllophaga (Phyllophaga) nudipennis Frey, 1975a: 215 Distribution: Ecuador

Phyllophaga nunezi Woodruff Phyllophaga nunezi Woodruff, 2005: 110 Distribution: Dominican Republic

Phyllophaga oaxaca Saylor Phyllophaga (Phytalus) oaxaca Saylor, 1940e: 116 Distributon: México

Phyllophaga oaxena Saylor Phyllophaga (Chirodines) oaxena Saylor, 1940e: 109 Phyllophaga (Listrochelus) oaxena Saylor; Blackwelder, 1944: 225 104 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phytalus) oaxena Saylor; Morón, 1986: 200 Distribution: México

Phyllophaga obliquestriata Saylor Phyllophaga (Listrochelus) obliquestriata Saylor, 1938e: 187 Distribution: México

Phyllophaga oblongula (Bates) Listrochelus oblongulus Bates 1888: 173 Phyllophaga oblongula (Bates); Blackwelder, 1944: 225 Phyllophaga (Listrochelus) oblongula (Bates); Morón, 1986: 202 Distribution: México

Phyllophaga obsoleta (Blanchard) Phytalus obsoletus Blanchard, 1851: 131 Phyllophaga obsoletus (Blanchard); Blackwelder, 1944: 225 Phyllophaga (Phytalus) obsoleta (Blanchard); Sanderson, 1958: 166 Distribution: Belize, Colombia, Costa Rica, El Salvador, Guatemala, Honduras, México, Nicaragua, United States, Venezuela.

Synonym: Phytalus laevigatus Blanchard, 1851: 131 (not Burmeister, 1855: 351) Phyllophaga laevigata (Blanchard); Blackwelder, 1944: 225

Synonym: Ancylonycha longicornis Burmeister, 1855: 329 (not Blanchard, 1851: 134)

Synonym: Phytalus longicornis Burmeister, 1855: 350 (not Blanchard, 1851: 134)

Synonym: Phyllophaga mecocerus Blackwelder, 1944: 225 (for Burmeister, 1855: 329)

Synonym: Phytalus vanalleri Schaeffer, 1927: 215 Phyllophaga (Phytalus) obsoleta vanalleri (Schaeffer); Saylor, 1939c: 164

Synonym: Phytalus oxypygus Burmeister, 1855: 350 Phyllophaga oxypyga (Burmeister); Blackwelder, 1944: 225

Phyllophaga ocozocuana Morón Phyllophaga (Phyllophaga) ocozocuana Morón, 2003a: 279 Distribution: México

Phyllophaga odomi Saylor Phyllophaga (Phyllophaga) odomi Saylor, 1943b: 107 Distribution: Panama, Venezuela

Phyllophaga ohausi (Moser) Phytalus ohausi Moser, 1921c: 260 Phyllophaga ohausi (Moser); Blackwelder, 1944: 225 Phyllophaga (Phytalus) ohausi (Moser); Frey, 1975a: 222 Distribution: Ecuador 105 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga okeechobea Robinson Phyllophaga okeechobea Robinson, 1948: 33 Distribution: United States

Phyllophaga omani Sanderson Phyllophaga (Phytalus) omani Sanderson, 1937b: 66 Phyllophaga (Phyllophaga) omani Sanderson; Sanderson, 1958: 166 Distribution: United States

Phyllophaga ome Morón Phyllophaga (Chirodines) ome Morón, 1991: 400 Distribution: México

Phyllophaga omiltemia (Bates) Phytalus omiltemius Bates, 1889: 400 Phyllophaga (Phytalus) omiltemia (Bates); Morón, 1986: 200 Distribution: Guatemala, México

Phyllophaga onchophora Chapin Phyllophaga onchophora Chapin, 1932: 184 Cnemarachis onchophora (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga onita Saylor Phyllophaga (Phyllophaga) onita Saylor, 1941a: 27 Distribution: México

Phyllophaga onoreana Morón Phyllophaga (Phyllophaga) onoreana Morón, 2003a: 229 Distribution: Ecuador

Phyllophaga opaca (Moser) Lachnosterna opaca Moser, 1918b: 164 Phyllophaga opaca (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) opaca (Moser); Sanderson, 1958: 172 Distribution: México, United States

Synonym: Phyllophaga iroides Fall, 1929a: 112

Phyllophaga opacicollis (Horn) Listrochelus opacicollis Horn, 1878: 145 Phyllophaga (Listrochelus) opacicollis (Horn); Saylor, 1940d: 92 Distribution: United States

Phyllophaga orosina (Moser) Lachnosterna orosina Moser, 1918a: 26 Phyllophaga orosina (Moser): Blackwelder, 1944: 225 106 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phyllophaga) orosina (Moser); Solís and Morón, 1998: 23 Distribution: Costa Rica

Phyllophaga ortizi Woodruff Phyllophaga ortizi Woodruff, 2005: 113 Distribution: Dominican Republic

Phyllophaga ovalis Cartwright Phyllophaga ovalis Cartwright, 1939: 353 Distribution: United States

Phyllophaga pachuca Saylor Phyllophaga (Phyllophaga) pachuca Saylor, 1943e: 272 Distribution: México

Phyllophaga pachypyga (Burmeister) Ancylonycha pachypyga Buremeister, 1855: 330 Phyllophaga pachypyga (Burmeister); Blackwelder, 1944; 225 Phyllophaga (Phyllophaga) pachypyga (Burmeister); Frey, 1975a: 214 Distribution: Brazil, Colombia, Peru, Venezuela

Synonym: Lachnosterna nigrofusca Moser, 1918a: 39 Phyllophaga nigrofusca (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) nigrofusca (Moser); Frey, 1975a: 214

Phyllophaga pallida (Horn) Phytalus pallidus Horn, 1885: 121 Phyllophaga (Phytalus) pallidus (Horn); Saylor, 1939c: 166 Phyllophaga (Phyllophaga) pallidus (Horn); Sanderson, 1958: 166 Distribution: México, United States

Phyllophaga pallidicornis (Moser) Lachnosterna pallidicornis Moser, 1921c: 251 Phyllophaga pallidicornis (Moser); Blackwelder, 1944: 225 Distribution: México

Phyllophaga pameana Morón Phyllophaga (Phytalus) pameana Morón, 2000: 48 Distribution: México

Phyllophaga panamana (Chapin) Chlaenobia panama Chapin, 1935a: 11 Phyllophaga panamana (Chapin); Blackwelder, 1944: 225 Phyllophaga (Chlaenobia) panamana (Chapin); Morón, 1986: 209 Distribution: Costa Rica, Panama

Phyllophaga panamensis (Moser) Lachnosterna panamensis Moser, 1921c: 250 107 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga panamensis (Moser); Blackwelder, 1944: 225 Distribution: Panama

Phyllophaga panicula Sanderson Phyllophaga (Cnemarachis) panicula Sanderson, 1951: 275 Distribution: Dominican Republic, Haiti

Phyllophaga panorpa Sanderson Phyllophaga panorpa Sanderson, 1950: 91 Distribution: United States

Phyllophaga papaloana Morón Phyllophaga (Phyllophaga) papaloana Morón, 2001a: 187 Distribution: México

Phyllophaga paraguayana (Moser) Lachnosterna paraguayana Moser, 1921c: 256 Phyllophaga paraguayana (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) paraguayana (Moser); Frey, 1975a: 207 Distribution: Paraguay

Phyllophaga paraguayensis (Moser) Phytalus paraguayensis Moser, 1918a: 74 Phyllophaga paraguayensis (Moser); Blackwelder, 1944: 225 Phyllophaga (Phytalus) paraguayensis (Moser); Frey, 1975a: 225 Distribution: Paraguay

Phyllophaga parallela (Blanchard) Ancylonycha parallela Blanchard, 1851: 137 Phyllophaga parallela (Blanchard); Chapin, 1932b; 205 Cnemarachis parallela (Blanchard); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga parcesetifera Chapin Phyllophaga parcesetifera Chapin, 1935b: 71 Cnemarachis parcesetifera (Chapin); Blackwelder, 1944: 223 Phyllophaga parcesetifera (Blanchard); Evans, 2003: 134 Distribution: Cuba

Phyllophaga parilis (Bates) Listrochelus parilis Bates, 1888: 172 Phyllophaga (Listrochelus) parilis (Bates); Saylor, 1940d: 101 Distribution: México, United States

Phyllophaga parumpunctata (Bates) Lachnosterna parumpunctata Bates, 1888: 191 Phyllophaga parumpunctata Bates; Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) parumpunctata (Bates); Morón, 1986: 203 108 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Phyllophaga parvicornis (Moser) Lachnosterna parvicornis Moser, 1924b: 155 Phyllophaga parvicornis (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) parvicornis (Moser); Morón, 1986: 195 Distribution: Guatemala

Phyllophaga parvidens hysteropyga Davis Phyllophaga parvidens var. hysteropyga Davis 1920: 336 Distribution: United States

Phyllophaga parvidens parvidens (LeConte) Lachnosterna parvidens LeConte, 1856: 259 Phyllophaga parvidens (LeConte); Glasgow, 1916: 373 Distribution: United States

Phyllophaga parvisetis (Bates) Lachnosterna parvisetis Bates, 1888: 196 Phyllophaga (Phyllophaga) parvisetis (Bates); Saylor, 1943d: 134 Distribution: Guatemala, Honduras, México, Nicaragua

Phyllophaga pastassana (Moser) Lachnosterna pastassana Moser, 1924b: 156 Phyllophaga pastassana (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) pastassana (Moser) Frey, 1975a: 214 Distribution: Ecuador

Phyllophaga paternoi Glasgow Phyllophaga paternoi Glasgow, 1925: 293 Distribution: United States

Phyllophaga patruelis (Chevrolat) Ancylonycha patruelis Chevrolat, 1865: 25 Phyllophaga patruelis (Chevrolat); Chapin, 1932b: 191 Cnemarachis patruelis (Chevrolat); Blackwelder, 1944: 223 Distribution: Cuba, Dominican Republic, Guadeloupe, Haiti, St. Kitts-Nevis

Synonym: Phyllophaga tenuicalcaris Chapin, 1932b: 192

Phyllophaga patrueloides Paulian Phyllophaga patrueloides Paulian, 1947: 56 (for Fleutiaux and Sallé, 1889: 49) Phyllophaga (Cnemarachis) patrueloides Paulian; Chalumeau and Gruner, 1976: 91 Distribution: Guadeloupe

Synonym: Phyllophaga patruelis Fleutiaux and Sallé, 1889: 49 (not Chevrolat, 1865: 25)

Phyllophaga pauliani Chalumeau and Gruner 109 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Cnemarachis) pauliani Chalumeau and Gruner, 1976: 95 Distribution: Guadeloupe

Phyllophaga pauloensis Frey Phyllophaga (Phytalus) pauloensis Frey, 1975a: 224 Distribution: Brazil

Phyllophaga pearliae Davis Phyllophaga pearliae Davis, 1920: 332 Distribution: United States

Phyllophaga peccata Blackwelder Phyllophaga peccatus Blackwelder, 1944: 225 (for Arrow, 1933b: 145) Phyllophaga (Chlaenobia) peccata Blackwelder; Morón, 1986: 209 Distribution: México

Synonym: Chlaenobia colimana Arrow, 1933b: 145 (not Moser, 1921c: 256)

Phyllophaga pectoralis (Blanchard) Ancylonycha pectoralis Blanchard, 1851: 138 Phyllophaga pectoralis (Blanchard); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) pectoralis (Blanchard); Frey, 1975a; 206 Distribution: Brazil

Phyllophaga penaella Frey Phyllophaga (Phyllophaga) penaella Frey, 1975a: 209 Distribution: Ecuador

Phyllophaga peninsulana (Moser) Lachnosterna peninsulana Moser, 1918a: 55 Phyllophaga peninsulana (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) peninsulana (Moser); Morón, 1986: 205 Distribution: México

Phyllophaga peninsularis Saylor Phyllophaga (Listrochelus) peninsularis Saylor, 1940d: 77 Distribution: México

Phyllophaga pentaphylla (Bates) Phytalus pentaphyllus Bates, 1888: 180 Phyllophaga pentaphylla (Bates); Blackwelder, 1944: 225 Phyllophaga (Phytalus) pentaphylla (Bates); Morón, 1982: 18 Distribution: México

Phyllophaga perdernales Woodruff Phyllophaga perdernales Woodruff, 2005: 117 Distribution: Dominican Republic

110 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga perlonga Davis Phyllophaga perlonga Davis, 1920: 329 Distribution: United States

Phyllophaga permagna (Moser) Lachnosterna permagna Moser, 1918a: 57 Phyllophaga permagna (Moser); Blackwelder, 1944: 224 Phyllophaga permagna (Moser); Sanderson, 1951: 259 Distribution: Dominican Republic, Haiti

Phyllophaga persimilis Chapin Phyllophaga persimilis Chapin, 1935b: 70 Cnemarachis persimilis (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga personata (Chapin) Chlaenobia personata Chapin, 1935a: 12 Phyllophaga personata (Chapin); Blackwelder, 1944: 225 Phyllophaga (Chlaenobia) personata (Chapin); Morón, 1986: 199 Distribution: México

Phyllophaga peruana (Moser) Lachnosterna peruana Moser, 1918a: 48 Phyllophaga peruana (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) peruana (Moser): Frey, 1975a: 217 Distribution: Bolivia, Peru

Synonym: Phyllophaga (Phyllophaga) limasa Saylor, 1948b: 353

Phyllophaga picadoana Morón and Solís Phyllophaga (Phyllophaga) picadoana Morón and Solís, 2000: 906 Distribution: Costa Rica

Phyllophaga picea (Blanchard) Phytalus piceus Blanchard, 1851: 130 Phyllophaga picea (Blanchard); Blackwelder, 1944: 225 Phyllophaga (Phytalus) picea (Blanchard): Frey, 1975a: 223 Distribution: Brazil

Phyllophaga piceola (Bates) Lachnosterna piceola Bates, 1888: 187 Phyllophaga piceola (Bates); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) piceola (Bates); Cano and Morón, 1998: 14 Distribution: Guatemala

Phyllophaga picipennis (Moser) Phytalus picipennis Moser, 1918a: 73 Phyllophaga picipennis (Moser); Blackwelder, 1944: 225 111 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phytalus) picipennis (Moser); Frey, 1975a: 225 Distribution: Bolivia

Phyllophaga piligera (Moser) Lachnosterna piligera Moser, 1918a: 30 Phyllophaga piligera (Moser); Blackwelder, 1944: 225 Distribution: Panama

Phyllophaga piliventris (Moser) Lachnosterna piliventris Moser, 1918a: 20 Phyllophaga piliventris (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) piliventris (Moser); Morón, 1986: 203 Distribution: México

Phyllophaga pilosipes Saylor Phyllophaga (Listrochelus) pilosipes Saylor, 1940d: 76 (for LeConte, 1863b: 78) Distribution: México

Synonym: Listrochelus puberulus LeConte, 1863b: 78 (not Jacquelin du Val, 1856: 130)

Phyllophaga pilositarsis Blackwelder Phyllophaga pilositarsis Blackwelder, 1944: 225 (for Chapin, 1935a: 16) Phyllophaga (Chlaenobia) pilositarsis Blackwelder; Morón, 1986: 209 Distribution: México

Synonym: Chlaenobia dissimilis Chapin, 1935a: 16 (not Chevrolat, 1865: 26)

Phyllophaga pilosula (Moser) Lachnosterna pilosula Moser, 1918a: 32 Phyllophaga pilosula (Moser); Blackwelder, 1944: 225 Phyllophaga (Phyllophaga) pilosula (Moser); Morón, 1986: 204 Distribution: México

Phyllophaga pilotoensis Garcia-Vidal Phyllophaga pilotoensis Garcia-Vidal, 1987: 304 Distributon: Cuba

Phyllophaga pilula (Moser) Lachnosterna pilula Moser, 1921c: 250 Phyllophaga pilula (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) pilula (Moser); Morón, 1986: 205 Distribution: México

Phyllophaga plaei (Blanchard) Ancylonycha plaei Blanchard, 1851: 137 Ancylonycha pleei Blanchard; Lacordaire, 1856: 286 [unjustified emendation] Cnemarachis plaei (Blanchard); Blackwelder, 1944: 223 Phyllophaga pleei [sic] (Blanchard); Paulian, 1947: 55 112 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Cnemarachis) pleei [sic] (Blanchard); Chalumeau and Gruner, 1976: 95 Distribution: Puerto Rico

Synonym: Phyllophaga borinquensis Blackwelder, 1944: 224 (for Moser, 1918a: 62)

Synonym: Lachnosterna portoricensis Moser, 1918a: 62 (not Smyth, 1917: 145)

Synonym: Phyllophaga portoricensis Smyth, 1917: 145 Cnemarachis portoricensis (Smyth); Saylor, 1942a: plate 17

Phyllophaga plairi (Saylor) Lachnosterna plairi (Saylor, 1937d: 31 Phyllophaga plairi (Saylor); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) plairi (Saylor); Morón, 1986: 204 Distribution: México

Phyllophaga planeta Reinhard Phyllophaga (Listrochelus) planeta Reinhard, 1950: 30 Distribution: United States

Phyllophaga platti Saylor Phyllophaga (Phytalus) platti Saylor, 1935h: 499 Distribution: México

Phyllophaga platyrhina (Bates) Phytalus platyrhinus Bates, 1888: 179 Phyllophaga platyrhina (Bates); Blackwelder, 1944: 226 Phyllophaga (Phytalus) platyrhinus (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga plena (Fall) Listrochelus plenus Fall, 1932: 199 Phyllophaga (Listrochelus) plena (Fall): Saylor, 1940d: 90 Distribution: México, United States

Phyllophaga pleroma Reinhard Phyllophaga pleroma Reinhard, 1940: 5 (for Reinhard, 1939: 49) Distribution: United States

Synonym: Phyllophaga plena Reinhard, 1939: 49 (not Fall, 1932: 199)

Phyllophaga poculifer (Bates) Phytalus poculifer Bates, 1888: 184 Phyllophaga poculifer (Bates); Blackwelder, 1944: 226 Phyllophaga (Phytalus) poculifer (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga pokornyiana Morón 113 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Phyllophaga) pokornyiana Morón, 1996a: 76 Distributon: México

Phyllophaga polita (Harold) Phytalus politus Harold, 1869b: 123 (for Burmeister, 1855: 351) Phyllophaga polita (Harold); Blackwelder, 1944: 226 Distribution: Brazil

Synonym: Phytalus laevigatus Burmeister, 1855: 351 (not Blanchard, 1850: 131) Phyllophaga (Phytalus) laevigatus (Burmeister); Frey, 1975a: 224

Phyllophaga polyphylla (Bates) Lachnosterna polyphylla Bates, 1888: 194 Phyllophaga polyphylla (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) polyphylla (Bates); Morón, 1986: 202 Distribution: México

Synonym: Lachnosterna heterophylla Bates, 1888: 194 Phyllophaga heterophylla (Bates); Blackwelder, 1944: 225

Phyllophaga porodera (Bates) Lachnosterna porodera Bates, 1888: 211 Phyllophaga porodera (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) porodera (Bates); Morón, 1986: 205 Distribution: México

Phyllophaga postrema (Horn) Lachnosterna postrema Horn, 1887b: 233 Phyllophaga postrema (Horn); Glasgow, 1916: 373 Distribution: United States

Synonym: Lachnosterna quadrata Smith, 1889a: 94 Phyllophaga quadrata (Smith); Glasgow, 1916: 373

Phyllophaga potrerillo Garcia-Vidal Phyllophaga potrerillo Garcia-Vidal, 1987: 304 Distribution: Cuba

Phyllophaga praesidii (Bates) Listrochelus praesidii Bates, 1888: 171 Phyllophaga praesidii (Bates); Blackwelder, 1944: 226 Phyllophaga (Listrochelus) praesidii (Bates); Morón, 1986: 201 Distribution: México

Phyllophaga praetermissa (Horn) Lachnosterna praetermissa Horn 1887b: 223 Phyllophaga praetermissa (Horn); Glasgow, 1916: 373 Distribution: United States 114 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Lachnosterna definita Smith, 1889b: 501 (for Horn, 1887b: 243)

Phyllophaga probaporra Sanderson Phyllophaga probaporra Sanderson, 1951: 263 Distribution: Dominican Republic

Phyllophaga profunda (Blanchard) Ancylonycha profunda Blanchard, 1851: 132 Lachnosterna profunda (Blanchard); Horn, 1887b: 257 Phyllophaga profunda (Blanchard), Glasgow, 1916: 371 Distribution: United States

Synonym: Lachnosterna biimpressa Smith, 1889a: 97

Synonym: Lachnosterna grandior Linell, 1896: 727

Phyllophaga prolixa (Bates) Phytalus prolixus Bates, 1888: 178 Phyllophaga prolixa (Bates): Blackwelder, 1944; 226 Phyllophaga (Phytalus) prolixa (Bates); Solís and Morón, 1998: 22 Distribution: Costa Rica, Nicaragua

Phyllophaga propinqua (Moser) Phytalus propinquus Moser, 1918a: 67 Phyllophaga propinqua (Moser); Blackwelder, 1944: 226 Phyllophaga (Phytalus) propinqua (Moser); Frey, 1975a: 224 Distribution: Brazil, Paraguay

Phyllophaga pruinipennis (Moser) Lachnosterna pruinipennis Moser, 1918a: 40 Phyllophaga pruinipennis (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) pruinipennis (Moser); Frey, 1975a: 209 Distribution: Colombia

Phyllophaga pruinosa (Blanchard) Phytalus pruinosus Blanchard 1851: 131 Phyllophaga pruinosa (Blanchard); Blackwelder, 1944: 226 Phyllophaga (Phytalus) pruinosa (Blanchard); Frey, 1975a: 222 Distribution: Brazil, Costa Rica, México

Phyllophaga prunina (LeConte) Lachnosterna prunina LeConte, 1856: 251 (for Melsheimer, 1846: 139) Phyllophaga prunina (LeConte); Glasgow, 1916: 371 Distribution: United States

Synonym: Ancylonycha pruinosa Melsheimer, 1846: 139 (not Blanchard, 1851: 131)

115 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga prununculina (Burmeister) Trichestes prununculina Burmeister, 1855: 360 Lachnosterna prununculina (Burmeister); Horn, 1887b: 223 Phyllophaga prununculina (Burmeister), Glasgow, 1916: 372 Distribution: United States

Synonym: Lachnosterna cerasina LeConte, 1856: 241

Phyllophaga pseudoatra Morón Phyllophaga (Phyllophaga) pseudoatra Morón, 2003a: 246 Distribution: México

Phyllophaga pseudocalcaris Saylor Phyllophaga pseudocalcaris Saylor, 1940e: 309 Cnemarachis pseudocalcaris (Saylor); Blackwelder, 1944: 223 Phyllophaga pseudocalcaris Sanderson, 1951: 265 Distribution: Dominican Republic

Phyllophaga pseudocara Morón Phyllophaga (Phyllophaga) pseudocarus Morón, 1999: 380 Distribution: México

Phyllophaga pseudofloridana Woodruff and Beck Phyllophaga pseudofloridana Woodruff and Beck, 1989: 150 Distribution: United States

Phyllophaga pseudomicans Chapin Phyllophaga pseudomicans Chapin, 1935b: 73 Cnemarachis pseudomicans (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga psiloptera Sanderson Phyllophaga psiloptera Sanderson, 1939a: 7 Distribution: United States

Phyllophaga puberea (Mannerheim) Melolontha puberea Mannerheim, 1829: 63 Phytalus pubereus (Mannerheim); Burmeister, 1855: 351 Phyllophaga puberea (Mannerheim); Blackwelder, 1944: 226 Phyllophaga (Phytalus) puberea (Mannerheim); Frey, 1975a: 223 Distribution: Brazil

Phyllophaga puberula (Jacquelin du Val) Ancylonycha puberula Jacquelin du Val, 1856: 130 Phyllophaga puberula (Jacquelin du Val); Chapin, 1932b: 189 Cnemarachis puberula (Jacquelin du Val); Blackwelder, 1944: 223 Distribution: Cuba

116 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga pubescens (Burmeister) Ancylonycha pubescens Burmeister, 1855: 334 Phyllophaga pubescens (Burmeister); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) pubescens (Burmeister); Frey, 1975a; 208 Distribution: French Guiana

Phyllophaga pubicauda (Bates) Lachnosterna pubicauda Bates, 1888: 188 Phyllophaga pubicauda (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) pubicauda (Bates); Morón, 1986: 202 Distribution: México

Phyllophaga pubicollis (Blanchard) Phytalus pubicollis Blanchard, 1851: 131 Lachnosterna pubicollis (Blanchard); Bates, 1889: 400 Phyllophaga (Phyllophaga) pubicollis (Blanchard); Saylor, 1943b: 109 Distribution: Guatemala, México

Synonym: Lachnosterna heynei Moser, 1918b: 164 Phyllophaga heynei (Moser); Blackwelder, 1944: 225

Phyllophaga pudorosa Reinhard, 1939 Phyllophaga pudorosa Reinhard, 1939: 55 Distribution: United States

Phyllophaga puerilis (Wickham) Lachnosterna puerilis Wickham, 1914: 459 Phyllophaga puerilis (Wickham); Evans, 2003: 144 Distribution: United States Remark: fossil species

Phyllophaga pulcher (Linell) Listrochelus pulcher Linell, 1896: 730 Phyllophaga (Listrochelus) pulcher (Linell); Saylor, 1940d: 88 Distribution: United States

Phyllophaga punctipennis (Blanchard) Ancylonycha punctipennis Blanchard, 1851: 135 Phyllophaga punctipennis (Blanchard); Evans, 2003: 145 Distribution: ?South America

Synonym: Ancylonycha puncticollis Burmeister, 1855: 321

Phyllophaga punctulata (Blanchard) Ancylonycha punctulata Blanchard, 1851: 132 Phyllophaga punctulata (Blanchard); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) punctulata (Blanchard); Frey, 1975a: 214 Distribution: Colombia 117 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga punctuliceps (Bates) Phytalus punctuliceps Bates, 1888: 175 Phyllophaga punctuliceps Bates; Blackwelder, 1944: 226 Phyllophaga (Phytalus) punctuliceps Bates; Cano and Morón, 1998: 11 Distribution: Guatemala, México

Phyllophaga punctulicollis (Bates) Lachnosterna punctulicollis Bates, 1888: 191 Phyllophaga punctulicollis (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) punctulicollis (Bates); Morón, 1986: 203 Distribution: México

Phyllophaga puntarenosa Morón and Solís Phyllophaga (Phyllophaga) puntarenosa Morón and Solís, 2001: 19 Distribution: Costa Rica

Phyllophaga pusillidens Fall Phyllophaga pusillidens Fall, 1937: 31 (for Fall, 1929a: 113)

Synonym: Phyllophaga microdon Fall, 1929a: 113 (not Bates, 1888: 211)

Phyllophaga quadriphylla Saylor Phyllophaga (Phyllophaga) quadriphylla Saylor, 1943c: 29 Distribution: México

Phyllophaga quercus (Knoch) Melolontha quercus Knoch, 1801: 72 Ancylonycha quercus (Knoch); Burmeister, 1855: 340 Endrosa quercus (Knoch); LeConte, 1856: 235 Lachnosterna quercus (Knoch); Horn, 1887b: 281 Phyllophaga quercus (Knoch), Glasgow, 1916: 371 Distribution: United States

Phyllophaga quetzala Morón Phyllophaga (Phyllophaga) quetzala Morón, 2001a: 171 Distribution: México

Phyllophaga quetzaloides Morón Phyllophaga (Phyllophaga) quetzaloides Morón, 2001a: 174 Distribution: México

Phyllophaga quiana Morón Phyllophaga (Phyllophaga) quiana Morón, 2003a: 283 Distribution: México

Phyllophaga rangelana Chapin Phyllophaga rangelana Chapin, 1935b: 71 118 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Cnemarachis rangelana (Chapin): Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga ratcliffeiana Morón Phyllophaga (Chlaenobia) ratcliffeiana Morón, 1992b: 29 Distribution: México

Phyllophaga ravida (Blanchard) Ancylonycha ravida Blanchard,1851: 136 Lachnosterna ravida (Blanchard); Bates, 1888: 191 Phyllophaga ravida (Blanchard); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) ravida (Blanchard); Sanderson, 1958: 164 Distribution: Belize, Guatemala, México, United States

Synonym: Lachnosterna heterodoxa Horn, 1887b: 289 Phyllophaga heterodoxa (Horn); Glasgow, 1916: 373.

Synonym: Lachnosterna ravida guatemalica Moser, 1918a: 24 Phyllophaga ravida guatemalica (Moser); Sanderson, 1942: 53

Synonym: Lachnosterna ravida longiclava Moser, 1918a: 24 Phyllophaga longiclava (Moser); Blackwelder, 1944: 225 Phyllophaga ravida longiclava (Moser); Morón, 1986: 64

Phyllophaga rawlinsi Woodruff Phyllophaga rawlinsi Woodruff, 2005: 126 Distribution: Dominican Republic

Phyllophaga raydoma Saylor Phyllophaga (Phyllophaga) raydoma Saylor, 1943b: 107 Distribution: Guatemala

Phyllophaga recorta Sanderson Phyllophaga recorta Sanderson, 1951: 268 Distribution: Dominican Republic, Haiti

Phyllophaga reevesi Saylor Phyllophaga reevesi Saylor, 1939b: 89 Distribution: United States

Phyllophaga regiomontana Morón Phyllophaga (Phyllophaga) regiomontana Morón, 2001a: 176 Distribution: México

Phyllophaga reinhardi Saylor Phyllophaga (Listrochelus) reinhardi Saylor, 1940d: 89 Distribution: United States

119 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga renodis Reinhard Phyllophaga renodis Reinhard, 1939: 48 Distribution: United States

Phyllophaga reticulata Frey Phyllophaga reticulata Frey, 1975c: 257 Distribution: México

Phyllophaga reventazona Saylor Phyllophaga reventazona Saylor, 1936d: 295 Phyllophaga (Phyllophaga) reventazona Saylor; Solís and Morón, 1998: 25 Distribution: Costa Rica

Phyllophaga rex Woodruff and Sanderson Phyllophaga rex Woodruff and Sanderson, 2005: 130 Distribution: Dominican Republic

Phyllophaga reyesiana Morón Phyllophaga (Chlaeniobia) reyesiana Morón, 1992b: 26 Distribution: México

Phyllophaga riverana Morón Phyllophaga (Listrochelus) riverana Morón, 2004: 91 Distribution: México

Phyllophaga riviera Reinhard Phyllophaga riviera Reinhard, 1950: 27 Distribution: United States

Phyllophaga rodriguezi (Bates) Chlaenobia rodriguezi Bates, 1889: 399 Phyllophaga rodriguezi (Bates); Blackwelder, 1944: 226 Phyllophaga (Chlaenobia) rodriguezi (Bates); Cano and Morón 1998: 9 Distribution: Guatemala

Phyllophaga rolbakeri Saylor Phyllophaga (Phyllophaga) rolbakeri Saylor, 1940e: 111 Distribution: México

Phyllophaga rolstoni Riley and Wolfe Phyllophaga (Phyllophaga) rolstoni Riley and Wolfe, 1997: 428 Distribution: United States

Phyllophaga romana (Saylor) Cnemarachis romana Saylor, 1946a: 1 Phyllophaga romana (Saylor); Sanderson, 1951: 261 Distribution: Dominican Republic

120 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga rorida (Burmeister) Ancylonycha rorida Burmeister, 1855: 330 Phyllophaga rorida (Burmeister); Blackwelder, 1944: 226 Distribution: Brazil

Phyllophaga rorulenta patens (Arrow) Lachnosterna patens Arrow. 1900: 175 Cnemarachis patens (Arrow); Blackwelder, 1944: 223 Phyllophaga rorulenta patens (Arrow); Chalumeau, 1980: 87 Distribution: St. Vincent

Phyllophaga rorulenta rorulenta (Burmeister) Ancylonycha rorulenta Burmeister, 1855: 331 Lachnosterna rorulenta (Burmeister); Bates, 1888: 196 Phyllophaga (Phyllophaga) rorulenta (Burmeister); Saylor, 1943d: 131 Distribution: Colombia, Costa Rica, Guatemala, Guyana, México, Panama, Trinidad, Venezuela

Synonym: Lachnosterna trinitatis Arrow, 1900: 182 Phyllophaga trinitatis (Arrow); Blackwelder, 1944; 226

Phyllophaga roscida (Burmeister) Ancylonycha roscida Burmeister, 1855: 340 Phyllopahaga roscida (Burmeister); Blackwelder, 1944: 226 Phyllopahaga (Phyllophaga) roscida (Burmeister); Frey, 1975a: 214 Distribution: Colombia

Phyllophaga rossi Saylor Phyllophaga (Phyllophaga) rossi Saylor, 1939d: 452 Distribution: United States

Phyllophaga rostrypyga (Bates) Lachnosterna rostrypyga Bates, 1889: 401 Phyllophaga rostrypyga (Bates); Blackwelder, 1944; 226 Phyllophaga (Phyllophaga) rostrypyga (Bates); Cano and Morón, 1998: 14 Distribution: Guatemala, México

Synonym: Lachnosterna castaneipennis Moser, 1918a: 19 Phyllophaga castaneipennis (Moser): Blackwelder, 1944: 224

Phyllophaga rubella (Bates) Lachnosterna rubella Bates, 1888: 209 Phyllophaga rubella (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) rubella (Bates); Morón, 1986: 204 Distribution: México

Phyllophaga rubiginosa (LeConte) Lachnosterna rubiginosa LeConte, 1856: 259 Phyllophaga rubiginosa (LeConte); Glasgow, 1916: 373 121 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: United States

Phyllophaga rubricosa Reinhard Phyllophaga rubricosa Reinhard, 1939: 47 Distribution: United States

Phyllophaga ruficollis (Moser) Lachnosterna ruficollis Moser, 1918a: 47 Phyllophaga ruficollis (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) ruficollis (Moser); Morón, Vallejo, and Restrepo-Giraldo, 2001: 33 Distribution: Colombia

Phyllophaga rufipes (Moser) Lachnosterna rufipes Moser, 1924b: 156 Phyllophaga rufipes (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) rufipes (Moser); Frey, 1975a: 212 Distribution: Ecuador, Venezuela

Phyllophaga rufithorax (Moser) Lachnosterna rufithorax Moser, 1921c: 248 Phyllophaga rufithorax (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) rufithorax (Moser); Morón, 1986: 203 Distribution: México

Phyllophaga rufiventris (Kirsch) Lachnosterna rufiventris Kirsch, 1885: 220 Phyllophaga rufiventris (Kirsch); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) rufiventris (Kirsch); Frey, 1975a: 205 Distribution: Ecuador

Phyllophaga rufotestacea (Moser) Lachnosterna rufotestacea Moser, 1918a: 23 Phyllophaga rufotestacea (Moser); Blackwelder, 1944: 226 Phyllophaga (Phytalus) rufotestacea (Moser); Cano and Morón, 1998: 10 Distribution: Guatemala

Phyllophaga rugicollis (Bates) Lachnosterna rugicollis Bates, 1888: 190 Phyllophaga rugicollis (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) rugicollis (Bates); Morón, 1986: 202 Distribution: México

Phyllophaga rugipennis (Schaufuss) Anyclonycha rugipennis Schaufuss, 1858: 318 Phyllophaga rugipennis (Schaufuss); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) rugipennis (Schaufuss); Cano and Morón, 1998: 14 Distribution: Colombia, Guatemala, México, Panama

122 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga rugithorax Saylor Phyllophaga (Phytalus) rugithorax Saylor, 1938e: 188 Distribution: México

Phyllophaga rugosa (Melsheimer) Ancylonycha rugosa Melsheimer, 1846: 140 Lachnosterna rugosa (Melsheimer); LeConte, 1856: 252 Phyllophaga rugosa (Melsheimer); Glasgow, 1916: 371 Distribution: Canada, United States

Phyllophaga rugulosa (Blanchard) Ancylonycha rugulosa Blanchard, 1851: 134 Lachnosterna rugulosa (Blanchard): Bates, 1888: 203 Phyllophaga rugulosa (Blanchard); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) rugulosa (Blanchard); Cano and Morón, 1998: 13 Distribution: Guatemala, México

Phyllophaga rustica Woodruff Phyllophaga rustica Woodruff, 2005: 137 Distribution: Dominican Republic

Phyllophaga rzedowskiana Aragón and Morón Phyllophaga (Chlaenobia) rzedowskiana Aragón and Morón, 2003: 560 Distribution: México

Phyllophaga sacoma Reinhard Phyllophaga sacoma Reinhard, 1939: 52 Distribution: United States

Phyllophaga saginata (Mannerheim) Melolontha saginata Mannerheim, 1829: 64 Phytalus saginatus (Mannerheim); Harold, 1869a: 1172 Phyllophaga saginata (Mannerheim); Blackwelder, 1944: 226 Distribution: Brazil

Phyllophaga saltana Frey Phyllophaga (Phyllophaga) saltana Frey, 1975a: 219 Distribution: Argentina

Phyllophaga sanbarthensis Chalumeau and Gruner Phyllophaga (Cnemarachis) sanbarthensis Chalumeau and Gruner, 1976: 97 Distribution: Saint Barthelemey, Saint Martin

Phyllophaga sanctipauli (Blanchard) Ancylonycha sancti-pauli Blanchard, 1851: 132 Phyllophaga sancti-pauli (Blanchard); Blackwelder, 1944: 226 Phyllophaga sanctipauli (Blanchard); Frey, 1975a; 215 Distribution: Argentina, Brazil 123 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga sandersoni Garcia-Vidal Phyllophaga sandersoni Garcia-Vidal, 1988: 104 Distribution: Cuba

Phyllophaga sandersoniana Morón Phyllophaga (Chlaenobia) sandersoniana Morón, 1992b: 24 Distribution: México

Phyllophaga sandersoniella Chalumeau and Gruner Phyllophaga (Cnemarachis) sandersoniella Chalumeau and Gruner, 1976: 92 Distribution: Guadeloupe

Phyllophaga sanjosicola Saylor Phyllophaga sanjosicola Saylor, 1935h: 500 Phyllophaga (Phyllophaga) sanjosicola Saylor; Solís and Morón, 1998: 23 Distribution: Costa Rica

Synonym: Phyllophaga (Phytalus) valeriana Saylor, 1934: 50

Phyllophaga santachloe Woodruff Phyllophaga santachloe Woodruff, 2005: 141 Distribution: Dominican Republic

Phyllophaga santaclarae Chapin Phyllophaga santaclarae Chapin, 1932b: 189 Cnemarachis santa clarae (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga santiaguensis Garcia-Vidal Phyllophaga santiaguensis Garcia-Vidal, 1987: 303 Distribution: Cuba

Phyllophaga saylori Sanderson Phyllophaga saylori Sanderson, 1965: 560 Phyllophaga (Phyllophaga) saylori Sanderson; Morón, 1986: 205 Distribution: México

Phyllophaga sayloriana Morón, Rivera, and López Phyllophaga (Phyllophaga) sayloriana Morón, Rivera, and López, 2002: 486 Distribution: México

Phyllophaga scabrifrons (Bates) Lachnosterna scabrifrons Bates, 1888: 187 Phyllophaga scabrifrons (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) scabrifrons (Bates); Cano and Morón, 1998: 14 Distribution: Belize, Guatemala

124 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Lachnosterna guatemalensis Moser, 1921c: 251 Phyllophaga guatemalensis (Moser); Blackwelder, 1944: 224

Phyllophaga scabripyga (Bates) Chlaenobia scabripyga Bates, 1888: 167 Phyllophaga scabripyga (Bates); Blackwelder, 1944: 226 Phyllophaga (Chlaenobia) scabripyga (Bates); Morón, 1986: 209 Distribution: Costa Rica, El Salvador, Guatemala, México

Phyllophaga scaramuzzai Garcia-Vidal Phyllophaga scaramuzzai Garcia-Vidal, 1988: 106 Distribution: Cuba

Phyllophaga schaefferi Saylor Phyllophaga schaefferi Saylor, 1937d: 321 (for Schaeffer, 1909: 382) Distrtibution: United States

Synonym: Lachnosterna georgiana Schaeffer, 1909: 382 (not Horn, 1885: 122) Phyllophaga georgiana (Schaeffer); Glasgow, 1916: 374

Synonym: Phyllophaga duvalus Robinson, 1938: 110 Phyllophaga duvala Robinson; Smith and Evans, 2005: 52

Phyllophaga schencklingi (Moser) Lachnosterna schencklingi Moser, 1918b: 163 Phyllophaga schenklingi (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) schenklingi (Moser); Morón, 1986: 202 Distribution: México

Phyllophaga schizorhina (Bates) Lachnosterna schizorhina Bates, 1888: 202 Phyllophaga schizorhina (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) schizorhina (Bates); Solís and Morón, 1998: 24 Distribution: Costa Rica, Nicaragua, Panama

Phyllophaga schizorhinoides Morón Phyllophaga (Phyllophaga) schizorhinoides Morón, 2003a: 221 Distribution: Costa Rica, Panama

Phyllophaga schneblei Frey Phyllophaga (Phyllophaga) schneblei Frey, 1975a: 208 Distribution: Colombia

Phyllophaga schusteriana Cano and Morón Phyllophaga (Chlaenobia) schusteriiana Cano and Morón, 2002: 356 Distribution: Guatemala

Phyllophaga schwarzi Chapin 125 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga schwarzi Chapin, 1932b: 184 Cnemarachis schwarzi (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga scissa (Bates) Lachnosterna scissa Bates, 1888: 202 Phyllophaga scissa (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) scissa (Bates); Morón, 1986: 204 Distribution: México

Phyllophaga scitula (Horn) Lachnosterna scitula Horn, 1887b: 256 Phyllophaga scitula (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga scoparia (LeConte) Listrochelus scoparius LeConte, 1856: 264 Phyllophaga (Listrochelus) scoparia (LeConte); Saylor, 1940d: 80 Distribution: México, United States

Phyllophaga scuticeps (Bates) Listrochelus scuticeps Bates, 1888: 171 Phyllophaga (Listrochelus) scuticeps (Bates); Saylor, 1940d: 94 Distribution: México, United States

Phyllophaga segregans (Bates) Lachnosterna segregans Bates, 1888: 206 Phyllophaga segregans (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) segregans (Bates); Morón, 1986: 205 Distribution: México

Phyllophaga senex (Horn) Listrochelus senex Horn, 1878: 148 Phyllophaga (Listrochelus) senex (Horn); Saylor, 1940d: 115 Distribution: United States

Phyllophaga senicula (Bates) Phytalus seniculus Bates, 1888: 181 Phyllophaga senicula (Bates); Blackwelder, 1944: 226 Phyllophaga (Phytalus) senicula (Bates); Morón, 1986: 213 Distribution: Guatemala, México

Phyllophaga sequoiana Saylor Phyllophaga sequoiana Saylor, 1936a: 2 Distribution: United States

Phyllophaga sericata (Erichson) Anycylonycha sericata Erichson, 1848: 563 126 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga sericata (Erichson); Evans, 2003: 155 Distribution: French Guiana

Phyllophaga serrana Morón and Cano Phyllophaga (Phyllophaga) serrana Morón and Cano, 2000: 240 Distribution: Guatemala

Phyllophaga serratipes Morón and Cano Phyllophaga (Phyllophaga) serratipes Morón and Cano, 2000: 237 Distribution: Guatemala

Phyllophaga setidorsis (Bates) Lachnosterna setidorsis Bates, 1888: 200 Phyllophaga setidorsis (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) setidorsis (Bates); Sanderson, 1958: 172 Distribution: Costa Rica, México

Phyllophaga setifera (Burmeister) Ancylonycha setifera Burmeister, 1855: 335 Lachnosterna setifera (Burmeister); Bates, 1888: 198 Phyllophaga (Phyllophaga) setifera (Burmeister); Sanderson, 1958: 166 Distribution: Belize, Costa Rica, Colombia, Guatemala, Honduras, México, Nicaragua, Panama, United States, Venezuela

Phyllophaga siboneyensis Garcia-Vidal Phyllophaga siboneyensis Garcia-Vidal, 1988: 103 Distribution: Cuba

Phyllophaga signaticollis (Burmeister) Anyclonycha signaticollis Burmeister, 1855: 337 Phyllophaga signaticollis (Burmeister); Chapin, 1932b: 190 Cnemarachis signaticollis (Burmeister); Blackwelder, 1944: 223 Distribution: Cuba

Synonym: Ancylonycha conflusa Jacquelin du Val, 1856: 131

Phyllophaga sinaloana Saylor Phyllophaga (Phytalus) sinaloana Saylor, 1935h: 499 Distribution: México

Phyllophaga sinicollis Saylor Phyllophaga (Phyllophaga) sinicollis Saylor, 1943e: 265 Distribution: México

Phyllophaga sinuaticeps (Moser) Lachnosterna sinuaticeps Moser, 1921c: 249 Phyllophaga sinuaticeps (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) sinuaticeps (Moser); Morón, 1986: 203 127 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Phyllophaga skelleyi Woodruff and Beck Phyllophaga skelleyi Woodruff and Beck, 1989: 156 Distribution: United States

Phyllophaga smithi (Arrow) Phytalus smithi Arrow, 1912: 455 Clemora smithi (Arrow); Saylor, 1942a: 160 Phyllophaga smithi (Arrow); Evans, 2003: 156 Distribution: Barbados, Trinidad [Mauritius]

Phyllophaga snowi Saylor Phyllophaga (Listrochelus) snowi Saylor, 1940d: 114 Distribution: México, United States

Phyllophaga sociata (Horn) Listrochelus sociatus Horn, 1878: 146 Phyllophaga sociatus (Horn); Luginbill and Painter, 1953: 35 Distribution: United States

Phyllophaga soctona Morón Phyllophaga (Phytalus) soctona Morón, 2000b: 37 Distribution: México

Phyllophaga sodialis Reinhard Phyllophaga sodialis Reinhard: 1940: 4 Distribution: United States

Phyllophaga solanophaga Morón Phyllophaga (Chlaenobia) solanophaga Morón, 1988: 57 Distribution: El Salvador

Phyllophaga solavegana Morón Phyllophaga (Phytalus) solavegana Morón, 2000b: 55 Distribution: México

Phyllophaga solisiana Morón Phyllophaga (Phyllophaga) solisiana Morón, 2003a: 259 Distribution: Costa Rica, Guatemala, Nicaragua

Phyllophaga sonora Saylor Phyllophaga (Phytalus) sonora Saylor, 1939c: 167 (for Horn, 1885: 122) Phyllophaga (Phyllophaga) sonora Saylor; Sanderson, 1958: 166 Distribution: México, United States

Synonym: Phytalus debilis Horn, 1885: 122 (not LeConte, 1856: 262)

128 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga soror Davis Phyllophaga soror Davis, 1920: 333 Distribution: United States

Phyllophaga spaethi (Nonfried) Lachnosterna spaethi Nonfried, 1891: 265 Phyllophaga (Phyllophaga) spaethi (Nonfried); Saylor, 1943d: 133 Distribution: Honduras

Phyllophaga speculifera (Chevrolat) Ancylonycha speculifera Chevrolat, 1865: 27 Phyllophaga speculifera (Chevrolat); Chapin, 1932b: 196 Cnemarachis speculifera (Chevrolat); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga spinicola Garcia-Vidal Phyllophaga (Cnemarachis) spinicola Garcia-Vidal, 1984: 324 Distribution: Cuba

Phyllophaga spinifemora Saylor Phyllophaga (Phyllophaga) spinifemora Saylor, 1940e: 307 Cnemarachis spinifemora (Saylor); Blackwelder, 1944; 223 Distribution: Jamaica

Phyllophaga spinitarsis (Moser) Lachnosterna spinitarsis Moser, 1921c: 253 Phyllophaga spinitarsis (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) spinitarsis (Moser); Morón, 1986: 205 Distribution: México

Phyllophaga spreta (Horn) Lachnosterna spreta Horn, 1887b: 250 Phyllophaga spreta (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga squamifera Frey Phyllophaga squamifera Frey, 1976a: 299 Distribution: Venezuela

Phyllophaga squamipilosa Saylor Phyllophaga (Tostegoptera) squamipilosa Saylor, 1936c: 280 Distribution: United States

Phyllophaga stehlei Chalumeau Phyllophaga stehlei Chalumeau, 1985: 28 Distribution: St. Martin

Phyllophaga stipitalis (Blanchard) 129 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ancylonycha stipitalis Blanchard, 1851: 136 Phyllophaga stipitalis (Blanchard); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) stipitalis (Blanchard); Morón, 1986: 205 Distribution: México

Phyllophaga stohleri Saylor Phyllophaga stohleri Saylor, 1938c: 130 Distribution: United States

Phyllophaga straminea (Bates) Phytalus stramineus Bates, 1888: 183 Phyllophaga straminea (Bates); Blackwelder, 1944: 226 Phyllophaiga (Phytalus) striaminea (Bates); Morón, 1986: 200 Distribution: México

Phyllophaga sturmi (Bates) Lachnosterna sturmi Bates, 1888: 197 Phyllophaga sturmi (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) sturmi (Bates); Morón, 1986: 195 Distribution: Guatemala, México

Phyllophaga stzotzilana Morón Phyllophaga (Phyllophaga) stzotzilana Morón, 2001a: 178 Distribution: México

Phyllophaga submetallica (Bates) Lachnosterna submetallica Bates, 1888: 204 Phyllophaga submetallica (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) submetallica (Bates); Morón, 1986: 242 Distribution: Guatemala

Phyllophaga submucida (LeConte) Lachnosterna submucida LeConte, 1856: 260 Phyllophaga submucida (LeConte); Glasgow, 1916: 373 Phyllophaga (Phyllophaga) submucida (LeConte); Morón, 1986: 204 Distribution: México, United States

Phyllophaga subnitida (Moser) Lachnosterna subnitida Moser, 1918a: 44 Phyllophaga subnitida (Moser); Blackwelder, 1944: 226 Distribution: Venezuela

Phyllophaga subopaca (Moser) Lachnosterna subopaca Moser, 1918a: 45 Phyllophaga subopaca (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) subopaca (Moser); Frey 1975a: 206 Distribution: Venezuela, Trinidad

130 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga subpruinosa (Casey) Lachnosterna subpruinosa Casey, 1884: 38 Phyllophaga subpruinosa (Casey); Glasgow, 1916: 379 Distribution: United States

Synonym: Phyllophaga deanii Luginbill, 1928: 78

Phyllophaga subrugosa (Moser) Lachnosterna subrugosa Moser, 1924b: 155 Phyllophaga subrugosa (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) subrugosa (Moser); Morón, 1986: 205 Distribution: México

Phyllophaga subtonsa (LeConte) Lachnosterna subtonsa LeConte, 1856: 254 Phyllophaga subtonsa (LeConte); Glasgow, 1916: 372 Distribution: United States

Phyllophaga suriana Morón Phyllophaga (Listrochelus) suriana Morón, 2002: 460 Distribution: México

Phyllophaga suttonana Reinhard Phyllophaga tristis suttonana Reinhard, 1939: 59 Phyllophaga suttonana Reinhard; Riley and Wolfe, 2003: 23 Distribution: United States

Phyllophaga suturalis (Chevrolat) Ancylonycha suturalis Chevrolat, 1865: 29 Phyllophaga suturalis (Chevrolat); Chapin, 1932b: 200 Cnemarachis suturalis (Chevrolat); Blackwelder, 1944: 223 Phyllophaga (Cnemarachis) suturalis (Chevrolat); Woodruff, 1961; 12 Distribution: Cuba

Phyllophaga sylvatica Sanderson Phyllophaga sylvatica Sanderson, 1942: 49 Distribution: United States

Phyllophaga tajimaroana Morón Phyllophaga (Phytalus) tajimaroana Morón, 2000b: 57 Distribution: México

Phyllophaga talamancana Morón and Solís Phyllophaga (Phyllophaga) talamancana Morón and Solís, 2001: 22 Distribution: Costa Rica

Phyllophaga tancitara Saylor Phyllophaga (Phytalus) tancitara Saylor, 1943a: 23 131 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Phyllophaga tapantina Morón and Solís Phyllophaga (Phyllophaga) tapantina Morón and Solís, 2001: 14 Distribution: Costa Rica

Phyllophaga tarsalis (Schaeffer) Listrochelus tarsalis Schaeffer, 1908: 319 Phyllophaga (Listrochelus) tarsalis (Schaeffer), Saylor, 1940c: 93 Distribution: United States

Phyllophaga taxodii Langston Phyllophaga taxodii Langston, 1924: 449 Distribution: United States

Phyllophaga tecta Cartwright Phyllophaga tecta Cartwright, 1944: 32 Distribution: United States

Phyllophaga tegenara Saylor Phyllophaga (Phytalus) tegenara Saylor, 1935h: 496 Distribution: Costa Rica

Phyllophaga tegulicolis Saylor Phyllophaga (Phytalus) tegulicolis Saylor, 1934: 49 Distribution: Honduras

Phyllophaga tejulpicas Saylor Phyllophaga (Phyllophaga) tejulpicas Saylor, 1943e: 264 Distribution: México, Panama

Phyllophaga tejupana Morón Phyllophaga (Phytalus) tejupana Morón, 2000b: 54 Distribution: México

Phyllophaga temascalis Saylor Phyllophaga (Phyllophaga) temascalis Saylor, 1941a: 26 Distribution: México

Phyllophaga temaxa Saylor Phyllophaga (Phyllophaga) temaxa Saylor, 1940e: 314 (for Bates, 1889: 404) Distribution: Guatemala, México

Synonym: Lachnosterna parilis Bates, 1889: 404 (not Bates, 1888: 172)

Phyllophaga temescaltepeca Saylor Phyllophaga (Phytalus) temescaltepeca Saylor, 1934: 49 Distribution: México 132 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga temora Saylor Phyllophaga (Phyllophaga) temora Saylor, 1943d: 135 Distribution: México

Phyllophaga tenuipilis (Bates) Lachnosterna tenuipilis Bates, 1888: 202 Phyllophaga tenuipilis (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) tenuipilis (Bates); Cano and Morón, 1998: 14 Distribution: Guatemala, México, Nicaragua

Phyllophaga teosinteophaga Morón and Rivera Phyllophaga (Phytalus) teosinteophaga Morón and Rivera, 1992: 84 Distribution: México

Phyllophaga tepanana Saylor Phyllophaga tepanana Saylor, 1938f: 341 Phyllophaga (Phyllophaga) tepanana Saylor; Cano and Morón, 1998: 14 Distribution: Guatemala

Phyllophaga terezinae Frey Phyllophaga (Phyllophaga) terezinae Frey, 1965b: 147 Distribution: Brazil

Phyllophaga terminalis (Saylor) Listrochelus terminalis Saylor, 1935g: 35 Phyllophaga terminalis (Saylor); Blackwelder, 1944: 226 Phyllophaga (Listrochelus) terminalis (Saylor); Morón, 1986: 266 Distribution: México

Phyllophaga testaceipennis (Blanchard) Ancylonycha testaceipennis Blanchard, 1851: 134 Lachnosterna testaceipennis (Blanchard); Bates, 1888: 195 Phyllophaga (Phyllophaga) testaceipennis (Blanchard); Saylor, 1943b: 106 Distribution: México, Guatemala, Panama

Phyllophaga tetracera Chapin Phyllophaga tetracera Chapin, 1932b: 193 Cnemarachis tetracera (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga tetraphylla (Moser) Lachnosterna tetraphylla Moser, 1918a: 65 Cnemarachis tetraphylla (Moser); Blackwelder, 1944: 223 Phyllophaga tetraphylla (Moser); Chalumeau, 1985: 26 Distribution: Jamaica

Phyllophaga texensis Saylor 133 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga (Listrochelus) texensis Saylor, 1940c: 106 Distribution: United States

Phyllophaga thoracica (Burmeister) Ancylonycha thoracica Burmeister, 1855: 332 Phyllophaga thoracica (Burmeister); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) thoracica (Burmeister); Frey, 1975a; 212 Distriburtion: Colombia, Venezuela

Phyllophaga tilarana Morón and Solís Phyllophaga (Phyllophaga) tilarana Morón and Solís, 2001: 12 Distriubtion: Costa Rica

Phyllophaga timida (Horn) Listrochelus timidus Horn, 1878: 146 Phyllophaga (Listrochelus) timida (Horn); Saylor, 1940c: 113 Distribution: México, United States

Phyllophaga tlanchinolensis Morón Phyllophaga (Phytalus) tlanchinolensis Morón, 2000b: 49 Distriubtion: México

Phyllophaga tojolabala Morón Phyllophaga (Phyllophaga) tojolabala Morón, 1999: 378 Distriubtion: México

Phyllophaga toni Woodruff Phyllophaga toni Woodruff, 2005: 143 Distribution: Dominican Republic

Phyllophaga torta (LeConte) Lachnosterna torta LeConte, 1856: 239 Phyllophaga torta (LeConte); Glasgow, 1916: 372 Phyllophaga (Phyllophaga) torta (LeConte); Morón, 1986: 244 Distribution: México, United States

Synonym: Lachnosterna dampfi Arrow, 1933b: 148

Phyllophaga totonis Saylor Phyllophaga (Phyllophaga) totonis Saylor, 1941d: 386 Distribution: Guatemala

Phyllophaga transversicollis (Moser) Lachnosterna transversicollis Moser, 1918a: 49 Phyllophaga transversicollis (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) transversicollis (Moser); Frey, 1975a; 211 Distribution: Colombia

134 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga trichia (Bates) Lachnosterna trichia Bates, 1888: 200 Phyllophaga trichia (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) trichia (Bates); Morón, 1986: 204 Distribution: Guatemala, México

Phyllophaga trichoderma (Moser) Lachnosterna trichoderma Moser, 1921c: 256 Phyllophaga trichoderma (Moser); Blackwelder, 1944: 226 Distribution: Paraguay

Phyllophaga trichodes (Bates) Lachnosterna trichodes Bates, 1888: 178 Phyllophaga trichodes (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) trichodes (Horn); Sanderson, 1958: 166 Phyllophaga (Phytalus) trichodes (Bates); Morón, 1986: 212 Distribution: México, United States

Synonym: Phytalus robustus Horn, 1885: 120 (not LeConte, 1856: 257)

Synonym: Phyllophaga (Phytalus) sandersonia Saylor, 1939c: 161 (for Horn, 1885: 120)

Phyllophaga tridens (Bates) Lachnosterna tridens Bates, 1888: 187 Phyllophaga tridens (Bates); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) tridens (Bates); Morón, 1986: 202 Distribution: México

Phyllophaga tridilonycha Saylor Phyllophaga (Phyllophaga) tridilonycha Saylor, 1943a: 27 Distribution: México

Phyllophaga trinitariensis Garcia-Vidal Phyllophaga trinitariensis Garcia-Vidal, 1987: 306 Distribution: Cuba

Phyllophaga tristis (Fabricius) Melolontha tristis Fabricius, 1781: 39 Trichestes tristis (Fabricius); Burmeister, 1855: 358 Lachnosterna tristis (Fabricius); LeConte, 1856: 260 Phyllophaga tristis (Fabricius); Glasgow, 1916: 370 Phyllophaga (Phyllophaga) tristis (Fabricius); Morón, 1986: 204 Distribution: Canada, México, United States

Synonym: Melolontha pilosicollis Knoch, 1801: 85 Trichesthes pilosicollis (Knoch); Blanchard, 1850: 141 Phyllophaga pilosicollis (Knoch); Evans, 2003: 165

135 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga triticophaga Morón and Salvadori Phyllophaga (Phytalus) triticophaga Morón and Salvadori, 1998: 370 Distribution: Brazil

Phyllophaga trochanter Saylor Phyllophaga (Listrochelus) trochanter Saylor, 1940c: 78 (for Horn, 1885: 123) Distribution: México, United States

Synonym: Listrochelus gracilis Horn, 1885: 123 (not Burmeister, 1855: 361)

Phyllophaga tsajumiana Morón Phyllophaga (Phyllophaga) tsajumiana Morón, 2001a: 184 Distribution: México

Phyllophaga tuberculifrons (Chevrolat) Ancylonycha tuberculifrons Chevrolat, 1865: 23 Phyllophaga tuberculifrons (Chevrolat); Chapin, 1932b: 186 Cnemarachis tuberculifrons (Chevrolat); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga tumulosa (Bates) Chlaenobia tumulosa Bates, 1888: 168 Phyllophaga tumulosa (Bates); Blackwelder, 1944: 226 Phyllophaga (Chlaenobia) tumulosa (Bates); Morón, 1986: 209 Distribution: Belize, El Salvador, Guatemala, Honduras, México, Nicaragua

Phyllophaga tusa (Horn) Lachnosterna tusa Horn, 1887b: 290 Phyllophaga tusa (Horn); Glasgow, 1916: 373 Distribution: United States

Phyllophaga tuxtleca Morón Phyllophaga (Phyllophaga) tuxtleca Morón, 2003a: 237 Distribution: México

Phyllophaga tzintzontliana Morón Phyllophaga (Chlaenobia) tzintzontliana Morón, 1992b: 18 Distribution: México

Phyllophaga ueiacayoca Morón Phyllophaga (Phyllophaga) ueiacayoca Morón, 1992a: 121 Distribution: México

Phyllophaga ulkei (Smith) Lachnosterna ulkei Smith, 1889a: 94 Phyllophaga ulkei (Smith); Glasgow, 1916: 373 Distribution: United States

136 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga umbrosa (Erichson) Ancylonycha umbrosa Erichson, 1847a: 101 Phyllophaga umbrosa (Erichson); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) umbrosa (Erichson); Frey, 1975a: 203 Distribution: Peru

Phyllophaga uniformis (Blanchard) Ancylonycha uniformis Blanchard, 1851: 133 Lachnosterna uniformis (Blanchard); LeConte, 1856: 245 Phyllophaga uniformis (Blanchard); Glasgow, 1916: 371 Distribution: United States

Synonym: Lachnosterna carolina Fall, 1912: 43

Phyllophaga uruguayana (Saylor) Lachnosterna (Phytalus) uruguayana Saylor, 1935f: 206 Phyllophaga uruguayana (Saylor); Blackwelder, 1944: 226 Phyllophaga (Phytalus) uruguayana (Saylor); Frey, 1975a: 226 Distribution: Uruguay

Phyllophaga valia Saylor Phyllophaga (Listrochelus) valia Saylor, 1940e: 113 Distribution: México

Phyllophaga vandinei Smyth Lachnosterna vandinei Smyth, 1917: 68 Cnemarachis vandinei (Smyth); Saylor, 1942a: 159 Phyllophaga vandinei (Smyth); Blackwelder, 1944: 223 Distribution: Puerto Rico

Synonym: Lachnosterna major Moser, 1918a: 59

Phyllophaga vandykei Saylor Phyllophaga vandykei Saylor, 1935g: 34 Distribution: Panama

Phyllophaga vazquezae Morón Phyllophaga (Phyllophaga) vazquezae Morón, 1995: 5 Distribution: México

Phyllophaga vehemens (Horn) Lachnosterna vehemens Horn, 1887b: 244 Phyllophaga vehemens (Horn); Glasgow, 1916: 373 Phyllophaga (Phyllophaga) vehemens (Horn) Distribution: United States

Phyllophaga venodiola Saylor Phyllophaga (Listrochelus) venodiola Saylor, 1938e: 185 137 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Phyllophaga vermiculata Chapin Phyllophaga vermiculata Chapin, 1932b: 195 Cnemarachis vermiculata (Chapin); Blackwelder, 1944: 223 Distribution: Cuba

Phyllophaga verruciventris (Moser) Lachnosterna verruciventris Moser, 1918a: 21 Phyllophaga verruciventris (Moser); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) verruciventris (Moser); Morón, 1986: 202 Distribution: México

Phyllophaga vestita (Moser) Phytalus vestitus Moser, 1918a: 70 Phyllophaga vestita (Moser); Blackwelder, 1944: 227 Phyllophaga (Phytalus) vestita (Moser); Frey, 1975a: 220 Distribution: Bolivia, Brazil

Phyllophaga vetula (Horn) Lachnosterna vetula Horn, 1887b: 274 Phyllophaga vetula (Horn); Glasgow, 1916: 373 Phyllophaga (Phyllophaga) vetula (Horn); Sanderson, 1958: 165 Distribution: México, United States

Synonym: Lachnosterna longipilosa Bates, 1888: 209

Phyllophaga vexata unituberculata (Bates) Chlaenobia unituberculata Bates, 1889: 399 Chlaenobia vexatus unituberculata Bates; Chapin, 1935: 14 Phyllophaga vexatus unituberculata (Bates); Blackwelder, 1944; 227 Phyllophaga (Chlaenobia) vexatus unituberculata (Bates); Morón, 1986: 208 Distribution: México

Phyllophaga vexata vexata (Horn) Phytalus vexatus Horn, 1885: 121 Chlaenobia vexata (Horn); Chapin, 1935: 13 Phyllophaga vexata (Horn); Blackwelder, 1944: 227 Phyllophaga (Chlaenobia) vexata (Horn); Morón, 1986: 208 Distribution: Guatemala, México, United States

Synonym: Phytalus cavifrons Linell, 1896: 729

Phyllophaga vicina (Moser) Lachnosterna vicina Moser, 1918a: 29 Phyllophaga (Phyllophaga) vicina (Moser); Saylor, 1943d: 136 Distribution: Costa Rica, Guatemala, Panama

138 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga vilifrons (LeConte) Lachnosterna vilifrons LeConte, 1856: 255 Lachnosterna villifrons LeConte; Horn, 1887b; 263 Phyllophaga vilifrons (LeConte); Glasgow, 1916: 372. Distribution: Canada, United States

Synonym: Lachnosterna hirticeps LeConte, 1856: 255

Phyllophaga villaclarensis Garcia-Vidal Phyllophaga villaclarensis Garcia-Vidal, 1987: 302 Distribution: Cuba

Phyllophaga vulpes (Arrow) Lachnosterna vulpes Arrow, 1913c: 463 Phyllophaga vulpes (Arrow); Blackwelder, 1944: 227 Phyllophaga (Phyllophaga) vulpecula [sic] (Arrow); Frey, 1975a: 217 Distribution: Brazil

Phyllophaga wickhami Saylor Phyllophaga (Listrochelus) wickami Saylor, 1940c: 107 Distribution: United States

Phyllophaga wittkugeli (Nonfried) Lachnosterna wittkugeli Nonfried, 1891: 264 Phyllophagha wittkugeli (Nonfried); Blackwelder, 1944; 227 Distribution: Honduras

Phyllophaga wolcotti Saylor Phyllophaga (Phyllophaga) wolcotti Saylor, 1940e: 307 Cnemarachis wolcotti (Saylor); Blackwelder, 1944: 223 Distribution: Puerto Rico

Phyllophaga woodruffi Warner and Morón Phyllophaga (Triodonyx) woodruffi Warner and Morón, 1992: 337 Distribution: México

Phyllophaga xanthe (Bates) Lachnosterna xanthe Bates, 1888: 189 Phyllophaga xanthe (Bates); Blackwelder, 1944: 227 Phyllophaga (Phyllophaga) xanthe (Bates); Morón, 1986: 202 Distribution: México

Phyllophaga xanthocoma (Bates) Phytalus xanthocomus Bates, 1888: 181 Phyllophaga xanthocoma (Bates); Blackwelder, 1944: 227 Phyllophaga (Phytalus) xanthocoma (Bates); Morón, 1986: 213 Distribution: Guatemala, México

139 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga xerophila Saylor Phyllophaga xerophila Saylor, 1939b: 89 Distribution: United States

Phyllophaga xkumuka Morón Phyllophaga (Phyllophaga) xkumuka Morón, 1999: 382 Distribution: México

Phyllophaga yaqui Saylor Phyllophaga (Listrochelus) yaqui Saylor, 1940e: 314 (for debilis Arrow, 1933b: 147) Distribution: México

Synonym: Listrochelus debilis Arrow, 1933b: 147 (not LeConte, 1856: 262) Phyllophaga debilis (Arrow); Evans, 2003: 170

Phyllophaga yaxbitana Morón Phyllophaga (Phytalus) yaxbitana Morón, 2000b: 39 Distribution: México

Phyllophaga yei Morón Phyllophaga (Chirodines) yei Morón, 1991: 402 Distribution: México

Phyllophaga yemasseei Cartwright Phyllophaga yemasseei Cartwright, 1944: 30 Distribution: United States

Phyllophaga yoloxana Morón Phyllophaga (Phyllophaga) yoloxana Morón, 2003a: 285 Distribution: México

Phyllophaga youngi Cartwright Phyllophaga youngi Cartwright, 1935: 102 Phyllophaga (Cnemarachis) youngi Cartwright; Woodruff, 1961: 12 Distribution: Bahamas, United States

Synonym: Phyllophaga (Phyllophaga) bahama Saylor, 1940e: 311

Phyllophaga yucana Saylor Phyllophaga yucana Saylor, 1937d: 321 (for Blanchard, 1851: 136) Distribution: Colombia, Venezuela

Synonym: Ancylonycha sericata Blanchard, 1851: 136 (not Erichson, 1848: 563) Phyllophaga sericata (Blanchard); Blackwelder, 1944: 226 Phyllophaga (Phyllophaga) sericata (Blanchard); Frey, 1975a: 212

Phyllophaga yucateca (Bates) Lachnosterna yucateca Bates, 1889: 404 140 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phyllophaga yucateca (Bates); Blackwelder, 1944: 227 Phyllophaga (Phyllophaga) yucateca (Bates); Morón, 1986: 204 Distribution: México

Phyllophaga yunqueana Chapin Phyllophaga yunqueana Chapin, 1935c: 70 Cnemarachis yunqueana (Chapin); Blackwelder, 1944: 223 Distribution: Puerto Rico

Phyllophaga zaragozana Morón Phyllophaga (Phyllophaga) zaragozana Morón, 2003a: 227 Distribution: México

Phyllophaga zarcoana Morón Phyllophaga (Phyllophaga) zarcoana Morón, 2003a: 275 Distribution: Guatemala

Phyllophaga zavalana Reinhard Phyllophaga zavalana Reinhard, 1946: 69 Distribution: United States

Phyllophaga zayasi Garcia-Vidal Phyllophaga (Cnemarachis) zayasi, Garcia-Vidal, 1978: 10 Distribution: Cuba

Phyllophaga zunilensis (Bates) Chirodines zunilensis Bates, 1888: 169 Phyllophaga (Chirodines) zunilensis (Bates); Saylor, 1942; 162 Distribution: Guatemala, Honduras, México

Genus PLECTRODES Horn Plectrodes Horn, 1867: 166. Type species Plectrodes pubescens Horn, by monotypy.

Plectrodes pubescens Horn Plectrodes pubescens Horn, 1867: 167 Distribution: United States

Genus POLYPHYLLA Harris Polyphylla Harris, 1841: 30. Type species Melolontha variolosa Hentz, by subsequent designation (Young, 1988: 21).

Synonym: Macronoxia Crotch, 1873: 61. Type species Melolontha occidentalis Linné, subsequent designation (Evans 2003: 171)..

Synonym: Polylamina Hardy, 1974: 6. Type species Polyphylla pubescens Cartwright, by original designation.

Keys to species: Fall, 1928: 34; Hardy, 1981: 299; Young, 1967: 285; 1988: 22, 23; Morón, 1994: 41 141 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Polyphylla aeolus La Rue Polyphylla aeolus La Rue, 1998: 24 Distribution: United States

Polyphylla anteronivea Hardy Polyphylla anteronivea Hardy, 1978a: 2 Distribution: United States

Polyphylla arguta Casey Polyphylla arguta Casey, 1914: 338 Distribution: United States

Polyphylla avittata Hardy Polyphylla avittata Hardy, 1978a: 1 Distribution: United States

Polyphylla barbata Cazier Polyphylla barbata Cazier, 1938: 161 Distribution: United States

Polyphylla brownae Young Polyphylla brownae Young, 1986: 47 Distribution: United States

Polyphylla cavifrons LeConte Polyphylla cavifrons LeConte, 1854: 222 Distribution: México, United States

Polyphylla comes Casey Polyphylla comes Casey, 1914: 351 Distribution: United States

Polyphylla concurrens Casey Polyphylla concurrens Casey, 1889: 169 Distribution: Honduras

Polyphylla conspersa Burmester Polyphylla conspersa Burmester, 1855: 407 Distribution: México

Polyphylla crinita LeConte Polyphylla crinita LeConte, 1856: 230 Distribution: Canada, México, United States

Synonym: Polyphylla crinita mystica Casey, 1914: 334

Synonym: Polyphylla incolumis Casey, 1914: 335 142 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Polyphylla ona Bloeker, 1939: 148

Synonym: Polyphylla pacifica Casey, 1895: 607

Synonym: Polyphylla ruficollis preversa Casey, 1914: 348

Synonym: Polyphylla santarosae Bloeker, 1939: 149

Polyphylla decemlineata (Say) Melolontha decemlineata Say, 1824: 246 Polyphylla decemlineata (Say): Burmeister, 1855: 407 Distribution: Canada, México, United States

Synonym: Polyphylla comstockiana Bloeker, 1939: 150

Synonym: Polyphylla decemlineata laticauda Casey, 1914: 345

Synonym: Polyphylla decemlineata parilis Casey, 1914: 345

Synonym: Polyphylla decemlineata reducta Casey, 1914: 345

Synonym: Polyphylla matrona Casey, 1914: 349

Synonym: Polyphylla potosiana Casey, 1914: 349

Synonym: Polyphylla ruficollis Casey, 1914: 346

Synonym: Polyphylla ruficollis castanea Casey, 1914: 347

Synonym: Polyphylla ruficollis oregona Casey, 1914: 347

Synonym: Polyphylla squamotecta Casey, 1914: 343

Polyphylla devestiva Young Polyphylla devestiva Young, 1966: 233 Distribution: United States

Polyphylla diffracta Casey Polyphylla diffracta Casey, 1892: 18 Distribution: México, United States

Synonym: Polyphylla adusta Casey, 1914: 331

Synonym: Polyphylla alleni Cazier, 1939b: 201

Synonym: Polyphylla diffracta arida Van Dyke, 1947: 161

143 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Polyphylla fuscula Fall, 1908: 161

Synonym: Polyphylla laevicauda Casey, 1914: 338

Synonym: Polyphylla opposita Casey, 1914: 330

Synonym: Polyphylla uteana Tanner, 1928: 276

Polyphylla donaldsoni Skelley Polyphylla donaldsoni Skelley, 2005: 150 Distribution: United States

Polyphylla erratica Hardy Polyphylla erratica Hardy, 1978: 4 Distribution: United States

Polyphylla gracilis Horn Polyphylla gracilis Horn, 1881: 7 Distribution: United States

Polyphylla hammondi LeConte Polyphylla hammondi LeConte, 1856: 228 Distribution: Canada, México, United States

Synonym: Polyphylla diffusa Casey, 1914: 329

Synonym: Polyphylla latifrons Casey, 1914: 339

Synonym: Polyphylla oblita Casey, 1914: 326

Synonym: Polyphylla oblita impigra Casey, 1914

Synonym: Polyphylla oklahomensis Hatch, 1926: 145

Synonym: Polyphylla pimalis Casey, 1914: 329

Synonym: Polyphylla proba Casey, 1914: 328

Synonym: Polyphylla rufescenta Tanner, 1928: 276

Synonym: Polyphylla sejuncta Casey, 1914: 328

Synonym: Polyphylla speciosa Casey, 1889: 168

Synonym: Polyphylla speciosa acomana Casey, 1914: 342

Synonym: Polyphylla squamicauda Casey, 1914: 324

144 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Polyphylla squamicauda molesta Casey, 1914: 324

Synonym: Polyphylla subvittata LeConte, 1856: 229

Synonym: Polyphylla subvittata bisinuata Casey, 1914: 327

Synonym: Polyphylla verecunda Casey, 1914: 325

Polyphylla hirsuta Van Dyke Polyphylla hirsuta Van Dyke, 1933: 116 Distribution: United States

Polyphylla mescalerensis Young Polyphylla mescalerensis Young, 1988: 64 Distribution: United States

Polyphylla modulata Casey Polyphylla modulata Casey, 1914: 333 Polyphylla decemlineata modulata Casey; Cazier, 1939b: 202 Distribution: United States

Polyphylla monahansensis Hardy Polyphylla monohansensis Hardy, 1978: 7 Distribution: United States

Polyphylla multimaculata Hardy Polyphylla multimaculata Hardy, 1981: 300 Distribution: México

Polyphylla nigra Casey Polyphylla crinita nigra Casey, 1914: 334 Polyphylla nigra Casey: Young, 1988: 67 Distribution: México, United States

Synonym: Polyphylla incolumis relicta Casey, 1914: 336

Synonym: Polyphylla incolumis robustula Casey, 1914: 336

Synonym: Polyphylla martini Bloeker, 1939: 150

Synonym: Polyphylla santacruzae Bloeker, 1939: 149

Polyphylla nubila Van Dyke Polyphylla nubila Van Dyke, 1947: 160 Distribution: United States

Polyphylla occidentalis (Linné) Scarabaeus occidentalis Linné, 1767: 555 145 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Melolontha occidentalis (Linné); Fabricius, 1775: 32 Polyphylla occidentalis (Linné); Burmeister, 1855: 408 Distribution: United States

Polyphylla petitii (Guérin-Méneville) Melolontha petitii Guérin-Méneville 1844a: 97 Polyphylla petitii Guérin-Méneville; Lacordaire, 1856: 293 Distribution: México, Nicaragua

Synonym: Polyphylla dubia Hatch, 1971: 477

Synonym: Melolontha leucogramma Blanchard, 1851: 161

Polyphylla pottsorum Hardy Polyphylla pottsorum Hardy, 1978: 6 Distribution: United States

Polyphylla pubescens Cartwright Polyphylla pubescens Cartwright, 1939: 362 Polylamina pubescens (Cartwright); Hardy, 1974: 6 Polyphylla pubescens Cartwright; Coca-Abia, 2000: 19 Distribution: United States

Polyphylla ratcliffei Young Polyphylla ratcliffei Young, 1986: 49 Distribution: United States

Polyphylla rugosipennis Casey Polyphylla rugosipennis Casey, 1914: 337 Distribution: México, United States

Synonym: Polyphylla peninsularis Van Dyke, 1943: 103

Polyphylla sobrina Casey Polyphylla sobrina Casey, 1914: 339 Distribution: United States

Polyphylla squamiventris Cazier Polyphylla squamiventris Cazier, 1939b: 199 Distribution: México, United States

Polyphylla stellata Young Polyphylla stellata Young, 1986: 50 Distribution: United States

Polyphylla variolosa (Hentz) Melolontha variolosa Hentz, 1830: 256 Polyphylla variolosa (Hentz); Harris, 1841: 30 146 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: United States

Polyphylla woodruffi Skelley Polyphylla woodruffi Skelley, 2005: 148 Distribution: United States

Genus THYCE LeConte Thyce LeConte, 1856: 232. Type species Thyce squamicollis LeConte, 1856, by monotypy.

Keys to species: Hardy, 1974: 23

Thyce deserta Hardy Thyce deserta Hardy, 1974: 23 Distribution: United States

Thyce squamicollis LeConte Thyce squamicollis LeConte, 1856: 232 Distribution: México, United States

Tribe DIPLOTAXINI Burmeister, 1855

Genus DIPLOTAXIS Kirby Diplotaxis Kirby, 1837: 129. Type species Diplotaxis tristis Kirby, by monotypy.

Synonym: Alobus LeConte, 1856: 273. Type species Alobus fulvus LeConte, by monotypy.

Synonym: Diazus LeConte, 1859: 9. Type species Diazus rudis LeConte, by monotypy.

Synonym: Orsonyx LeConte, 1856: 265. Type species Orsonyx anxius LeConte, by monotypy.

Keys to species: Blatchley, 1910: 954 (Indiana); Fall, 1909: 11 (United States); Vaurie, 1956: 2 (eastern United States); Vaurie, 1958: 282 (revision); 1960: 180 (revision); 1963: 69 (Baja California); Ritcher, 1966: 277 (larvae); Lago et al., 1979: 34 (North Dakota); Delgado, 1990: 69; Ratcliffe, 1991: 172 (Nebraska); Delgado and Rivera, 1992: 328 (puberea group); Delgado and Capistran, 1993: 268 (trapezifera group); Morón, 1994: 40 (Hidalgo); Harpootlian, 2001: 73 (South Carolina).

Diplotaxis abnormis Fall Diplotaxis abnormis Fall, 1909: 78 Distribution: México, United States

Diplotaxis academia Vaurie Diplotaxis academia Vaurie, 1960: 242 Distribution: México

Diplotaxis aenea Blanchard Diplotaxys [sic] aenea Blanchard, 1851: 172 Diplotaxis aenea Blanchard; Burmeister, 1855: 536 Distribution: México, United States 147 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Liogenys pauperata Burmeister, 1855: 16 Diplotaxis pauperata (Burmeister); Evans, 2003: 179

Diplotaxis aequalis Cazier Diplotaxis aequalis Cazier, 1940c: 129 Distribution: United States

Diplotaxis aereomicans Moser Diplotaxis aereomicans Moser, 1918d: 316 Distribution: México

Diplotaxis alphamartinezi Delgado Diplotaxis alphamartinezi Delgado, 1990: 63 Distribution: México

Diplotaxis alutacea Bates Diplotaxis alutacea Bates, 1889: 398 Distribution: Guatemala

Diplotaxis ambigua Fall Diplotaxis ambigua Fall, 1909: 66 Distribution: México, United States

Diplotaxis amecameca Vaurie Diplotaxis amecameca Vaurie, 1960: 263 Distribution: México

Diplotaxis angularis LeConte Diplotaxis angularis LeConte, 1856: 268 Distribution: México, United States

Synonym: Diplotaxis producta Fall, 1932: 195

Diplotaxis angustula Moser Diplotaxis angustula Moser, 1918d: 299 Distribution: El Salvador, Guatemala, México

Diplotaxis anneae McCleve Diplotaxis anneae McCleve, 1993: 44 Distribution: México

Diplotaxis anthracina Fall Diplotaxis anthracina Fall, 1909: 71 Distribution: México

Diplotaxis anxius (LeConte) Orsonyx anxius LeConte, 1856: 266 148 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis anxius (LeConte); Fall, 1909: 70 Distribution: United States

Diplotaxis arctifrons Bates Diplotaxis arctifrons Bates, 1888: 163 Distribution: México

Diplotaxis arizonica Schaeffer Diplotaxis arizonica Schaeffer, 1907: 63 Distribution: México, United States

Diplotaxis assimilis Vaurie Diplotaxis assimilis Vaurie, 1960: 410 Distribution: México

Diplotaxis atlantis Fall Diplotaxis atlantis Fall, 1909: 44 Distribution: United States

Synonym: Diplotaxis nigra Cartwright, 1939: 361

Diplotaxis atramentaria Bates Diplotaxis atramentaria Bates, 1888: 163 Distribution: México

Diplotaxis atratula LeConte Diplotaxis atratula LeConte, 1856: 270 Distribution: México, United States

Diplotaxis aulacochela Cazier Diplotaxis aulacochela Cazier, 1940c: 131 Distribution: México, United States

Diplotaxis aurata Bates Diplotaxis aurata Bates, 1889: 398 Distribution: Guatemala

Diplotaxis aurora Wickham Diplotaxis aurora Wickham, 1914: 294 Distribution: United States Remark: fossil species

Diplotaxis australis Vaurie Diplotaxis australis Vaurie, 1960: 310 Distribution: México

Diplotaxis bakeri Cazier Diplotaxis bakeri Cazier, 1940b: 139 149 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Synonym: Diplotaxis volatica Cazier, 1940b: 140

Diplotaxis barbarae Vaurie and Cazier Diplotaxis barbarae Vaurie and Cazier, 1955: 13 Distribution: México

Diplotaxis basalis Fall Diplotaxis basalis Fall, 1909: 73 Distribution: United States

Diplotaxis belfragei Fall Diplotaxis belfragei Fall, 1909: 43 Distribution: México, United States

Synonym: Diplotaxis belfragei sinuatua Fall, 1909: 44

Synonym: Diplotaxis cartwrighti Cazier, 1940b: 144

Synonym: Diplotaxis protensa Fall, 1932: 193

Synonym: Diplotaxis stabilis Cazier, 1940b: 145

Diplotaxis beyeri Schaeffer Diplotaxis beyeri Schaeffer, 1907: 66 Distribution: México, United States

Diplotaxis bicolor Bates Diplotaxis trapezifera var. bicolor Bates, 1887: 159 Diplotaxis bicolor Bates; Vaurie, 1960: 215 Distribution: Guatemala, México

Diplotaxis bidentata LeConte Diplotaxis bidentata LeConte, 1856: 271 Distribution: United States

Diplotaxis bifida Vaurie Diplotaxis bifida Vaurie, 1960: 249 Distribution: México

Diplotaxis blanchardi Vaurie Diplotaxis blanchardi Vaurie, 1956: 4 Distribution: United States

Diplotaxis boops Bates Diplotaxis boops Bates, 1888: 165 Distribution: México, United States 150 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis bowditchi Fall Diplotaxis bowditchi Fall, 1909: 25 Distribution: México

Diplotaxis brachyptera Vaurie Diplotaxis brachyptera Vaurie, 1960: 257 Distribution: México, United States

Diplotaxis brevicollis LeConte Diplotaxis brevicollis LeConte, 1856: 267 Distribution: Canada, México, United States

Synonym: Ancylonycha nigropicea Walker, 1866: 322 Diplotaxis nigropicea (Walker); Evans, 2003: 183

Synonym: Diplotaxis transversicollis Moser, 1918d: 309

Diplotaxis brevicornis Cazier Diplotaxis brevicornis Cazier, 1940c: 136 Distribution: México, United States

Diplotaxis brevidens LeConte Diplotaxis brevidens LeConte, 1856: 272 Distribution: México, United States

Synonym: Diplotaxis laeviscutata Moser, 1918d: 313

Diplotaxis brevipilosa Moser Diplotaxis brevipilosa Moser, 1918d: 302 Distribution: Guatemala

Diplotaxis brevisetosa Linell Diplotaxis brevisetosa Linell, 1896: 400 Distribution: United States

Diplotaxis caelestis Delgado and Capistran Diplotaxis caelestis Delgado and Capistran, 1993: 269 Distribution: México

Diplotaxis carbonata LeConte Diplotaxis carbonata LeConte, 1856: 270 Distribution: México, United States

Synonym: Diplotaxis arcuata Fall, 1909: 75

Synonym: Diplotaxis levicula Casey, 1885a: 178

151 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis carinata Vaurie Diplotaxis carinata Vaurie, 1958: 373 Distribution: México

Diplotaxis carinifrons Bates Diplotaxis carinifrons Bates, 1889: 398 Distribution: Guatemala, México

Diplotaxis catarinas Vaurie and Cazier Diplotaxis catarinas Vaurie and Cazier, 1955: 20 Distribution: México

Diplotaxis cavifrons Moser Diplotaxis cavifrons Moser, 1918d: 304 Distribution: Guatemala, México

Diplotaxis chiricahuae Fall Diplotaxis chiricahuae Fall, 1909: 86 Distribution: México, United States

Diplotaxis circulans Vaurie Diplotaxis circulans Vaurie, 1960: 368 Distribution: México

Diplotaxis clypeata Bates Diplotaxis clypeata Bates, 1887: 157 Distribution: México

Synonym: Diplotaxis albosetosa Moser, 1921b: 180

Diplotaxis commixta Vaurie Diplotaxis commixta Vaurie, 1960: 339 Distribution: México

Diplotaxis completa Cazier Diplotaxis completa Cazier, 1940b: 141 Distribution: México, United States

Diplotaxis concava Vaurie Diplotaxis concava Vaurie, 1960: 213 Distribution: México

Diplotaxis conformis Fall Diplotaxis conformis Fall, 1909: 34 Distribution: United States

Diplotaxis confusa Fall Diplotaxis confusa Fall, 1909: 88 152 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Diplotaxis connata Schaeffer Diplotaxis connata Schaeffer, 1905: 158 Distribution: United States

Diplotaxis consentanea Bates Diplotaxis trapezifera var. consentanea Bates, 1887: 159 Diplotaxis consentanea Bates; Vaurie, 1960: 228 Distribution: México

Diplotaxis contracta Bates Diplotaxis contracta Bates, 1888: 166 Distribution: México

Diplotaxis convexilabrum Vaurie and Cazier Diplotaxis convexilabrum Vaurie and Cazier, 1955: 12 Distribution: México

Diplotaxis corbula Vaurie Diplotaxis corbula Vaurie, 1960: 400 Distribution: México, United States

Diplotaxis coriacea Bates Diplotaxis coriacea Bates, 1888: 161 Distribution: México

Diplotaxis corrosa Bates Diplotaxis corrosa Bates, 1888: 161 Distribution: México

Synonym: Diplotaxis corrosa pachuana Bates, 1888: 161

Diplotaxis corvina LeConte Diplotaxis corvina LeConte, 1856: 272 Distribution: México, United States

Diplotaxis costanera Vaurie Diplotaxis costanera Vaurie, 1958: 351 Distribution: México

Diplotaxis cribratella Bates Diplotaxis cribratella Bates, 1889: 399 Distribution: México

Synonym: Diplotaxis ennea Vaurie and Cazier, 1955: 7

Diplotaxis cribraticollis Blanchard 153 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxys [sic] cribraticollis Blanchard, 1851: 171 Diplotaxis cribraticollis Blanchard; Harold, 1869a: 1157 Distribution: México

Diplotaxis cribriceps Bates Diplotaxis cribriceps Bates, 1889: 398 Distribution: México

Diplotaxis cribulosa cribulosa LeConte Diplotaxis cribulosa LeConte, 1856: 270 Distribution: México, United States

Synonym: Diplotaxis popino Casey, 1885a: 179

Diplotaxis cribulosa sinaloa Vaurie Diplotaxis cribulosa sinaloa Vaurie, 1958: 331 Distribution: México

Diplotaxis crinigera Bates Diplotaxis crinigera Bates, 1888: 161 Distribution: El Salvador, Guatemala, México

Diplotaxis crucis Vaurie Diplotaxis crucis Vaurie, 1960: 229 Distribution: México

Diplotaxis curvaticeps Fall Diplotaxis curvaticeps Fall, 1909: 77 Distribution: México, United States

Diplotaxis dahli Cazier Diplotaxis dahli Cazier, 1940a: 251 Distribution: United States

Diplotaxis decima Vaurie and Cazier Diplotaxis decima Vaurie and Cazier, 1955: 10 Distribution: México

Diplotaxis denigrata Bates Diplotaxis trapezifera var. denigrata Bates, 1889: 397 Diplotaxis denigrata Bates; Vaurie, 1960: 208 Distribution: Costa Rica, El Salvador, Honduras, México

Diplotaxis dentella Fall Diplotaxis dentella Fall, 1909: 41 Distribution: México, United States

Diplotaxis denticeps Bates 154 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis trapezifiera denticeps Bates 1887: 159 Diplotaxis denticeps Bates, Vaurie, 1958: 287 Distribution: México, United States

Diplotaxis deserta Fall Diplotaxis deserta Fall, 1932b: 196 Distribution: United States

Diplotaxis dissona Vaurie Diplotaxis dissona Vaurie, 1960: 234 Distribution: México

Diplotaxis dubia LeConte Diplotaxis dubia LeConte, 1856: 269 Distribution: United States

Diplotaxis durango Vaurie Diplotaxis durango Vaurie, 1960: 248 Distribution: México

Diplotaxis elongata Vaurie Diplotaxis elongata Vaurie, 1960: 405 Distribution: México

Diplotaxis errans Fall Diplotaxis errans Fall, 1909: 84 Distribution: United States

Synonym: Diplotaxis ungula Cazier, 1940b: 137

Diplotaxis exstans Vaurie Diplotaxis exstans Vaurie, 1960: 335 Distribution: México

Diplotaxis femoralis Vaurie Diplotaxis femoralis Vaurie, 1960: 224 Distribution: México

Diplotaxis fimbriata Fall Diplotaxis fimbriata Fall, 1909: 56 Distribution: México, United States

Synonym: Diplotaxis arta Cazier, 1940b: 136

Diplotaxis fissilabris Fall Diplotaxis fissilabris Fall, 1909: 32 Distribution: México, United States

155 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis fissilis Vaurie and Cazier Diplotaxis fissilis Vaurie and Cazier, 1955: 23 Distribution: México

Diplotaxis flexa Vaurie Diplotaxis flexa Vaurie, 1960: 239 Distribution: México

Diplotaxis fossifrons Moser Diplotaxis fossifrons Moser, 1918d: 314 Distribution: México

Diplotaxis fossipalpa Fall Diplotaxis fossipalpa Fall, 1909: 57 Distribution: México, United States

Synonym: Diplotaxis villosipes Fall, 1932: 193

Diplotaxis foveicollis Moser Diplotaxis foveicollis Moser, 1918d: 315 Distribution: México

Diplotaxis frondicola (Say) Melolontha frondicola Say, 1825: 198 Diplotaxis frondicola (Say); LeConte, 1856: 269 Distribution: United States

Synonym: Diplotaxis testacea Burmeister, 1855: 263

Diplotaxis fulva (LeConte) Alobus fulvus LeConte, 1856: 273 Diplotaxis fulva (LeConte); Fall, 1909: 80 Distribution: United States

Diplotaxis geos McCleve Diplotaxis geos McCleve, 1993: 47 Distribution: México

Diplotaxis glabrimargo Vaurie and Cazier Diplotaxis glabrimargo Vaurie and Cazier, 1955: 22 Distribution: México

Diplotaxis guatemalica Moser Diplotaxis guatemalica Moser, 1918d: 306 Distribution: Costa Rica, Guatemala

Diplotaxis hallei Vaurie Diplotaxis hallei Vaurie, 1958: 291 156 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Diplotaxis harperi Blanchard Diplotaxys [sic] harperi Blanchard, 1851: 171 Diplotaxis harperi (Blanchard); LeConte, 1856: 269 Distribution: United States

Diplotaxis haydenii LeConte Diplotaxis haydenii LeConte, 1856: 272 Distribution: México, United States

Synonym: Diplotaxis impressifrons Cazier, 1940c: 127

Synonym: Diplotaxis innoxia LeConte, 1856: 273

Synonym: Diplotaxis levicoxa Fall, 1909: 39

Diplotaxis hebes Bates Diplotaxis hebes Bates, 1888: 164 Distribution: México

Synonym: Diplotaxis sericeiventris Moser, 1918d: 308

Diplotaxis hirsuta Vaurie Diplotaxis hirsuta Vaurie, 1958: 341 Distribution: Guatemala, México

Diplotaxis hirtipes Vaurie Diplotaxis hirtipes Vaurie, 1960: 204 Distribution: México

Diplotaxis icarus McCleve Diplotaxis icarus McCleve, 1993: 48 Distribution: México

Diplotaxis iguala Vaurie Diplotaxis iguala Vaurie, 1960: 251 Distribution: México

Diplotaxis illustris Fall Diplotaxis illustris Fall, 1909: 58 Distribution: México, United States

Diplotaxis impar Vaurie and Cazier Diplotaxis impar Vaurie and Cazier, 1955: 18 Distribution: México

Diplotaxis incisa Vaurie 157 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis incisa Vaurie, 1960: 221 Distribution: México

Diplotaxis incuria Fall Diplotaxis incuria Fall, 1932b: 196 Distribution: United States

Diplotaxis indigena Vaurie and Cazier Diplotaxis indigena Vaurie and Cazier, 1955: 6 Distribution: México

Diplotaxis ingenua Fall Diplotaxis ingenua Fall, 1909: 65 Distribution: México, United States

Synonym: Diplotaxis fusca Moser, 1918d: 312

Synonym: Diplotaxis futilis Fall, 1932: 194

Synonym: Diplotaxis rufocastanea Moser, 1918d: 307

Diplotaxis insignis LeConte Diplotaxis insignis LeConte, 1862: 346 Distribution: United States

Synonym: Diplotaxis falli Saylor, 1935b: 35

Diplotaxis jacala Vaurie Diplotaxis jacala Vaurie, 1958: 308 Distribution: México

Diplotaxis jamaicensis Cazier Diplotaxis jamaicensis Cazier, 1952: 3 Distribution: Jamaica

Diplotaxis jardeli Delgado and Rivera Diplotaxis jardeli Delgado and Rivera, 1992: 325 Distribution: México

Diplotaxis juquilensis Bates Diplotaxis juquilensis Bates, 1888: 162 Distribution: México

Diplotaxis knausii Schaeffer Diplotaxis knausii Schaeffer, 1907: 64 Distribution: México, United States

Diplotaxis kuschei Vaurie 158 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis kuschei Vaurie, 1960: 337 Distribution: México

Diplotaxis laevivertex Bates Diplotaxis laevivertex Bates, 1889: 397 Distribution: México

Diplotaxis languida LeConte Diplotaxis languida LeConte, 1878: 402 Distribution: United States

Diplotaxis latispina Vaurie Diplotaxis latispina Vaurie, 1960: 218 Distribution: México

Diplotaxis lengii Fall Diplotaxis lengii Fall, 1909: 74 Distribution: United States

Diplotaxis levicosta Fall Diplotaxis levicosta Fall, 1909: 87 Distribution: México, United States

Diplotaxis liberta (Germar) Melolontha liberta Germar, 1824: 123 Diplotaxis liberta (Germar); Fall, 1909: 46 Distribution: Martinique, United States

Synonym: Diplotaxys [sic] ebenina Blanchard, 1851: 170 Diplotaxis ebenina Blanchard; Burmeister, 1855: 262

Synonym: Diplotaxys [sic] georgiae Blanchard, 1851: 171 Diplotaxis georgiae Blanchard; Harold, 1869a: 1157

Synonym: Melolontha moesta Say, 1825: 197 (not Germar, 1824) Diplotaxys [sic] moesta (Say); Blanchard, 1851: 170 Diplotaxis moesta (Say); Harold, 1869a: 1157

Diplotaxis macronycha Fall Diplotaxis macronycha Fall, 1909: 55 Distribution: United States

Diplotaxis macrotarsus Vaurie Diplotaxis macrotarsus Vaurie, 1960: 219 Distribution: El Salvador, Guatemala, México

Diplotaxis magna Bates Diplotaxis magna Bates, 1888: 165 159 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México, United States

Synonym: Diplotaxis costulata Fall, 1909: 36

Synonym: Diplotaxis impressicollis Moser, 1918d: 309

Diplotaxis marginicollis Fall Diplotaxis marginicollis Fall, 1909: 67 Distribution: México, United States

Diplotaxis marina Vaurie Diplotaxis marina Vaurie, 1960: 247 Distribution: México

Diplotaxis martinezi Delgado and Capistran Diplotaxis martinezi Delgado and Capistran, 1993: 267 Distribution: México

Diplotaxis mascula Vaurie Diplotaxis mascula Vaurie, 1960: 238 Distribution: México

Diplotaxis maura Fall Diplotaxis maura Fall; 1909: 72 Distribution: México, United States

Diplotaxis maya Vaurie Diplotaxis maya Vaurie, 1958: 310 Distribution: Guatemala, México

Diplotaxis mediafusca Vaurie Diplotaxis mediafusca Vaurie, 1960: 203 Distribution: México

Diplotaxis megapleura Vaurie Diplotaxis megapleura Vaurie, 1959: 110 (nomen nudum) Diplotaxis megapleura Vaurie, 1960: 209 Distribution: México

Diplotaxis mentalis Fall Diplotaxis mentalis Fall, 1909: 33 Distribution: México, United States

Diplotaxis metallescens Bates Diplotaxis metallescens Bates, 1888: 164 Distribution: México

Diplotaxis mexcala Vaurie 160 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis mexcala Vaurie, 1960: 341 Distribution: México

Diplotaxis microchele Vaurie Diplotaxis microchele Vaurie, 1960: 346 Distribution: México

Diplotaxis microtrichia Moser Diplotaxis microtrichia Moser, 1921b: 179 Diplotaxis microtichia Moser; Vaurie, 1958: 305 Distribution: México

Diplotaxis mima Vaurie and Cazier Diplotaxis mima Vaurie and Cazier, 1955: 8 Distribution: México

Diplotaxis mimosae Fall Diplotaxis mimosae Fall, 1909: 22 Distribution: México, United States

Diplotaxis misella Fall Diplotaxis misella Fall, 1909: 91 Distribution: México, United States

Diplotaxis missionaria Vaurie Diplotaxis missionaria Vaurie, 1960: 311 Distribution: México, United States

Diplotaxis mistura Vaurie Diplotaxis mistura Vaurie, 1960: 401 Distribution: El Salvador, Honduras, México

Diplotaxis moerens moerens LeConte Diplotaxis moerens LeConte, 1856: 268 Distribution: México, United States

Diplotaxis moerens peninsularis Fall Diplotaxis peninsularis Fall, 1909: 70 Dipltoxis moerens peninsularis Fall; Vaurie, 1958: 323 Distribution: México

Diplotaxis monticola Delgado Diplotaxis monticola Delgado, 2001b: 139 Distribution: México

Diplotaxis muricata Schaeffer Diplotaxis muricata Schaeffer, 1907: 62 Distribution: México, United States 161 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Diplotaxis hispida Fall, 1909: 29

Synonym: Diplotaxis sparsesetosa Schaeffer, 1907: 63

Diplotaxis mus Fall Diplotaxis mus Fall, 1932: 192 Distribution: México, United States

Diplotaxis nigriventris Bates Diplotaxis nigriventris Bates, 1887: 160 Distribution: México

Diplotaxis nitidicollis Blanchard Diplotaxys [sic] nitidicollis Blanchard, 1851: 171 Diplotaxis nitidicollis Blanchard; Harold, 1869a: 1157 Distribution: México

Diplotaxis obregon Vaurie Diplotaxis obregon Vaurie, 1958: 332 Distribution: México

Diplotaxis obscura LeConte Diplotaxis obscura LeConte, 1859: 9 Distribution: Canada, United States

Synonym: Ancylonycha consequens Walker, 1866: 322 Diplotaxis consequens (Walker); Evans, 2003: 196

Synonym: Diplotaxis obscura compacta Fall, 1909: 48

Synonym: Diplotaxis obscura prominens Fall, 1909: 47

Diplotaxis ohausi Moser Diplotaxis ohausi Moser, 1921b: 181 Distribution: Guatemala, Honduras, México

Diplotaxis pacata LeConte Diplotaxis pacata LeConte, 1856: 272 Distribution: El Salvador, México, United States

Diplotaxis pala Vaurie Diplotaxis pala Vaurie, 1960: 222 Distribution: México

Diplotaxis parallela Fall Diplotaxis parallela Fall, 1909: 46 Distribution: United States 162 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Diplotaxis parkeri Cazier, 1940c: 133

Diplotaxis parpolita Vaurie Diplotaxis parpolita Vaurie, 1960: 238 Distribution: México

Diplotaxis parvicollis Fall Diplotaxis parvicollis Fall, 1909: 38 Distribution: México, United States

Synonym: Diplotaxis acononicus Cazier, 1940c: 131

Synonym: Diplotaxis labrata Moser, 1918d: 311

Diplotaxis parvula Burmeister Diplotaxis parvula Burmeister, 1855: 264 Distribution: México

Diplotaxis patyvaurieae Delgado Diplotaxis patyvaurieae Delgado, 1990: 67 Distribution: México

Diplotaxis persisae Cazier Diplotaxis persisae Cazier, 1940c: 133 Distribution: México, United States

Diplotaxis pilifera (Burmeister) Liogenys pilifera Burmeister, 1855: 16 Diplotaxis pilifera (Burmeister); Bates, 1887: 157 Distribution: México

Diplotaxis pilipennis Moser Diplotaxis pilipennis Moser, 1918d: 300 Distribution: México

Diplotaxis pinalica Fall Diplotaxis pinalica Fall, 1932: 196 Distribution: United States

Synonym: Diplotaxis microps Cazier, 1940c: 134

Synonym: Diplotaxis recticanthus Cazier, 1940c: 135

Diplotaxis planidens Fall Diplotaxis planidens Fall, 1909: 65 Distribution: México, United States

163 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis polita Fall Diplotaxis polita Fall, 1909: 83 Distribution: México

Diplotaxis poropyge Bates Diplotaxis poropyge Bates, 1887: 160 Distribution: Costa Rica, El Salvador, Guatemala, México, Panama

Synonym: Diplotaxis subrugosa Moser, 1918d: 297

Diplotaxis profunda Vaurie and Cazier Diplotaxis profunda Vaurie and Cazier, 1955: 17 Distribution: México

Diplotaxis puberea (Bates) Liogenys pubereus Bates, 1887: 156 Diplotaxis puberea (Bates); Vaurie, 1958: 304 Distribution: México

Diplotaxis puberula LeConte Diplotaxis puberulus LeConte, 1863b: 76 Diplotaxis puberula LeConte; Harold, 1869a: 1158 Distribution: México, United States

Synonym: Diplotaxis villosa Fall, 1909: 26

Diplotaxis pubipes Schaeffer Diplotaxis pubipes Schaeffer, 1907: 65 Distribution: México, United States

Diplotaxis pumila Fall Diplotaxis pumila Fall, 1909: 82 Distribution: United States

Diplotaxis punctata LeConte Diplotaxis punctata LeConte, 1856: 270 Distribution: México, United States

Synonym: Diplotaxis acerba Fall, 1909: 64

Synonym: Diplotaxis fulgida Cazier, 1940c: 130

Diplotaxis punctatorugosa Blanchard Diplotaxys [sic] punctatorugosa Blanchard, 1851: 171 Diplotaxis punctatorugosa Blanchard; Burmeister, 1855: 263 Distribution: United States

Synonym: Diplotaxis densicollis Fall, 1909: 52 164 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Diplotaxis excavata LeConte, 1856: 267

Synonym: Diplotaxis frontalis LeConte, 1856: 268

Diplotaxis puncticeps Moser Diplotaxis puncticeps Moser, 1918d: 310 Distribution: México, United States

Diplotaxis puncticollis Moser Diplotaxis puncticollis Moser, 1918d: 303 Distribution: El Salvador, Guatemala, México

Diplotaxis punctipennis LeConte Diplotaxis punctipennis LeConte, 1856: 268 Distribution: United States

Diplotaxis punctulata Horn Diplotaxis punctulata Horn, 1894: 397 Distribution: México

Diplotaxis pygidialis Vaurie Diplotaxis pygidialis Vaurie, 1960: 320 Distribution: México

Diplotaxis residua Fall Diplotaxis residua Fall, 1909: 34 Distribution: United States

Diplotaxis rex Vaurie Diplotaxis rex Vaurie, 1958: 378 Distribution: United States

Diplotaxis rita Vaurie Diplotaxis rita Vaurie, 1958: 368 Distribution: Guatemala, México

Diplotaxis roberti Vaurie Diplotaxis roberti Vaurie, 1960: 223 Distribution: México

Diplotaxis rockefelleri Vaurie and Cazier Diplotaxis rockefelleri Vaurie and Cazier, 1955: 5 Distribution: México

Diplotaxis rosae Vaurie and Cazier Diplotaxis rosae Vaurie and Cazier, 1955: 15 Distribution: México 165 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis rotunda Vaurie Diplotaxis rotunda Vaurie, 1960: 395 Distribution: United States

Diplotaxis rudis (LeConte) Diazus rudis LeConte, 1859: 10 Diplotaxis rudis (LeConte); Vaurie, 1958: 379 Distribution: United States

Diplotaxis rufa Linell Diplotaxis rufa Linell, 1896: 725 Distribution: United States

Diplotaxis rufiola Fall Diplotaxis rufiola Fall, 1909: 63 Distribution: México, United States

Synonym: Diplotaxis fulva Robinson, 1946: 58 (not LeConte, 1856: 273)

Diplotaxis rufipes Moser Diplotaxis rufipes Moser, 1921b: 180 Distribution: México

Diplotaxis rugosifrons Moser Diplotaxis rugosifrons Moser, 1918d: 301 Distribution: México

Diplotaxis rugosipennis Blanchard Diplotaxys [sic] rugosipennis Blanchard, 1851: 172 Diplotaxis rugosipennis Blanchard; Harold, 1869a: 1158 Distribution: México

Diplotaxis saltensis Vaurie Diplotaxis saltensis Vaurie, 1960: 233 Distribution: México

Diplotaxis saylori Cazier Diplotaxis saylori Cazier, 1940c: 128 Distribution: United States

Diplotaxis schaefferi Fall Diplotaxis schaefferi Fall, 1909: 32 Distribution: United States

Diplotaxis selanderi Vaurie Diplotaxis selanderi Vaurie, 1958: 354 Distribution: México 166 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis semifoveata Fall Diplotaxis semifoveata Fall, 1909: 38 Distribution: United States

Diplotaxis sierrae Fall Diplotaxis sierrae Fall, 1909: 45 Distribution: United States

Diplotaxis simillima Moser Diplotaxis simillima Moser, 1921b: 180 Distribution: México

Diplotaxis simplex Blanchard Diplotaxys [sic] simplex Blanchard, 1851: 172 Diplotaxis simplex Blanchard; Harold, 1969a: 1157 Distribution: México, United States

Synonym: Diplotaxis cephalotes Fall, 1909: 67

Synonym: Diplotaxis sinuaticeps Bates, 1888: 162

Diplotaxis simplicipes Wickham Diplotaxis simplicipes Wickham, 1912: 25 Distribution: United States Remark: fossil species

Diplotaxis sinuans Vaurie Diplotaxis sinuans Vaurie, 1960: 336 Distribution: México

Diplotaxis sonora Vaurie Diplotaxis sonora Vaurie, 1960: 217 Distribution: México

Diplotaxis sordida (Say) Melolontha sordida Say, 1825: 197 Diplotaxis sordida (Say); LeConte, 1856: 266 Distribution: Canada, United States

Synonym: Diplotaxis carbonaria Burmeister, 1855: 262

Synonym: Diplotaxis rugosioides Schaeffer, 1907: 61

Diplotaxis sparsa Vaurie Diplotaxis sparsa Vaurie, 1960: 245 Distribution: México

167 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis spina Vaurie Diplotaxis spina Vaurie, 1958: 307 Distribution: México

Diplotaxis squamisetis Delgado and Capistran Diplotaxis squamisetis Delgado and Capistran 1992: 578 Distribution: México

Diplotaxis statura Cazier Diplotaxis statura Cazier, 1940b: 143 Distribution: México, United States

Diplotaxis subangulata LeConte Diplotaxis subangulata LeConte, 1856: 271 Distribution: Canada, México, United States

Synonym: Diplotaxis californica Schaeffer, 1907: 66

Diplotaxis subcostata Blanchard Diplotaxys [sic] subcostata Blanchard, 1851: 171 Diplotaxis subcostata Blanchard; LeConte, 1856: 267 Distribution: United States

Synonym: Diplotaxis castanea Burmeister, 1855: 262

Diplotaxis subrugata Moser Diplotaxis subrugata Moser, 1918d: 298 Distribution: México

Diplotaxis sulcatula Fall Diplotaxis sulcatula Fall, 1909: 60 Distribution: México, United States

Diplotaxis superflua Vaurie Diplotaxis superflua Vaurie, 1960: 231 Distribution: México

Diplotaxis tarascana Vaurie Diplotaxis tarascana Vaurie, 1958: 352 Distribution: México

Diplotaxis tarsalis Schaeffer Diplotaxis tarsalis Schaeffer, 1907: 64 Distribution: México, United States

Synonym: Diplotaxis mexicana Moser, 1918d: 305

Diplotaxis tehuana Vaurie 168 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis tehuana Vaurie, 1960: 212 Distribution: México, Nicaragua

Diplotaxis tenebrosa Fall Diplotaxis tenebrosa Fall, 1909: 35 Distribution: Canada, United States

Diplotaxis tenuis LeConte Diplotaxis tenuis LeConte, 1856: 271 Distribution: United States

Diplotaxis tepicana Moser Diplotaxis tepicana Moser, 1918d: 300 Distribution: México

Diplotaxis texana LeConte Diplotaxis texana LeConte, 1856: 268 Distribution: United States

Diplotaxis thoracica Fall Diplotaxis thoracica Fall, 1909: 75 Distribution: México, United States

Synonym: Diplotaxis pinguescens Fall, 1909: 72

Diplotaxis trapezifera Bates Diplotaxis trapezifera Bates, 1887: 159 Distribution: Belize, Costa Rica, El Salvador, Guatemala, México, Nicaragua

Diplotaxis trementina Fall Diplotaxis trementina Fall, 1909: 69 Distribution: United States

Diplotaxis tristis Kirby Diplotaxis tristis Kirby, 1837: 130 Distribution: Canada, United States

Synonym: Diplotaxis corpulenta Burmeister, 1855: 263

Diplotaxis truncatula LeConte Diplotaxis truncatula LeConte, 1856: 269 Distribution: México, United States

Synonym: Diplotaxis consors LeConte, 1856: 269

Synonym: Diplotaxis morula LeConte, 1856: 270

Diplotaxis tumida Vaurie 169 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Diplotaxis tumida Vaurie, 1960: 227 Distribution: México

Diplotaxis turgidula Vaurie Diplotaxis turgidula Vaurie, 1960: 406 Distribution: México

Diplotaxis tzitzio Vaurie Diplotaxis tzitzio Vaurie, 1960: 338 Distribution: México

Diplotaxis ungulata Cazier Diplotaxis ungulatus Cazier, 1940c: 132 Diplotaxis ungulata Cazier; Vaurie, 1960: 384 Distribution: United States

Diplotaxis urbana Vaurie Diplotaxis urbana Vaurie, 1960: 286 Distribution: United States

Diplotaxis vandykei Vaurie Diplotaxis vandykei Vaurie, 1958: 382 Distribution: United States

Diplotaxis veracruzana Vaurie Diplotaxis veracruzana Vaurie, 1960: 404 Distribution: México

Diplotaxis xalapensis Delgado and Capistran Diplotaxis xalapensis Delgado and Capistran 1992: 574 Distribution: México

Diplotaxis yucateca Vaurie Diplotaxis yucateca Vaurie, 1960: 226 Distribution: Costa Rica, El Salvador, México

Diplotaxis zapoteca Vaurie Diplotaxis zapoteca Vaurie, 1958: 355 Distribution: México

Diplotaxis zeteki Vaurie Diplotaxis zeteki Vaurie, 1958: 311 Distribution: Panama

Genus HOMALOCHILUS Blanchard Homalochilus Blanchard, 1851: 172. Type species Homalochilus niger Blanchard, subsequent designation (Evans 2003: 206).

170 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Keys to species: Gutiérrez, 1952: 213

Homalochilus niger Blanchard Homalochilus niger Blanchard, 1851: 172 Distribution: Argentina

Homalochilus punctatostriatus Blanchard Homalochilus punctatostriatus Blanchard, 1851: 172 Distribution: Argentina

Synonym: Astanea bicolor Ohaus, 1911a: 557

Genus HOMOLIOGENYS Gutiérrez Homoliogenys Gutiérrez, 1952: 216. Type species Liogenys tarsalis Moser, by original designation.

Homoliogenys tarsalis Moser Liogenys tarsalis Moser, 1921a: 54 Homoliogenys tarsalis (Moser); Gutiérrez, 1952: 216 Distribution: Brazil

Genus LIOGENYS Guérin-Méneville Liogenys Guérin-Méneville, 1831b: pl. 3. Type species Melolontha palpalis Eschscholtz, by monotypy.

Synonym: Amphicrania Dejean, 1833: 163. Type species Melolontha palpalis Eschscholtz, by monotypy.

Synonym: Peritryssus Reitter, 1918: 77. Type species Peritryssus excisus Reitter by monotypy.

Keys to species: Gutiérrez , 1951: 132; Frey, 1969a: 38

Liogenys acutidens Moser Liogenys acutidens Moser, 1919b: 14 Distribution: Brazil

Liogenys bicuspis Moser Liogenys bicuspis Moser, 1919b: 14 Distribution: Brazil

Liogenys bidentata Burmeister Liogenys bidentata Burmeister, 1855: 13 Distribution: Brazil

Liogenys bidenticeps Moser Liogenys bidenticeps Moser, 1919b: 13 Distribution: Brazil

Liogenys bidentula Fairmaire 171 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Liogenys bidentulus Fairmaire, 1892: 243 Liogenys bidentula Fairmaire; Blackwelder, 1944: 227 Distribution: Argentina

Liogenys bilobata Frey Liogenys bilobatus Frey, 1969a: 58 Distribution: Brazil

Liogenys boliviensis Moser Liogenys boliviensis Moser, 1919b: 11 Distribution: Bolivia

Liogenys calcarata Frey Liogenys calcaratus Frey, 1970b: 283 Distribution: Argentina

Liogenys cartwrighti Frey Liogenys cartwrighti Frey, 1969a: 57 Distribution: Argentina

Liogenys concolor Blanchard Liogenys concolor Blanchard, 1851: 167 Distribution: Brazil

Liogenys corumbana Moser Liogenys corumbanus Moser, 1921b: 139 Liogenys corumbana Moser; Blackwelder, 1944: 227 Distribution: Brazil

Liogenys densata Frey Liogenys densatus Frey, 1969a: 52 Distribution: Argentina

Liogenys densicollis Moser Liogenys densicollis Moser, 1921b: 140 Distribution: Argentina

Synonym: Liogenys cribricollis Moser, 1921b: 140 Liogenys densicollis var. cribricollis Moser; Frey, 1969a: 38

Liogenys denticeps Blanchard Liogenys denticeps Blanchard, 1851: 167 Distribution: Bolivia

Liogenys denticulata Moser Liogenys denticulatus Moser, 1918b: 98 Liogenys denticulata Moser; Blackwelder, 1944: 227 Distribution: Argentina 172 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Liogenys diodon Burmeister Liogenys diodon Burmeister, 1855: 15 Distribution: Brazil

Synonym: Liogenys caviceps Frey, 1964c: 692

Liogenys elegans Nonfried Liogenis [sic] elegans Nonfried, 1891: 262 Liogenys elegans Nonfried; Moser, 1921b: 140 Distribution: Brazil

Synonym: Liogenys brasiliensis Moser, 1919b: 12

Liogenys excisa Reitter Peritryssus excisus Reitter, 1918: 77 Liogenys excisus (Reitter); Keith and Lacroix, 2003: 48 Liogenys excisa (Reitter); Keith, 2004: 154 Distribution: Neotropical (adventive in Sicile)

Liogenys ferrugata (Mannerheim) Geniates ferrugatus Mannerheim, 1829: 62 Liogenys ferrugata (Mannerheim); Ohaus, 1917: 52 Distribution: Brazil

Liogenys flaveola Moser Liogenys flaveolus Moser, 1924a: 122 Liogenys flaveola Moser; Blackwelder, 1944: 227 Distribution: Argentina

Liogenys flavicollis Frey Liogenys flavicollis Frey, 1964c: 693 Distribution: Argentina

Liogenys flavida Moser Liogenys flavidus Moser, 1918b: 106 Liogenys flavida Moser; Blackwelder, 1944: 227 Distribution: Argentina

Liogenys forcipata Frey Liogenys forcipatus Frey, 1970b: 282 Distribution: Argentina

Liogenys forsteri Frey Liogenys forsteri Frey, 1975c: 260 Distribution: Argentina

Liogenys fulvescens Blanchard 173 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Liogenys fulvescens Blanchard, 1851: 167 Distribution: Uruguay

Liogenys fusca Blanchard Liogenys fuscus Blanchard, 1851: 168 Liogenys fusca Blanchard; Blackwelder, 1944: 227 Distribution: Argentina, Bolivia, Brazil

Synonym: Liogenys argentinus Moser, 1918b: 97 Liogenys argentina Moser; Blackwelder, 1944: 227

Synonym: Liogenys cuyabanus Moser, 1919b: 12 Liogenys cuyabana Moser; Blackwelder, 1944: 227 Liogenys cuyabanis [sic] Moser; Frey, 1969a: 55

Liogenys gebieni Moser Liogenys gebieni Moser, 1921b: 140 Distribution: Colombia

Liogenys grandis Philippi and Philippi Liogenys grandis Philippi and Philippi, 1864: 322 Distribution: Chile

Liogenys hirtipennis Frey Liogenys hirtipennis Frey, 1969a: 53 Distribution: Brazil

Liogenys hirta Gutiérrez Liogenys hirtus Gutiérrez, 1951: 139 Distribution: Chile

Liogenys kadleci Frey Liogenys kadleci Frey, 1970b: 281 Distribution: Argentina

Liogenys kuntzeni Moser Liogenys kuntzeni Moser, 1921b: 139 Distribution: Chile

Liogenys laminiceps Moser Liogenys laminiceps Moser, 1919b: 16 Distribution: Brazil

Liogenys latipalpa Moser Liogenys latipalpus Moser, 1919b: 10 Distribution: Bolivia

Liogenys latitarsis Moser 174 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Liogenys latitarsis Moser, 1918b: 99 Distribution: Argentina

Liogenys leechi Frey Liogenys leechi Frey, 1967b: 419 Distribution: Peru

Liogenys macropelma Bates Liogenys macropelma Bates, 1887: 155 Distribution: Panama

Liogenys mendozana incisa Frey Liogenys mendozanus incisus Frey, 1969a: 52 Distribution: Argentina

Liogenys mendozana mendozana Moser Liogenys mendozana Moser, 1918b: 102 Liogenys mendozanus Moser; Frey, 1969a: 38 Distribution: Argentina

Liogenys micropyga Burmeister Liogenys micropyga Burmeister, 1855: 15 Distribution: Brazil

Liogenys minuta Moser Liogenys minutus Moser, 1924a: 122 Liogenys minuta Moser; Blackwelder, 1944; 227 Distribution: Argentina

Liogenys morio Burmeister Liogenys morio Burmeister, 1855: 16 Distribution: Argentina

Liogenys moseri Frey Liogenys moseri Frey, 1969a: 60 Distribution: Brazil

Liogenys nigrofusca Moser Liogenys nigrofuscus Moser, 1918b: 103 Liogenys nigrofusca Moser; Blackwelder, 1944: 227 Distribution: Argentina

Synonym: Liogenys longicrus Moser, 1918b: 100

Liogenys obesa Burmeister Liogenys obesa Burmeister, 1855: 15 Distribution: Brazil

175 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Liogenys obesina Frey Liogenys obesinus Frey, 1969a: 59 Distribution: Argentina

Liogenys obesula Gutiérrez Liogenys obesulus Gutiérrez , 1951: 135 Distribution: Chile

Liogenys obscura Blanchard Liogenys obscurus Blanchard, 1851: 167 Liogenys obscura Blanchard; Blackwelder, 1944: 227 Distribution: Argentina

Synonym: Liogenys bruchi Moser, 1924a: 122

Liogenys opacicollis Fairmaire Liogenys opacicollis Fairmaire, 1892: 243 Distribution: Argentina

Liogenys opacipennis Frey Liogenys opacipennis Frey, 1969a: 58 Distribution: Bolivia

Liogenys opthalmica Frey Liogenys opthalmicus Frey, 1973a: 256 Distribution: Paraguay

Liogenys pallens Blanchard Liogenys pallens Blanchard, 1851: 167 Distribution: Argentina, Uruguay

Liogenys pallidicornis Blanchard Liogenys pallidicornis Blanchard, 1851: 167 Distribution: Brazil

Liogenys palpalis (Eschscholtz) Melolontha palpalis Eschscholtz, 1822: 17 Liogenys palpalis (Eschscholtz); Guérin-Méneville, 1838: 85 Distribution: Chile

Synonym: Liogenys castaneus Guérin-Méneville, 1831b: pl.3

Synonym: Liogenys reichei Germain, 1903b: 51

Liogenys parallela Frey Liogenys parallelus Frey, 1965b: 144 Distribution: Bolivia

176 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Liogenys parva Blanchard Liogenys parvus Blanchard, 1851: 168 Liogenys parva Blanchard; Blackwelder, 1944: 228 Distribution: Argentina, Uruguay

Liogenys penai Gutiérrez Liogenys peñai Gutiérrez, 1951:141 Distribution: Chile

Liogenys peritryssoidea Keith Liogenys peritryssoidea Keith, 2004: 195 (for Moser, 1919: 15) Distribution: Brazil

Synonym: Liogenys excisus Moser, 1919b: 15 (not Reitter, 1918: 78) Liogenys excisa Moser; Blackwelder, 1944; 227

Liogenys pilosipennis Moser Liogenys pilosipennis Moser, 1918b: 101 Distribution: Brazil

Liogenys puberea cuprascens Bates Liogenys puberea var. cuprascens Bates, 1887: 156 Distribution: México

Liogenys puberea puberea Bates Liogenys pubereus Bates, 1887: 155 Liogenys puberea Bates; Blackwelder, 1944: 228 Distribution: México

Liogenys pubisternis Bates Liogenys pubisternis Bates, 1887: 155 Distribution: México

Liogenys punctaticollis (Blanchard) Hilarianus punctaticollis Blanchard, 1851: 169 Liogenys punctaticollis (Blanchard); Frey, 1974: 331 Distribution: Brazil

Synonym: Liogenys palmata Burmeister, 1855: 13 Liogenys palmatus Burmeister; Frey, 1974f: 331

Synonym: Liogenys tibialis Moser, 1918b: 107

Liogenys quadridens (Fabricius) Melolontha quadridens Fabricius, 1798: 131 Liogenys quadridens (Fabricius); Burmeister, 1855: 14 Distribution: Brazil, Colombia, Guyana, Panama, Venezuela

177 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Liogenys quadridentata Blanchard Liogenys quadridentatus Blanchard, 1851: 168 Liogenys quadridentata Blanchard; Blackwelder, 1944: 227 Distribution: Uruguay

Liogenys rufocastanea Moser Liogenys rufocastaneus Moser, 1918b: 103 Liogenys rufocastanea Moser; Blackwelder, 1944: 228 Distribution: Brazil, Paraguay

Liogenys rufoflava Moser Liogenys rufoflavus Moser, 1918b: 105 Liogenys rufoflava Moser; Blackwelder, 1944: 228 Distribution: Argentina

Liogenys rugosicollis Frey Liogenys rugosicollis Frey, 1969a: 63 Distribution: Argentina

Liogenys sanctaecrucis Blanchard Liogenys santae-crucis [sic] Blanchard, 1851: 167 Liogenys sanctae crucis Blanchard; Harold, 1869a: 1140 Liogenys sanctae-crucis Blanchard; Blackwelder, 1944: 228 Liogenys sanctaecrucis Blanchard; Frey, 1969a: 37 Distribution: Bolivia

Liogenys seabrai Martínez Liogenys seabrai Martínez, 1957: 51 Distribution: Brazil

Liogenys sinuaticeps Moser Liogenys sinuaticeps Moser, 1918b: 104 Distribution: Brazil

Liogenys spiniventris Moser Liogenys spiniventris Moser, 1918b: 108 Distribution: Brazil

Liogenys suturalis (Blanchard) Hilarianus suturalis Blanchard, 1851: 169 Liogenys suturalis (Blanchard); Frey, 1969a: 54 Distribution: Argentina, Brazil

Liogenys testaceipennis Moser Liogenys testaceipennis Moser, 1918b: 109 Distribution: Brazil

Liogenys unicolor Evans 178 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Liogenys unicolor Evans, 2003 (for Blanchard, 1851: 170) Distribution: Brazil

Synonym: Hilarianus concolor Blanchard, 1851: 170 Liogenys concolor (Blanchard); Frey, 1974f: 331 (not Blanchard, 1851: 167)

Liogenys vicina Frey Liogenys vicinus Frey, 1969a: 62 Distribution: Argentina

Liogenys wagenknechti Gutiérrez Liogenys wagenknechti Gutiérrez, 1951: 137 Distribution: Chile

Liogenys xanthocera Harold Liogenys xanthocerus Harold, 1869b: 123 (for Blanchard, 1851: 168) Liogenys xanthocera Harold; Blackwelder, 1944: 228 Distribution: Argentina

Synonym: Liogenys pallidicornis Blanchard, 1851: 168 (not Blanchard, 1851: 167)

Liogenys zischkai Frey Liogenys zischkai Frey, 1965b: 146 Distribution: Bolivia

Genus PACHRODEMA Blanchard Pachrodema Blanchard, 1851: 168. Type species Pachrodema castanea Blanchard, subsequent designation (Evans 2003: 215).

Pachrodema abnormis Moser Pachrodema abnormis Moser, 1918d: 324 Distribution: Argentina

Pachrodema castanea Blanchard Pachrodema castanea Blanchard, 1851: 168 Distribution: Chile

Pachrodema cuprascens Burmeister Pachrodema cuprascens Burmeister, 1855: 260 Distribution: Argentina

Pachrodema flaveola Moser Pachrodema flaveola Moser, 1918d: 322 Distribution: Argentina

Pachrodema lucida Berg Pachrodema lucida Berg, 1881: 100 Distribution: Argentina 179 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Pachrodema picea Nonfried Pachrodema picea Nonfried, 1894: 115 Distribution: Argentina

Pachrodema piceola Moser Pachrodema piceola Moser, 1918d: 322 Distribution: Argentina

Pachrodema pruinosa Moser Pachrodema pruinosa Moser, 1918d: 323 Distribution: Argentina

Pachrodema substriata Blanchard Pachrodema substriata Blanchard, 1851: 169 Distribution: Argentina

Genus PACUVIA Curtis Pacuvia Curtis, 1845: 453. Type species Pacuvia castanea Curtis, by monotypy.

Keys to species: Gutiérrez, 1951: 143

Pacuvia castanea Curtis Pacuvia castanea Curtis, 1845: 453 Liogenys castaneus (Curtis); Philippi, 1887: 688 Distribution: Chile

Synonym: Liogenys gayanus Solier, 1851: 100

Pacuvia philippiana Gutiérrez Pacuvia philippiana Gutiérrez, 1951: 144 Distribution: Chile

Genus PSEUDODIPLOTAXIS Nonfried Pseudodiplotaxis Nonfried, 1894: 117. Type species Pseudodiplotaxis albosetosa Nonfried, by monotypy.

Pseudodiplotaxis albosetosa Nonfried Pseudodiplotaxis albosetosa Nonfried, 1894: 118 Distribution: Brazil

Tribe PACHYDEMINI Reitter, 1902

Keys to genera: Martínez, 1958: 100; 1975: 228.

Genus ACYCLOCHILUS Ohaus Acyclochilus Ohaus, 1909: 433. Type species Acylochilus strumosus Ohaus, by subsequent designation (Martínez, 1975: 243). 180 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Subgenus: Acylochoides Martínez, 1975: 231 (as subgenus). Type species Acylochilus persimilis Martínez, by original designation.

Acyclochilus assimilis Ohaus Acyclochilus assimilis Ohaus, 1910: 177 Acyclochilus (Acyclochilus) assimilis Ohaus; Martínez, 1975: 244 Distribution: Argentina

Acyclochilus curvidens Ohaus Acyclochilus curvidens Ohaus, 1910: 177 Acyclochilus (Acyclochilus) curvidens Ohaus; Martínez, 1975: 244 Distribution: Argentina

Acyclochilus ottianus Ohaus Acyclochilus ottianus Ohaus, 1909: 435 Acyclochilus (Acyclochilus) ottianus Ohaus; Martínez, 1975: 244 Distribution: Argentina

Acyclochilus persimilis Martínez Acyclochilus persimilis Martínez, 1959b: 61 Acyclochilus (Acyclochoides) persimilis Martínez; Martínez, 1975: 233 Distribution: Argentina

Acyclochilus strumosus Ohaus Acyclochilus strumosus Ohaus, 1909: 434 Acyclochilus (Acyclochilus) strumosus Ohaus; Martínez, 1975: 244 Distribution: Argentina

Genus ANAHI Martínez Anahi Martínez, 1958: 102. Type species Anahi guaraniticus Martínez, by original designation.

Anahi guaraniticus Martínez Anahi guaraniticus Martínez, 1958: 106 Distribution: Paraguay

Genus WARWICKIA Smith and Evans Warwickia Smith and Evans, 2005: 42. Type species Benedictia pilosa Sanderson, by original designation.

Synonym: Benedictia Sanderson, 1939a: 1 (not Dybowski, 1875: 4 [Mollusca]). Type species Benedictia pilosa Sanderson, by original designation.

Warwickia pilosa Sanderson Benedictia pilosa Sanderson, 1939a: 2 Warwickia pilosa (Sanderson); Smith & Evans, 2005: 42 Distribution: México, United States

181 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Genus BURMEISTERIELLUS Berg Burmeisteriellus Berg, 1898: 17. Type species Burmeisteria mirablilis Schickendantz, by monotypy.

Synonym: Burmeisteria Schickendantz, 1868: 101 (not Gray, 1865)

Burmeisteriellus mirabilis (Schickendantz) Burmeisteria mirabilis Schickendantz, 1868: 102 Burmeisteriellus mirabilis (Schickendantz); Berg, 1898: 17 Distribution: Argentina

Genus CASTANOCHILUS Ohaus Castanochilus Ohaus, 1909: 436. Type species Castanochilus bruchianus Ohaus, by monotypy.

Castanochilus bruchianus Ohaus Castanochilus bruchianus Ohaus, 1909: 436 Distribution: Argentina

Genus FAARGIA Martínez Faargia Martínez, 1982: 89. Type species Myloxena gentilii Martínez, by original designation.

Faargia gentillii (Martínez) Myloxena gentillii Martínez, 1975: 239 Faargia gentillii (Martínez); Martínez, 1982: 91 Distributon: Argentina

Genus LEURETRA Erichson Leuretra Erichson, 1847a: 100. Type species Leuretra pectoralis Erichson, by monotypy.

Leuretra pectoralis Erichson Leuretra pectoralis Erichson, 1847a: 101 Distribution: ?Chile, Peru

Genus LICHNIOPS Gutiérrez Lichniops Gutiérrez, 1946b: 49. Type species Lichniops barberoi Gutiérrez , by original designation.

Lichniops barberoi Gutiérrez Lichniops barberoi Gutiérrez, 1946b: 50 Distribution: Argentina, Bolivia, Paraguay

Genus LICHNIOPSOIDES Martínez Lichniopsoides Martínez, 1953: 3. Type species Lichniopsoides breyeri Martínez, by original designation.

Lichniopsoides breyeri Martínez Lichniopsoides breyeri Martínez, 1953: 3 Distribution: Argentina

Genus LONGICRURA Frey 182 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Longicrura Frey, 1974c: 139. Type species Longicrura fusca, by monotypy.

Longicrura fusca Frey Longicrura fusca Frey, 1974c: 139 Distribution: Brazil

Genus LUISPENAIA Martínez Luispenaia Martínez, 1972a:86. Type species Luispenaia paradoxa Martínez, by original designation.

Luispenaia paradoxa Martínez Luispenaia paradoxa Martínez, 1972a: 89 Distribution: Chile

Genus MADINIELLA Chalumeau and Gruner Madiniella Chalumeau and Gruner, 1976: 102. Type species Madiniella christinae Chalumeau and Gruner, by original designation.

Madiniella christinae Chalumeau and Gruner Madiniella christinae Chalumeau and Gruner,1976: 102 Distribution: Martinique

Genus MYLOXENA Berg Myloxena Berg, 1881: 100. Type species Myloxena vestita Berg, by monotypy.

Myloxena bruchiana Ohaus Myloxena bruchiana Ohaus, 1909: 440 Distribution: Argentina

Myloxena patagonica Ohaus Myloxena patagonica Ohaus, 1909: 440 Distribution: Argentina

Myloxena vestita Berg Myloxena vestita Berg, 1881: 100 Distribution: Argentina

Myloxena vicina Moser Myloxena vicina Moser, 1926: 207 Distribution: Argentina

Genus MYLOXENOIDES Martínez Myloxenoides Martínez, 1975: 233. Type species Myloxenoides tandilensis Martínez, by original designation.

Myloxenoides tandilensis Martínez Myloxenoides tandilensis Martínez, 1975: 235 Distribution: Argentina

183 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Genus NEOGUTIERREZIA Martínez Neogutierrezia Martínez, 1953: 2. Type species Neogutierrezia mirablis Martínez, by original designation.

Keys to species: Martínez, 1973: 30

Neogutierrezia araucana Martínez Neoguttierrezia araucana Martínez, 1973: 36 Distribution: Argentina

Neogutierrezia miriabilis affinis Martínez Neoguttierrezia miriablis affinis Martínez, 1973: 35 Distribution: Argentina

Neogutierrezia miriablis miriablis Martínez Neoguttierrezia miriablis Martínez, 1953: 2 Neoguttierrezia miriablis miriablis Martínez; Martínez, 1973: 31 Distribution: Argentina

Genus PARAPETIIA Martínez Parapetiia Martínez, 1961: 1. Type species Parapetiia camba Martínez, by original designation.

Parapetiia camba Martínez Parapetiia camba Martínez, 1961: 4 Distribution: Bolivia

Genus PENTACORYNA Moser Pentacoryna Moser, 1926: 207. Type species Pentacoryna bruchi Moser, by monotypy.

Pentacoryna bruchi Moser Pentacoryna bruchi Moser, 1926: 208 Distribution: Argentina

Genus PHOBETUS LeConte Phobetus LeConte, 1856: 227. Type species Phobetus comatus LeConte, by monotypy.

Keys to species: Barrett, 1935: 52; Cazier, 1937b: 81; Hardy, 1978b: 51.

Phobetus chearyi Hardy Phobetus chearyi Hardy, 1973: 128 Distribution: México

Phobetus ciliatus Barrett Phobetus ciliatus Barrett, 1935: 51 Distribution: United States

Phobetus comatus LeConte Phobetus comatus LeConte, 1856: 227 184 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Tryssus comatus (LeConte); LeConte, 1857:38 Phobetus comatus LeConte; LeConte, 1862: 140 Distribution: Canada, United States

Synonym: Phobetus centralis Casey, 1909: 282 Phobetus comatus var. centralis Casey; Cazier, 1937b: 83

Synonym:Phobetus comatus robinsoni Saylor, 1939a: 54

Synonym:Phobetus sloopi Barrett, 1933: 130 Phobetus comatus var. sloopi Barrett; Cazier, 1937b: 83

Synonym:Phobetus leachi Barrett, 1935: 51

Phobetus desertus Blom and Clark Phobetus desertus Blom and Clark, 1984: 305 Distribution: México

Phobetus humeralis Cazier Phobetus humeralis Cazier, 1937b: 85 Distribution: United States

Phobetus mojavus Barrett Phobetus mojavus Barrett, 1933: 130 Distribution: United States

Phobetus palpalis Saylor Phobetus palpalis Saylor, 1936b: 1 Distribution: United States

Phobetus panamintensis Hardy Phobetus panamintensis Hardy, 1978b: 50 Distribution: United States

Phobetus saylori Cazier Phobetus saylori Cazier, 1937a: 116 Distribution: United States

Phobetus sleeperi Hardy Phobetus sleeperi Hardy, 1978b: 50 Distribution: México

Phobetus testaceus LeConte, 1862: 346 Phobetus testaceus LeConte, 1862: 346 Distribution: United States

Synonym: Rhizotrogus collactus Walker, 1866: 321

185 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Genus PSEUDOLIOGENYS Moser Pseudoliogenys Moser, 1919c: 358. Type species Pseudoliogenys flavidus Moser, by monotypy.

Pseudoliogenys flavidus Moser Pseudoliogenys flavidus Moser, 1919c: 359 Distribution: Argentina

Genus PTYOPHIS Redtenbacher Ptyophis Redtenbacher, 1868: 69. Type species Ptyophis macrophylla Redtenbacher, by monotypy.

Synonym: Tetraphyllus Philippi, 1864: 448 (preoccupied). Type species Tetraphyllus paulseni Philippi, by monotypy.

Ptyophis paulseni (Philippi) Tetraphyllus paulseni Philippi, 1864: 448 Ptyophis paulseni (Philippi); Gutiérrez, 1950: 276 Distribution: Chile

Synonym: Ptyophis macrophylla Redtenbacher, 1868: 70

Tribe MACRODACTYLINI Burmeister, 1855 Dichelonyciini LeConte and Horn, 1883

Genus AGAOCNEMIS Moser Agaocnemis Moser, 1918b: 115. Type species Agaocnemis pruina Moser, by monotypy.

Agaocnemis pruina Moser Agaocnemis pruina Moser, 1918b: 116 Distribution: Brazil

Genus ALVARINUS Blanchard Alvarinus Blanchard, 1850: 123. Type species Alvarinus hilarii Blanchard, subsequent designation (Evans 2003: 224).

Alvarinus hilarii Blanchard Alvarinus hilarii Blanchard, 1850: 123 Distribution: Brazil

Alvarinus pallidipennis Blanchard Alvarinus pallidipennis Blanchard, 1850: 123 Distribution: Brazil

Synonym: Plectris rectangula Frey, 1967a: 40

Alvarinus submetallicus Blanchard Alvarinus submetallicus Blanchard, 1850: 123 Distribution: Argentina, Brazil

186 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Philochloenia chalcea Burmeister, 1855: 30 Plectris chalcea Burmeister; Frey, 1967a: 84

Alvarinus subsericeus Blanchard Alvarinus subsericeus Blanchard, 1850: 123 Distribution: Bolivia

Genus ANCISTROSOMA Curtis Ancistrosoma Curtis, 1835: 307. Type species Ancistrosoma klugii Curtis, by monotypy.

Keys to species: Arrow, 1913b: 425

Ancistrosoma argentinum Moser Ancistrosoma argentinum Moser, 1921b: 179 Distribution: Argentina

Ancistrosoma arrowi Ley Ancistrosoma arrowi Ley, 1923: 222 Distribution: Colombia

Ancistrosoma blanchardi Sallé Ancistrosoma blanchardi Sallé, 1886: 465 Distribution: Venezuela

Ancistrosoma buckleyi Sallé Ancistrosoma buckleyi Sallé, 1887: CXXVII Distribution: Venezuela

Ancistrosoma farinosum Sallé Ancystrosoma farinosum Sallé, 1849: 300 Ancistrosoma farinosa [sic] Sallé; Blanchard, 1850: 124 Distribution: Venezuela

Ancistrosoma flavovittatum Blanchard Ancistrosoma flavovittata Blanchard, 1850: 124 Ancistrosoma flavovittatum Blanchard; Harold, 1869a: 1155 Distribution: Colombia

Ancistrosoma hilare Arrow Ancistrosoma hilare Arrow, 1913b: 427 Distribution: Colombia, Peru

Ancistrosoma intermedium Arrow Ancistrosoma intermedium Arrow, 1913b: 426 Ancistrosoma intermedia [sic] Arrow; Blackwelder, 1944: 234 Distribution: Peru

Ancistrosoma klugii Curtis 187 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ancistrosoma klugii Curtis, 1835: 308 Ancistrosoma klugi Curtis; Harold, 1869a: 1155 Distribution: Peru

Ancistrosoma melolonthoides Sallé Ancistrosoma melolontoides Sallé, 1886: 468 Ancistrosoma melolonthoides Sallé; Dalla Torre, 1913: 338 Distribution: Venezuela

Ancistrosoma reductum Frey Ancystrosoma [sic] reductum Frey, 1964a: 132 Distribution: Peru

Ancistrosoma rufipes (Latreille) Melolontha rufipes Latreille, 1813: 94 Ancistrosoma rufipes (Latreille); Blanchard, 1850: 124 Distribution: Colombia

Ancistrosoma tobagoensis Arrow Ancistrosoma tobagoensis Arrow, 1913b: 428 Distribution: Tobago

Ancistrosoma trinitatis Arrow Ancistrosoma trinitatis Arrow, 1913b: 429 Distribution: Trinidad

Ancistrosoma vittigerum Erichson Ancistrosoma vittigera Erichson, 1847a: 103 Ancistrosoma vittigerum Erichson; Sallé, 1886: 466 Distribution: Ecuador

Genus ANOMALOCHILUS Blanchard Anomalochilus Blanchard, 1850: 123. Type species Anomalochilus singularis Blanchard, by monotypy.

Anomalochilus pilosellus Moser Anomalochilus pilosellus Moser, 1919b: 41 Distribution: Brazil

Anomalochilus singularis Blanchard Anomalochilus singularis Blanchard, 1850: 123 Distribution: Brazil

Genus ANOMONYX Saylor Anomonyx Saylor, 1940b: 46. Type species Anomalonyx uruguayensis Moser, by monotypy.

Synonym: Anomalonyx Moser, 1921b: 156 (preoccupied). Type species Anomalonyx uruguayensis Moser, by monotypy. 188 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Anomonyx uruguayensis (Moser) Anomalonyx uruguayensis Moser, 1921b: 156 Anomonyx uruguayensis (Moser); Saylor, 1940b: 46 Distribution: Uruguay

Genus ANOPLOSIAGUM Blanchard Anoplosiagum Blanchard, 1850: 119. Type species Anoplosiagum rufipenne (Fabricius), by subsequent designation (Lacrodaire, 1856: 264).

Keys to species: Chapin, 1932b: 174

Anoplosiagum antennale Moser Anoplosiagum antennale Moser, 1921b: 143 Distribution: Brazil

Anoplosiagum castaneum (Burmeister) Hadrocerus castaneum Burmeister, 1855: 23 Anoplosiagum castaneum (Burmeister); Harold, 1869a: 1142 Distribution: Brazil

Anoplosiagum cubensis Garcia-Vidal Anoplosiagum cubensis Garcia-Vidal, 1983: 330 Distribution: Cuba

Anoplosiagum flavicollis Chapin Anoplosiagum flavicollis Chapin, 1935b: 68 Distribution: Cuba

Anoplosiagum fulvum Moser Anoplosiagum fulvum Moser, 1918b: 119 Distribution: Brazil

Anoplosiagum gebieni Moser Anoplosiagum gebieni Moser, 1921b: 142 Distribution: Brazil

Anoplosiagum kuntzeni Moser Anoplosiagum kuntzeni Moser, 1921b: 142 Distribution: Colombia

Anoplosiagum macrophyllum Moser Anoplosiagum macrophyllum Moser, 1921b: 142 Distribution: Paraguay

Anoplosiagum oteroi Garcia-Vidal Anoplosiagum oteroi Garcia-Vidal, 1983: 332 Distribution: Cuba 189 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Anoplosiagum pallidulum Blanchard Anoplosiagum pallidulum Blanchard, 1850: 120 Distribution: Cuba

Anoplosiagum pauliani Chalumeau Anoplosiagum pauliani Chalumeau, 1979: 229 Distribution: Cuba

Anoplosiagum peltatum Chapin Anoplosiagum peltatum Chapin, 1932b: 176 Distribution: Cuba

Anoplosiagum pentaphyllum (Burmeister) Hadrocerus pentaphyllus Burmeister, 1855: 25 Anoplosiagum pentaphyllum (Burmeister); Harold, 1869a: 1143 Distribution: Colombia

Anoplosiagum rufipenne (Fabricius) Melolontha rufipennis Fabricius, 1801: 167 Anoplosiagum rufipenne (Fabricius); Blanchard, 1850: 119 Distribution: Brazil

Synonym: Melolontha filitarsis Germar, 1824: 125 Philochloenia filitarsis (Germar); Dejean, 1833: 180 Hadrocerus filitarsis (Germar); Burmeister, 1855: 25

Anoplosiagum rufum Chapin Anoplosiagum rufum Chapin, 1932b: 177 Distribution: Cuba

Anoplosiagum rutilum (Chevrolat) Clavipalpus rutilus Chevrolat, 1865: 22 Anoplosiagum rutilum (Chevrolat); Chapin, 1932b; 178 Distribution: Cuba

Anoplosiagum scabrosum Chapin Anoplosiagum scabrosum Chapin, 1932b: 175 Distribution: Cuba

Anoplosiagum scaramuzzai Chapin Anoplosiagum scaramuzzai Chapin, 1935b: 67 Distribution: Cuba

Anoplosiagum simplicipes Chapin Anoplosiagum simplicipes Chapin, 1932b: 175 Distribution: Cuba

190 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Anoplosiagum sjostedti Moser Anoplosiagum sjostedti Moser, 1921a: 54 Distribution: Brazil

Anoplosiagum sulcatulum setosellum Moser Anoplosiagum sulcatulum var. setosellum Moser, 1924a: 123 Distribution: Brazil

Anoplosiagum sulcatulum sulcatulum Blanchard Anoplosiagum sulcatulum Blanchard, 1850: 119 Distribution: Brazil

Anoplosiagum sulcipenne Moser Anoplosiagum sulcipenne Moser, 1918b: 118 Distribution: Brazil

Anoplosiagum swezeyi Garcia-Vidal Anoplosiagum swezeyi Garcia-Vidal, 1983: 333 Distribution: Cuba

Anoplosiagum truncatum Frey Anoplosiagum truncatum Frey, 1973a: 257 Distribution: Brazil

Anoplosiagum turquinensis Garcia-Vidal Anoplosiagum turquinensis Garcia-Vidal, 1983: 334 Distribution: Cuba

Anoplosiagum variabile Chevrolat Anoplosiagum variabile Chevrolat, 1865: 22 Distribution: Cuba

Anoplosiagum versicolor (Burmeister) Hadrocerus versicolor Burmeister, 1855: 24 Anoplosiagum versicolor (Burmeister); Harold, 1869a: 1143 Distribution: Brazil, Colombia

Anoplosiagum vicinum Frey Anoplosiagum vicinum Frey, 1973a: 257 Distribution: Brazil

Anoplosiagum villosicolle Blanchard Anoplosiagum villosicolle Blanchard, 1850: 119 Distribution: Brazil

Anoplosiagum zayasi Garcia-Vidal Anoplosiagum zayasi Garcia-Vidal, 1983: 335 Distribution: Cuba 191 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Genus ASTAENOPLIA Martínez Astaenoplia Martínez, 1957: 47. Type species Astaenoplia miserabilis Martínez, by original designation.

Astaenoplia miserabilis Martínez Astaenoplia miserabilis Martínez, 1957: 48 Distribution: Argentina

Genus ASTAENOSIAGUM Martínez Astaenosiagum Martínez, 1957: 50. Type species Schizochelus longipes Philippi, by original designation.

Astaenosiagum longipes (Philippi) Schizochelus longipes Philippi, 1861: 738 Astaenosiagum longipes (Philippi); Martínez, 1957: 51 Distribution: Chile

Genus AULANOTA Moser Aulanota Moser, 1924b: 157. Type species Aulanota sulcipennis Moser, by monotypy.

Aulanota sulcipennis Moser Aulanota sulcipennis Moser, 1924b: 157. Distribution: Brasil

Genus BARYBAS Blanchard Barybas Blanchard, 1850: 94. Type species Barybas nana Blanchard, by subsequent designation (Bates, 1887: 150).

Synonym: Microcrania Burmeister, 1855: 75. Type species Philochloenia compacta Erichson, subsequent designation (Evans 2003: 230).

Barybas aenescens Moser Barybas aenescens Moser, 1926: 206 Distribution: French Guiana

Barybas aenea Moser Barybas aeneus Moser, 1919a: 18 Barybas aenea Moser; Blackwelder, 1944: 233 Distribution: Venezuela

Barybas aurita Bates Barybas auritus Bates, 1887: 150 Barybas aurita Bates; Blackwelder, 1944: 233 Distribution: Panama

Barybas bituberculata Moser Barybas bituberculatus Moser, 1921b: 173 192 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Barybas bituberculata Moser; Blackwelder, 1944; 233 Distribution: Venezuela

Barybas boliviana Moser Barybas bolivianus Moser, 1919a: 19 Barybas boliviana Moser; Blackwelder, 1944: 233 Distribution: Bolivia

Barybas callosipyga Frey Barybas callosipygus Frey, 1969b: 387 Distribution: Brazil

Barybas callosipennis Frey Barybas callosipennis Frey, 1972b: 96 Distribution: Brazil

Barybas chacoensis Moser Barybas chacoensis Moser, 1918b: 142 Distribution: Bolivia

Barybas clypeata Moser Barybas clypeatus Moser, 1918b: 141 Barybas clypeata Moser; Blackwelder, 1944: 233 Distribution: Bolivia

Barybas compacta (Erichson) Philochloenia compacta Erichson, 1847a: 103 Microcrania compacta (Erichson); Burmeister, 1855: 76 Barybas compactus (Erichson); Dalla Torre, 1913: 334 (error) Barybas compacta (Erichson); Blackwelder, 1944: 233 Distribution: Peru

Barybas crassa Moser Barybas crassus Moser, 1921b: 172 Barybas crassa Moser; Blackwelder, 1944: 233 Distribution: Bolivia

Barybas curta Burmeister Barybas curta Burmeister, 1855: 76 Distribution: Colombia

Barybas ferruginea Moser Barybas ferrugineus Moser, 1921b: 173 Barybas ferruginea Moser; Blackwelder, 1944: 233 Distribution: Brazil

Barybas fraterna Moser Barybas fraternus Moser, 1921b: 172 193 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Barybas fraterna Moser; Blackwelder, 1944: 233 Distribution: Brazil

Barybas idiota Bates Barybas idiota Bates, 1887: 151 Distribution: Brazil, Nicaragua

Barybas insulana Moser Barybas insulanus Moser, 1919a: 17 Barybas insulana Moser; Blackwelder, 1944: 233 Distribution: Trinidad

Barybas kulzeri Frey Barybas kulzeri Frey, 1970b: 284 Distribution: Venezuela

Barybas latesquamosa Frey Barybas latesquamosus Frey, 1964c: 701 Distribution: Venezuela

Barybas maculata Moser Barybas maculatus Moser, 1919a: 20 Barybas maculata Moser; Blackwelder, 1944: 233 Distribution: Bolivia

Barybas murina Burmeister Microcrania murina Burmeister, 1855: 77 Barybas murina (Burmeister); Dalla Torre, 1913: 334 Distribution: Colombia

Barybas nana Blanchard Barybas nanus Blanchard, 1850: 94 Barybas nana Blanchard; Blackwelder, 1944: 233 Distribution: Brazil

Barybas oxygenys Bates Barybas oxygenys Bates, 1887: 151 Distribution: Panama

Barybas parvula Moser Barybas parvulus Moser, 1919a: 20 Barybas parvula Moser; Blackwelder, 1944: 233 Distribution: Bolivia

Barybas peruana Moser Barybas peruana Moser, 1918b: 141 Barybas peruana Moser; Blackwelder, 1944: 233 Distribution: Peru 194 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Barybas scopifer (Burmeister) Microcrania scopifer Burmeister, 1855: 77 Barybas scopifer (Burmeister); Dalla Torre, 1913: 334 Distribution: Brazil

Synonym: Barybas excisiceps Frey, 1973a: 259

Barybas setifer Moser Barybas setifer Moser, 1921b: 172 Distribution: Venezuela

Barybas sparsesetosa Frey Barybas sparsesetosus Frey, 1973a: 261 Distribution: Brazil

Barybas squamiger Frey Barybas squamiger Frey, 1967b: 423 Distribution: Peru

Barybas sublobata Bates Barybas sublobatus Bates, 1887: 151 Barybas sublobata Bates; Blackwelder, 1944; 233 Distribution: Brazil

Barybas uniformis Moser Barybas uniformis Moser, 1921b: 171 Distribution: Boliva

Barybas variegata Blanchard Barybas variegatus Blanchard, 1850: 94 Barybas variegata Blanchard; Blackwelder, 1944: 233 Distribution: Brazil

Barybas variipennis Moser Barybas variipennis Moser, 1921b: 173 Distribution: Venezuela

Genus BYRASBA Harold Byrasba Harold, 1869b: 122. Type species Barybas volvulus Burmeister, by monotypy.

Synonym: Barybas Burmeister, 1855: 20 (not Blanchard, 1850: 94). Type species Barybas volvulus Burmeister, by monotypy.

Byrasba volvula (Burmeister) Barybas volvulus Burmeister, 1855: 20 Byrasba volvula (Burmeister); Harold, 1869a: 1142 Distribution: Brazil 195 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Genus CALODACTYLUS Blanchard Calodactylus Blanchard, 1850: 91. Type species Calodactylus tibialis Blanchard, by monotypy.

Synonym: Dioplia Burmeister, 1855: 70. Type species Dioplia sulphurea Burmeister, subsequent designation (Evans 2003: 234).

Calodactylus abendrothii Kirsch Calodactylus abendrothii Kirsch, 1874: 341 Calodactylus abendrothi Kirsch; Blackwelder, 1944: 227 (incorrect subsequent spelling) Distribution: Peru

Calodactylus aequatorialis Moser Calodactylus aequatorialis Moser, 1918b: 97 Distribution: Ecuador

Calodactylus amabilis Frey Calodactylus amabilis Frey, 1969b: 376 Distribution: Brazil

Calodactylus boliviensis Moser Calodactylus boliviensis Moser, 1918b: 96 Distribution: Bolivia

Calodactylus heterosquamosus Frey Calodactylus heterosquamosus Frey, 1973a: 255 Distribution: Peru

Calodactylus niveosignatus Moser Calodactylus niveosignatus Moser, 1921b: 138 Distribution: Brazil

Calodactylus schmalzi Ohaus Calodactylus schmalzi Ohaus, 1913a: 48 Distribution: Brazil

Calodactylus sulphureus (Burmeister) Dioplia sulphureus Burmeister, 1855: 71 Calodactylus sulphureus Burmeister; Distribution: Brazil

Calodactylus tibialis Blanchard Calodactylus tibialis Blanchard, 1850: 91 Distribution: Brazil

Synonym: Dioplia heteroclita Burmeister, 1855: 71 Calodactylus heteroclitus (Burmeister); Harold, 1869a: 1140

196 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Calodactylus vittipennis Moser Calodactylus vittipennnis Moser, 1921b: 138 Distribution: Bolivia

Genus CANESTERA Saylor Canestera Saylor, 1938a: 72. Type species Canestera marshalli Saylor, by monotypy.

Canestera marshalli Saylor Canestera marshalli Saylor, 1938a: 73 Distribution: Guyana

Genus CERASPIS Le Peletier and Audinet-Serville Ceraspis Le Peletier and Audinet-Serville, 1828: 370. Type species Melolontha bivulnerata Germar, a senor synonym of Ceraspis pruinosa Le Peletier and Audinet-Serville, by subsequent designation (Lacordaire, 1856: 252)

Synonym: Faula Blanchard, 1850: 124. Type species Faula cornuta Blanchard, subsequent designation (Evans 2003: 236).

Subgenus: Isoceraspis Ohaus, 1911b: 669 (as subgenus). Type species Ceraspis duckei Kolbe, by monotypy.

Keys: Frey, 1962: 3; Delgado and Morón, 1987: 51 (México) ; Delgado, 2001a: 169 (México).

Ceraspis albipennis Frey Ceraspis albipennis Frey, 1973a: 276 Distribution: Brazil

Ceraspis albovaria Blanchard Ceraspis albovaria Blanchard, 1850: 93 Distribution: Brazil

Ceraspis alvarengai Frey Ceraspis alvarengai Frey, 1973a: 278 Distribution: Brazil

Ceraspis amazonica Frey Ceraspis amazonica Frey, 1962: 34 Distribution: Brazil

Ceraspis amoena Frey Ceraspis amoena Frey, 1962: 46 Distribution: Brazil

Ceraspis bicolor Moser Ceraspis bicolor Moser, 1919a: 26 Distribution: Brazil

197 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ceraspis bivittata Burmeister Ceraspis bivittata Burmeister, 1855: 100 Distribution: Colombia

Ceraspis bivulnerata (Germar) Melolontha bivulnerata Germar, 1824: 126 Ceraspis bivulnerata (Germar); Burmeister, 1855: 92 Distribution: Brazil, Guyana

Synonym: Ceraspis albida Guérin-Méneville, 1831a: pl. 24 bis

Synonym: Ceraspis pruinosa Le Peletier and Audinet-Serville, 1828: 371

Ceraspis brittoni Frey Ceraspis brittoni Frey, 1962: 57 Distribution: Brazil

Ceraspis brunneipennis (Bates) Faula brunneipennis Bates, 1887: 137 Ceraspis brunneipennis (Bates); Dalla Torre, 1913: 336 Distribution: Costa Rica

Ceraspis bufo Frey Ceraspis bufo Frey, 1962:44 Distribution: Brazil

Ceraspis burmeisteri Frey Ceraspis burmeisteri Frey, 1962: 38 Distribution: Brazil

Ceraspis castaneipennis Blanchard Ceraspis castaneipennis Blanchard, 1850: 92 Distribution: Brazil

Ceraspis centralis (Sharp) Faula centralis Sharp, 1877: 131 Ceraspis centralis (Sharp); Dalla Torre, 1913: 337 Distribution: Costa Rica, Nicaragua, Panama

Ceraspis cinerea Moser Ceraspis cinerea Moser, 1921b: 177 Distribution: Brazil

Synonym: Ceraspis lepidiota Moser, 1921b: 177

Ceraspis citrina Blanchard Ceraspis citrina Blanchard, 1850: 92 Distribution: Brazil 198 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Ceraspis pisicolor Burmeister, 1855: 96 Ceraspis citrina var. pisicolor Burmeister; Frey, 1962: 44

Ceraspis clypealis Frey Ceraspis clypealis Frey, 1962: 15 Distribution: Brazil

Ceraspis colon Burmeister Ceraspis colon Burmeister, 1855: 93 Distribution: Brazil

Ceraspis conspersa Burmeister Ceraspis conspersa Burmeister, 1855: 96 Distribution: Brazil

Ceraspis convexicollis Frey Ceraspis convexicollis Frey, 1969b: 400 Distribution: Brazil

Ceraspis cornuta (Blanchard) Faula cornuta Blanchard, 1850: 124 Ceraspis cornuta (Blanchard); Dalla Torre, 1913: 337 Distribution: Brazil

Synonym: Ceraspis rufipes Burmeister, 1855: 100

Ceraspis costulata Frey Ceraspis costulata Frey, 1965b: 150 Distribution: Colombia

Ceraspis decora Gory Ceraspis decora Gory, 1831: pl. 24 bis Distribution: Brazil

Synonym: Ceraspis variabilis Burmeister, 1855: 94

Ceraspis diversa Frey Ceraspis diversa Frey, 1962: 55 Distribution: Brazil

Ceraspis dorsata Burmeister Ceraspis dorsata Burmeister, 1855: 98 Distribution: Brazil

Ceraspis dorsopicta Frey Ceraspis dorsopicta Frey, 1972b: 97 Distribution: Brazil 199 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ceraspis duckei Ohaus Ceraspis (Isoceraspis) duckei Ohaus 1911: 669 Distribution: Brazil

Ceraspis elegans Nonfried Ceraspis elegans Nonfried, 1892a: 223 Distribution: Brazil, Honduras

Ceraspis elongata Frey Ceraspis elongata Frey, 1962: 35 Distribution: Brazil

Ceraspis farinosa Burmeister Ceraspis farinosa Burmeister, 1855: 97 Distribution: Brazil

Ceraspis femorata Frey Ceraspis femorata Frey, 1969b: 401 Distribution: Brazil

Ceraspis flava Blanchard Ceraspis flava Blanchard, 1850: 93 Distribution: Brazil

Ceraspis fulva Blanchard Ceraspis fulva Blanchard, 1850: 93 Distribution: Brazil

Ceraspis gibbicollis Blanchard Ceraspis gibbicollis Blanchard, 1850: 92 Distribution: Brazil

Ceraspis globicollis Frey Ceraspis globicollis Frey, 1962: 30 Distribution: Brazil

Ceraspis griseosquamosa Moser Ceraspis griseo-squamosa Moser, 1921: 178 Distribution: Brazil

Ceraspis guttata Blanchard Ceraspis guttata Blanchard, 1850: 93 Distribution: Brazil

Synonym: Ceraspis guttata var. crucifera Frey, 1962: 54

Synonym: Ceraspis vitticollis Moser, 1919a: 24 200 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ceraspis hispida Bates Faula hispida Bates, 1887: 137 Ceraspis hispida (Bates); Dalla Torre, 1913: 337 Distribution: Costa Rica, Panama

Ceraspis imitatrix Nonfried Ceraspis imitatrix Nonfried, 1892a: 224 Distribution: Honduras

Ceraspis immaculata Burmeister Ceraspis immaculata Burmeister, 1855: 101 Distribution: Colombia

Ceraspis innotata (Blanchard) Faula innotata Blanchard, 1850: 125 Ceraspis innotata (Blanchard); Dalla Torre, 1913: 337 Distribution: Peru

Ceraspis insularis (Arrow) Faula insularis Arrow, 1903d: 520 Ceraspis insularis (Arrow): Dalla Torre, 1913: 337 Distribution: St. Vincent

Ceraspis jaliscoensis Delagado and Navarette-Heredia Ceraspis jaliscoensis Delagado and Navarette-Heredia, 2004: 2 Distribution: Mexico

Ceraspis klenei Brenske Ceraspis klenei Brenske, 1890a: 34 Ceraspis kleinei [sic] Brenske; Blackwelder, 1944: 234 Distribution: Ecuador

Ceraspis kuntzeni Moser Ceraspis kuntzeni Moser, 1921b: 178 Distribution: Brazil

Ceraspis lepida Frey Ceraspis lepida Frey, 1973a: 274 Distribution: Brazil

Ceraspis leucosoma Blanchard Ceraspis leucosoma Blanchard, 1850: 93 Distribution: Brazil

Ceraspis lineata (Waterhouse) Faula lineata Waterhouse, 1879: 421 Ceraspis lineata (Waterhouse); Dalla Torre, 1913: 337 201 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Colombia

Ceraspis linharensis Frey Ceraspis linharensis Frey, 1973a: 273 Distribution: Brazil

Ceraspis lurida Frey Ceraspis lurida Frey, 1962: 16 Distribution: Brazil

Ceraspis macrophylla Moser Ceraspis macrophylla Moser, 1919a: 29 Distribution: Colombia

Ceraspis martinezi Frey Ceraspis martinezi Frey, 1962: 30 Distribution: Brazil

Ceraspis melanoleuca Le Peletier and Audinet-Serville Ceraspis melanoleuca Le Peletier and Audinet-Serville, 1828: 371 Distribution: Brazil

Ceraspis mexicana (Harold) Faula mexicana Harold, 1863: 175 Ceraspis mexicana (Harold); Harold, 1869a: 1155 Distribution: Guatemala, México

Ceraspis mixta Blanchard Ceraspis mixta Blanchard, 1850: 92 Distribution: Brazil

Ceraspis modesta Burmeister Ceraspis modesta Burmeister, 1855: 99 Distribution: Colombia

Ceraspis moseri Frey Ceraspis moseri Frey, 1962: 19 Distribution: Colombia

Ceraspis mustela Frey Ceraspis mustela Frey, 1962: 31 Distribution: Brazil

Ceraspis mutica Moser Ceraspis mutica Moser, 1921b: 178 Distribution: Brazil

Ceraspis nitida Frey 202 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ceraspis nitida Frey, 1962: 28 Distribution: Bolivia, Brasil, Peru

Ceraspis nivea Le Peletier and Audinet-Serville Ceraspis nivea Le Peletier and Audinet-Serville, 1828: 371 Distribution: Brazil

Synonym: Ceraspis brenskei Moser, 1919a: 23

Ceraspis niveipennis Gistel Ceraspis niveipennis Gistel, 1857: 91 Distribution: Brazil

Ceraspis oaxacaensis Delgado Ceraspis oaxacaensis Delgado, 2001a: 167 Distribution: México

Ceraspis oblonga Moser Ceraspis oblonga Moser, 1919a: 25 Distribution: Brazil

Ceraspis obscura Blanchard Ceraspis obscura Blanchard, 1850: 94 Distribution: French Guiana, Surinam

Ceraspis ocellata Frey Ceraspis ocellata Frey, 1962:17 Distribution: Panama

Ceraspis ohausi Moser Ceraspis ohausi Moser, 1921b: 179 Distribution: Ecuador

Ceraspis opacipennis (Moser) Faula opacipennis Moser, 1919a: 22 Ceraspis opacipennis (Moser); Blackwelder, 1944; 234 Distribution: Brazil

Ceraspis ornata Frey Ceraspis ornata Frey, 1962: 58 Distribution: Brazil

Ceraspis pallida Blanchard Ceraspis pallida Blanchard, 1850: 93 Distribution: Brazil

Ceraspis pauperata Burmeister Ceraspis pauperata Burmeister, 1855: 98 203 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Ceraspis penai Frey Ceraspis penai Frey, 1964c: 691 Distribution: Peru

Ceraspis pereirae Frey Ceraspis pereirae Frey, 1962: 40 Distribution: Argentina

Ceraspis pilatei (Harold) Faula pilatei Harold, 1863: 174 Ceraspis pilatei (Harold); Harold, 1869a: 1155 Distribution: Guatemala, Honduras, México

Synonym: Ancistrosoma baui Nonfried, 1890: 76

Synonym: Faula velutina Bates, 1887: 137 Ceraspis velutina (Bates); Dalla Torre, 1913: 338

Ceraspis plaumanni Frey Ceraspis plaumanni Frey, 1962: 49 Distribution: Brazil

Ceraspis pulchra Frey Ceraspis pulchra Frey, 1962: 46 Distribution: Brazil

Ceraspis quadrifoliata Moser Ceraspis quadrifoliata Moser, 1919a: 28 Distribution: Colombia

Ceraspis quadrimaculata (Blanchard) Faula quadrimaculata Blanchard 1850: 124 Ceraspis quadrimaculata (Blanchard); Dalla Torre, 1913: 337 Distribution: Colombia, Venezuela

Synonym: Ceraspis quadrinotata Burmeister, 1855: 101

Ceraspis quadripustulata (Blanchard) Faula quadripustulata Blanchard, 1850: 124 Ceraspis quadripustulata (Blanchard); Burmeister, 1855: 100 Distribution: Colombia

Ceraspis rotundicollis Frey Ceraspis rotundicollis Frey, 1973a: 277 Distribution: Brazil

204 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ceraspis rubiginosa (Latreille) Melolontha rubiginosa Latreille, 1813: 364 Ceraspis rubiginosa (Latreille); Dejean, 1833: 163 Distribution: Peru

Synonym: Ceraspis peruviana Guérin-Méneville, 1831b: pl. 3

Ceraspis ruehli Brenske Ceraspis rühli Brenske, 1890a: 34 Distribution: Ecuador

Ceraspis ruficollis Frey Ceraspis ruficollis Frey, 1962: 23 Distribution: Colombia

Ceraspis rufoscutellata Moser Ceraspis rufoscutellata Moser, 1919a: 27 Distribution: Peru

Ceraspis setiventris Moser Ceraspis setiventris Moser, 1921b: 176 Distribution: Brazil

Ceraspis signata Blanchard Ceraspis signata Blanchard, 1850: 93 Distribution: Brazil

Synonym: Ceraspis internata Burmeister, 1855: 96

Ceraspis sparsesetosa Frey Ceraspis sparsesetosa Frey, 1972b: 97 Distribution: Bolivia

Ceraspis squamulata Moser Ceraspis squamulata Moser, 1924a: 131 Distribution: Paraguay

Ceraspis squamulifera (Moser) Faula squamulifera Moser, 1919a: 21 Ceraspis squamulifera (Moser); Blackwelder, 1944: 234 Distribution: Colombia, Ecuador, Peru

Ceraspis striata Frey Ceraspis striata Frey, 1973a: 276 Distribution: Brazil

Ceraspis subvittata Moser Ceraspis subvittata Moser, 1921b: 176 205 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Ceraspis sulcicollis Moser Ceraspis sulcicollis Moser, 1921b: 177 Distribution: Brazil

Ceraspis tenuisquamosa Frey Ceraspis tenuisquamosa Frey, 1962: 39 Distribution: Brazil

Ceraspis tibialis Blanchard Ceraspis tibialis Blanchard, 1850: 92 Distribution: Brazil

Ceraspis unguicularis Moser Ceraspis unguicularis Moser, 1919a: 30 Distribution: Brazil

Ceraspis variegata (Perty) Hoplia variegata Perty, 1833: 52 Ceraspis variegata (Perty); Harold, 1869a: 1155 Distribution: Brazil

Ceraspis ventralis Frey Ceraspis ventralis Frey, 1962: 40 Distribution: Brazil

Ceraspis vestita Blanchard Ceraspis vestita Blanchard, 1850: 94 Distribution: Brazil, Surinam

Ceraspis vittata Moser Ceraspis vitticollis vittata Moser, 1919a: 25 Ceraspis vittata Moser; Blackwelder, 1944: 234 Distribution: Brazil

Ceraspis vulpes Frey Ceraspis vulpes Frey, 1962: 39 Distribution: Brazil

Ceraspis zikani Moser Ceraspis zikani Moser, 1924: 131 Distribution: Brazil

Genus CERATOLONTHA Arrow Ceratolontha Arrow, 1948a: 372. Type species Ceratolontha venezuelae Arrow, by monotypy.

Ceratolontha venezuelae Arrow 206 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ceratolontha venezuelae Arrow, 1948a: 372 Distribution: Venezuela

Genus CHARIODACTYLUS Moser Chariodactylus Moser, 1919b: 45. Type species Chariodactylus chacoensis Moser, subsequent designation (Evans 2003: 247).

Chariodactylus chacoenis Moser Chariodactylus chacoenis Moser, 1919b: 45 Distribution: Bolivia

Chariodactylus sublaevicollis Moser Chariodactylus sublaevicollis Moser, 1919b: 46 Distribution: Peru

Genus CHARIODEMA Blanchard Chariodema Blanchard, 1850: 117. Type species Philochloenia virescens Blanchard, subsequent designation (Evans 2003: 247).

Chariodema amoena Kirsch Chariodema amoena Kirsch, 1865: 49 Distribution: Colombia

Chariodema bogotensis Kirsch Chariodema bogotensis Kirsch, 1865: 49 Distribution: Colombia

Chariodema chlorodera Blanchard Chariodema chlorodera Blanchard, 1850: 117 Distribution: Colombia

Chariodema costatipennis Blanchard Chariodema costatipennis Blanchard, 1850: 117 Distribution: Brazil

Chariodema pallens Blanchard Chariodema pallens Blanchard, 1850: 118 Distribution: Brazil

Chariodema subcostata Blanchard Chariodema subcostata Blanchard, 1850: 117 Distribution: Brazil

Chariodema suturalis Blanchard Chariodema suturalis Blanchard, 1850: 117 Distribution: Brazil

Chariodema tolimana Moser 207 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Chariodema tolimana Moser, 1918b: 140 Distribution: Colombia

Chariodema villosa Blanchard Chariodema villosa Blanchard, 1850: 118 Distribution: Bolivia

Chariodema virescens (Blanchard) Phylochlaenia virescens Blanchard, 1842: 11 Chariodema virescens (Blanchard); Blanchard, 1850: 117 Distribution: Bolivia

Chariodema vittata Blanchard Chariodema vittata Blanchard, 1850: 117 Distribution: Bolivia

Chariodema xylina Blanchard Chariodema xylina Blanchard, 1850: 117 Distribution: Colombia

Genus CHREMASTODUS Solier Chremastodus Solier, 1851: 103. Type species Chremastodus pubescens Solier, subsequent designation (Evans 2003: 248). note: misspelled as Cremastodus Solier by Dalla Torre (1913: 328)

Chremastodus marmoratus (Curtis) Macrodactylus marmoratus Curtis, 1845: 455 Chremastodus marmoratus (Curtis); Solier, 1851: 104 Distribution: Chile

Chremastodus pubescens Solier Chremastodus pubescens Solier, 1851: 103 Distribution: Chile

Genus CLAVIPALPUS Laporte Clavipalpus Laporte, 1832: 406. Type species Clavipalpus dejeani Laporte, by monotypy.

Synonym: Amphicrania Burmeister, 1855: 26. Type species Amphicrania ursina Burmeister, subsequent designation (Evans 2003: 249).

Clavipalpus aequatorialis Moser Clavipalpus aequatorialis Moser, 1918b: 120 Distribution: Ecuador

Clavipalpus antisanae Bates Clavipalpus antisanae Bates, 1891: 27 Distribution: Ecuador 208 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Clavipalpus basalis Moser Clavipalpus basalis Moser, 1918b: 121 Distribution: Ecuador

Clavipalpus blanchardi Kirsch Clavipalpus blanchardi Kirsch, 1885: 215 Distribution: Colombia

Clavipalpus castaneus Moser Clavipalpus castaneus Moser, 1924a: 123 Distribution: Ecuador

Clavipalpus cochleatus (Burmeister) Amphicrania cochleata Burmeister, 1855: 27 Clavipalpus cochleatus (Burmeister); Harold, 1869a: 1143 Distribution: Brazil

Clavipalpus dejeanii Laporte Clavipalpus dejeanii Laporte, 1832: 406 Distribution: Brazil

Clavipalpus hirsutus Kirsch Clavipalpus hirsutus Kirsch, 1885: 216 Distribution: Ecuador

Clavipalpus hirticollis Frey Clavipalpus hirticollis Frey, 1969b: 377 Distribution: Brazil

Clavipalpus orbignyanus Blanchard Clavipalpus orbignyanus Blanchard, 1850: 120 Distribution: Bolivia

Clavipalpus peruanus Moser Clavipalpus peruanus Moser, 1918b: 119 Distribution: Peru

Clavipalpus rimbachi Moser Clavipalpus rimbachi Moser, 1924: 123 Distribution: Ecuador

Clavipalpus sinuatus Kirsch Clavipalpus sinuatus Kirsch, 1885: 215 Distribution: Ecuador

Clavipalpus spadiceus (Burmeister) Amphicrania spadicea Burmeister, 1855: 27 209 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Clavipalpus spadiceus Burmeister; Harold, 1869a: 1143 Distribution: Peru

Clavipalpus tenuis Arrow Clavipalpus tenuis Arrow, 1903a: 251 Distribution: Brazil

Clavipalpus ursinus Blanchard Clavipalpus ursinus Blanchard, 1850: 120 Distribution: Colombia

Synonym: Amphicrania ursina Burmeister, 1855: 27

Clavipalpus variolosus (Burmeister) Amphicrania variolosa Burmeister, 1855: 28 Clavipalpus variolosus (Burmeister); Harold, 1869a: 1143 Distribution: Guiana

Clavipalpus vestitus Moser Clavipalpus vestitus Moser, 1924a: 123 Distribution: Ecuador

Clavipalpus whymperi chimborazonus Bates Clavipalpus whymperi var. chimborazonus Bates, 1891: 28 Distribution: Ecuador

Clavipalpus whymperi whymperi Bates Clavipalpus whymperi Bates, 1891: 28 Distribution: Ecuador

Genus COENONYCHA Horn, 1876 Coenonycha Horn, 1876: 192. Type species Dichelonyx rotundata LeConte, by original designation.

Keys: Cazier and McClay, 1943: 10; Hatch, 1971: 470; Evans and Smith, 1986; 85; Evans and D’Hotman, 1988: 206

Coenonycha acuta Cazier Coenonycha acuta Cazier, 1943: 14 Distribution: United States

Coenonycha ampla Cazier Coenonycha ampla Cazier, 1943: 14 Distribution: United States

Coenonycha barri Cazier Coenonycha barri Cazier, 1943: 19 Distribution: United States

210 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Coenonycha bowelsi Cazier Coenonycha bowelsi Cazier, 1943: 20 Distribution: United States

Coenonycha clementina Casey Coenonycha clementina Casey, 1909: 281 Distribution: United States

Coenonycha clypeata McClay Coenonycha clypeata McClay, 1943: 22 Distribution: United States

Coenonycha crispata McClay Coenonycha crispata McClay, 1943: 26 Distribution: United States

Coenonycha dimorpha Evans Coenonycha dimorpha Evans, 1986: 83 Distribution: United States

Coenonycha fuga Cazier Coenonycha fuga Cazier, 1943: 12 Distribution: United States

Coenonycha fulva McClay Coenonycha fulva McClay, 1943: 23 Distribution: United States

Coenonycha fusca McClay Coenonycha fusca McClay, 1943: 24 Distribution: United States

Coenonycha globosa McClay Coenonycha globosa McClay, 1943: 25 Distribution: United States

Coenonycha hageni Cazier Coenonycha hageni Cazier, 1943: 15 Distribution: United States

Coenonycha lurida Cazier Coenonycha lurida Cazier, 1943: 19 Distribution: United States

Coenonycha mediata Cazier Coenonycha mediata Cazier, 1943: 21 Distribution: United States

211 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Coenonycha ochreata Evans Coenonycha ochreata Evans, 1986: 82 Distribution: United States

Coenonycha ovatis McClay Coenonycha ovatis McClay, 1943: 27 Distribution: United States

Coenonycha ovipennis Horn Coenonycha ovipennis Horn, 1876: 192 Distribution: United States

Coenonycha pallida Cazier Coenonycha pallida Cazier, 1943: 18 Distribution: United States

Coenonycha parvula Fall Coenonycha parvula Fall, 1901: 293 Distribution: United States

Coenonycha pascuensis Potts Coenonycha pascuensis Potts, 1945: 141 Distribution: United States

Coenonycha purshiae Cazier Coenonycha purshiae Cazier, 1943: 13 Distribution: United States

Coenonycha pygmaea Smith Coenonycha pygmaea Smith, 1986: 85 Distribution: United States

Coenonycha rotundata (LeConte) Dichelonycha [sic] rotundata LeConte 1856: 281 Coenonycha rotundata (LeConte); Horn, 1876: 192: 293 Distribution: México, United States

Synonym: Coenonycha rufobrunnea Howden, 1969: 55

Coenonycha rubida McClay Coenonycha rubida McClay, 1943: 21 Distribution: United States

Coenonycha santacruzae Evans Coenonycha santacruzae Evans, 1986: 82 Distribution: United States

Coenonycha saylori Cazier 212 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Coenonycha saylori Cazier, 1943: 20 Distribution: United States

Coenonycha scotti McClay Coenonycha scotti McClay, 1943: 22 Distribution: United States

Coenonycha sleeperi Evans Coenonycha sleeperi Evans, 1988: 208 Distribution: México

Coenonycha socialis Horn Coenonycha socialis Horn, 1876: 192 Distribution: México

Coenonycha stohleri Saylor Coenonycha stohleri Saylor, 1935d: 102 Distribution: United States

Coenonycha testacea Cazier Coenonycha testacea Cazier, 1937c: 127 Distribution: United States

Coenonycha tingi Cazier Coenonycha tingi Cazier, 1937c: 126 Distribution: United States

Coenonycha utahensis McClay Coenonycha utahensis McClay, 1943: 24 Distribution: United States

Genus CORMINUS Burmeister Corminus Burmeister, 1855: 39. Type species Corminus canescens Burmeister, subsequent designation (Evans 2003: 254).

Corminus bahianus Moser Corminus bahianus Moser, 1919b: 39 Distribution: Brazil

Corminus brasiliensis Moser Corminus brasiliensis Moser, 1919b: 36 Distribution: Brazil

Corminus canescens Burmeister Corminus canescens Burmeister, 1855: 40 Distribution: Brazil

Corminus guayaquilanus Moser 213 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Corminus guayaquilanus Moser, 1921b: 156 Distribution: Ecuador

Corminus luridipennis Burmeister Corminus luridipennis Burmeister, 1855: 40 Distribution: Brazil

Corminus maniculatus Burmeister Corminus maniculatus Burmeister, 1855: 40 Distribution: Brazil

Corminus oblongus Moser Corminus oblongus Moser, 1919b: 38 Distribution: Brazil

Corminus parvulus Moser Corminus parvulus Moser, 1919b: 40 Distribution: Brazil

Corminus rufofucsus Moser Corminus rufofucsus Moser, 1924b: 162 Distribution: Brazil

Corminus setulosus Moser Corminus setulosus Moser, 1919b: 37 Distribution: Brazil

Corminus testaceipennis Moser Corminus testaceipennis Moser, 1921b: 155 Distribution: Brazil

Corminus varians Moser Corminus varians Moser, 1921b: 155 Distribution: Brazil

Genus CTENOTIS Burmeister Ctenotis Burmeister, 1855: 20. Type species Ctenotis obesa Burmeister, by monotypy.

Ctenotis obesa Burmeister Ctenotis obesa Burmeister, 1855: 21 Distribution: Brazil

Synonym: Phyllophaga (Phytalus) alvarengai Frey, 1964c: 697

Genus CTILOCEPHALA Burmeister Ctilocephala Burmeister, 1855: 78. Type species Ctilocephala pellucens Burmeister, by monotypy.

Ctilocephala abdominalis Moser 214 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ctilocephala abdominalis Moser, 1918b: 148 Distribution: Colombia

Ctilocephala amazonica Moser Ctilocephala amazonica Moser, 1918b: 145 Distribution: Brazil

Ctilocephala asperula (Perty) Serica asperula Perty, 1833: 48 Hercitis asperula (Perty); Burmeister, 1855: 80 Barybas asperula (Perty); Lacordaire, 1856: 260 Microcrania asperula (Perty); Harold, 1869a: 1151 Barybas asperulus (Perty); Dalla Torre, 1913: 334 Ctilocephala asperula (Perty); Moser, 1918b: 144 Distribution: Brazil

Ctilocephala brasiliensis Moser Ctilocephala brasiliensis Moser, 1918b: 145 Distribution: Brazil

Ctilocephala breviceps Moser Ctilocephala breviceps Moser, 1918b: 147 Distribution: Brazil

Ctilocephala cayennensis Moser Ctilocephala cayennensis Moser, 1921a: 56 Distribution: French Guiana

Ctilocephala grisea Moser Ctilocephala grisea Moser, 1918b: 144 Distribution: Colombia

Ctilocephala kuntzeni Moser Ctilocephala kuntzeni Moser, 1921b:176 Distribution: Bolivia

Ctilocephala minuta Moser Ctilocephala minuta Moser, 1918b: 146 Distribution: Brazil

Ctilocephala ohausi Moser Ctilocephala ohausi Moser, 1924a: 127 Distribution: Ecuador

Ctilocephala pellucens Burmeister Ctilocephala pellucens Burmeister, 1855: 78 Distribution: Colombia

215 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Ctilocephala sarana Moser Ctilocephala sarana Moser, 1921b: 175 Distribution: Bolivia

Genus DASYUS Le Peletier and Audinet-Serville Dasyus Le Peletier and Audinet-Serville, 1828: 369. Type species Dasyus collaris Le Peletier and Audinet-Serville, by monotypy.

Dasyus collaris collaris Le Peletier and Audinet-Serville Dasyus collaris Le Peletier and Audinet-Serville, 1828: 370 Distribution: Brazil

Synonym: Dasyus collaris var. luridipennis Moser, 1918b: 95

Dasyus collaris nigellus Burmeister Dasyus nigellus Burmeister, 1855: 72 Dasyus collaris var. nigellus Burmeister; Moser. 1918b: 95 Distribution: Brazil

Dasyus variabilis Moser Dasyus variabilis Moser, 1918b: 95 Distribution: Brazil

Genus DEMODEMA Blanchard Demodema Blanchard, 1850: 121. Type species Demodema fallax Blanchard, by subsequent designation (Lacordaire, 1856: 258).

Demodema aulai Bruch Demodema aulai Bruch, 1909: 349 Distribution: Argentina

Demodema bonariensis Bruch Demodema bonariensis Bruch, 1909: 350 Distribution: Argentina

Demodema brevitarsis Blanchard Philochlaenia [sic] brevitarsis Blanchard, 1850: 122 Demodema brevitarsis (Blanchard); Lacordaire, 1856: 258 Distribution: Brazil

Demodema cinerascens Blanchard Demodema cinerascens Blanchard, 1850: 121 Distribution: Bolivia

Demodema comata Blanchard Philochlaenia [sic] comata Blanchard, 1850: 122 Demodema comata (Blanchard); Lacordaire, 1856: 258 Distribution: Argentina, Paraguay 216 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Demodema distincta Bruch Demodema distincta Bruch, 1909: 351 Distribution: Argentina

Demodema fallax Blanchard Demodema fallax Blanchard, 1850: 121 Distribution: Brazil

Demodema paraguayensis Moser Demodema paraguayensis Moser, 1921b: 156 Distribution: Paraguay

Genus DICHELONYX Harris Dichelonyx Harris, 1827: 7. Type species Melolontha linearis Gyllenhal, a junior synonym of M. elongatula Schoenherr, by monotypy.

Synonym: Philochloenia Dejean, 1833: 163. Type species Melolontha elongata Fabricius, 1792 by subsequent designation (Smith & Evans 2005: 40)

Synonym: Anaeretes Dejean, 1836: 181. Type species Melolontha elongata Fabricius (not Olivier, 1789). note: Dichelonycha Kirby, 1837 is a misspelling; see Pate, 1946: 68.

Keys to species: Horn, 1876: 186 (revision); Fall, 1901: 280 (revision); Blatchley, 1910: 951 (Indiana); Saylor, 1945b: 141 (revision); Brown, 1946: 107 (eastern United States); Hatch, 1971: 468 (Pacific Northwest); Lago et al., 1979: 68 (North Dakota); Harpootlian, 2001: 94 (South Carolina) (see also Cornell, 1972: 28).

Dichelonyx albicollis Burmeister Dichelonycha [sic] albicollis Burmeister, 1855: 74 Dichelonyx albicollis Burmeister; Leng, 1920: 258 Distribution: Canada, United States

Dichelonyx backii arizonica Barrett Dichelonyx arizonica Barrett, 1933: 132 Dichelonyx backii arizonica Barrett; Saylor, 1945b: 149 Distribution: United States

Dichelonyx backii backii Kirby Dichelonycha [sic] backii Kirby, 1837: 134 Dichelonyx backi Kirby; Leng, 1920: 258 Distribution: Canada, United States

Dichelonyx canadensis Horn Dichelonycha [sic] canadensis Horn, 1876: 188 Dichelonyx canadensis Horn; Leng, 1920: 257 217 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Canada, United States

Dichelonyx clypeata Horn Dichelonycha [sic] clypeata Horn, 1876: 190 Dichelonyx clypeata Horn; Leng, 1920: 258 Distribution: United States

Dichelonyx crumbi Hatch Dichelonyx crumbi Hatch, 1971: 469 Distribution: United States

Dichelonyx decolorata Fall Dichelonycha [sic] decolorata Fall, 1901: 290 Dichelonyx decolorata Fall; Leng, 1920: 258 Distribution: United States

Dichelonyx diluta Fall Dichelonycha [sic] diluta Fall, 1901: 283 Dichelonyx diluta Fall; Leng, 1920: 258 Distribution: Canada, United States

Dichelonyx elongatula (Schoenherr) Melolontha elongatula Schoenherr, 1817: 210 Anaeretes elongatula (Schoenherr); Dejean, 1833: 181 Dichelonyx elongatula (Schoenherr); Brown, 1946: 106 Distribution: Canada, United States

Synonym: Melolontha elongata Fabricius, 1792: 170 (not Olivier, 1789) Anaeretes elongata (Fabricius); Dejean, 1833: 181 Dichelonycha [sic] elongata (Fabricius); Blanchard, 1850: 116 Dichelonyx elongata (Fabricius); Leng, 1920: 257

Synonym: Melolontha hexagona Germar, 1824: 124

Synonym: Dichelonycha [sic] virescens Kirby, 1837: 134

Dichelonyx fulgida crotchii Horn Dichelonycha [sic] crotchii Horn, 1876: 189 Dichelonyx fulgida crotchi (Horn); Distribution: United States

Dichelonyx fulgida fulgida LeConte Dichelonycha [sic] fulgida LeConte, 1856: 280 Dichelonyx fulgida (LeConte); Leng, 1920: 258 Dichelonyx fulgida fulgida (LeConte): Saylor, 1945b: 151 Distribution: Canada, United States

Dichelonyx fulgida mormona Fall 218 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Dichelonycha [sic] mormona Fall, 1907: 249 Dichelonyx mormona Fall; Leng, 1920: 258 Dichelonyx fulgida mormona (Fall): Saylor, 1945b: 151 Distribution: United States

Dichelonyx fuscula LeConte Dichelonycha [sic] fuscula LeConte, 1856: 281 Dichelonyx fuscula LeConte; Leng, 1920: 258 Distribution: United States

Dichelonyx kirbyi Brown Dichelonyx kirbyi Brown, 1946: 107 Distribution: Canada, United States

Dichelonyx lateralis Fall Dichelonycha [sic] lateralis Fall, 1901a: 292 Dichelonyx lateralis Fall; Leng, 1920: 258 Distribution: United States

Dichelonyx linearis (Gyllenhal) Melolontha linearis Gyllenhal, 1817: 186 Dichelonyx linearis (Gyllenhal); Harris, 1827: 7 Dichelonycha [sic] linearis (Gyllenhal); Kirby, 1837: 134 Distribution: Canada, United States

Synonym: Dichelonycha [sic] testacea Kirby, 1837: 135 Dichelonyx testacea (Kirby); Leng, 1920: 258

Dichelonyx muscula Fall Dichelonycha [sic] muscula Fall, 1901: 286 Dichelonyx muscula Fall; Leng, 1920: 258 Distribution: United States

Dichelonyx nana Fall Dichelonycha [sic] nana Fall, 1901: 286 Dichelonyx nana Fall; Leng, 1920: 258 Distribution: United States

Dichelonyx oregona Van Dyke Dichelonyncha [sic] oregona Van Dyke, 1918: 11 Dichelonyx oregona Van Dyke; Leng, 1920: 258 Distribution: United States

Dichelonyx pallens LeConte Dichelonycha [sic] pallens LeConte,1860: 283 Dichelonyx pallens LeConte; Leng, 1920: 258 Distribution: United States

219 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Dichelonycha [sic] longiclava Fall, 1901: 287 Dichelonyx longiclava Fall; Leng, 1920: 258

Dichelonyx picea Horn Dichelonycha [sic] picea Horn, 1894: 396 Dichelonyx picea Horn; Leng, 1920: 258 Distribution: México

Dichelonyx pusilla LeConte Dichelonycha [sic] pusilla LeConte,1856: 282 Dichelonyx pusilla LeConte; Leng, 1920: 258 Distribution: México, United States

Dichelonyx robusta Fall Dichelonycha [sic] robusta Fall, 1901: 288 Dichelonyx robusta Fall; Leng, 1920: 258 Distribution: United States

Dichelonyx subvittata LeConte Dichelonycha [sic] subvittata LeConte,1856: 279 Dichelonyx subvittata LeConte; Leng, 1920: 257 Distribution: Canada, United States

Dichelonyx sulcata arizonensis Saylor Dichelonyx arizonensis Saylor, 1933: 158 Dichelonyx sulcata arizonensis Saylor; Saylor, 1945b: 156 Distribution: United States

Dichelonyx sulcata sulcata LeConte Dichelonycha [sic] sulcata LeConte,1856: 281 Dichelonyx sulcata LeConte; Leng, 1920: 258 Distribution: United States

Dichelonyx testaceipennis Fall Dichelonycha [sic] testaceipennis Fall, 1907: 248 Dichelonyx testaceipennis Fall; Leng, 1920: 258 Distribution: United States

Dichelonyx truncata LeConte Dichelonycha [sic] truncata LeConte, 1856: 281 Dichelonyx truncata LeConte; Leng, 1920: 258 Distribution: United States

Dichelonyx vaga Fall Dichelonycha [sic] vaga Fall, 1901: 285 Dichelonyx vaga Fall; Leng, 1920: 258 Distribution: United States

220 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Dichelonyx valida LeConte Dichelonycha [sic] valida LeConte, 1856: 281 Dichelonyx valida LeConte; Leng, 1920: 258 Distribution: United States

Dichelonyx vandykei Saylor Dichelonyx vandykei Saylor, 1932: 284 Distribution: United States

Dichelonyx vicina columbiana Hopping Dichelonyx columbiana Hopping, 1931: 236 Dichelonyx vicina columbiana Hopping; Saylor, 1939d: 457 Distribution: Canada, United States

Dichelonyx vicina vicina Fall Dichelonycha [sic] vicina Fall, 1901: 291 Dichelonyx vicina Fall; Leng, 1920: 258 Distribution: Canada, United States

Synonym: Dichelonyx deserta Hopping, 1931: 236

Genus DICRANIA Le Peletier and Audinet-Serville Dicrania Le Peletier and Audinet-Serville, 1828: 371. Type species Dicrania nigra Le Peletier and Audinet-Serville, by monotypy.

Synonym: Monocrania Laporte, 1832: 410. Type species Monocrania luridipennis Laporte, subsequent designation (Evans 2003: 263).

Keys to species: Frey, 1972a: 68

Dicrania appendiculata (Mannerheim) Melolontha appendiculata Mannerheim, 1829: 67 Monocrania appendiculata (Mannerheim); Burmeister, 1855; 70 Dicrania appendiculata (Mannerheim); Lacordaire, 1856: 250 Distribution: Brazil

Dicrania argentina Moser Dicrania argentina Moser, 1919a: 13 Dicrania argentinus Moser; Frey, 1972a: 73 [error] Distribution: Argentina

Dicrania castaneipennis Chevrolat Dicrania castaneipennis Chevrolat, 1844: 103 Distribution: Brazil

Synonym: Dicrania badia Burmeister, 1855: 69

Dicrania cuibana Frey 221 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Dicrania cuibana Frey, 1965b: 143 Distribution: Brazil

Dicrania ebenina Blanchard Dicrania ebenina Blanchard, 1850: 95 Distribution: Brazil

Dicrania fasciculata Blanchard Dicrania fasciculata Blanchard, 1850: 95 Distribution: Brazil

Dicrania femorata (Mannerheim) Melolontha femorata Mannerheim, 1829: 66 Monocrania femorata (Mannerheim); Burmeister, 1855: 70 Dicrania femorata (Mannerheim); Lacordaire, 1856: 250 Distribution: Brazil

Dicrania flavipennis Kraatz Dicrania flavipennis Kraatz, 1896: 189 Distribution: Paraguay

Dicrania flavoscutellata Laporte Dicrania flavoscutellata Laporte, 1832: 408 Distribution: Argentina, Brazil, Paraguay

Synonym: Dicrania kulzeri Frey, 1970b: 286

Synonym: Monocrania nigricans Laporte, 1832: 410 Dicrania nigricans (Laporte); Lacordaire, 1856: 249

Synonym: Dicrania scutellaris Chevrolat, 1844: 102

Dicrania fraudulenta Frey Dicrania fraudulenta Frey, 1972a: 73 Distribution: Paraguay

Dicrania hirsuta Frey Dicrania hirsuta Frey, 1970b: 285 Distribution: Brazil

Dicrania hirtipes Laporte Dicrania hirtipes Laporte, 1832: 409 Distribution:

Dicrania laevipennis Moser Dicrania laevipennis Moser, 1921b: 171 Distribution: Colombia, French Guiana

222 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Dicrania lasiopus Burmeister Dicrania lasiopus Burmeister, 1855: 69 Distribution: Brazil

Dicrania luridipennis (Laporte) Monocrania luridipennis Laporte, 1832: 410 Dicrania luridipennis (Laporte); Lacordaire, 1856: 249 Distribution: Brazil

Dicrania martinezi Frey Dicrania martinezi Frey, 1972a: 71 Distribution: Peru

Dicrania metzi Kraatz Dicrania (Monocrania) metzii Kraatz, 1895: 319 Dicrania metzi Kraatz; Dalla Torre, 1913: 333 Distribution: Brazil

Dicrania morio Kraatz Dicrania morio Kraatz, 1895: 317 Distribution: Brazil

Dicrania moseri Frey Dicrania moseri Frey, 1972a: 74 Distribution: Brazil

Dicrania nigra Le Peletier and Audinet-Serville Dicrania nigra Le Peletier and Audinet-Serville, 1828: 372 Distribution: Brazil

Synonym: Dicrania brasiliensis Laporte, 1832: 409

Synonym: Dicrania dichroa Perty, 1833: 51

Synonym: Dicrania nigra rubricollis Le Peletier and Audinet-Serville, 1828: 372

Dicrania nigriceps Guérin-Méneville Dicrania nigriceps Guérin-Méneville, 1844a: 102 Distribution: Brazil, Paraguay

Dicrania nitida Frey Dicrania nitida Frey, 1972a: 78 Distribution: Brazil

Dicrania notaticollis Moser Dicrania notaticollis Moser, 1921b: 171 Distribution: Brazil

223 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Dicrania ohausi Moser Dicrania ohausi Moser, 1921b: 170 Distribution: Brazil

Dicrania ornaticollis Frey Dicrania ornaticollis Frey, 1972a: 79 Distribution: Brazil

Dicrania parvula Frey Dicrania parvula Frey, 1970b: 287 Distribution: Bolvia

Dicrania pellita Eschscholtz Dicrania pellita Eschscholtz, 1822: 15 Distribution: Brazil

Dicrania pentyphylla Moser Dicrania pentyphylla Moser, 1919a: 16 Distribution: Brazil

Dicrania peruana Frey Dicrania peruana Frey, 1972a: 82 Distribution: Peru

Dicrania piligera Blanchard Dicrania piligera Blanchard, 1850: 95 Distribution: Bolivia

Dicrania pilosa Moser Dicrania pilosa Moser, 1919a: 15 Distribution: Brazil

Dicrania plaumanni Frey Dicrania plaumanni Frey, 1972a: 74 Distribution: Bolivia

Dicrania popei Frey Dicrania popei Frey, 1972a: 72 Distribution: Brazil

Dicrania rugosipennis Blanchard Dicrania rugosipennis Blanchard, 1850: 95 Distribution: Brazil

Dicrania santarema Frey Dicrania santarema Frey, 1972a: 81 Distribution: Brazil

224 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Dicrania setosicollis Frey Dicrania setosicollis Frey, 1972a: 69 Distribution: Brazil

Dicrania signaticollis Moser Dicrania signaticollis Moser, 1919a: 13 Distribution: Brazil

Dicrania similis Frey Dicrania similis Frey, 1972a: 70 Distribution: Brazil

Dicrania subvestita Guérin-Méneville Dicrania subvestita Guérin-Méneville, 1844a: 102 Distribution: Brazil

Synonym: Dicrania squamulata Moser, 1919a: 14

Dicrania unicolor fulvicollis Kraatz Dicrania (Monocrania) unicolor var. fulvicollis Kraatz, 1895: 316 Distribution: Brazil

Dicrania unicolor unicolor Kraatz Dicrania (Monocrania) unicolor Kraatz, 1895: 316 Dicrania unicolor Kraatz; Frey, 1972a: 73 Distribution: Argentina, Paraguay

Dicrania varicolor Frey Dicrania varicolor Frey, 1972a: 75 Distribution: Guyana

Dicrania velutina Laporte Dicrania velutina Laporte, 1832: 409 Distribution: Brazil

Synonym: Dicrania quadracristata Guérin-Méneville, 1844a: 101

Genus EUBARYBAS Gutiérrez Eubarybas Gutiérrez, 1952: 216. Type species Eubarybas asper Gutiérrez by original designation.

Eubarybas asper Gutiérrez Eubarybas asper Gutiérrez, 1952: 216 Distribution: Venezuela

Genus GAMA Blanchard Gama Blanchard, 1850: 118. Type species Gama grandicornis Blanchard, by subsequent designation (Lacordaire, 1856: 265).

225 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Harpodactyla Burmeister, 1855: 17. Type species Harpodactyla grandicornis Burmeister (a junior homonym and synonym of Gama grandicornis Blanchard), subsequent designation (Evans 2003: 268).

Gama aenea (Moser) Harpodactyla aenea Moser, 1924a: 121 Gama aenea (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama aeneotincta Moser Gama aeneotincta Moser, 1918b: 112 Distribution: Brazil

Gama aeneoviridis (Moser) Harpodactyla aeneoviridis Moser, 1918c: 333 Gama aeneoviridis (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama aenescens (Moser) Harpodactyla aenescens Moser, 1918c: 333 Gama aenescens (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama angustula (Moser) Harpodactyla angustula Moser, 1921b: 141 Gama angustula (Moser) Distribution: Brazil

Gama aurichalcea (Burmeister) Harpodactyla aurichalcea Burmeister, 1855: 19 Gama aurichalcea (Burmeister); Harold, 1869a: 1141 Distribution: Brazil

Gama breviceps (Moser) Harpodactylus breviceps Moser, 1918b: 113 Gama breviceps (Moser); Distribution: Brazil

Gama espiritosantensis (Moser) Harpodactyla esperitosantensis Moser, 1919b: 18 Gama espiritosantensis (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama fulvipennis (Moser) Harpodactylus fulvipennis Moser, 1918c: 331 Gama fulvipennis (Moser); Blackwelder, 1944: 228 Distribution: Brazil

226 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Gama fusca Blanchard Gama fusca Blanchard, 1850: 119 Distribution: Brazil

Gama fuscoaenea Moser Gama fuscoaenea Moser, 1921b: 140 Distribution: Brazil

Gama grandicornis Blanchard Gama grandicornis Blanchard, 1850: 118 Distribution: Brazil, Colombia

Synonym: Harpodactyla grandicornis Burmeister, 1855: 18

Gama hamifera (Moser) Plectris hamifera Moser, 1921b: 169 Gama hamifera (Moser); Frey, 1967a: 6 Distribution: Brazil

Synonym: Gama rotundiceps Frey, 1964a: 130

Gama lepichaeta (Moser) Harpodactyla lepichaeta Moser, 1919b: 18 Gama lepichaeta (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama lutea Moser Harpodactyla lutea Moser, 1924b: 157 Gama lutea (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama metallescens (Moser) Harpodactyla metallescens Moser, 1918c: 330 Gama metallescens (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama musiva (Burmeister) Philochloenia musiva Burmeister, 1855: 30 Harpodactyla musiva (Burmeister); Moser, 1919b: 17 Gama musiva (Burmeister); Blackwelder, 1944: 228 Distribution: Brazil

Gama ohausi (Moser) Harpodactyla ohausi Moser, 1921b: 141 Gama ohausi (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama pallida Blanchard 227 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Gama pallida Blanchard, 1850: 118 Distribution: Argentina

Gama pilosa (Moser) Harpodactyla pilosa Moser, 1918c: 334 Gama pilosa (Moser); Blackwelder, 1944; 228 Distribution: Brazil

Gama pulverulenta (Burmeister) Harpodactyla pulverulenta Burmeister, 1855: 19 Gama pulverulenta (Burmeister); Harold, 1869a: 1141 Distribution: Brazil

Gama quadrifoliata (Moser) Harpodactyla quadrifoliata Moser, 1918b: 114 Gama quadrifoliata (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama relucens Blanchard Gama relucens Blanchard, 1850: 118 Distribution: Venezuela

Gama rufoflava (Moser) Harpodactyla rufoflava Moser, 1919b: 18 Gama rufoflava (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama rugipennis (Moser) Harpodactyla rugipennis Moser, 1918c: 335 Gama rugipennis (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama squamiventris Steinheil Gama squamiventris Steinheil, 1874: 558 Distribution: Argentina

Gama viridescens Moser Gama viridescens Moser, 1918b: 114 Distribution: Brazil

Gama viridiaenea (Moser) Harpodactyla viridiaenea Moser, 1918c: 332 Gama viridiaenea (Moser); Blackwelder, 1944: 228 Distribution: Brazil

Gama viridifusca Moser Gama viridifusca Moser, 1918b: 111 Distribution: Brazil 228 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Genus GASTROHOPLUS Moser Gastrohoplus Moser, 1921b: 165. Type species Gastrohoplus mirabilis Moser, by monotypy.

Gastrohoplus mirabilis Moser Gastrohoplus mirabilis Moser, 1921b: 165 Distribution: Brazil

Genus GYMNOPYGE Linell Gymnopyge Linell, 1896: 723. Type species Gymnopyge hopliaeformis Linell, by original designation.

Gymnopyge coquilletti Linell Gymnopyge coquilletti Linell, 1896: 724 Distribution: United States

Gymnopyge hirsuta Cazier Gymnopyge hirsuta Cazier, 1939a: 17 Distribution: United States

Gymnopyge hopliaeformis Linell Gymnopyge hopliaeformis Linell, 1896: 724 Distribution: United States

Gymnopyge pygmaea Linell Gymnopyge pygmaea Linell, 1896: 724 Distribution: United States

Genus HADROCERUS Guérin-Méneville Hadrocerus Guérin-Méneville, 1838: 83. Type species Hadrocerus castaneipennis Guérin-Méneville, by monotypy. (for Pachycerus)

Synonym: Pachycerus Guérin-Méneville, 1831b: pl. 3 (preoccupied). Type species Pachycerus castaneipennis Guérin-Méneville, by monotypy.

Hadrocerus castaneipennis (Guérin-Méneville) Pachycerus castaneipennis Guérin-Méneville, 1831b: pl. 3. Hadrocerus castaneipennis (Guérin-Méneville) 1838: 83 Distribution: Brazil

Genus HAMATOPLECTRIS Frey Hamatoplectris Frey, 1967a: 126. Type species Hamatoplectris rosettae Frey, by original designation.

Hamatoplectris caracana Frey Hamatoplectris caracana Frey, 1969b: 399 Distribution: Brazil

Hamatoplectris rosettae Frey Hamatoplectris rosettae Frey, 1967a: 126 229 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Hamatoplectris spinifer Frey Hamatoplectris spinifer Frey, 1967a: 127 Distribution: Brazil

Genus HERCITIS Burmeister Hercitis Burmeister, 1855: 79. Type species Hercitis pygmaea Burmeister, by monotypy.

Hercitis flavosquamosa Moser Hercitis flavosquamosa Moser, 1924a: 128 Distribution: Brazil

Hercitis pygmaea Burmeister Hercitis pygmaea Burmeister, 1855: 79 Distribution: Brazil

Hercitis testacea Moser Hercitis testacea Moser, 1918b: 143 Distribution: Brazil

Hercitis zikani Moser Hercitis zikani Moser, 1921b: 174 Distribution: Brazil

Genus HIERITIS Burmeister Hieritis Burmeister, 1855: 41. Type species Hieritis macrocera Burmeister, by monotypy.

Hieritis macrocera Burmeister Hieritis macrocera Burmeister, 1855: 41 Distribution: Brazil

Genus HILARIANUS Blanchard Hilarianus Blanchard, 1851: 169. Type species Hilarianus anguliceps Blanchard, subsequent designation (Evans 2003: 273).

Hilarianus anguliceps Blanchard Hilarianus anguliceps Blanchard, 1851: 169 Hilarianus angulicollis Blanchard; Frey, 1974f: 331 (misspelling) Distribution: Brazil

Hilarianus ovalis Blanchard Hilarianus ovalis Blanchard, 1851: 170 Distribution: Brazil

Hilarianus rufinus Blanchard Hilarianus rufinus Blanchard, 1851: 169 Distribution: Brazil 230 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Hilarianus suboblongus Blanchard Hilarianus suboblongus Blanchard, 1851: 170 Distribution: Brazil

Hilarianus uniformis Blanchard Hilarianus uniformis Blanchard, 1851: 170 Distribution: Brazil

Genus ISONYCHUS Mannerheim Isonychus Mannerheim, 1829: 69. Type species Isonychus sulphureus Mannerheim, original designation.

Synonym: Colporhina Curtis, 1845: 454. Type species Colporhina bifoveolata Curtis, by monotypy.

Keys: Frey, 1964b: 320; 1965a: 113; 1970a: 124; Morón, 1994: 44

Isonychus aenescens Moser Isonychus aenescens Moser, 1919b: 42 Distribution: Bolivia

Isonychus aequatorialis Moser Isonychus aequatorialis Moser, 1924a: 124 Distribution: Ecuador

Isonychus albicinctus (Mannerheim) Omaloplia albicincta Mannerheim,1829: 68 Isonichus [sic] albicinctus (Mannerheim); Dejean, 1833: 182 Isonychus albicinctus (Mannerheim); Blanchard, 1850: 89 Distribution: Brazil

Isonychus albofasciatus Blanchard Isonychus albofasciatus Blanchard, 1850: 88 Distribution: Brazil

Synonym: Isonychus impluviatus Burmeister, 1855: 46

Isonychus albosignatus Moser Isonychus albosignatus Moser, 1921b: 158 Distribution: Brazil

Isonychus alienus Frey Isonychus alienus Frey, 1970a: 138 Distribution: Brazil

Isonychus angosturanus Moser Isonychus angosturanus Moser, 1924a: 126 Distribution: Venezuela 231 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus arbusticola Erichson Isonychus arbusticola Erichson, 1847a: 102 Distribution: Peru

Isonychus argentinus Moser Isonychus argentinus Moser, 1919b: 41 Distribution: Argentina

Isonychus arizonensis Howden Isonychus arizonensis Howden, 1959: 59 Distribution: México, United States

Isonychus bahianus Frey Isonychus bahianus Frey, 1974f: 331 Distribution: Brazil

Isonychus bimaculatus Burmeister Isonychus bimaculatus Burmeister, 1855: 55 Distribution: Brazil

Synonym: Isonychus sulcatus Moser, 1918b: 137

Isonychus bistrigus Burmeister (emendation) Isonychus bistriga Burmeister, 1855: 47 Distribution: Brazil

Isonychus bivittatus Burmeister Isonychus bivittatus Burmeister, 1855: 53 Distribution: Colombia

Isonychus boliviensis Moser Isonychus boliviensis Moser, 1918c: 349 Distribution: Bolivia

Isonychus braumeisteri Frey Isonychus braumeisteri Frey, 1969b: 383 Distribution: Venezuela

Isonychus bruchiformis (Germar) Melolontha bruchiformis Germar, 1829: 128 Isonychus bruchiformis (Germar); Burmeister, 1855: 52 Distribution: Brazil

Synonym: Isonychus fuscomaculatus Moser, 1921b: 159

Isonychus burmeisteri Dalla Torre Isonychus burmeisteri Dalla Torre, 1913: 326 (for Burmeister, 1855: 53) 232 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Colombia

Synonym: Isonychus concolor Burmeister, 1855: 53 (not Blanchard, 1850: 85)

Isonychus callosipygus Frey Isonychus callosipygus Frey, 1965a: 123 Distribution: Brazil

Isonychus castaneus Burmeister Isonychus castaneus Burmeister, 1855: 49 Distribution: Colombia, Venezuela

Isonychus catharinae Blanchard Isonychus catharinae Blanchard, 1850: 88 Distribution: Brazil

Isonychus caudiculatus Moser Isonychus caudiculatus Moser, 1918c: 347 Distribution: Brazil

Isonychus cervicapra Frey Isonychus cervicapra Frey, 1965a: 120 Distribution: Peru

Isonychus cervinalis Frey Isonychus cervinalis Frey, 1965a: 121 Distribution: Peru

Isonychus cervinodes Frey Isonychus cervinodes Frey, 1965a: 119 Distribution: Peru

Isonychus cervinus Erichson Isonychus cervinus Erichson, 1847a: 103 Distribution: Bolivia

Synonym: Isonychus abdominalis Moser, 1918c: 342

Synonym: Isonychus fuscescens Blanchard, 1850: 87

Isonychus chacoensis Moser Isonychus chacoensis Moser, 1918c: 344 Distribution: Bolivia, Ecuador

Isonychus chiriquinus Bates Isonychus chiriquinus Bates, 1887: 149 Distribution: Colombia, Panama

233 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus cinereus Blanchard Isonychus cinereus Blanchard, 1850: 87 Distribution: Brazil

Isonychus costaricensis Moser Isonychus costaricensis Moser, 1918c: 349 Distribution: Costa Rica

Isonychus crinitus Blanchard Isonychus crinitus Blanchard, 1850: 88 Distribution: Colombia, Venezuela

Isonychus denudatus Blanchard Isonychus denudatus Blanchard, 1850: 87 Distribution: Argentina, Colombia, Uruguay

Synonym: Isonychus tenuivestis Moser, 1921b: 163

Isonychus discolor Moser Isonychus discolor Moser, 1924a: 126 Distribution: Ecuador

Isonychus egregius Frey Isonychus egregius Frey, 1965a: 122 Distribution: Peru

Isonychus elegans Blanchard Isonychus elegans Blanchard, 1850: 86 Distribution: Brazil

Isonychus elongatus Frey Isonychus elongatus Frey, 1965a: 118 Distribution: Bolivia

Isonychus erectepilosus Frey Isonychus erectepilosus Frey, 1970a: 143 Distribution: Brazil

Isonychus fasciatipennis Moser Isonychus fasciatipennis Moser, 1924a: 125 Distribution: Venezuela

Isonychus fasciolatus Blanchard Isonychus fasciolatus Blanchard, 1850: 87 Distribution: Brazil, Guyana

Isonychus flaviventris Moser Isonychus flaviventris Moser, 1921b: 165 234 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Ecuador

Isonychus flavopilosus Moser Isonychus flavopilosus Moser, 1921b: 161 Distribution: Ecuador

Isonychus fraternus Moser Isonychus fraternus Moser, 1918c: 346 Distribution: Bolivia

Isonychus fraudulentus Frey Isonychus fraudulentus Frey, 1969b: 381 Distribution: Peru

Isonychus fulvescens Blanchard Isonychus fulvescens Blanchard, 1850: 88 Distribution: Colombia

Isonychus fulvipennis Moser Isonychus fulvipennis Moser, 1921b: 161 Distribution: Ecuador

Isonychus fuscipennis Blanchard Isonychus fuscipennis Blanchard, 1850: 85 Distribution: Brazil

Isonychus gracilipes Blanchard Isonychus gracilipes Blanchard, 1850: 86 Distribution: Brazil

Isonychus gracilis Burmeister Isonychus gracilis Burmeister, 1855: 33 Distribution: Colombia

Isonychus granarius Burmeister Isonychus granarius Burmeister, 1855: 48 Distribution: Brazil, Surinam

Isonychus granuliventris Frey Isonychus granuliventris Frey, 1970a: 145 Distribution: Brazil

Isonychus griseolus Moser Isonychus griseolus Moser, 1921b: 160 Distribution: Brazil

Isonychus griseopilosus Moser Isonychus griseopilosus Moser, 1921b: 162 235 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Isonychus griseus Mannerheim Isonychus griseus Mannerheim, 1829: 74 Isonichus [sic] griseus Mannerheim; Dejean, 1833: 182 Distribution: Brazil

Isonychus guayanensis Frey Isonychus guayanensis Frey, 1970a: 144 Distribution: Guyana

Isonychus hiekei Frey Isonychus hiekei Frey, 1965a: 118 Distribution: Ecuador

Isonychus hirsutus Bates Isonychus hirsutus Bates, 1887: 148 Distribution: Panama

Isonychus kulzeri Frey Isonychus kulzeri Frey, 1967b: 421 Distribution: Ecuador

Isonychus kuntzeni Moser Isonychus kuntzeni Moser, 1921b: 164 Distribution: Ecuador

Isonychus laevipygus Frey Isonychus laevipygus Frey, 1964b: 331 Distribution: Colombia

Isonychus leechi Frey Isonychus leechi Frey, 1969b: 380 Distribution: Colombia

Isonychus limbatus Burmeister Isonychus limbatus Burmeister, 1855: 51 Distribution: Brazil (not México)

Isonychus lindemannae Frey Isonychus lindemannae Frey, 1974e: 279 Distribution: Brazil

Isonychus lineatus Burmeister Isonychus lineatus Burmeister, 1855: 51 Distribution: French Guiana, Guyana

Isonychus lineola Blanchard 236 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus lineola Blanchard, 1850: 87 Distribution: Brazil

Isonychus lituratus Blanchard Isonychus lituratus Blanchard, 1850: 86 Distribution: Colombia, Venezuela

Synonym: Isonychus concolor Blanchard, 1850: 85 Isonychus concolor var. concolor Blanchard; Frey, 1964b: 322

Synonym: Isonychus tristis Moser, 1924a: 126 Isonychus concolor var. tristis Blanchard; Frey, 1964b: 322

Synonym: Isonychus villosus Blanchard, 1850: 86 Isonychus concolor var. villosus Blanchard; Frey, 1964b: 322

Isonychus lojanus Frey Isonychus lojanus Frey, 1967b: 420 Distribution: Ecuador

Isonychus maculatus Waterhouse Isonychus maculatus Waterhouse, 1874: 203 Distribution: Colombia

Isonychus maculipennis Moser Isonychus maculipennis Moser, 1918c: 344 Distribution: Bolivia

Isonychus marmoratus Blanchard Isonychus marmoratus Blanchard, 1850: 88 Distribution: Colombia, Venezuela

Synonym: Isonychus flavofasciatus Moser, 1918c: 339

Synonym: Isonychus principalis Burmeister, 1855: 43

Isonychus marmoreus Burmeister Isonychus marmoreus Burmeister, 1855: 46 Distribution: Brazil

Isonychus microsquamosus Frey Isonychus microsquamosus Frey, 1970a: 140 Distribution: Brazil

Isonychus minutus (Fabricius) Trichius minutus Fabricius, 1801: 134 Isonychus minutus (Fabricius); Burmeister, 1855: 52 Distribution: Brazil, French Guiana 237 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus murinus Blanchard Isonychus murinus Blanchard, 1850: 85 Distribution: Brazil

Isonychus mus Burmeister Isonychus mus Burmeister, 1855: 45 Distribution: Brazil

Isonychus mutans Frey Isonychus mutans Frey, 1970a: 142 Distribution: Brazil

Isonychus nebeculus Frey Isonychus nebeculus Frey, 1969b: 378 Distribution: Peru

Isonychus neglectus Moser Isonychus neglectus Moser, 1918c: 337 Distribution: México

Isonychus nigripes Moser Isonychus nigripes Moser, 1921b: 164 Distribution: Brazil

Isonychus nitens Moser Isonychus nitens Moser, 1921b: 163 Distribution: Peru

Isonychus nitidus Burmeister Isonychus nitidus Burmeister, 1855: 49 Distribution: Colombia, Venezuela

Isonychus nubeculus Frey Isonychus nubeculus Frey, 1969b: 378 Distribution: Peru

Isonychus nubilis Burmeister Isonychus nubilis Burmeister, 1855: 48 Distribution: Brazil

Synonym: Isonychus ruficeps Moser, 1924a: 125

Isonychus nudipennis Frey Isonychus nudipennis Frey, 1967b: 422 Distribution: Ecuado

Isonychus obesulus Burmeister 238 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus obesulus Burmeister, 1855: 55 Distribution: Brazil

Isonychus oblongoguttatus Moser Isonychus oblongoguttatus Moser, 1921b: 162 Distribution: Brazil

Isonychus oblongomaculatus Moser Isonychus oblongomaculatus Moser, 1921b: 158 Distribution: Brazil

Isonychus obsoletus Blanchard Isonychus obsoletus Blanchard, 1850: 87 Distribution: Colombia, Venezuela

Synonym: Isonychus fucatus Burmeister, 1855: 45

Isonychus ocellatus Burmeister Isonychus ocellatus Burmeister, 185: 46 Distribution: Guatemala, México

Isonychus ochreaceus Blanchard Isonychus ochreaceus Blanchard, 1850: 87 Distribution: Brazil

Isonychus ohausi Moser Isonychus ohausi Moser, 1921b: 163 Distribution: Brazil, Ecuador

Isonychus ornatipennis Moser Isonychus ornatipennis Moser, 1921b: 158 Distribution: Brazil

Isonychus ovinus Erichson Isonychus ovinus Erichson, 1847a: 103 Distribution: Peru

Isonychus paganus Blanchard Isonychus paganus Blanchard, 1850: 86 Distribution: Colombia

Isonychus paradoxus Bates Isonychus paradoxus Bates, 1887: 150 Distribution: Panama

Isonychus parallelus Frey Isonychus parallelus Frey, 1970a: 144 Distribution: Brazil 239 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus paranus Moser Isonychus paranus Moser, 1921b: 159 Distribution: Brazil

Isonychus parvulus Moser Isonychus parvulus Moser, 1918c: 345 Distribution: Bolivia

Isonychus pauloensis Frey Isonychus pauloensis Frey, 1970a: 143 Distribution: Brazil

Isonychus pavonii Erichson Isonychus pavonii Erichson, 1847a: 102 Distribution: Peru

Isonychus penai Frey Isonychus penai Frey, 1966: 330 Distribution: Ecuador

Isonychus pereirai Frey Isonychus pereirai Frey, 1970a: 140 Distribution: Brazil

Isonychus peruanus Moser Isonychus peruanus Moser, 1921b: 157 Distribution: Peru

Isonychus phlaeopterus Blanchard Isonychus phlaeopterus Blanchard, 1850: 88 Isonychus phoenicopterus Blanchard; Blackwelder, 1944: 231 (misspelling) Distribution: Brazil

Isonychus pictus Sharp Isonychus pictus Sharp, 1877: 131 Distribution: Costa Rica, Nicaragua

Isonychus pilicollis Moser Isonychus pilicollis Moser, 1924a: 125 Distribution: Ecuador

Isonychus pilosus Evans Isonychus pilosus Evans, 2003 (for Burmeister, 1855: 43) Distribution: Colombia

Synonym: Isonychus crinitus Burmeister, 1855: 43 (not Blanchard, 1850: 88)

240 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus piperitus Bates Isonychus ocellatus var. piperitus Bates, 1887: 148 Isonychus piperatus Bates; Morón et al., 1997: 258 Distribution: México

Isonychus podicalis Moser Isonychus podicalis Moser, 1918c: 340 Distribution: Colombia

Isonychus politus Burmeister Isonychus politus Burmeister, 1855: 49 Distribution: Brazil

Isonychus prasinus Nonfried Isonychus prasinus Nonfried, 1892a: 223 Distribution: Honduras

Isonychus puchellus Moser Isonychus puchellus Moser, 1918c: 348 Distribution: Peru

Isonychus religiosus Evans Isonychus religiosus Evans, 2003 (for Burmeister, 1855: 54) Distribution: Colombia

Synonym: Isonychus paganus Burmeister, 1855: 54 (not Blanchard, 1850: 86)

Isonychus rosettae Frey Isonychus rosettae Frey, 1969: 380 Distribution: Peru

Isonychus rugicollis Burmeister Isonychus rugicollis Burmeister, 1855: 55 Distribution: Brazil

Isonychus saltanus Frey Isonychus saltanus Frey, 1969b: 382 Distribution: Argentina

Isonychus saylori Frey Isonychus saylori Frey, 1969b: 380 Distribution: Peru

Isonychus schneblei Frey Isonychus schneblei Frey, 1964b: 329 Distribution: Colombia

Isonychus scutellaris Moser 241 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus scutellaris Moser, 1918c: 341 Distribution: Colombia

Isonychus setifer Moser Isonychus setifer Moser, 1918c: 338 Distribution: Bolivia

Isonychus similis Frey Isonychus similis Frey, 1973a: 260 Distribution: Peru

Isonychus simplex Frey Isonychus simplex Frey, 1976b: 377 Distribution: Brazil

Isonychus simulator Frey Isonychus simulator Frey, 1969: 380 Distribution: Peru

Isonychus soricinus Blanchard Isonychus soricinus Blanchard, 1850: 85 Distribution: Brazil

Isonychus squamifer Blanchard Isonychus squamifer Blanchard, 1850: 86 Distribution: Colombia

Synonym: Isonychus jaspideus Burmeister, 1855: 54

Isonychus squamulosus Frey Isonychus squamulosus Frey, 1970a: 138 Distribution: Brazil

Isonychus striatipennis Moser Isonychus striatipennis Moser, 1918c: 338 Distribution: México

Isonychus striolatus Frey Isonychus striolatus Frey, 1970a: 141 Distribution: Brazil

Isonychus submaculatus Moser Isonychus submaculatus Moser, 1921b: 157 Distribution: Brazil

Isonychus sulcicollis Moser Isonychus sulcicollis Moser, 1921b: 160 Distribution: Brazil 242 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus sulphureus seminiger Moser Isonychus sulphureus var. seminiger Moser, 1918c: 346 Distribution: Brazil

Isonychus sulphureus sulphureus Mannerheim Isonychus sulphureus Mannerheim, 1829: 73 Isonichus [sic] sulphureus Mannerheim; Dejean 1833: 182 Distribution: Brazil

Synonym: Isonychus aurantiacus Burmeister, 1855: 50

Synonym: Isonychus plagiatus Burmeister, 1855: 51

Isonychus suturalis Mannerheim Isonychus suturalis Mannerheim, 1829: 74 Distribution: Brazil

Isonychus tesellatus Burmeister Isonychus tesellatus Burmeister, 1855: 50 Distribution: Brazil

Isonychus tomentosus Burmeister Isonychus tomentosus Burmeister, 1855: 48 Distribution: México

Isonychus unicolor Blanchard Isonychus unicolor Blanchard, 1850: 88 Distribution: Brazil

Isonychus unidens Frey Isonychus unidens Frey, 1972b: 98 Distribution: Brazil

Isonychus uniformis Moser Isonychus uniformis Moser, 1920: 23 Distribution: Costa Rica

Isonychus ursus Moser Isonychus ursus Moser, 1918c: 341 Distribution: Colombia

Isonychus varians Blanchard Isonychus varians Blanchard, 1850: 86 Distribution: Colombia

Isonychus variegatus (Germar) Melolontha variegatus Germar, 1824: 128 243 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Isonychus variegatus (Germar); Burmeister, 1855: 536 Distribution: Brazil

Synonym: Colporhina bifoveolata Curtis, 1845: 454

Synonym: Isonychus degluptus Germar, 1824: 623

Isonychus variipennis Moser Isonychus variipennis Moser, 1918c: 343 Distribution: Bolivia

Isonychus ventralis Moser Isonychus ventralis Moser, 1921b: 159 Distribution: Brazil

Isonychus vestitus (Laporte) Serica vestita Laporte, 1840: 148 Isonychus vestitus (Laporte); Burmeister, 1855: 54 Distribution: Brazil

Isonychus vicinus Moser Isonychus vicinus Moser, 1818: 345 Distribution: Bolivia

Isonychus vittatus Burmeister Isonychus vittatus Burmeister, 1855: 47 Distribution: México

Isonychus vittiger Blanchard Isonychus vittiger Blanchard, 1850: 86 Distribution: Brazil

Isonychus vittipennis Moser Isonychus vittipennis Moser, 1921b: 164 Distribution: Brazil

Isonychus zikani Moser Isonychus zikani Moser, 1921b: 160 Distribution: Brazil

Genus ISSACARIS Fairmaire Issacaris Fairmaire, 1889: 117. Type species Issacaris petalophora Fairmaire (a junior synonym of Dicrania aeneobrunnea Philippi), by monotypy.

Keys to species: Gutiérrez, 1952: 219

Issacaris bullocki Gutiérrez Issacaris bullocki Gutiérrez, 1952: 221 244 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Chile

Issacaris petalophora Fairmaire Issacaris petalophora Fairmaire, 1889: 117 Distribution: Chile

Synonym: Dicrania aeneobrunnea Philippi, 1861: 738 Issacaris aeneobrunnea (Philippi); Gutiérrez, 1946a: 15

Issacaris setosiventris Gutiérrez Issacaris setosiventris Gutiérrez, 1952: 219 Distribution: Chile

Genus MACRODACTYLUS Dejean Macrodactylus Dejean, 1821: 58. Type species Melolontha subspinosa Fabricius, by subsequent designation (Evans 2003: 290).

Synonym: Stenothorax Harris, 1827: 8 (as subgenus). Type species Melolontha subspinosa Fabricius, by monotypy.

Keys to species: Horn, 1876: 184; Carrillo and Gibson, 1960, 26; Morón, 1994: 44; Arce-Pérez and Morón, 2000a: 150.

Macrodactylus aeneicollis Moser Macrodactylus aeneicollis Moser, 1918c: 353 Distribution: Argentina

Macrodactylus aeneipes Moser Macrodactylus aeneipes Moser, 1918b: 138 Distribution: Argentina

Macrodactylus aeneus Kirsch Macrodactylus aeneus Kirsch, 1885: 219 Distribution: Bolivia

Macrodactylus aequatorialis Moser Macrodactylus aequatorialis Moser, 1918c: 352 Distribution: Ecuador

Macrodactylus affinis Laporte Macrodactylus affinis Laporte, 1840: 147 Distribution: Brazil

Macrodactylus amoenus Frey Macrodactylus amoenus Frey, 1975c: 256 Distribution: Ecuador

Macrodactylus angustatus (Palisot de Beauvois) 245 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Melolontha angustata Palisot de Beauvois, 1805: 30 Macrodactylus angustatus (Palisot de Beauvois); LeConte, 1856: 278 Distribution: Canada, United States

Synonym: Macrodactylus setulosus LeConte, 1856: 277

Macrodactylus annulitarsis Blanchard Macrodactylus annulitarsis Blanchard, 1850: 90 Distribution: Venezuela

Macrodactylus argentinus Moser Macrodactylus argentinus Moser, 1918c: 352 Distribution: Argentina

Macrodactylus batesi Arce-Pérez and Morón Macrodactylus batesi Arce-Pérez and Morón, 2000a: 161 Distribution: México

Macrodactylus bicolor Moser Macrodactylus bicolor Moser, 1921b: 166 Distribution: Brazil

Macrodactylus bilineolatus Moser Macrodactylus bilineolatus Moser, 1919a: 7 Distribution: Peru

Macrodactylus bistriatus Moser Macrodactylus bistriatus Moser, 1919a: 8 Distribution: Peru

Macrodactylus bolivianus Moser Macrodactylus bolivianus Moser, 1918c: 350 Distribution: Bolivia

Macrodactylus brenskei Moser Macrodactylus brenskei Moser, 1918c: 354 Distribution: Peru

Macrodactylus carrilloi Arce-Pérez and Morón Macrodactylus carrilloi Arce-Pérez and Morón, 2000a: 223 Distribution: México

Macrodactylus championi Bates Macrodactylus championi Bates, 1887: 146 Distribution: Guatemala, México

Macrodactylus chilensis Solier Macrodactylus chilensis Solier, 1851: 115 246 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Chile

Macrodactylus cinereus Blanchard Macrodactylus cinereus Blanchard, 1850: 90 Distribution: Peru

Macrodactylus columbianus Dalla Torre Macrodactylus columbianus Dalla Torre, 1913: 328 (for Burmeister, 1855: 58) Distribution: Colombia

Synonym: Macrodactylus cinereus Burmeister, 1855: 58 (not Blanchard, 1850: 90)

Macrodactylus conjunctus Moser Macrodactylus conjunctus Moser, 1921a: 56 Distribution: Colombia

Macrodactylus costaricensis Moser Macrodactylus costaricensis Moser, 1919a: 12 Distribution: Costa Rica

Macrodactylus costulatus Bates Macrodactylus costulatus Bates, 1887: 143 Distribution: Belize, Costa Rica, Guatemala, Honduras, México, Nicaragua

Synonym: Macrodactylus costulatus var. rufipennis Bates, 1887: 143

Macrodactylus crassipes Philippi Macrodactylus crassipes Philippi, 1864: 444 Distribution: Chile

Synonym: Macrodactylus nigrinus Philippi, 1864: 443

Macrodactylus cupreus Blanchard Macrodactylus cupreus Blanchard, 1850: 91 Distribution: Bolivia

Macrodactylus curtipilis Frey Macrodactylus curtipilis Frey, 1969b: 385 Distribution: Brazil

Macrodactylus cyanonigrus Evans Macrodactylus cyanonigrus Evans, 2003 (for Moser, 1919a: 9) Distribution: Colombia

Synonym: Macrodactylus nigrocyaneus Moser, 1919a: 9 (not Bates, 1887: 145)

Macrodactylus dimidiatus Guérin-Méneville Macrodactylus dimidiatus Guérin-Méneville, 1844b: pl. 147 247 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Guatemala, Honduras, México

Synonym: Macrodactylus semicoeruleus Burmeister, 1855: 62

Macrodactylus discicollis Moser Macrodactylus discicollis Moser, 1919a: 1 Distribution: Brazil

Macrodactylus discipennis Moser Macrodactylus discipennis Moser, 1918c: 356 Distribution: Peru

Macrodactylus disparilis Moser Macrodactylus disparilis Moser, 1919a: 1 Distribution: Bolivia

Macrodactylus dorsatus Burmeister Macrodactylus dorsatus Burmeister, 1855: 59 Distribution: Brazil

Macrodactylus duckei Ohaus Macrodactylus duckei Ohaus, 1913b: 206 Distribution: Brazil

Macrodactylus elegans Nonfried Macrodactylus elegans Nonfried, 1894: 142 Distribution: Ecuador

Macrodactylus excellens Kirsch Macrodactylus excellens Kirsch, 1865: 47 Distribution: Colombia

Macrodactylus farinosus Philippi Macrodactylus farinosus Philippi, 1864: 442 Distribution: Chile

Macrodactylus felix Kirsch Macrodactylus felix Kirsch, 1885: 219 Distribution: Colombia

Macrodactylus fulvescens Bates Macrodactylus fulvescens Bates, 1887: 140 Distribution: México

Synonym: Macrodactylus fulvescens var. nigritarsis Bates, 1887: 141 Macrodactylus nigritarsis Bates; Arce-Peréz and Morón, 2001: 166

Synonym: Macrodactylus rhomboderus Bates, 1887: 141 248 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Macrodactylus fulvipennis Blanchard Macrodactylus fulvipennis Blanchard, 1850: 91 Distribution: Brazil

Synonym: Macrodactylus luridipennis Burmeister, 1855: 62

Macrodactylus fulvipes Moser Macrodactylus fulvipes Moser, 1918b: 139 Distribution: Paraguay

Macrodactylus gracilis Moser Macrodactylus gracilis Moser, 1918c: 351 Distribution: Bolivia

Macrodactylus griseus Moser Macrodactylus griseus Moser, 1918c: 351 Distribution: Argentina

Macrodactylus haenschi Moser Macrodactylus haenschi Moser, 1919a: 9 Distribution: Ecuador

Macrodactylus hondurensis Arce-Pérez and Morón Macrodactylus hondurensis Arce-Pérez and Morón, 2005: 32 Distribution: Honduras

Macrodactylus impressus Bates Macrodactylus impressus Bates, 1887: 143 Distribution: México

Macrodactylus infuscatus infuscatus Bates Macrodactylus infuscatus Bates, 1887: 139 Distribution: México

Macrodactylus infuscatus vicinus Bates Macrodactylus infuscatus var. vicinus Bates, 1887: 139 Distribution: México

Macrodactylus limbatus Blanchard Macrodactylus limbatus Blanchard, 1850: 91 Distribution: Colombia, Venezuela

Synonym: Macrodactylus pauperatus Burmeister, 1855: 62

Macrodactylus lineaticollis Bates Macrodactylus lineaticollis Bates, 1887: 140 Distribution: Guatemala, México 249 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Macrodactylus lineatus Chevrolat Macrodactylus lineatus Chevrolat, 1834: 79 Distribution: México

Macrodactylus longicollis Latrielle Macrodactylus longicollis Latrielle, 1811: 202 Distribution: México

Synonym: Macrodactylus nigrocyaneus Bates, 1887: 145

Macrodactylus longipes Burmeister Macrodactylus longipes Burmeister, 1855: 61 Distribution: Brazil

Macrodactylus manantlecus Arce-Pérez and Morón Macrodactylus manantlecus Arce-Pérez and Morón, 2000a: 163 Distribution: México

Macrodactylus marginicollis Moser Macrodactylus marginicollis Moser, 1919a: 11 Distribution: Peru

Macrodactylus marianus Moser Macrodactylus marianus Moser, 1924a: 124 Distribution: Colombia

Macrodactylus meridanus Moser Macrodactylus meridanus Moser, 1926: 205 Distribution: Venezuela

Macrodactylus mexicanus Burmeister Macrodactylus mexicanus Burmeister, 1855: 58 Distribution: México

Macrodactylus montanus Arce-Pérez and Morón Macrodactylus montanus Arce-Pérez and Morón, 2000a: 212 Distribution: Honduras, México

Macrodactylus murinus Bates Macrodactylus murinus Bates, 1887: 147 Distribution: México

Macrodactylus nigricornis Moser Macrodactylus nigricornis Moser, 1919b: 43 Distribution: Brazil

Macrodactylus nigripes Bates 250 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Macrodactylus nigripes Bates, 1887: 140 Distribution: México

Macrodactylus nitidicollis Blanchard Macrodactylus nitidicollis Blanchard, 1850: 89 Distribution: Colombia, Venezuela

Synonym: Macrodactylus bilineatus Burmeister, 1855: 61

Macrodactylus nitididorsis Frey Macrodactylus nitididorsis Frey, 1976b: 379 Distribution: Brazil

Macrodactylus nobilis Frey Macrodactylus nobilis Frey, 1963: 554 Distribution: Bolivia

Macrodactylus obscuritarsis Moser Macrodactylus obscuritarsis Moser, 1926: 205 Distribution: Paraguay

Macrodactylus ocreatus Bates Macrodactylus ocreatus Bates, 1887: 141 Distribution: México

Macrodactylus ohausi Moser Macrodactylus ohausi Moser, 1921a: 55 Distribution: Ecuador

Macrodactylus ovaticollis Bates Macrodactylus ovaticollis Bates, 1887: 146 Distribution: Colombia, Panama

Macrodactylus pallens Blanchard Macrodactylus pallens Blanchard, 1850: 90 Distribution: Brazil

Synonym: Macrodactylus flavicornis Burmeister, 1855: 58

Macrodactylus paraguayensis Moser Macrodactylus paraguayensis Moser, 1921b: 166 Distribution: Paraguay

Macrodactylus penai Frey Macrodactylus penai Frey, 1972b: 101 Distribution: Ecuador

Macrodactylus peruanus Moser 251 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Macrodactylus peruanus Moser, 1919a: 2 Distribution: Peru

Macrodactylus pexus Kirsch Macrodactylus pexus Kirsch, 1865: 48 Distribution: Colombia

Macrodactylus pluto Wickham Macrodactylus pluto Wickham, 1912: 24 Distribution: United States Remark: fossil species

Macrodactylus pokornyanus Morón Macrodactylus pokornyanus Arce-Pérez and Morón, 2000a: 214 Distribution: México

Macrodactylus praecellens Moser Macrodactylus praecellens Moser, 1919a: 6 Distribution: Ecuador

Macrodactylus propheticus Wickham Macrodactylus propheticus Wickham, 1912: 25 Distribution: United States Remark: fossil species

Macrodactylus pulchellus Moser Macrodactylus pulchellus Moser, 1918c: 355 Distribution: Colombia

Macrodactylus pulchripes Blanchard Macrodactylus pulchripes Blanchard, 1850: 90 Distribution: Colombia

Macrodactylus pumilio Burmeister Macrodactylus pumilio Burmeister, 1855: 58 Distribution: Brazil

Macrodactylus rufescens Bates Macrodactylus rufescens Bates, 1887: 145 Distribution: Guatemala, México

Macrodactylus sapphirinus Moser Macrodactylus sapphirinus Moser, 1919a: 10 Distribution: Peru

Macrodactylus scutellaris Frey Macrodactylus scutellaris Frey, 1972b: 101 Distribution: Brazil 252 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Macrodactylus sericeicollis Bates Macrodactylus sericeicollis Bates, 1887: 142 Distribution: Guatemala, Honduras, Nicaragua

Macrodactylus sericinus Bates Macrodactylus sericinus Bates, 1887: 144 Distribution: Nicaragua

Macrodactylus signatipennis Moser Macrodactylus signatipennis Moser, 1919a: 4 Distribution: Bolivia

Macrodactylus silaonus Bates Macrodactylus silaonus Bates, 1887: 140 Distribution: México

Macrodactylus silarutus Evans Macrodactylus silarutus Evans, 2003 (for Laporte, 1840: 147) Distribution: Brazil

Synonym: Macrodactylus suturalis Laporte, 1840: 147 (not Mannerheim, 1829: 74)

Macrodactylus sparsesetosus Frey Macrodactylus sparsesetosus Frey, 1969b: 386 Distribution: Brazil

Macrodactylus squamiger Frey Macrodactylus squamiger Frey, 1963: 553 Distribution: Colombia

Macrodactylus suavis Bates Macrodactylus suavis Bates, 1887: 144 Distribution: Costa Rica, Nicaragua, Panama

Macrodactylus subaeneus Burmeister Macrodactylus subaeneus Burmeister, 1855: 63 Distribution: Brazil

Macrodactylus submarginatus Bates Macrodactylus submarginatus Bates, 1887: 142 Distribution: México

Macrodactylus subspinosus (Fabricius) Melolontha subspinosa Fabricius, 1775: 39 Melolontha (Stenothorax) subspinosa (Fabricius); Harris, 1827: 8 Macrodactylus subspinosus (Fabricius); Laporte, 1840: 147 Distribution: United States 253 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Macrodactylus barbatus Fitch, 1865: 199

Synonym: Melolontha elongata Herbst, 1790: 145 Melolontha (Stenothorax) elongata Herbst; Harris, 1827: 8

Synonym: Macrodactylus polyphagus Burmeister, 1855: 57

Macrodactylus subvittatus Burmeister Macrodactylus subvittatus Burmeister, 1855: 59 Distribution: Colombia

Macrodactylus sulcicollis Moser Macrodactylus sulcicollis Moser, 1919a: 5 Distribution: Peru

Macrodactylus sulphureus Blanchard Macrodactylus sulphureus Blanchard, 1850: 90 Distribution: Colombia

Macrodactylus surianus Arce-Pérez and Morón Macrodactylus surianus Arce-Pérez and Morón, 2000a: 216 Distribution: México

Macrodactylus suturalis Mannerheim Macrodactylus suturalis Mannerheim, 1829: 74 Distribution: Brazil

Synonym: Macrodactylus haemhorrhous Perty, 1833: 51

Macrodactylus sylphis Bates Macrodactylus sylphis Bates, 1887: 144 Distribution: Costa Rica, Nicaragua, Panama

Macrodactylus tenuilineatus Guérin-Méneville Macrodactylus tenuilineatus Guérin-Méneville, 1843: 21 Distribution: Colombia

Synonym: Macrodactylus flavolineatus Guérin-Méneville, 1843: 21

Macrodactylus thoracicus Kirsch Macrodactylus thoracicus Kirsch, 1885: 218 Distribution: Ecuador, Panama

Macrodactylus tibialis Arce-Pérez and Morón Macrodactylus tibialis Arce-Pérez and Morón, 2005: 29 Distribution: Honduras, Nicaragua

254 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Macrodactylus uniformis Horn Macrodactylus uniformis Horn, 1876: 185 Distribution: México, United States

Macrodactylus variipes Bates Macrodactylus variipes Bates, 1887: 138 Distribution: México

Macrodactylus velutinus Frey Macrodactylus velutinus Frey, 1976b: 380 Distribution: Brazil

Macrodactylus virens Bates Macrodactylus virens Bates, 1887: 147 Distribution: México

Macrodactylus vittipennis Moser Macrodactylus vittipennis Moser, 1919a: 3 Distribution: Peru

Macrodactylus zischkai Frey Macrodactylus zischkai Frey, 1966: 331 Distribution: Bolivia

Macrodactylus zunilensis Bates Macrodactylus zunilensis Bates, 1887: 142 Distribution: Guatemala

Genus MALLOTARSUS Blanchard Mallotarsus Blanchard, 1850: 119. Type species Mallotarsus spadiceus Blanchard, by monotypy.

Mallotarsus spadiceus Blanchard Mallotarsus spadiceus Blanchard, 1850: 119 Distribution: Brazil

Genus MANODACTYLUS Moser Manodactylus Moser, 1919b: 43. Type species Manodactylus gaujoni Moser, by monotypy.

Manodactylus gaujoni Moser Manodactylus gaujoni Moser, 1919b: 44 Distribution: Ecuador

Genus MANONYCHUS Moser Manonychus Moser, 1919c: 346. Type species Manonychus unguicularis Moser, by monotypy.

Manonychus birabeni Martínez Manonychus birabeni Martínez, 1959b: 59 Description: Brazil 255 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Manonychus conipygus Frey Manonychus conipygus Frey, 1976b: 378 Description: Brazil

Manonychus densicollis Frey Manonychus densicollis Frey, 1974f: 320 Description: Brazil

Manonychus martinezi Frey Manonychus martinezi Frey, 1974f: 320 Description: Brazil

Manonychus rosettae Frey Manonychus rosettae Frey, 1976b: 377 Description: Brazil

Manonychus unguicularis Moser Manonychus unguicularis Moser, 1919c: 346 Description: Brazil

Genus MANOPUS Laporte Manopus Laporte, 1840: 147. Type species Manopus biguttatus Laporte, by monotypy.

Manopus biguttatus Laporte Manopus biguttatus Laporte, 1840: 147 Distribution: Colombia

Manopus simillimus Moser Manopus simillimus Moser, 1926: 207 Distribution: Colombia

Genus OEDICHIRA Burmeister Oedichira Burmeister, 1855: 80. Type species Oedichira pachydactyla Burmeister, subsequent designation (Evans 2003: 303).

Oedichira crassimana Burmeister Oedichira crassimana Burmeister, 1855: 81 Distribution: Brazil

Oedichira pachydactyla Burmeister Oedichira pachydactyla Burmeister, 1855: 81 Distribution: Brazil

Genus PACHYLOTOMA Blanchard Pachylotoma Blanchard, 1850: 121. Type species Pachylotoma viridis Blanchard, by monotypy.

Pachylotoma viridis Blanchard 256 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Pachylotoma viridis Blanchard, 1850: 121 Distribution: South America

Genus PAULOSAWAYA Martínez and D'Andretta Paulosawaya Martínez and D'Andretta, 1955: 346. Type species Paulosawaya ornatissima Martínez and d'Andretta, by original designation.

Paulosawaya ornatissima Martínez and D'Andretta Paulosawaya ornatissima Martínez and D'Andretta, 1955: 348 Distribution: Ecuador

Genus PECTINOSOMA Arrow Pectinosoma Arrow, 1913b: 430. Type species Pectinosoma elongata Arrow, by monotypy.

Pectinosoma elongata Arrow Pectinosoma elongata Arrow, 1913b: 430 Distribution: Guyana

Genus PLECTRIS Le Peletier and Audinet-Serville Plectris Le Peletier and Audinet-Serville, 1828: 369. Type species Plectris tomentosa Le Peletier and Audinet-Serville, by monotypy.

Synonym: Euryaspis Blanchard, 1851: 130. Type species Euryaspis gaudichaudii Blanchard, by monotypy.

Synonym: Junkia Dalla Torre, 1913: 310 (for Nonfried, 1894: 11). Type species Trichoderma ceylanica Nonfried, by monotypy.

Synonym: Pseudoserica Guérin-Méneville, 1838: 86. Type species Pseudoserica marmorea Guérin- Méneville, by monotypy.

Synonym: Trichoderma Nonfried, 1894: 11 (preoccupied). Type species Trichoderma ceylanica Nonfried, by monotypy.

Keys to species: Frey, 1967a: 6; Chalumeau, 1982: 330.

Plectris abdominalis Frey Plectris abdominalis Frey, 1967a: 31 Distribution: Brazil

Plectris aberrans Frey Plectris aberrans Frey, 1964c: 698 Distribution: Peru

Plectris abnormalis (Moser) Philochlaenia [sic] abnormalis Moser, 1919b: 24 Plectris abnormalis (Moser); Frey, 1967a: 52 Distribution: Brazil 257 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris acreensis Frey Plectris acreensis Frey, 1973a: 269 Distribution: Brazil

Plectris acutestosa Frey Plectris acutestosa Frey, 1974f: 327 Distribution: Brazil

Plectris aenea Blanchard Plectris aenea Blanchard, 1851: 129 Distribution: Colombia

Plectris aeneicollis (Moser) Philochlaenia [sic] aeneicollis Moser, 1918b: 127 Plectris aeneicollis (Moser); Frey, 1967a: 88 Distribution: Brazil

Plectris aeneofusca (Moser) Philochlaenia [sic] aeneofusca Moser, 1918c: 336 Plectris aeneofusca (Moser); Frey, 1967a: 103 Distribution: Ecuador

Plectris aeneomicans Frey Plectris aeneomicans Frey, 1967a: 94 Distribution: Brazil

Plectris aeneorufa (Moser) Philochlaenia [sic] aeneorufa Moser, 1918b: 122 Plectris aeneorufa (Moser); Frey, 1967a: 84 Distribution: Argentina, Brazil

Plectris aenescens (Moser) Philochlaenia [sic] aenescens Moser, 1918b: 135 Plectris aenescens (Moser); Frey, 1967a; 118 Distribution: Brazil

Plectris aequatorialis (Moser) Philochlaenia [sic] aequatorialis Moser 1921b: 143 Plectris aequatorialis (Moser); Frey, 1967a: 104 Distribution: Ecuador

Synonym: Philochlaenia [sic] witti Moser, 1924b: 160 Plectris witli [sic] Moser; Frey, 1967a: 104

Plectris aerata Burmeister Plectris aerata Burmeister, 1855: 87 Distribution: Brazil 258 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris afflicta Blanchard Plectris afflicta Blanchard, 1850: 127 Distribution: Brazil

Synonym: Philochloenia setifera Burmeister, 1855: 35 Plectris setifera (Burmeister); Evans, 2003: 306

Plectris ahena Burmeister Plectris ahena Burmeister, 1855: 32 Distribution: Brazil

Synonym: Philochlaenia [sic] virescens Blanchard, 1850: 122 (not Blanchard, 1842: 11) Plectris virescens (Blanchard); Frey, 1967a: 114

Plectris alboscutata Frey Plectris alboscutata Frey, 1967a: 100 Distribution: Brazil

Plectris albovittata Frey Plectris albovittatus Frey, 1967a: 118 Plectris albovittata Frey; Frey, 1974d: 245 Distribution: Brazil

Plectris aliena Chapin Plectris aliena Chapin, 1934a: 34 Distribution: Paraguay [Australia, United States]

Plectris alternata Frey Plectris alternata Frey, 1967a: 65 (for Burmeister, 1855: 36) Distribution: Brazil, Colombia

Synonym: Philochloenia augerona Burmeister, 1855: 36 (not Blanchard, 1850: 127) Plectris augerona (Burmeister); Evans, 2003: 307

Plectris alvarengai Frey Plectris alvarengai Frey, 1967a: 52 Distribution: Brazil

Plectris amabilis Frey Plectris amabilis Frey, 1967a: 67 Distribution: Venezuela

Plectris ambigua Frey Plectris ambigua Frey, 1967a: 95 Distribution: Brazil

Plectris ambitiosa (Blanchard) 259 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Philochlaenia [sic] ambitiosa Blanchard, 1850: 121 Plectris ambitiosa (Blanchard); Frey, 1967a: 75 Distribution: Brazil

Plectris amoena Frey Plectris amoena Frey, 1973a: 272 Distribution: Brazil

Plectris andicola Frey Plectris andicola Frey, 1969b: 387 Distribution: Bolivia

Plectris andina Frey Plectris andina Frey, 1967a: 54 Distribution: Bolivia

Plectris angerona Blanchard Plectris angerona Blanchard, 1850: 127 Distribution: Colombia

Plectris angusta Frey Plectris angusta Frey, 1974f: 330 Distribution: Brazil

Plectris anomala (Kirsch) Philochlaenia [sic] anomala Kirsch, 1885: 217 Plectris anomala (Kirsch); Frey, 1967a: 79 Distribution: Ecuador

Plectris antennalis Moser Plectris antennalis Moser, 1924a: 161 Distribution: Brazil

Plectris antennata Frey Plectris antennata Frey, 1969b: 394 Distribution: Brazil

Plectris apicalis (Moser) Philochlaenia [sic] apicalis Moser, 1921b: 145 Plectris apicalis (Moser); Frey, 1967a; 118 Distribution: Ecuador

Plectris archidonensis Moser Plectris archidonensis Moser, 1924a: 161 Distribution: Ecuador

Plectris argentata Frey Plectris argentata Frey, 1967a: 76 260 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris argentina (Bruch) Philochloenia argentina Bruch, 1909: 343 Plectris argentinus (Bruch); Frey, 1967a; 36 Distribution: Argentina

Plectris atubana Frey Plectris atubana Frey, 1972b: 93 Distribution: Brazil

Plectris bahiensis Frey Plectris bahiensis Frey, 1975b: 253 Distribution: Brazil

Plectris balthasari Frey Plectris balthasari Frey, 1967a: 59 Distribution: Brazil

Plectris baraudi Frey Plectris baraudi Frey, 1967a: 77 Distribution: Ecuador

Plectris barbitarsis (Moser) Philochlaenia [sic] barbitarsis Moser, 1919b: 26 Plectris barbitarsis (Moser); Frey, 1967a: 49 Distribution: Venezuela

Plectris barda Blanchard Plectris barda Blanchard, 1850: 127 Distribution: Brazil

Synonym: Plectris texta Burmeister Plectris texta Burmeister, 1855: 34

Plectris bicolor Frey Plectris bicolor Frey, 1967a: 96 Distribution: Brazil

Plectris bilobata (Moser) Philochlaenia [sic] bilobata Moser, 1921b: 148 Plectris bilobata (Moser); Frey, 1967a: 103 Distribution: Bolivia

Synonym: Plectris stöckleini Frey, 1967a: 124 Plectris stoeckleini Frey; Frey, 1974d: 256

Plectris binotata Blanchard 261 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris binotata Blanchard, 1850: 127 Distribution: Brazil

Synonym: Plectris cuprescens Blanchard, 1850: 127

Plectris blanchardi Frey Plectris blanchardi Frey, 1967a: 86 (for Moser 1926: 206) Distribution: Argentina

Synonym: Plectris argentina Moser 1926: 206 (not Bruch, 1909: 343)

Plectris boliviensis (Moser) Philochlaenia [sic] boliviensis Moser, 1921b: 153 Plectris boliviensis (Moser); Frey, 1967a: 42 Distribution: Bolivia

Plectris bonariensis Frey Plectris bonariensis Frey, 1967a: 92 Distribution: Argentina

Plectris breiti Frey Plectris breiti Frey, 1967a: 69 Distribution: Brazil

Plectris breviceps Frey Plectris breviceps Frey, 1967a: 91 Distribution: Brazil

Plectris brevicollis (Moser) Philochlaenia [sic] brevicollis Moser, 1919b: 31 Plectris brevicollis (Moser); Frey, 1967a: 52 Distribution: Brazil

Plectris brevipes (Blanchard) Philochlaenia [sic] brevipes Blanchard, 1850: 122 Plectris brevipes (Blanchard); Evans, 2003: 310 Distribution: Brazil

Plectris brevipilosa Moser Plectris brevipilosa Moser, 1924a: 128 Distribution: Brazil

Plectris brevis Blanchard Plectris brevis Blanchard, 1851: 129 Distribution: Colombia

Plectris brevisetosa Moser Plectris brevisetosa Moser, 1918b: 153 262 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris brittoni Frey Plectris brittoni Frey, 1967a: 101 Distribution: Brazil

Plectris bruchi Moser Plectris bruchi Moser, 1924a: 128 Distribution: Argentina

Plectris burmeisteri (Kirsch) Philochlaenia [sic] burmeisteri Kirsch, 1871: 362 Plectris burmeisteri (Kirsch); Frey, 1967a: 38 Distribution: Colombia

Plectris calcarata Frey Plectris calcarata Frey, 1967a: 29 Distribution: Brazil

Plectris caliginosa Blanchard Plectris caliginosa Blanchard, 1850: 126 Distribution: Brazil

Synonym: Plectris cervina Burmeister, 1855: 84

Synonym: Plectris latifrons Blanchard, 1850: 125

Plectris calva Frey Plectris calva Frey, 1969b: 391 Distribution: Brazil

Plectris candezei Frey Plectris candezei Frey, 1967a: 54 Distribution: Brazil, Guyana, Peru

Plectris caniventris (Burmeister) Philochloenia caniventris Burmeister, 1855: 38 Plectris caniventris (Burmeister); Frey, 1967a; 117 Distribution: Brazil

Plectris castaneipennis (Moser) Philochlaenia [sic] castaneipennis Moser, 1921b: 150 Plectris castaneipennis (Moser); Frey, 1967a: 79 Distribution: Bolivia

Plectris cayennensis Moser Plectris cayennensis Moser, 1921b: 152 Distribution: French Guiana 263 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris centralis (Bruch) Philochloenia centralis Bruch, 1909: 342 Plectris centralis (Bruch); Frey, 1967a: 36 Distribution: Argentina

Plectris ceylanica (Nonfried) Trichoderma ceylanica Nonfried, 1894: 11 Junkia ceylanica (Nonfried); Dalla Torre, 1913: 310 Plectris ceylanica (Nonfried); Frey, 1967a: 79 Distribution: Brazil

Plectris ciliata Frey Plectris ciliata Frey, 1967a: 27 Distribution: Brazil

Plectris cinerascens Moser Plectris cinerascens Moser, 1918b: 161 Distribution: Brazil

Plectris cinereopilosa (Moser) Philochlaenia [sic] cinereopilosa Moser, 1924b: 161 Plectris cinereopilosa (Moser); Evans, 2003: 312 Distribution: Brazil

Plectris clypealis Frey Plectris clypealis Frey, 1975b: 252 Distribution: Ecuador

Plectris clypeata Burmeister Plectris clypeata Burmeister, 1855: 83 Distribution: Brazil

Plectris collarti Frey Plectris collarti Frey, 1967a: 11 Distribution: Uruguay

Plectris comata (Blanchard) Brahmina comata Blanchard, 1851; 140 Plectris comata (Blanchard); Frey, 1967a: 8 Distribution: Argentina, Paraguay

Plectris conformis Frey Plectris conformis Frey, 1967a: 63 Distribution: Colombia

Plectris contaminata Blanchard Plectris contaminata Blanchard, 1850: 128 264 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris cordobana (Moser) Philochlaenia [sic] cordobana Moser, 1924b: 158 Plectris cordobana (Moser); Frey, 1967a: 89 Distribution: Argentina

Plectris corrientensis (Moser) Philochlaenia [sic] corrientensis Moser, 1919b: 19 Plectris corrientensis (Moser); Moser, 1926: 206 Distribution: Argentina, Brazil, Paraguay

Synonym: Rhinaspis nitidula Moser, 1921b: 170 Plectris nitidula (Moser); Evans, 2003: 313

Plectris corumbana Moser Plectris corumbana Moser, 1921b: 169 Distribution: Brazil

Plectris corvicoana (Moser) Philochlaenia [sic] corvicoana Moser, 1919b: 34 Plectris corvicoana (Moser); Frey, 1967a: 52 Distribution: Bolivia

Plectris costulata Frey Plectris costulata Frey, 1967a: 34 Distribution: Brazil

Plectris coxalis (Moser) Philochlaenia [sic] coxalis Moser, 1921b: 147 Plectris coxalis (Moser); Frey, 1967a: 105 Distribution: Ecuador

Plectris crassa Blanchard Plectris crassa Blanchard, 1850: 125 Distribution: Brazil

Plectris crassesetosa Frey Plectris crassesetosa Frey, 1967a: 107 Distribution: Brazil

Plectris cucullata Burmeister Plectris cucullata Burmeister, 1855: 90 Distribution: Brazil

Plectris curta Burmeister Plectris curta Burmeister, 1855: 88 Distribution: Brazil 265 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris curticollis Frey Plectris curticollis Frey, 1967a: 11 Distribution: Brazil

Plectris curtipilis Frey Plectris curtipilis Frey, 1967a: 58 Distribution: Colombia

Plectris curtisetis Frey Plectris curtisetis Frey, 1969b: 393 Distribution: Brazil

Plectris cuyana (Bruch) Philochloenia cuyana Bruch, 1909: 347 Plectris cuyana (Bruch); Frey, 1967a: 79 Distribution: Argentina

Plectris cylindrica Burmeister Plectris cylindrica Burmeister, 1855: 86 Distribution: Brazil

Synonym: Philochlaenia [sic] gracilicornis Moser, 1919c: 347 Plectris gracilicornis (Moser); Evans, 2003: 314

Plectris cylindriformis Frey Plectris cylindriformis Frey, 1967a: 120 Distribution: Venezuela

Plectris decipiens Burmeister Plectris decipiens Burmeister, 1855: 88 Distribution: Argentina, Brazil, Paraguay

Plectris decolorata Blanchard Plectris decolorata Blanchard, 1850: 126 Distribution: Brazil

Synonym: Plectris eucalyptra Burmeister, 1855: 87

Plectris delicatula (Blanchard) Philochlaenia [sic] delicatula Blanchard, 1850: 122 Plectris delicatula (Blanchard); Frey, 1967a: 83 Distribution: Brazil

Plectris denominata Frey Plectris denominata Frey, 1967a: 116 (for Burmeister, 1855: 32) Distribution: Brazil

266 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Plectris tomentosa Burmeister, 1855: 32 (not Le Peletier and Audinet-Serville, 1828: 369)

Plectris densaticollis Frey Plectris densaticollis Frey, 1967a: 72 Distribution: Brazil

Plectris densehirsuta Frey Plectris densehirsuta Frey, 1967a: 91 Distribution: Uruguay

Plectris densevestita Moser Plectris densevestita Moser, 1924a: 160 Distribution: Brazil

Plectris depressicollis Frey Plectris depressicollis Frey, 1967a: 96 Distribution: Brazil

Plectris desiderata Evans Plectris desiderata Evans, 2003 (for Burmeister, 1855: 89) Distribution: Brazil

Synonym: Plectris ambitiosa Burmeister, 1855: 89 (not Blanchard, 1850: 121)

Plectris devillei Frey Plectris devillei Frey, 1967a: 93 Distribution: Ecuador

Plectris difformis Frey Plectris difformis Frey, 1973a: 265 Distribution: Brazil

Plectris dimorpha Frey Plectris dimorpha Frey, 1976b: 382 Distribution: Brazil

Plectris duplopilosa (Moser) Philochlaenia [sic] duplopilosa Moser, 1921b: 148 Plectris duplopilosa (Moser); Frey, 1967a: 114 Distribution: Brazil

Plectris elongata Burmeister Plectris elongata Burmeister, 1855: 86 Distribution: Brazil

Synonym: Plectris excisiceps Moser, 1918b: 158

Plectris emarginata (Moser) 267 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Philochlaenia [sic] emarginata Moser, 1921a: 55 Plectris emarginata (Moser); Frey, 1967a: 49 Distribution: Brazil

Plectris endroedii Frey Plectris endrödii Frey, 1967a: 60 Plectris endroedii Frey; Frey, 1974: 248 Distribution: Brazil

Plectris eucalypti Frey Plectris eucalypti Frey, 1973a: 264 Distribution: Brazil

Plectris eusquamosa Frey Plectris eusquamosa Frey, 1975b: 254 Distribution: Brazil

Plectris excisiceps (Moser) Philochlaenia [sic] excisiceps Moser, 1921b: 153 Plectris excisiceps (Moser); Frey, 1967a: 42 Distribution: French Guiana

Plectris exigua Frey Plectris exigua Frey, 1973a: 267 Distribution: Brazil

Plectris farinosa Burmeister Plectris farinosa Burmeister, 1855: 87 Distribution: Brazil

Plectris fassli (Moser) Philochlaenia [sic] fassli Moser, 1919b: 30 Plectris fassli (Moser); Frey, 1967a: 52 Distribution: Colombia

Plectris festiva (Burmeister) Philochloenia festiva Burmeister, 1855: 37 Plectris festiva (Burmeister); Frey, 1967a: 64 Distribution: Colombia, Venezuela

Plectris flavicornis (Burmeister) Philochloenia flavicornis Burmeister, 1855: 30 Plectris flavicornis (Burmeister); Evans, 2003: 317 Distribution: Brazil

Plectris flavohirta Blanchard Plectris flavohirta Blanchard, 1851: 129 Distribution: Guyana 268 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris fulgida Frey Plectris fulgida Frey, 1973a: 268 Distribution: Brazil

Plectris fulva Blanchard Plectris fulva Blanchard, 1851: 129 Distribution: Brazil, Colombia

Synonym: Plectris grenadensis Blanchard, 1851: 129

Plectris fungicola Arrow Plectris fungicola Arrow, 1900: 177 Distribution: St. Vincent

Plectris fusca Moser Plectris fusca Moser, 1818b: 162 Distribution: Brazil

Plectris fuscoaenea Moser Plectris fuscoaenea Moser, 1921b: 167 Distribution: Brazil

Plectris fuscoviridis (Moser) Philochlaenia [sic] fuscoviridis Moser, 1918b: 129 Plectris fuscoviridis (Moser); Frey, 1967a: 75 Distribution: Brazil

Plectris fuscula (Moser) Philochlaenia [sic] fuscula Moser, 1918b: 133 Plectris fuscula (Moser); Frey, 1967a: 118 Distribution: Venezuela

Plectris gaudichaudii (Blanchard) Euryaspis gaudichaudii Blanchard, 1851: 130 Plectris gaudichaudi (Blanchard); Dalla Torre, 1913: 335 Distribution: Brazil

Synonym: Phyllophaga (Phytalus) malleata Frey, 1964c: 696 Plectris malleatus [sic] Frey; Frey, 1975a: 220

Plectris gebieni (Moser) Philochlaenia [sic] gebieni Moser, 1921b: 150 Plectris gebieni (Moser); Frey, 1967a: 118 Distribution: Brazil

Plectris glabrata Frey Plectris glabrata Frey, 1967a: 33 269 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris glabripennis Frey Plectris glabripennis Frey, 1967a: 7 Distribution: Brazil

Plectris globulicollis Frey Plectris globulicollis Frey, 1973a: 270 Distribution: Brazil

Plectris goetzi Frey Plectris goetzi Frey, 1967a: 111 Distribution: Colombia

Plectris gracilicornis Moser Plectris gracilicornis Moser, 1918b: 154 Distribution: Brazil

Plectris grisea Moser Plectris grisea Moser, 1921b: 167 Distribution: Brazil

Plectris griseohirta Frey Plectris griseohirta Frey, 1967a: 87 (for Moser, 1921b: 149) Distribution: Argentina, Brazil, Colombia

Synonym: Philochlaenia [sic] griseovestita Moser, 1921b: 149 (not Moser, 1921b: 167) Plectris griseovestita (Moser); Evans, 2003: 319

Plectris griseopilosa (Moser) Philochlaenia [sic] griseopilosa Moser, 1921b: 147 Plectris griseopilosa (Moser); Frey, 1967a: 117 Distribution: Colombia

Plectris griseosetosa Moser Plectris griseosetosa Moser, 1924a: 159 Distribution: Colombia

Synonym: Philochlaenia [sic] columbiana Moser, 1919b: 27 Plectris columbiana (Moser); Evans, 2003: 319

Plectris griseovestita Moser Plectris griseovestita Moser, 1921b: 167 Distribution: Argentina

Plectris guayrana Frey Plectris guayrana Frey, 1967a: 100 Distribution: Paraguay 270 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris gutierrezi Frey Plectris gutierrezi Frey, 1967a: 97 Distribution: Brazil

Plectris hellmichi Frey Plectris hellmichi Frey, 1967a: 110 Distribution: Colombia

Plectris herteli Frey Plectris herteli Frey, 1967a: 76 Distribution: Colombia

Plectris hispidula Frey Plectris hispidula Frey, 1974f: 323 Distribution: Brazil

Plectris huedepohli Frey Plectris huedepohli Frey, 1967a: 29 Distribution: Brazil

Plectris imitans Frey Plectris imitans Frey, 1967a: 109 Distribution: Colombia

Plectris incana (Burmeister) Philochloenia incana Burmeister, 1855: 31 Plectris incana (Burmeister); Frey, 1967a: 89 Distribution: Brazil

Plectris indigens Blanchard Plectris indigens Blanchard, 1850: 127 Plectris indigena Blanchard; Frey, 1967: 41 Distribution: Brazil

Synonym: Philochlaenia [sic] duplosetosa Moser, 1919b: 22 Plectris duplosetosa (Moser); Evans, 2003: 320

Plectris inopinata Frey Plectris inopinata Frey, 1973a: 263 Distribution: Brazil

Plectris integrata (Bates) Pseudoserica integrata Bates, 1887: 154 Plectris integrata (Bates); Dalla Torre, 1913: 335 Distribution: Panama

Plectris intermixta Frey 271 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris intermixta Frey, 1967a: 39 Distribution: Argentina

Plectris juengeri Frey Plectris jüngeri Frey, 1969b: 390 Plectris juengeri Frey; Frey, 1974d: 250 Distribution: Brazil

Plectris juncea (Burmeister) Philochloenia juncea Burmeister, 1855: 39 Plectris juncea (Burmeister); Frey, 1967a: 49 Distribution: Brazil

Plectris junceana Frey Plectris junceana Frey, 1969b: 392 Distribution: Brazil

Plectris kirschi Frey Plectris kirschi Frey, 1967a: 67 (for Burmeister, 1855: 37) Distribution: Brazil

Synonym: Philochloenia tessellata Burmeister, 1855: 37 (not Burmeister, 1855: 84) Plectris tessellata (Burmeister); Evans, 2003: 321

Plectris kochi Frey Plectris kochi Frey, 1967a: 123 Distribution: Peru

Plectris kriegi Frey Plectris kriegi Frey, 1973a: 271 Distribution: Paraguay

Plectris kulzeri Frey Plectris kulzeri Frey, 1967a: 12 Distribution: Colombia

Plectris kuntzeni Moser Plectris kuntzeni Moser, 1921b: 169 Distribution: Brazil

Plectris laevipennis Frey Plectris laevipennis Frey, 1967a: 78 Distribution: Brazil

Plectris laevis Frey Plectris laevis Frey, 1967a: 76 Distribution: Ecuador

272 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris laeviscutata (Moser) Philochlaenia [sic] laeviscutata Moser, 1918b: 122 Plectris laeviscutata (Moser); Frey, 1967a: 83 Distribution: Brazil, Paraguay

Plectris lanata Frey Plectris lanata Frey, 1964c: 699 Distribution: Peru

Plectris laticeps Blanchard Plectris laticeps Blanchard, 1850: 127 Distribution: Brazil

Plectris lepida Burmeister Plectris lepida Burmeister, 1855: 38 Distribution: Brazil

Plectris lignicola Arrow Plectris lignicola Arrow, 1900: 177 Distribution: St. Vincent

Plectris lignicolor Blanchard Plectris lignicolor Blanchard, 1850: 126 Distribution: Argentina, Bolivia

Plectris ligulata Frey Plectris ligulata Frey, 1969: 396 Distribution: Brazil

Synonym: Plectris bahiana Frey, 1974f: 326

Plectris lindneri Frey Plectris lindneri Frey, 1967a: 40 Distribution: Paraguay

Plectris lineatocollis (Blanchard) Philochlaenia [sic] lineatocollis Blanchard, 1850: 121 Plectris lineatocollis (Blanchard); Frey, 1967a: 113 Distribution: Brazil

Synonym: Plectris vittata Burmeister, 1855: 90

Plectris lobaticeps Frey Plectris lobaticeps Frey, 1967a: 16 Distribution: Brazil

Plectris lobaticollis Frey Plectris lobaticollis Frey, 1967a: 68 273 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris lojana Moser Plectris lojana Moser, 1924a: 160 Distribution: Ecuador

Plectris longeantennata Frey Plectris longeantennata Frey, 1967a: 14 Distribution: Brazil

Plectris longiclava Frey Plectris longiclava Frey, 1976b: 383 Distribution: Brazil

Plectris longicornis (Burmeister) Philochloenia longicornis Burmeister, 1855: 29 Plectris longicornis (Burmeister); Frey, 1967a: 33 Distribution: Venezuela

Plectris longitarsis (Bates) Pseudoserica longitarsis Bates, 1887: 154 Plectris longitarsis (Bates); Dalla Torre, 1913: 336 Distribution: Brazil

Plectris longula Moser Plectris longula Moser, 1918b: 156 Distribution: Brazil

Plectris luctuosa Frey Plectris luctuosa Frey, 1967a: 53 Distribution: Brazil

Plectris maculata (Moser) Philochlaenia [sic] maculata Moser, 1919b: 33 Plectris maculata (Moser); Frey, 1967a: 67 Distribution: Brazil

Plectris maculicollis (Arrow) Philochlaenia [sic] maculicollis Arrow, 1925: 216 Plectris maculicollis (Arrow); Frey, 1967a: 85 Distribution: Paraguay

Plectris maculifera Frey Plectris maculifera Frey, 1976b: 384 Distribution: Brazil

Plectris maculipennis Moser Plectris maculipennis Moser, 1924a: 129 274 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris maculipyga Moser Plectris maculipyga Moser, 1918b: 152 Distribution: Brazil

Plectris maculosa (Moser) Philochlaenia [sic] maculosa Moser, 1918b: 132 Plectris maculosa (Moser); Frey, 1967a: 106 Distribution: Brazil

Plectris magdalenae Frey Plectris magdalenae Frey, 1967a: 109 Distribution: Colombia

Plectris mandli Frey Plectris mandli Frey, 1967a: 60 Distribution: Brazil

Plectris marmorea (Guérin-Méneville) Serica marmorea Guérin-Méneville, 1831b: pl. 3 Pseudoserica marmorea Guérin-Méneville, 1838: 87 Plectris marmorea (Guérin-Méneville); Blanchard, 1850: 128 Distribution: Brazil

Plectris martinezi Frey Plectris martinezi Frey, 1967a: 65 (for Moser, 1918b: 135) Distribution: Brazil

Synonym: Philochlaenia [sic] maculipennis Moser, 1918b: 135 (not Moser, 1924a: 129) Plectris maculipennis (Moser); Evans, 2003: 325

Plectris martinicensis Chalumeau Plectris martinicensis Chalumeau, 1982: 331 Distribution: Martinique

Plectris meridana (Moser) Philochlaenia [sic] meridana Moser, 1918b: 131 Plectris meridana (Moser); Frey, 1967a: 114 Distribution: Venezuela

Plectris metallescens (Moser) Philochlaenia [sic] metallescens Moser, 1918b: 124 Plectris metallescens (Moser); Frey, 1967a: 82 Distribution: Brazil

Plectris metallica (Moser) Philochlaenia [sic] metallica Moser, 1918b: 126 275 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris metallica (Moser); Frey, 1967a: 89 Distribution: Colombia

Plectris micans (Kirsch) Philochlaenia [sic] micans Kirsch, 1885: 217 Pseudoserica micans (Kirsch); Bates, 1887: 153 Plectris micans (Kirsch); Frey, 1967a: 105 Distribution: Ecuador, Panama

Plectris minuta (Moser) Philochlaenia [sic] minuta Moser, 1920: 23 Plectris minuta (Moser); Frey, 1967a: 114 Distribution: Brazil

Plectris molesta (Kirsch) Philochlaenia [sic] molesta Kirsch 1874a: 341 Philochloenia modesta (Kirsch); Nonfried, 1892: 269 Plectris molesta (Kirsch); Frey, 1967a: 49 Distribution: Peru

Plectris montana Frey Plectris montana Frey, 1967a: 112 Distribution: Venezuela

Plectris moseri Saylor Plectris moseri Saylor, 1935b: 36 (for Moser, 1918b: 157) Distribution: Brazil

Synonym: Plectris amazonica Moser, 1918b: 157 (not Bates, 1887: 153)

Plectris murina (Blanchard) Philochlaenia [sic] murina Blanchard, 1850: 122 Plectris murina (Blanchard); Frey, 1967a: 83 Distribution: Brazil

Synonym: Philochloenia vittata Burmeister, 1855: 30 Plectris vittata (Burmeister); Evans, 2003: 326

Plectris mus Blanchard Plectris mus Blanchard, 1851: 129 Distribution: Brazil

Plectris muscula Frey Plectris muscula Frey, 1967a: 121 Distribution: Guyana

Plectris neglecta Blanchard Plectris neglecta Blanchard, 1850: 128 276 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris nigrita Moser Plectris nigrita Moser, 1918b: 151 Distribution: Paraguay

Plectris nigritula (Moser) Philochlaenia [sic] nigritula Moser, 1918b: 123 Plectris nigritula (Moser); Frey, 1967a: 85 Distribution: Brazil

Plectris nitidicollis Frey Plectris nitidicollis Frey, 1967a: 99 Distribution: Brazil

Plectris niveoscutata Moser Plectris niveoscutata Moser, 1918b: 149 Distribution: Brazil

Plectris nuda Frey Plectris nuda Frey, 1967a: 34 Distribution: Brazil

Plectris nudicollis Frey Plectris nudicollis Frey, 1967a: 123 Distribution: Bolivia

Plectris obsoleta Blanchard Plectris obsoleta Blanchard, 1850: 128 Distribution: Brazil, Colombia, Costa Rica, México

Synonym: Plectris minor Moser, 1919b: 32

Plectris obtusa (Burmeister) Philochloenia obtusa Burmeister, 1855: 35 Plectris obtusa (Burmeister); Frey, 1967a: 50 Distribution: Brazil

Plectris obtusiodes Frey Plectris obtusiodes Frey, 1967a: 61 Distribution: Brazil

Plectris obtusior Frey Plectris obtusior Frey, 1967a: 62 Distribution: Brazil

Plectris ocularis Frey Plectris ocularis Frey, 1967a: 14 277 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris ohausi (Bruch) Philochloenia ohausi Bruch, 1909: 346 Plectris ohausi (Bruch); Frey, 1967a: 81 Distribution: Brazil

Plectris ohausiana (Moser) Philochlaenia [sic] ohausiana Moser, 1921b: 144 Plectris ohausiana (Moser); Frey, 1967a: 105 Distribution: Ecuador

Plectris ohausiella Frey Plectris ohausiella Frey, 1967a: 8 (for Moser, 1921b: 166) Distribution: Argentina

Synonym: Plectris ohausi Moser, 1921b: 166 (not Bruch, 1909: 346)

Plectris olivierai Frey Plectris olivierai Frey, 1974f: 324 Distribution: Brazil

Plectris opacula albosquamosa Frey Plectris opacula albosquamosa Frey, 1974f: 329 Distribution: Brazil

Plectris opacula opacula Moser Plectris opacula Moser, 1918b: 150 Distribution: Brazil

Plectris ophthalmica Frey Plectris ophthalmica Frey, 1967a: 26 Distribution: Amazonas

Plectris ornaticeps Frey Plectris ornaticeps Frey, 1967a: 92 Distribution: Argentina

Plectris ornatipennis (Moser) Philochlaenia [sic] ornatipennis Moser, 1921b: 154 Plectris ornatipennis (Moser); Frey, 1967a: 66 Distribution: Brazil

Plectris orocuensis Frey Plectris orocuensis Frey, 1967a: 93 Distribution: Colombia

Plectris palpalis (Moser) 278 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Philochlaenia [sic] palpalis Moser, 1918b: 128 Plectris palpalis (Moser); Frey, 1967a: 82 Distribution: Brazil

Plectris panamaensis Frey Plectris panamaensis Frey, 1967a: 124 Distribution: Panama

Plectris paraguayensis Moser Plectris paraguayensis Moser, 1924a: 130 Distribution: Argentina, Paraguay

Plectris paraensis Frey Plectris paraensis Frey, 1967a: 119 Distribution: Brazil, Guyana

Plectris parallela Frey Plectris parallela Frey, 1976b: 382 Distribution: Brazil

Plectris paranensis (Moser) Philochlaenia [sic] paranensis Moser, 1921b: 150 Plectris paranensis (Moser); Frey, 1967a: 79 Distribution: Argentina

Plectris parcesetosa Frey Plectris parcesetosa Frey, 1976b: 386 Distribution: Brazil

Plectris parumsetosa Frey Plectris parumsetosa Frey, 1969b: 395 Distribution: Brazil

Plectris pauloana Moser Plectris pauloana Moser, 1924a: 130 Distribution: Brazil

Plectris pavida (Burmeister) Philochloenia pavida Burmeister, 1855: 32 Plectris pavida (Burmeister); Frey, 1967a: 105 Distribution: Colombia, Venezuela

Synonym: Philochloenia aeruginosa Burmeister, 1855: 32 Pseudoserica aeruginosa (Burmeister); Bates, 1887: 153 Plectris aeruginosa (Burmeister); Dalla Torre, 1913: 335

Plectris pedestris Frey Plectris pedestris Frey, 1967a: 31 279 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris pelliculata (Perty) Serica pelliculata Perty, 1833: 48 Distribution: Brazil

Plectris penaella Frey Plectris penaella Frey, 1967a: 57 Distribution: Peru

Plectris penai Frey Plectris penai Frey, 1967a: 101 Distribution: Peru

Plectris pentaphylla (Moser) Philochlaenia [sic] pentaphylla Moser, 1918b: 136 Plectris pentaphylla (Moser); Frey, 1967a: 65 Distribution: Brazil, French Guiana

Plectris pereirai Frey Plectris pereirai Frey, 1967a: 90 Distribution: Brazil

Plectris perplexa Blanchard Plectris perplexa Blanchard, 1850: 127 Distribution: Brazil

Plectris pexa (Germar) Melolontha pexa Germar, 1824: 123 Plectris pexa (Germar); Blanchard, 1850: 125 Distribution: Brazil

Plectris picea Evans Plectris picea Evans, 2003 (for Moser, 1924: 158) Distribution: Brazil

Synonym: Philochlaenia [sic] nigrita Moser, 1924b: 158 Plectris nigrita (Moser); Frey, 1967a: 35 (not Moser, 1918b: 151)

Plectris pilicollis (Moser) Philochlaenia [sic] pilicollis Moser, 1921b: 146 Plectris pilicollis (Moser); Frey, 1967a: 79 Distribution: Ecuador

Plectris pilifera (Moser) Philochlaenia [sic] pilifera Moser, 1919b: 23 Plectris pilifera (Moser); Frey, 1967a: 52 Distribution: Brazil 280 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris pilosa (Moser) Philochlaenia [sic] pilosa Moser, 1921b: 152 Plectris pilosa (Moser); Frey, 1967a: 89 Distribution: Brazil

Plectris pilosula (Moser) Philochlaenia [sic] pilosula Moser, 1921b: 149 Plectris pilosula (Moser); Frey, 1967a: 117 Distribution: Brazil

Plectris pinsdorfi Frey Plectris pinsdorfi Frey, 1967a: 98 Distribution: Brazil

Plectris piottii (Bruch) Philochloenia piottii Bruch, 1909: 344 Plectris piottii (Bruch); Frey, 1967a: 36 Distribution: Argentina

Plectris plaumanni Frey Plectris plaumanni Frey, 1967a: 38 Distribution: Brazil

Plectris plaumanniella Frey Plectris plaumanniella Frey, 1967a: 44 Distribution: Brazil

Plectris podicalis Moser Plectris podicalis Moser, 1921b: 168 Distribution: Brazil

Plectris postnotata Frey Plectris postnotata Frey, 1967a: 94 Distribution: Brazil

Plectris praecellens Frey Plectris praecellens Frey, 1969b: 398 Distribution: Brazil

Plectris primaria (Burmeister) Philochloenia primaria Burmeister, 1855: 29 Plectris primaria (Burmeister); Frey, 1967a: 74 Distribution: Brazil

Plectris prolata Evans Plectris prolata Evans, 2003 (for Moser, 1918b: 134) Distribution: Colombia 281 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Synonym: Philochlaenia [sic] longula Moser, 1918b: 134 Plectris longula (Moser); Frey, 1967a: 22 (not Moser, 1918b: 156)

Plectris pruina (Burmeister) Philochloenia pruina Burmeister, 1855: 33 Plectris pruina (Burmeister); Frey, 1967a; 118 Distribution: Brazil

Plectris pubens (Moser) Philochlaenia [sic] pubens Moser, 1921b: 151 Plectris pubens (Moser); Frey, 1967a: 88 Distribution: Colombia

Plectris pubera (Burmeister) Philochloenia pubera Burmeister, 1855: 31 Plectris pubera (Burmeister); Frey, 1967a: 88 Distribution: Venezuela

Synonym: Philochlaenia [sic] rufa Moser, 1921b: 151 Plectris rufa (Moser); Evans, 2003: 332

Plectris puberoides Frey Plectris puberoides Frey, 1976b: 388 Distribution: Brazil

Plectris pubescens Blanchard Plectris pubescens Blanchard, 1850: 126 Distribution: Brazil

Plectris pusio Frey Plectris pusio Frey, 1967a: 55 Distribution: Brazil

Plectris putida Frey Plectris putida Frey, 1967a: 19 Distribution: Bolivia

Plectris pygmaea Frey Plectris pygmaea Frey, 1973a: 263 Distribution: Brazil

Plectris quinqueflabellata Moser Plectris quinqueflabellata Moser, 1926: 205 Distribution: Colombia

Plectris reitteri Frey Plectris reitteri Frey, 1967a: 17 282 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Plectris reticulata Frey Plectris reticulata Frey, 1967a: 62 Distribution: Brazil

Plectris riodejaneiroensis Evans Plectris riodejaneiroensis Evans, 2003 (for Moser, 1921b: 151) Distribution: Brazil

Synonym: Philochlaenia [sic] kuntzeni Moser, 1921b: 151 Plectris kuntzeni (Moser); Frey, 1967a: 88 (not Moser, 1921b: 169)

Plectris riveti Frey Plectris riveti Frey, 1967a: 119 Distribution: Ecuador

Plectris roeri Frey Plectris roeri Frey, 1967a: 12 Distribution: Brazil

Plectris rorida (Burmeister) Philochloenia rorida Burmeister, 1855: 34 Plectris rorida (Burmeister); Frey, 1967a: 42 Distribution: Brazil

Plectris rubescens Blanchard Plectris rubescens Blanchard, 1850: 128 Distribution: Brazil

Synonym: Pseudoserica amazonica Bates, 1887: 153 Plectris amazonica (Bates); Blackwelder, 1944; 233

Synonym: Philochloenia puta Burmeister, 1855: 36 Plectris puta (Burmeister); Evans, 2003: 334

Synonym: Philochlaenia [sic] tijucana Moser, 1919b: 20 Plectris tijucana (Moser); Evans, 2003: 334

Plectris ruffoi Frey Plectris ruffoi Frey, 1972b: 94 Distribution: Brazil

Plectris ruficollis Frey Plectris ruficollis Frey, 1976b: 385 Distribution: Brazil

Plectris rufina (Moser) 283 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Philochlaenia [sic] rufina Moser, 1926: 204 Plectris rufina (Moser); Frey, 1967a: 89 Distribution: Venezuela

Plectris rugiceps (Moser) Philochlaenia [sic] rugiceps Moser, 1921b: 152 Plectris rugiceps (Moser); Frey, 1967a: 52 Distribution: Brazil

Plectris rugipennis (Moser) Philochlaenia [sic] rugipennis Moser, 1921b: 146 Plectris rugipennis (Moser); Frey, 1967a: 106 Distribution: Bolivia

Plectris rugulosa Blanchard Plectris rugulosa Blanchard, 1850: 128 Distribution: Brazil

Plectris rugulosipennis Blanchard Plectris rugulosipennis Blanchard, 1851: 129 Distribution: Brazil

Plectris santaecrucis Frey Plectris santaecrucis Frey, 1972b: 94 Distribution: Bolvia

Plectris santosana Frey Plectris santosana Frey, 1967a: 16 Distribution: Brazil

Plectris sarana (Moser) Philochlaenia [sic] sarana Moser, 1921b: 154 Plectris sarana (Moser); Frey, 1967a: 52 Distribution: Bolivia

Plectris schereri Frey Plectris schereri Frey, 1967a: 27 Distribution: Bolivia

Plectris schneblei Frey Plectris schneblei Frey, 1967a: 111 Distribution: Colombia

Plectris scopulata (Burmeister) Philochloenia scopulata Burmeister, 1855: 33 Plectris scopulata (Burmeister); Frey, 1967a: 83 Distribution: Brazil

284 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris sculpterata Frey Plectris sculpterata Frey, 1967a: 73 Distribution: Venezuela

Plectris sculptipennis Frey Plectris sculptipennis Frey, 1974f: 326 Distribution: Peru

Plectris scutalis Blanchard Plectris scutalis Blanchard, 1850: 128 Distribution: Brazil

Plectris scutellaris Blanchard Plectris scutellaris Blanchard, 1850: 126 Distribution: Brazil

Synonym: Plectris securifera Moser, 1919b: 47

Plectris sequela Evans Plectris sequela Evans, 2003:336 (for Moser, 1921b: 148) Distribution: Brazil

Synonym: Philochlaenia [sic] corumbana Moser, 1921b: 148 Plectris corumbana (Moser); Frey, 1967a; 118 (not Moser, 1921b: 169)

Plectris sericea (Moser) Philochlaenia [sic] sericea Moser, 1921b: 145 Plectris sericea (Moser); Frey, 1967a: 104 Distribution: Venezuela

Plectris setifera Burmeister Plectris setifera Burmeister, 1855: 85 Distribution: Brazil

Plectris setisparsa (Bates) Pseudoserica setisparsa Bates, 1887: 153 Plectris setisparsa (Bates); Frey, 1967a: 52 Distribution: Guatemala, Nicaragua

Plectris setiventris Moser Plectris setiventris Moser, 1918b: 155 Distribution: Brazil

Plectris setosa Moser Plectris setosa Moser, 1924a: 129 Distribution: Brazil

Plectris setosella (Moser) 285 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Philochlaenia [sic] setosella Moser, 1919b: 35 Plectris setosella (Moser); Frey, 1967a: 48 Distribution: Brazil

Plectris setulifera (Moser) Philochlaenia [sic] setulifera Moser, 1919b: 36 Plectris setulifera (Moser); Frey, 1967a: 48 Distribution: Brazil

Plectris signaticollis Frey Plectris signaticollis Frey, 1974f: 325 Distribution: Brazil

Plectris signativentris Moser Plectris signativentris Moser, 1918b: 160 Distribution: Brazil

Plectris similis Frey Plectris similis Frey, 1973a: 273 Distribution: Brazil

Plectris sinuaticeps (Moser) Philochlaenia [sic] sinuaticeps Moser, 1921b: 146 Plectris sinuaticeps (Moser); Frey, 1967a: 52 Distribution: Brazil

Plectris sordida (Burmeister) Philochloenia sordida Burmeister, 1855: 37 Plectris sordida (Burmeister); Frey, 1967a: 64 Distribution: Brazil

Plectris sororia (Moser) Philochlaenia [sic] sororia Moser, 1919b: 21 Plectris sororia (Moser); Frey, 1967a: 47 Distribution: Brazil

Synonym: Philochlaenia [sic] piliventris Moser, 1919b: 22 Plectris piliventris (Moser); Evans, 2003: 337

Plectris sparsecrinita Frey Plectris sparsecrinita Frey, 1976b: 385 Distribution: Brazil

Plectris sparsepilosa Frey Plectris sparsepilosa Frey, 1967a: 97 Distribution: Venezuela

Plectris sparsepunctata Frey 286 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris sparsepunctata Frey, 1967a: 71 Distribution: Colombia, Venezuela

Plectris sparsesetosa Moser Plectris sparsesetosa Moser, 1924b: 159 Distribution: French Guiana

Plectris spatulata Frey Plectris spatulata Frey, 1967a: 30 Distribution: Brazil

Plectris splendens Frey Plectris splendens Frey, 1967a: 89 Distribution: Ecuador

Plectris splendida Frey Plectris splendida Frey, 1967a: 70 Distribution: Venezuela

Plectris squalida Frey Plectris squalida Frey, 1967a: 102 Distribution: Brazil

Plectris squamiger Frey Plectris squamiger Frey, 1967a: 98 Distribution: Brazil

Plectris squamisetis Frey Plectris squamisetis Frey, 1967a: 39 Distribution: Brazil

Plectris subaenea Moser Plectris subaenea Moser, 1918b: 159 Distribution: Brazil

Plectris subcarinata Frey Plectris subcarinata Frey, 1967a: 25 Distribution: Brazil

Plectris subcostata Blanchard Plectris subcostata Blanchard, 1850: 126 Distribution: Brazil

Plectris subdepressa Blanchard Plectris subdepressa Blanchard, 1850: 126 Distribution: Brazil

Plectris subglabra (Moser) 287 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Philochlaenia [sic] subglabra Moser, 1918b: 125 Plectris subglabra (Moser); Frey, 1967a: 74 Distribution: Brazil

Plectris sulcicollis (Moser) Philochlaenia [sic] sulcicollis Moser, 1919a: 29 Plectris sulcicollis (Moser); Frey, 1967a: 41 Distribution: Brazil

Plectris suturalis Burmeister Plectris suturalis Burmeister, 1855: 91 Distribution: Brazil

Plectris tarsalis (Moser) Philochlaenia [sic] tarsalis Moser, 1919b: 25 Plectris tarsalis (Moser); Frey, 1967a: 52 Distribution: Brazil

Plectris tenebrosa Frey Plectris tenebrosa Frey, 1967a: 121 Distribution: Peru

Plectris tenueclava Frey Plectris tenueclava Frey, 1976b: 387 Distribution: Brazil

Plectris tenuevestita Frey Plectris tenuevestita Frey, 1967a: 13 Distribution: Paraguay

Plectris tessellata Burmeister Plectris tessellata Burmeister, 1855: 84 Distribution: Brazil

Plectris tetraphylla Frey Plectris tetraphylla Frey, 1967a: 58 Distribution: Brazil

Plectris teutoniensis Frey Plectris teutoniensis Frey, 1969b: 389 Distribution: Brazil

Plectris tolimana (Moser) Philochlaenia [sic] tolimana Moser, 1921b: 144 Plectris tolimana (Moser); Frey, 1967a: 106 Distribution: Colombia, Peru

Plectris tomentosa Le Peletier and Audinet-Serville 288 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris tomentosa Le Peletier and Audinet-Serville, 1828: 369 Distribution: Brazil

Plectris tricostata (Burmeister) Philochloenia tricostata Burmeister, 1855: 35 Plectris tricostata (Burmeister); Frey, 1967a; 49 Distribution: Brazil

Plectris tristis Frey Plectris tristis Frey, 1967a: 13 Distribution: Argentina

Plectris truncata (Blanchard) Philochlaenia [sic] truncata Blanchard, 1850: 122 Plectris truncata (Blanchard); Frey, 1967a: 74 Distribution: Brazil

Plectris tuberculata (Moser) Philochlaenia [sic] tuberculata Moser, 1919b: 28 Plectris tuberculata (Moser); Frey, 1967a: 47 Distribution: Colombia, Peru, Venezuela

Plectris tucumana (Bruch) Philochloenia tucumana Bruch, 1909: 345 Plectris tucumana (Bruch); Frey, 1967a: 80 Distribution: Argentina

Plectris umbilicata Frey Plectris umbilicata Frey, 1967a: 55 Distribution: Brazil

Plectris umbrata Frey Plectris umbrata Frey, 1967a: 56 Distribution: Brazil

Plectris unidens Frey Plectris unidens Frey, 1967a: 26 Distribution: Brazil

Plectris validior Burmeister Plectris validior Burmeister, 1855: 89 Distribution: Argetina

Plectris variegata (Moser) Philochlaenia [sic] variegata Moser, 1919b: 32 Plectris variegata (Moser); Frey, 1967a: 65 Distribution: Brazil

289 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris variipennis (Moser) Philochlaenia [sic] variipennis Moser, 1921b: 154 Plectris variipennis (Moser); Frey, 1967a: 64 Distribution: Brazil

Synonym: Philochloenia brevis Burmeister, 1855: 38 (not Blanchard, 1851: 129) Plectris brevis (Burmeister); Evans, 2003: 341

Plectris vauriella Frey Plectris vauriella Frey, 1967a: 108 Distribution: Bolivia

Plectris vestita (Burmeister) Philochloenia vestita Burmeister, 1855: 31 Distribution: Brazil

Plectris vicina Frey Plectris vicina Frey, 1967a: 28 Distribution: Brazil

Plectris vilis (Burmeister) Philochloenia vilis Burmeister, 1855: 33 Plectris vilis (Burmeister); Frey, 1967a: 118 Distribution: Colombia

Plectris villiersi Frey Plectris villiersi Frey, 1967a: 107 Distribution: Ecuador

Plectris violascens Blanchard Plectris violascens Blanchard, 1851: 129 Distribution: Brazil

Plectris viridifusca (Moser) Philochlaenia [sic] viridifusca Moser, 1918b: 130 Plectris viridimicans (Moser); Frey, 1967a: 89 Distribution: Brazil

Plectris viridimicans (Moser) Philochlaenia [sic] viridimicans Moser, 1918b: 128 Plectris viridimicans (Moser); Frey, 1967a: 85 Distribution: French Guiana

Plectris vitticollis Moser Plectris vitticollis Moser, 1918b: 149 Distribution: Brazil

Plectris vittipennis Frey 290 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Plectris vittipennis Frey, 1972b: 95 Distribution: Ecuador

Plectris vonvolxemi Frey Plectris vonvolxemi Frey, 1967a: 45 Distribution: Brazil

Plectris wittmeri Frey Plectris wittmeri Frey, 1967a: 57 Distribution: Argentina

Plectris witzgalli Frey Plectris witzgalli Frey, 1973a: 266 Distribution: Brazil

Plectris wolfrumi Frey Plectris wolfrumi Frey, 1969b: 388 Distribution: Brazil

Plectris yungasa Frey Plectris yungasa Frey, 1975b: 252 Distribution: Bolivia

Plectris zikani Moser Plectris zikani Moser, 1924a: 124 Distribution: Brazil

Plectris zischkaella Frey Plectris zischkaella Frey, 1967a: 122 Distribution: Bolivia

Plectris zischkai Frey Plectris zischkai Frey, 1967a: 38 Distribution: Bolivia

Genus PRISTEROPHORA Harold Pristerophora Harold, 1869b: 123 (for Solier, 1851: 101). Type species Prionophora picipennis Solier, by monotypy.

Synonym: Prionophora Solier, 1851: 101 (preoccupied). Type species Prionophora picipennis Solier, by monotypy.

Pristerophora flavipennis (Philippi) Prionophora flavipennis Philippi, 1864: 436 Pristerophora flavipennis (Philippi); Dalla Torre, 1913: 331 Distribution: Chile

Pristerophora picipennis (Solier) 291 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Prionophora picipennis Solier, 1851: 102 Pristerophora picipennis (Solier); Harold, 1869a: 1149 Distribution: Chile

Genus PSEUDODICRANIA Gutiérrez Pseudodicrania Gutiérrez, 1950: 275. Type species Dicrania aeneobrunnea Philippi, by original designation.

Pseudodicrania aeneobrunnea (Philippi) Dicrania aeneobrunnea Philippi, 1861: 738 Pseudodicrania aeneobrunnea (Philippi); Gutiérrez, 1950: 276 Distribution: Chile

Genus PSEUDOHERCITIS Moser Pseudohercitis Moser, 1921b: 174. Type species Pseudohercitis viridiaenea Moser, subsequent designation (Evans 2003: 343).

Pseudohercitis corumbana Moser Pseudohercitis corumbana Moser, 1921b: 175 Distribution: Brazil

Pseudohercitis nudipennis Frey Pseudohercitis nudipennis Frey, 1972b: 99 Distribution: Brazil

Pseudohercitis umbonata Gutiérrez Pseudohercitis umbonata Gutiérrez, 1952: 217 Distribution: Venezuela

Pseudohercitis viridiaenea Moser Pseudohercitis viridiaenea Moser, 1921b: 174 Distribution: Brazil

Genus PSEUDOLEURETRA Martínez and D'Andretta Pseudoleuretra Martínez and D'Andretta, 1956: 351. Type species Pseudoleuretra bokermanni Martínez and D'Andretta, by original designation.

Pseudoleuretra bokermanni Martínez and D'Andretta Pseudoleuretra bokermanni Martínez and D'Andretta, 1956: 352 Distribution: Ecuador

Genus RHINASPIS Perty Rhinaspis Perty, 1833: 46. Type species Rhinaspis schrankii Perty, a junior synonym of Melolontha aenea Billberg, by monotypy.

Rhinaspis aenea (Billberg) Melolontha aenea Billberg, 1820: 385 Rhinaspis aenea (Billberg); Burmeister, 1855: 66 292 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: Brazil

Synonym: Rhinaspis metallica Brullé, 1834: 38

Synonym: Rhinaspis schrankii Perty, 1833: 47

Rhinaspis aeneocuprea Moser Rhinaspis aeneocuprea Moser, 1921b: 170 Distribution: Brazil

Rhinaspis dilaticornis Moser Rhinaspis dilaticornis Moser, 1924a: 127 Distribution: Argentina

Rhinaspis fusca Moser Rhinaspis fusca Moser, 1924a: 127 Distribution: Brazil

Rhinaspis micans Burmeister Rhinaspis micans Burmeister, 1855: 66 Distribution: Brazil

Rhinaspis murina Burmeister Rhinaspis murina Burmeister, 1855: 66 Distribution: Brazil

Rhinaspis ohausi Moser Rhinaspis ohausi Moser, 1921b: 169 Distribution: Brazil

Rhinaspis rugicollis Moser Rhinaspis rugicollis Moser, 1924a: 127 Distribution: Brazil

Rhinaspis setosa Frey Rhinaspis setosa Frey, 1974f: 322 Distribution: Brazil

Rhinaspis umbilicata Frey Rhinaspis umbilicata Frey, 1973a: 258 Distribution: Brazil

Genus RHINASPOIDES Moser Rhinaspoides Moser, 1919c: 348. Type species Rhinaspoides aeneofusca Moser, by monotypy.

Rhinaspoides aeneofusca Moser Rhinaspoides aeneofusca Moser, 1919c: 348 Distribution: Brazil 293 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Genus SCHIZOCHELUS Blanchard Schizochelus Blanchard, 1850: 89. Type species Schizochelus flavescens Blanchard, subsequent designation (Evans 2003: 346).

Schizochelus bicoloripes Blanchard Schizochelus bicoloripes Blanchard, 1850: 89 Distribution: Brazil

Schizochelus breviventris Philippi Schizochelus breviventris Philippi, 1864: 445 Distribution: Chile

Schizochelus flavescens Blanchard Schizochelus flavescens Blanchard, 1850: 89 Distribution: Brazil

Schizochelus modestus Philippi Schizochelus modestus Philippi, 1861: 738 Distribution: Chile

Schizochelus serratus Philippi Schizochelus serratus Philippi, 1864: 444 Distribution: Chile

Schizochelus ursulus Philippi Schizochelus ursulus Philippi, 1864: 446 Distribution: Chile

Schizochelus vestitus Philippi Schizochelus vestitus Philippi, 1864: 447 Distribution: Chile

Genus ULOMENES Blanchard Ulomenes Blanchard, 1850: 125. Type species Ulomenes hypocrita Blanchard, by monotypy.

Ulomenes hypocrita Blanchard Ulomenes hypocrita Blanchard, 1850: 125 Distribution: Brazil

Tribe HOPLIINI Burmeister, 1855

Genus HOPLIA Illiger Hoplia Illiger, 1803: 226. Type species Scarabaeus farinosus Linné, by subsequent designation (Medvedev, 1952: 244).

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Keys to species: LeConte, 1880: 191 (United States); Blatchley, 1910: 948 (Indiana); Boyer, 1940: 6 (Canada, United States); Hardy, 1977: 5 (Canada, United States); Lago et al., 1979: 74 (North Dakota); Morón, 1994: 39 (Hidalgo); Harpootlian, 2001: 96 (South Carolina).

Hoplia albisparsa Bates Hoplia albisparsa Bates, 1887: 135 Distribution: México

Hoplia argentata Nonfried Hoplia argentata Nonfried, 1891: 259 Distribution: Honduras

Synonym: Hoplia tricolor Nonfried, 1895: 288

Hoplia argyritis Bates Hoplia argyritis Bates, 1887: 133 Distribution: Costa Rica, Nicaragua

Hoplia asperula Bates Hoplia asperula Bates, 1887: 131 Distribution: México

Hoplia callipyge LeConte Hoplia callipyge LeConte, 1856: 285 Distribution: Canada, México, United States

Synonym: Hoplia cazieri Boyer, 1940: 8

Synonym:Hoplia convexula LeConte, 1856: 285

Synonym: Hoplia humboldtensis Boyer, 1940: 12

Synonym: Hoplia irrorata LeConte, 1857: 40 (not Blanchard, 1850; Burmeister, 1855)

Synonym: Hoplia lecontei Dalla Torre, 1913: 376 (for LeConte, 1856: 285)

Synonym: Hoplia mutata Harold, 1869a: 1113 (for LeConte, 1857: 40)

Synonym: Hoplia oregona LeConte, 1856: 284

Synonym: Hoplia oregona deserticola Boyer, 1940: 20

Synonym: Hoplia pubicollis LeConte, 1856: 285 (not Kuster, 1849)

Synonym: Hoplia utahensis Boyer, 1940: 24

Hoplia cinereonebulosa Nonfried Hoplia cinereo-nebulosa Nonfried, 1895: 288 295 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Hoplia cretacea Bates Hoplia cretacea Bates, 1887: 132 Distribution: México

Hoplia dispar LeConte Hoplia dispar LeConte, 1880: 192 Distribution: United States

Hoplia disparilis Bates Hoplia disparilis Bates, 1887: 132 Distribution: Guatemala

Hoplia equina LeConte Hoplia equina LeConte, 1880: 193 Distribution: United States

Hoplia festiva Burmeister Hoplia festiva Burmeister, 1844: 193 Distribution: México

Hoplia floridana Fisher Hoplia floridana Fisher, 1918: 140 Distribution: United States

Hoplia gilleti Hardy Hoplia gilleti Hardy, 1977: 28 Distribution: United States

Hoplia guatemalensis Bates Hoplia guatemalensis Bates, 1887: 135 Distribution: Guatemala

Hoplia hirta LeConte Hoplia hirta LeConte, 1880: 193 Distribution: United States

Hoplia horrida Moser Hoplia horrida Moser, 1918d: 342 Distribution: Honduras

Hoplia inops Bates Hoplia inops Bates, 1887: 133 Distribution: Guatemala, México

Hoplia jalapana Moser Hoplia jalapana Moser, 1918d: 341 296 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Hoplia laticollis LeConte Hoplia laticollis LeConte, 1856: 284 Distribution: Canada, United States

Hoplia limbata LeConte Hoplia limbata LeConte, 1856: 286 Distribution: ?Neotropical [see Hardy, 1977: 30]

Hoplia lineata Moser Hoplia lineata Moser, 1921b: 181 Distribution: México

Hoplia lurida Moser Hoplia lurida Moser, 1918d: 340 Distribution: México

Hoplia mexicana Harold Hoplia mexicana Harold, 1869a: 1113 (for Burmeister, 1855: 484) Distribution: México

Synonym: Hoplia irrorata Burmeister, 1855: 484 (not Blanchard, 1850; LeConte, 1857: 40)

Hoplia modesta Haldeman Hoplia modesta Haldeman, 1843: 304 Distribution: Canada, United States

Synonym: Hoplia modesta var. barbata Blatchley, 1910: 950 Hoplia barbata Blatchley; Boyer, 1940: 15

Synonym: Hoplia helvola Melsheimer, 1846: 142

Synonym: Hoplia monticola Melsheimer, 1846: 141

Synonym: Hoplia singularis Burmeister, 1844: 192

Hoplia mucorea (Germar) Melolontha mucorea Germar, 1824: 129 Hoplia mucorea (Germar), Burmeister, 1844: 193 Distribution: United States

Synonym: Hoplia meridionalis Boyer, 1940: 14

Hoplia oblonga Gyllenhal Hoplia oblonga Gyllenhal, 1817: 68 Distribution: ?New World [see Hardy, 1977: 31]

297 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Hoplia ochracea Burmeister Hoplia ochracea Burmeister, 1844: 198 Distribution: Brazil

Hoplia pisicolor Burmeister Hoplia pisicolor Burmeister, 1844: 194 Distribution: Brazil

Hoplia rotunda Bates Hoplia rotunda Bates, 1887: 134 Distribution: Guatemala

Hoplia sackenii LeConte Hoplia sackenii LeConte, 1880: 192 Distribution: Canada, United States

Hoplia squamifera Burmeister Hoplia squamifera Burmeister, 1844: 198 Distribution: Guatemala, México, Panama

Hoplia striatipennis Wickham Hoplia striatipennis Wickham, 1914: 457 Distribution: United States Remark: fossil species

Hoplia subcostata Bates Hoplia subcostata Bates, 1887: 134 Distribution: Guatemala

Hoplia surata Bates Hoplia surata Bates, 1887: 135 Distribution: Costa Rica

Hoplia teapensis Bates Hoplia teapensis Bates, 1887: 133 Distribution: México

Hoplia trifasciata Say Hoplia trifasciata Say, 1825: 200 Distribution: Canada, United States

Synonym: Hoplia ferrisi Boyer, 1940: 10

Synonym: Hoplia primaria Burmeister, 1844: 192

Synonym: Hoplia tristis Melsheimer, 1846: 141

Hoplia trivialis Harold 298 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Hoplia trivialis Harold, 1869a: 1114 (for LeConte, 1856: 285) Distribution: United States

Synonym: Hoplia debilis LeConte, 1856: 285 (not Burmeister, 1844)

Hoplia vidua Moser Hoplia vidua Moser, 1918d: 339 Distribution: Honduras

INCERTAE SEDIS-GENERA

Genus ACOMA Casey Acoma Casey, 1889:165. Type species Acoma brunnea Casey, by monotypy.

Keys to species: Howden, 1958b: 379

Acoma arizonica Brown Acoma arizonica Brown, 1929: 212 Distribution: United States

Acoma brunnea Casey Acoma brunnea Casey, 1889: 167 Distribution: United States

Acoma cazieri Saylor Acoma cazieri Saylor, 1948a: 341 Distribution: México

Acoma confusa Van Dyke Acoma confusa Van Dyke, 1928: 160 Distribution: México

Acoma conjuncta Howden Acoma conjuncta Howden, 1962: 1153 Distribution: United States

Acoma dilemma Saylor Acoma dilemma Saylor, 1948a: 342 Distribution: México

Acoma diminiata Howden Acoma diminiata Howden, 1958b: 396 Distribution: United States

Acoma evansi Howden Acoma evansi Howden, 1962: 1148 Distribution: México

299 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Acoma gibsoni Howden Acoma gibsoni Howden, 1962: 1149 Distribution: México

Acoma glabrata Cazier Acoma glabrata Cazier, 1953: 5 Distribution: México, United States

Synonym: Acoma nigrita Cazier, 1953: 7

Acoma granulifrons Howden Acoma granulifrons, Howden, 1958b: 392 Distribution: México

Acoma incognita Howden Acoma incognita Howden, 1958b: 388 Distribution: México

Acoma knulli Howden Acoma knulli Howden, 1958b: 395 Distribution: United States

Acoma leechi Cazier, 1953: 7 Acoma leechi Cazier, 1953: 7 Distribution: México

Acoma martini Howden Acoma martini Howden, 1962: 1152 Distribution: México

Acoma mimica Howden Acoma mimica Howden, 1962: 1147 Distribution: México

Acoma minuta Cazier Acoma minuta Cazier, 1953: 10 Distribution: México

Acoma mixta Howden Acoma mixta Howden, 1958b: 384 Distribution: United States

Acoma ochlera Howden Acoma ochlera Howden, 1958b: 391 Distribution: México

Acoma parva Howden Acoma parva Howden, 1958b: 390 300 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Distribution: México

Acoma robusta Van Dyke Acoma robusta Van Dyke, 1928: 159 Distribution: México

Acoma rossi Saylor Acoma rossi Saylor, 1948a: 342 Distribution: México

Acoma rufula Howden Acoma rufula Howden, 1958b: 397 Distribution: Arizona

Acoma seticollis Howden Acoma seticollis Howden, 1958b: 400 Distribution: México

Acoma sexfoliata Saylor Acoma sexfoliata Saylor, 1948a: 342 Distribution: México

Acoma stathamae Cazier Acoma stathamae Cazier, 1953: 3 Distribution: México

Genus CHAUNOCOLUS Saylor Chaunocolus Saylor, 1937b: 35. Type species Chaunocolus cornutus Saylor, by monotypy.

Chaunocolus cornutus Saylor Chaunocolus cornutus Saylor, 1937b: 35 Distribution: México

Genus CHNAUNANTHUS Burmeister Chnaunanthus Burmeister, 1844: 31. Type species Chnaunanthus discolor Burmeister, by monotypy.

Synonym: Acratus Horn, 1867: 165 (not Lacordaire, 1867). Type species Acratus flavipennis Horn, by monotypy.

Synonym: Pseudacratus Dalla Torre, 1912a: 7 (for Horn, 1867: 165). Type species Acratus flavipennis Horn, by monotypy.

Keys to species: Horn, 1894: 394; Saylor, 1937e: 534

Chnaunanthus chapini Saylor Chnaunanthus chapini Saylor, 1937e: 535 Distribution: United States

301 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Chnaunanthus discolor Burmeister Chnaunanthus discolor Burmeister, 1844: 32 Distribution: México

Chnaunanthus flavipennis (Horn) Acratus flavipennis Horn, 1867: 166 Pseudacratus flavipennis (Horn); Dalla Torre, 1912a: 7 Chnaunanthus flavipennis (Horn); Saylor, 1937e: 535 Distribution: United States

Synonym: Chnaunanthus palmeri Horn, 1894: 393

Genus DIAPHYLLA Erichson Diaphylla Erichson, 1847a: 102. Type species Diaphylla hispida Erichson, by monotypy.

Diaphylla granulata Philippi Diaphylla granulata Philippi, 1864: 439 Distribution: Chile

Diaphylla hispida Erichson Diaphylla hispida Erichson, 1847a: 102 Distribution: Peru

Diaphylla lampropyga Philippi Diaphylla lampropyga Philippi, 1864: 438 Distribution: Chile

Diaphylla luctuosa Philippi and Philippi Diaphylla luctuosa Philippi and Philippi, 1864: 323 Distribution: Chile

Diaphylla ornata Philippi Diaphylla ornata Philippi, 1864: 439 Distribution: Chile

Genus PHYTHOLAEMA Blanchard Phytholaema Blanchard, 1851: 218. Type species Areoda mutabilis Solier, by monotypy.

Synonym: Lacris Fairmaire and Germain, 1860: 268. Type species Lacris dilutipes Fairmaire and Germain, by monotypy.

Synonym: Melicurus Germain, 1905: 470. Type species Phytholaema herrmanni herrmanni Germain, by subsequent designation (Smith, 2002: 393). note: Areoda Solier, 1851: 92 (not MacLeay, 1819) is a misapplication.

Keys to species: Saylor, 1937a: 8

302 CHECKLIST OF THE NEW WORLD MELOLONTHINAE-EVANS AND SMITH

Phytholaema dilutipes (Fairmaire and Germain) Lacris dilutipes Fairmaire and Germain, 1860: 268 Phytholaema dilutipes Fairmaire and Germain; Martínez, 1975: 227 Distribution: Chile

Synonym: Phytolaema [sic] elaphocera Redtenbacher, 1868: 65

Synonym: Phytoloema [sic] flavipes Philippi, 1861: 735 Phytholaema flavipes Philippi; Saylor, 1937a: 11

Phytholaema herrmanni herrmanni Germain, 1901: 10 Phytoloema [sic] herrmanni Germain, 1901: 10 Phytholaema herrmanni Germain; Saylor 1937a: 10 Distribution: Chile

Phytholaema herrmanni pallida Saylor Phytholaema herrmanni pallida Saylor, 1937a: 10 Distribution: Chile

Synonym: Phytholaema herrmanni peccans Blackwelder, 1944: 221 (in error for Porter, 1939: 131)

Phytholaema mutabilis (Solier) Areoda mutabilis Solier, 1851: 93 Phytholaema mutabilis (Solier); Blanchard, 1851: 219 Phytolaema [sic] mutabilis (Solier); Lacordaire, 1856: 226 Phytoloema [sic] mutabilis (Solier); Germain, 1901: 16 Distribution: Chile

INCERTAE SEDIS-SPECIES

Dichelonyx violaceipennis (Blanchard) Dichelonycha [sic] violaceipennis Blanchard, 1850: 116 Dichelonyx violaceipennis (Blanchard); Saylor, 1945b: 137 Distribution: South America

Philochloenia femorata (Nonfried) Bolax femoratus Nonfried, 1894: 113 Philochlaenia [sic] femoratus (Nonfried); Ohaus, 1917: 9 Distribution: Brazil

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Aragón, A., Morón, M.A., Tapia-Rojas, A.M., and Rojas-Garcia, R. (2001) Fauna de Coleoptera Melolonthidae en el Rancho “La Joya,” Atlixco, Puebla, México. Acta Zoologica Mexicana, n.s., 83, 143-164.

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Arrow, G.J. (1927) A note on the coleopterous Genus Aserica (Melolonthinae). Proceedings of the Entomological Society of Washington, 29(3), 69-70.

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Blackwelder, R.E. (1939) Fourth supplement, 1933 to 1938 (inclusive) to the Leng catalogue of Coleoptera of America, north of Mexico. John D. Sherman, Mt. Vernon, USA, 146 pp.

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Frey, G. (1970a) Betimmumgstabelle der Isonychus-Arten aus Guayana, Brasilien und Argentinien (Col. Melol. Macrodactylini). Entomologischen Arbeiten aus dem Museum G. Frey, 21, 123- 147.

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Hardy, A.R. (1987) In: Hardy, A.R. and Andrews, F.G. Studies in the Coleoptera of western sand dunes. 3. Remarks upon Serica (Scarabaeidae) from some Nevada sand dunes with descriptions of two new taxa. The Coleopterists Bulletin, 41(2), 173-179. von Harold, E. (1863) Note sur les espèces mexicaines du genre Phanaeus et descriptions de quelques espèces nouvelles de coléoptères mexicains. Annales de la Société Entomologique de France, (4)3, 161-176. von Harold, E. (1869a) Scarabaeidae, pp. 976-1346. In: Gemminger, M. and Harold, E. Catalogus Coleopterorum. bucusque descriptorum, synonymicus et systematics, autotribus. Tome IV. Scarabaeidae. Munich. von Harold, E. (1869b) Abanderungen vergebener Namen. Coleopterische, 5, 122-125.

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Howden, H.F. (1959) A new species of Isonychus Mannerheim from southern Arizona (Coleoptera: Scarabaeidae). The Canadian Entomologist, 91(1), 59-61.

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Young, R.M. (1988) A monograph of the genus Polyphylla Harris in American north of Mexico (Coleoptera: Scarabaeidae: Melolonthinae). Bulletin of the University of Nebraska State Museum, 11(2), 1-115.

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